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<document id="9993B3D2C214929EBAA5AE908D3215F6" ID-CLB-Dataset="299666" ID-DOI="10.11646/zootaxa.5474.5.1" ID-GBIF-Dataset="f6a45a12-d3b6-44a7-997e-75eb5ebe9901" ID-ISSN="1175-5326" ID-Zenodo-Dep="12712066" ID-ZooBank="3DDAA7F0-27D0-43E8-AAFB-E56F0E24FB3F" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1720373654954" checkinUser="plazi" docAuthor="Pisanty, Gideon &amp; Wood, Thomas James" docDate="2024" docId="077587FDB85F643FFF6CDDC5EC53FC26" docLanguage="en" docName="zootaxa.5474.5.1.pdf" docOrigin="Zootaxa 5474 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5474.5.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Andrena remota Pisanty &amp; Wood 2024, sp. nov." docType="treatment" docVersion="5" lastPageNumber="471" masterDocId="FB4CFF85B84E642BFFFBDB78E95AFFCB" masterDocTitle="The early-diverging subgenera of the bee genus Andrena (Hymenoptera: Andrenidae) in the Old World" masterLastPageNumber="488" masterPageNumber="451" pageNumber="468" updateTime="1720652268470" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F3209D1195405A27E69E424C7F0673B0">The early-diverging subgenera of the bee genus Andrena (Hymenoptera: Andrenidae) in the Old World</mods:title>
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<mods:namePart id="33C84E79E597DBB473F9E418CA109C25">Pisanty, Gideon</mods:namePart>
<mods:affiliation id="305F0BBEFCCD59A5198F57727802DC34">The Steinhardt Museum of Natural History, Tel Aviv University, Tel Aviv 6997801, Israel.</mods:affiliation>
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<mods:namePart id="63FD037F2597CE995658AF5EEC92E71E">Wood, Thomas James</mods:namePart>
<mods:affiliation id="2B393A036919B9100095E7E7565DB399">Naturalis Biodiversity Center, Darwinweg 2, 2333 CR, Leiden, the Netherlands.</mods:affiliation>
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<mods:title id="2B68841C4F7CC03BE85A501A8B448CEA">Zootaxa</mods:title>
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<mods:part id="835C20AE94C90BB7B9C7EECE2ED59E32">
<mods:date id="EA217488416444FF6FE34CAB107F96F1">2024</mods:date>
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<mods:number id="080FCF5D3FE1B038C5E426FB577AD923">2024-07-02</mods:number>
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<mods:classification id="7B71FFBC02464737266B91C2489DCB84">journal article</mods:classification>
<mods:identifier id="863AFCFEA3308B2E6621B0B2A2930E09" type="CLB-Dataset">299666</mods:identifier>
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<treatment id="077587FDB85F643FFF6CDDC5EC53FC26" ID-DOI="http://doi.org/10.5281/zenodo.12710348" ID-GBIF-Taxon="235032126" ID-Zenodo-Dep="12710348" LSID="urn:lsid:plazi:treatment:077587FDB85F643FFF6CDDC5EC53FC26" httpUri="http://treatment.plazi.org/id/077587FDB85F643FFF6CDDC5EC53FC26" lastPageId="20" lastPageNumber="471" pageId="17" pageNumber="468">
<subSubSection id="C7C66560B85F643AFF6CDDC5EB8BF913" box="[151,721,1725,1752]" pageId="17" pageNumber="468" type="nomenclature">
<paragraph id="8F6336EBB85F643AFF6CDDC5EB8BF913" blockId="17.[151,721,1725,1789]" box="[151,721,1725,1752]" pageId="17" pageNumber="468">
<heading id="D42B8187B85F643AFF6CDDC5EB8BF913" bold="true" box="[151,721,1725,1752]" fontSize="11" level="1" pageId="17" pageNumber="468" reason="1">
<emphasis id="BDA8EAF9B85F643AFF6CDDC5EB8BF913" bold="true" box="[151,721,1725,1752]" pageId="17" pageNumber="468">
<taxonomicName id="48DC4D68B85F643AFF6CDDC5EB36F913" authority="Wood" authorityName="Pisanty &amp; Wood" authorityYear="2024" box="[151,620,1725,1752]" class="Insecta" family="Andrenidae" genus="Andrena" isHigherUncertain="true" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="remota" status="sp. nov.">
<emphasis id="BDA8EAF9B85F643AFF6CDDC5E85BF913" bold="true" box="[151,257,1725,1752]" italics="true" pageId="17" pageNumber="468">Andrena</emphasis>
(
<emphasis id="BDA8EAF9B85F643AFEE9DDC6E8ECF913" bold="true" box="[274,438,1725,1752]" italics="true" pageId="17" pageNumber="468">incertae sedis</emphasis>
)
<emphasis id="BDA8EAF9B85F643AFE3CDDC6EB46F913" bold="true" box="[455,540,1726,1752]" italics="true" pageId="17" pageNumber="468">remota</emphasis>
Wood
</taxonomicName>
<taxonomicNameLabel id="A69B5782B85F643AFD88DDC6EB8BF913" box="[627,721,1726,1752]" pageId="17" pageNumber="468" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C7C66560B85F6438FF6CDD9AE8D2FB6B" lastPageId="19" lastPageNumber="470" pageId="17" pageNumber="468" type="description">
<paragraph id="8F6336EBB85F643AFF6CDD9AE874F936" blockId="17.[151,721,1725,1789]" box="[151,302,1762,1789]" pageId="17" pageNumber="468">
(
<figureCitation id="17E72A6EB85F643AFF64DD9AE87CF936" box="[159,294,1762,1789]" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="17" pageNumber="468">Figs. 59–64</figureCitation>
)
</paragraph>
<paragraph id="8F6336EBB85F643AFF6CDC52E804F88F" blockId="17.[151,1436,1834,2040]" box="[151,350,1834,1860]" pageId="17" pageNumber="468">
<emphasis id="BDA8EAF9B85F643AFF6CDC52E9B4F88F" bold="true" box="[151,238,1834,1860]" pageId="17" pageNumber="468">Female</emphasis>
(
<figureCitation id="17E72A6EB85F643AFF05DC52E808F88F" box="[254,338,1834,1860]" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="17" pageNumber="468">Fig. 62</figureCitation>
).
</paragraph>
<paragraph id="8F6336EBB85F643AFF3CDC36E8B2F8A3" blockId="17.[151,1436,1834,2040]" box="[199,488,1870,1896]" pageId="17" pageNumber="468">
<emphasis id="BDA8EAF9B85F643AFF3CDC36E83AF8A3" bold="true" box="[199,352,1870,1896]" pageId="17" pageNumber="468">Body length:</emphasis>
<quantity id="48249B0EB85F643AFE9CDC36E8B9F8A3" box="[359,483,1870,1896]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="17" pageNumber="468" unit="mm" value="12.5" valueMax="13.0" valueMin="12.0">12–13 mm</quantity>
.
</paragraph>
<paragraph id="8F6336EBB85F643AFF3CDC09EC2BF833" blockId="17.[151,1436,1834,2040]" pageId="17" pageNumber="468">
<emphasis id="BDA8EAF9B85F643AFF3CDC09E87AF840" bold="true" box="[199,288,1905,1931]" pageId="17" pageNumber="468">Colour.</emphasis>
Body predominantly dark; terga 1–4 with apical margins and ventrolateral segments broadly lightened hyaline-orange, tergum 5 entirely orange (
<figureCitation id="17E72A6EB85F643AFD8EDCEEEA70F87B" box="[629,810,1942,1968]" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="17" pageNumber="468">Figs. 60, 62, 63</figureCitation>
). Legs with apical tarsal segments of fore and mid legs and hind tibiae and tarsi lightened bright orange (
<figureCitation id="17E72A6EB85F643AFD45DCC2EA1CF81F" box="[702,838,1978,2004]" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="17" pageNumber="468">Figs. 62, 63</figureCitation>
). Flagellomeres 4–12 ventrally lightened by presence of grey-brown scales (
<figureCitation id="17E72A6EB85F643AFE6ADCA6EB41F833" box="[401,539,2014,2040]" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="17" pageNumber="468">Figs. 61, 62</figureCitation>
). Wings hyaline to weakly infuscate, veins and stigma dark brown (
<figureCitation id="17E72A6EB85F643AFAEBDCA6EC3FF833" box="[1296,1381,2014,2040]" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="17" pageNumber="468">Fig. 62</figureCitation>
).
</paragraph>
<caption id="DBA36663B85C6439FF6CDED0EBFEFA2F" ID-DOI="http://doi.org/10.5281/zenodo.12712088" ID-Zenodo-Dep="12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="18" pageNumber="469" startId="18.[151,264,1448,1472]" targetBox="[165,1422,181,1424]" targetPageId="18" targetType="figure">
<paragraph id="8F6336EBB85C6439FF6CDED0EBFEFA2F" blockId="18.[151,1436,1448,1508]" pageId="18" pageNumber="469">
<emphasis id="BDA8EAF9B85C6439FF6CDED0E80CFA0B" bold="true" box="[151,342,1448,1472]" pageId="18" pageNumber="469">FIGURES 59–64.</emphasis>
<taxonomicName id="48DC4D68B85C6439FEA5DED0EBF0FA0B" authorityName="Pisanty &amp; Wood" authorityYear="2024" box="[350,682,1448,1472]" class="Insecta" family="Andrenidae" genus="Andrena" isHigherUncertain="true" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="remota" status="sp. nov.">
<emphasis id="BDA8EAF9B85C6439FEA5DED0E8EEFA0B" box="[350,436,1448,1472]" italics="true" pageId="18" pageNumber="469">Andrena</emphasis>
(
<emphasis id="BDA8EAF9B85C6439FE3EDED1EB09FA0B" box="[453,595,1448,1472]" italics="true" pageId="18" pageNumber="469">incertae sedis</emphasis>
)
<emphasis id="BDA8EAF9B85C6439FD98DED1EBF0FA0B" box="[611,682,1449,1472]" italics="true" pageId="18" pageNumber="469">remota</emphasis>
</taxonomicName>
<emphasis id="BDA8EAF9B85C6439FD49DED0EA5FFA0B" bold="true" box="[690,773,1448,1472]" pageId="18" pageNumber="469">
<taxonomicNameLabel id="A69B5782B85C6439FD49DED0EA5FFA0B" box="[690,773,1448,1472]" pageId="18" pageNumber="469" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle.
</paragraph>
</caption>
<paragraph id="8F6336EBB85C6439FF3CDD54E8BAF83E" blockId="18.[151,1437,1579,2037]" pageId="18" pageNumber="469">
<emphasis id="BDA8EAF9B85C6439FF3CDD54E80DF98D" bold="true" box="[199,343,1580,1606]" pageId="18" pageNumber="469">Pubescence.</emphasis>
Head with moderately long minutely plumose hairs, hairs pale on gena, clypeus, and around antennal insertions, becoming dark brown on paraocular areas, frons, and vertex (
<figureCitation id="17E72A6EB85C6439FBB5DD37EC56F9A2" box="[1102,1292,1615,1642]" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="18" pageNumber="469">Figs. 59, 61, 62</figureCitation>
). Mesosoma with long minutely plumose hairs, hairs of mesepisternum, propodeum, metanotum, posterior margin of scutellum, and anterior margin of scutum white, remaining hairs of scutum and scutellum black (
<figureCitation id="17E72A6EB85C6439FBA4DDEFEC4EF97A" box="[1119,1300,1687,1714]" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="18" pageNumber="469">Figs. 59, 62, 64</figureCitation>
). Metasoma with scattered short golden hairs on tergal discs, sterna with pale to golden plumose hairs forming long fringes on marginal areas. Terminal fringe orange to orange-brown (
<figureCitation id="17E72A6EB85C6439FCEDDDA7EA9CF932" box="[790,966,1759,1785]" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="18" pageNumber="469">Figs. 60, 62, 63</figureCitation>
). Facial foveae with very dense short dark brown hair (
<figureCitation id="17E72A6EB85C6439FEDCDC7BE8EDF8D6" box="[295,439,1795,1822]" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="18" pageNumber="469">Figs. 59, 61</figureCitation>
). Propodeal corbicula incomplete, dorsoposterior fringe composed of moderately long finely plumose white hairs, corbicula surface scattered short whitish hairs (
<figureCitation id="17E72A6EB85C6439FC23DC5FED71F88A" box="[984,1067,1831,1857]" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="18" pageNumber="469">Fig. 62</figureCitation>
). Legs with variable pubescence; fore and mid femora with whitish to light brownish hairs, outer surface of fore and mid tibiae with short dark brown pubescence, fore and mid tarsi with dense fringes of bristles, anteriorly orange, posteriorly dark brown (
<figureCitation id="17E72A6EB85C6439FAC3DC17E9ECF866" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="18" pageNumber="469">Figs. 62, 63</figureCitation>
). Flocculus poorly developed, incomplete, pure white. Femoral and tibial scopae composed of simple, relatively thick sparse hairs, femoral scopae whitish, tibial scopae becoming more golden, dorsally with dark brown hairs as basitibial plate (
<figureCitation id="17E72A6EB85C6439FEB0DCA3E889F83E" box="[331,467,2011,2037]" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="18" pageNumber="469">Figs. 62, 63</figureCitation>
).
</paragraph>
<paragraph id="8F6336EBB85D6438FF3CDBEEEB37FD07" blockId="19.[151,1437,150,2013]" pageId="19" pageNumber="470">
<emphasis id="BDA8EAF9B85D6438FF3CDBEEE85DFF7B" bold="true" box="[199,263,150,176]" pageId="19" pageNumber="470">Head</emphasis>
(
<figureCitation id="17E72A6EB85D6438FEE3DBEEE88AFF7B" box="[280,464,150,177]" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="19" pageNumber="470">Figs. 59, 61, 62</figureCitation>
). 1.1 times broader than long (
<figureCitation id="17E72A6EB85D6438FCD4DBEEEADEFF7B" box="[815,900,150,176]" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="19" pageNumber="470">Fig. 61</figureCitation>
). Mandible bidentate. Galea finely shagreened, moderately shining. Labral process rounded rectangular, 3.5 times wider than long, surface with weak latitudinal wrinkles, weakly shining, anterior margin broadly but shallowly emarginate. Clypeus domed, medially slightly flattened, this appearing slightly 3-faced; underlying surface polished and shining, with only weak shagreen laterally; punctation deep but irregular, punctures separated by 0.5–2 puncture diameters, punctures of variable size, forming slightly raised impunctate longitudinal mid-line. Lower half of paraocular areas shagreened and very densely punctate, weakly shining (
<figureCitation id="17E72A6EB85D6438FDD3DA16EB25FE43" box="[552,639,366,392]" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="19" pageNumber="470">Fig. 61</figureCitation>
). Malar area linear (
<figureCitation id="17E72A6EB85D6438FC8BDA16ED5BFE43" box="[880,1025,366,392]" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="19" pageNumber="470">Figs. 61, 62</figureCitation>
). Supraclypeal area weakly domed, shagreened and weakly shining. Facial foveae moderately long, dorsally occupying ½ space between lateral ocellus and compound eye, separated from lateral ocellus by 1.5 times its diameter; foveae deeply impressed, slightly broadening medially before narrowing ventrally, ventrally extending only to lower margin of antennal insertions, at broadest clearly exceeding width of flagellum; separated from compound eye by narrow shiny cuticular strip (
<figureCitation id="17E72A6EB85D6438FA9FDA86E9B8FDF7" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="19" pageNumber="470">Figs. 59, 61</figureCitation>
). Flagellomere 1 equaling length of 2+3+4. Frons shagreened, weakly shining, densely but finely punctate (
<figureCitation id="17E72A6EB85D6438FF64D93EE9AEFDAB" box="[159,244,582,608]" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="19" pageNumber="470">Fig. 61</figureCitation>
). Vertex immediately behind lateral ocelli very weakly shagreened, almost shining, otherwise shagreened and weakly shining; weakly sculptured areas posterolateral to the lateral ocelli impunctate, forming shining strips. Ocelloccipital distance 1.5 times diameter of lateral ocellus (
<figureCitation id="17E72A6EB85D6438FCA0D9F6EAB1FD63" box="[859,1003,654,681]" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="19" pageNumber="470">Figs. 59, 61</figureCitation>
). Genal area subequal to diameter of compound eye, posterior margin rounded.
</paragraph>
<paragraph id="8F6336EBB85D6438FF3CD9AEEDB6FC27" blockId="19.[151,1437,150,2013]" pageId="19" pageNumber="470">
<emphasis id="BDA8EAF9B85D6438FF3CD9AEE81FFD3B" bold="true" box="[199,325,726,752]" pageId="19" pageNumber="470">Mesosoma</emphasis>
(
<figureCitation id="17E72A6EB85D6438FEAFD9AEE884FD3B" box="[340,478,726,753]" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="19" pageNumber="470">Figs. 59, 64</figureCitation>
). Dorsolateral angle of pronotum weakly but distinctly elevated, lateral carina absent. Scutum and scutellum weakly shagreened, more or less shining, deeply and densely punctate, punctures separated by &lt;0.5–1 puncture diameter (
<figureCitation id="17E72A6EB85D6438FE07D866EB0EFCF2" box="[508,596,798,825]" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="19" pageNumber="470">Fig. 59</figureCitation>
). Mesepisternum shagreened, surface with shallow punctures, punctures separated by 1 puncture diameter. Propodeum finely shagreened, dorsolateral parts shallowly punctate, propodeal triangle delineated by finely raised rim; entire surface weakly shining (
<figureCitation id="17E72A6EB85D6438FC5BD81EEAA9FC4B" box="[928,1011,870,896]" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="19" pageNumber="470">Fig. 64</figureCitation>
). Inner side of hind femur not carinate. Inner hind tibial spur not broadened basally, straight, slightly curved near apex. Hind pretarsal claw with strong inner tooth. Stigma slender. Submarginal cross vein 1 meets marginal cell 8 vein widths from stigma. Recurrent vein 1 meets second submarginal cell slightly beyond its middle (
<figureCitation id="17E72A6EB85D6438FCC6D8AAEACBFC27" box="[829,913,978,1004]" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="19" pageNumber="470">Fig. 62</figureCitation>
). Nervulus strongly postfurcal.
</paragraph>
<paragraph id="8F6336EBB85D6438FF3CD88EEB22FBB7" blockId="19.[151,1437,150,2013]" pageId="19" pageNumber="470">
<emphasis id="BDA8EAF9B85D6438FF3CD88EE819FBDB" bold="true" box="[199,323,1014,1040]" pageId="19" pageNumber="470">Metasoma</emphasis>
(
<figureCitation id="17E72A6EB85D6438FEAFD88EEB53FBDB" box="[340,521,1014,1040]" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="19" pageNumber="470">Figs. 60, 62, 63</figureCitation>
). Tergal discs finely shagreened, weakly shining, surface with scattered irregular punctures, punctures separated by 1–3 puncture diameters. Marginal areas not depressed, sculpture not differentiated from tergal discs (
<figureCitation id="17E72A6EB85D6438FE9EDF46E8AAFB93" box="[357,496,1086,1112]" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="19" pageNumber="470">Figs. 60, 63</figureCitation>
). Pygidial plate with strongly raised longitudinal ridge medially, remaining surface otherwise featureless, shagreened and dull.
</paragraph>
<paragraph id="8F6336EBB85D6438FF3CDFFEE8D2FB6B" blockId="19.[151,1437,150,2013]" box="[199,392,1158,1184]" pageId="19" pageNumber="470">
<emphasis id="BDA8EAF9B85D6438FF3CDFFEE851FB6B" bold="true" box="[199,267,1158,1184]" pageId="19" pageNumber="470">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C7C66560B85D643FFF3CDFD2EBD7FE8A" lastPageId="20" lastPageNumber="471" pageId="19" pageNumber="470" type="diagnosis">
<paragraph id="8F6336EBB85D6438FF3CDFD2EA1BFAB3" blockId="19.[151,1437,150,2013]" pageId="19" pageNumber="470">
<emphasis id="BDA8EAF9B85D6438FF3CDFD2E818FB0F" bold="true" box="[199,322,1194,1220]" pageId="19" pageNumber="470">Diagnosis.</emphasis>
<taxonomicName id="48DC4D68B85D6438FEB1DFD2EB5AFB0F" authorityName="Pisanty &amp; Wood" authorityYear="2024" box="[330,512,1194,1220]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="remota">
<emphasis id="BDA8EAF9B85D6438FEB1DFD2EB5AFB0F" box="[330,512,1194,1220]" italics="true" pageId="19" pageNumber="470">Andrena remota</emphasis>
</taxonomicName>
is surprisingly similar to members of the subgenus
<emphasis id="BDA8EAF9B85D6438FBA8DFD3ED3EFB0E" box="[1107,1124,1195,1221]" italics="true" pageId="19" pageNumber="470">A</emphasis>
. (
<taxonomicName id="48DC4D68B85D6438FB86DFD2EC74FB0F" box="[1149,1326,1194,1220]" pageId="19" pageNumber="471" rank="subGenus" subGenus="Warnckandrena">
<emphasis id="BDA8EAF9B85D6438FB86DFD2EC74FB0F" box="[1149,1326,1194,1220]" italics="true" pageId="19" pageNumber="470">Warnckandrena</emphasis>
</taxonomicName>
), in large part due to the largely finely shagreened propodeum which lacks a clear horizontal basal area, the compound eyes which slightly converge dorsally, the deeply impressed facial foveae which are separated from the compound eyes by a narrow slightly elevated shining strip, the broad process of the labrum, the pronotum with a weak angle, flagellomere 1 equaling the length of flagellomeres 2+3+4, the slender stigma, the moderately large body size (for an
<taxonomicName id="48DC4D68B85D6438FF42DE26E842FAB3" authorityName="Confalonierii Guiglia" authorityYear="1929" box="[185,280,1374,1400]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="genus">
<emphasis id="BDA8EAF9B85D6438FF42DE26E842FAB3" box="[185,280,1374,1400]" italics="true" pageId="19" pageNumber="470">Andrena</emphasis>
</taxonomicName>
), and the lack of any other distinctive characters.
</paragraph>
<paragraph id="8F6336EBB85D6438FF3CDEFBED9BF8CF" blockId="19.[151,1437,150,2013]" pageId="19" pageNumber="470">
However, unlike
<taxonomicName id="48DC4D68B85D6438FE6BDEFAEB1BFA57" box="[400,577,1410,1436]" pageId="19" pageNumber="471" rank="subGenus" subGenus="Warnckandrena">
<emphasis id="BDA8EAF9B85D6438FE6BDEFAEB1BFA57" box="[400,577,1410,1436]" italics="true" pageId="19" pageNumber="470">Warnckandrena</emphasis>
</taxonomicName>
,
<taxonomicName id="48DC4D68B85D6438FDA8DEFAEA51FA57" authorityName="Pisanty &amp; Wood" authorityYear="2024" box="[595,779,1410,1436]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="remota">
<emphasis id="BDA8EAF9B85D6438FDA8DEFAEA51FA57" box="[595,779,1410,1436]" italics="true" pageId="19" pageNumber="470">Andrena remota</emphasis>
</taxonomicName>
has the scutum and scutellum with shallow but distinct and dense punctation that clearly contrasts the underlying surface which is shagreened and shining (
<figureCitation id="17E72A6EB85D6438FAC4DEDEECCDFA0A" box="[1343,1431,1446,1473]" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="19" pageNumber="470">Fig. 59</figureCitation>
;
<taxonomicName id="48DC4D68B85D6438FF6CDEB2E812FA2F" box="[151,328,1482,1508]" pageId="19" pageNumber="471" rank="subGenus" subGenus="Warnckandrena">
<emphasis id="BDA8EAF9B85D6438FF6CDEB2E812FA2F" box="[151,328,1482,1508]" italics="true" pageId="19" pageNumber="470">Warnckandrena</emphasis>
</taxonomicName>
has the scutum and scutellum largely shagreened and dull with superficial punctation that disappears into the background sculpture,
<figureCitation id="17E72A6EB85D6438FDFDDE96EB15F9C3" box="[518,591,1518,1544]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1944,1968]" captionTargetBox="[253,1335,179,1918]" captionTargetId="figure-17@5.[253,1335,179,1918]" captionTargetPageId="5" captionText="FIGURES 1–9. Andrena (Warnckandrena) curiosa (Morawitz), female: 1, flagellum; 2, vertex and mesosoma; 3, metasoma; 4, profile of propodeum; 5, propodeal triangle; 6, flocculus and scopa; 7, forewing venation; 8, habitus; 9, head." figureDoi="http://doi.org/10.5281/zenodo.12712068" httpUri="https://zenodo.org/record/12712068/files/figure.png" pageId="19" pageNumber="470">Fig. 2</figureCitation>
), the mesepisternum has shallow but distinct punctures, surface weakly shining (
<taxonomicName id="48DC4D68B85D6438FF01DD6AE8F1F9E7" box="[250,427,1554,1580]" pageId="19" pageNumber="471" rank="subGenus" subGenus="Warnckandrena">
<emphasis id="BDA8EAF9B85D6438FF01DD6AE8F1F9E7" box="[250,427,1554,1580]" italics="true" pageId="19" pageNumber="470">Warnckandrena</emphasis>
</taxonomicName>
has the mesepisternum shagreened and almost dull, with obscure punctures), the nervulus is strongly postfurcal (interstitial to antefurcal in
<taxonomicName id="48DC4D68B85D6438FD3EDD4EEA2CF99B" box="[709,886,1590,1616]" pageId="19" pageNumber="471" rank="subGenus" subGenus="Warnckandrena">
<emphasis id="BDA8EAF9B85D6438FD3EDD4EEA2CF99B" box="[709,886,1590,1616]" italics="true" pageId="19" pageNumber="470">Warnckandrena</emphasis>
</taxonomicName>
), the tergal punctation is irregular and scattered, punctures separated by 1–3 puncture diameters (
<figureCitation id="17E72A6EB85D6438FD4EDD22EA52F9BF" box="[693,776,1626,1652]" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="19" pageNumber="470">Fig. 60</figureCitation>
;
<taxonomicName id="48DC4D68B85D6438FCEFDD22EA9FF9BF" box="[788,965,1626,1652]" pageId="19" pageNumber="471" rank="subGenus" subGenus="Warnckandrena">
<emphasis id="BDA8EAF9B85D6438FCEFDD22EA9FF9BF" box="[788,965,1626,1652]" italics="true" pageId="19" pageNumber="470">Warnckandrena</emphasis>
</taxonomicName>
has the tergal punctation fine and regular, typically separated by around 1 puncture diameter,
<figureCitation id="17E72A6EB85D6438FD2DDD06EA43F953" box="[726,793,1662,1688]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1944,1968]" captionTargetBox="[253,1335,179,1918]" captionTargetId="figure-17@5.[253,1335,179,1918]" captionTargetPageId="5" captionText="FIGURES 1–9. Andrena (Warnckandrena) curiosa (Morawitz), female: 1, flagellum; 2, vertex and mesosoma; 3, metasoma; 4, profile of propodeum; 5, propodeal triangle; 6, flocculus and scopa; 7, forewing venation; 8, habitus; 9, head." figureDoi="http://doi.org/10.5281/zenodo.12712068" httpUri="https://zenodo.org/record/12712068/files/figure.png" pageId="19" pageNumber="470">Fig. 3</figureCitation>
), the pygidial plate with clear elevated longitudinal ridge medially (
<taxonomicName id="48DC4D68B85D6438FEF2DDDAE8E0F977" box="[265,442,1698,1724]" pageId="19" pageNumber="471" rank="subGenus" subGenus="Warnckandrena">
<emphasis id="BDA8EAF9B85D6438FEF2DDDAE8E0F977" box="[265,442,1698,1724]" italics="true" pageId="19" pageNumber="470">Warnckandrena</emphasis>
</taxonomicName>
has the pygidial plate flat over its entire surface), and the marginal areas and lateral parts of the terga are broadly lightened hyaline-orange (
<figureCitation id="17E72A6EB85D6438FD23DDBEEACBF92B" box="[728,913,1734,1760]" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="19" pageNumber="470">Figs. 60, 62, 63</figureCitation>
;
<taxonomicName id="48DC4D68B85D6438FC64DDBEED0AF92B" box="[927,1104,1734,1760]" pageId="19" pageNumber="471" rank="subGenus" subGenus="Warnckandrena">
<emphasis id="BDA8EAF9B85D6438FC64DDBEED0AF92B" box="[927,1104,1734,1760]" italics="true" pageId="19" pageNumber="470">Warnckandrena</emphasis>
</taxonomicName>
has the terga almost entirely dark, at most with the apical rims narrowly lightened hyaline-brown,
<figureCitation id="17E72A6EB85D6438FC5ADD92ED50F8CF" box="[929,1034,1770,1796]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1944,1968]" captionTargetBox="[253,1335,179,1918]" captionTargetId="figure-17@5.[253,1335,179,1918]" captionTargetPageId="5" captionText="FIGURES 1–9. Andrena (Warnckandrena) curiosa (Morawitz), female: 1, flagellum; 2, vertex and mesosoma; 3, metasoma; 4, profile of propodeum; 5, propodeal triangle; 6, flocculus and scopa; 7, forewing venation; 8, habitus; 9, head." figureDoi="http://doi.org/10.5281/zenodo.12712068" httpUri="https://zenodo.org/record/12712068/files/figure.png" pageId="19" pageNumber="470">Figs. 3, 8</figureCitation>
,
<figureCitation id="17E72A6EB85D6438FBECDD92EDEEF8CF" box="[1047,1204,1770,1796]" captionStart="FIGURES 10–17" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[263,1335,181,1920]" captionTargetId="figure-33@6.[252,1335,181,1920]" captionTargetPageId="6" captionText="FIGURES 10–17. Andrena (Warnckandrena) female habitus (10, 12, 14, 16) and heads (11, 13, 15, 17): 10–11, A. innesi Gribodo; 12–13, A. papio sp. nov.; 14–15, A. ramlehiana Pérez; 16–17, A. otoolei sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12712072" httpUri="https://zenodo.org/record/12712072/files/figure.png" pageId="19" pageNumber="470">10, 12, 14, 16</figureCitation>
).
</paragraph>
<paragraph id="8F6336EBB85D643FFF3CDC76EBD7FE8A" blockId="19.[151,1437,150,2013]" lastBlockId="20.[151,1437,151,1005]" lastPageId="20" lastPageNumber="471" pageId="19" pageNumber="470">
Separation from
<taxonomicName id="48DC4D68B85D6438FE7CDC76EB25F8E3" authorityName="melittoides Friese" authorityYear="1899" box="[391,639,1806,1832]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="subGenus" sensu="stricto" subGenus="Melittoides">
<emphasis id="BDA8EAF9B85D6438FE7CDC76E8BCF8E3" box="[391,486,1806,1832]" italics="true" pageId="19" pageNumber="470">Andrena</emphasis>
(
<emphasis id="BDA8EAF9B85D6438FE03DC76EB2CF8E3" box="[504,630,1806,1832]" italics="true" pageId="19" pageNumber="470">Melittoides</emphasis>
)
</taxonomicName>
<taxonomicNameLabel id="A69B5782B85D6438FD73DC77EA4FF8E3" box="[648,789,1806,1833]" pageId="19" pageNumber="470" sensu="stricto">sensu stricto</taxonomicNameLabel>
can be made by some of the same characters;
<taxonomicName id="48DC4D68B85D6438FAD6DC77ECC6F8E3" authorityName="Pisanty &amp; Wood" authorityYear="2024" box="[1325,1436,1807,1833]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="remota">
<emphasis id="BDA8EAF9B85D6438FAD6DC77ECC6F8E3" box="[1325,1436,1807,1833]" italics="true" pageId="19" pageNumber="470">A. remota</emphasis>
</taxonomicName>
has the scutum and scutellum with shallow but distinct and dense punctation that clearly contrasts the underlying surface which is shagreened and shining (
<figureCitation id="17E72A6EB85D6438FD8FDC2EEB93F8BA" box="[628,713,1878,1905]" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="19" pageNumber="470">Fig. 59</figureCitation>
;
<taxonomicName id="48DC4D68B85D6438FD2CDC2FEA29F8BB" authorityName="melittoides Friese" authorityYear="1899" box="[727,883,1878,1905]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="melittoides">
<emphasis id="BDA8EAF9B85D6438FD2CDC2FEA29F8BB" box="[727,883,1878,1905]" italics="true" pageId="19" pageNumber="470">A. melittoides</emphasis>
</taxonomicName>
has the scutum and scutellum finely shagreened and shining with extremely shallow and obscure punctures), by the longer ocelloccipital distance that is 1.5 times the diameter of a lateral ocellus (
<taxonomicName id="48DC4D68B85D6438FDFDDCE7EBC5F873" authorityName="melittoides Friese" authorityYear="1899" box="[518,671,1950,1977]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="melittoides">
<emphasis id="BDA8EAF9B85D6438FDFDDCE7EBC5F873" box="[518,671,1950,1977]" italics="true" pageId="19" pageNumber="470">A. melittoides</emphasis>
</taxonomicName>
has the ocelloccipital distance slightly but distinctly shorter than the diameter of a lateral ocellus), by the pygidial plate with clear elevated longitudinal ridge medially (
<taxonomicName id="48DC4D68B85D6438FAFADCBBECC6F817" authorityName="melittoides Friese" authorityYear="1899" box="[1281,1436,1986,2013]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="melittoides">
<emphasis id="BDA8EAF9B85D6438FAFADCBBECC6F817" box="[1281,1436,1986,2013]" italics="true" pageId="19" pageNumber="470">A. melittoides</emphasis>
</taxonomicName>
has the pygidial plate flat over its entire surface), by the colouration of the terga which in dorsal view have reddish-yellow markings restricted to the tergal margins, including the margins of T3–4 which are clearly lightened hyaline-orange (
<figureCitation id="17E72A6EB85A643FFF0EDBA7E8F5FF32" box="[245,431,223,249]" captionStart="FIGURES 59–64" captionStartId="18.[151,264,1448,1472]" captionTargetBox="[165,1422,181,1424]" captionTargetId="figure-33@18.[165,1422,181,1424]" captionTargetPageId="18" captionText="FIGURES 59–64. Andrena (incertae sedis) remota sp. nov., female holotype: 59, vertex and mesosoma; 60, metasoma; 61, head; 62, habitus; 63, scopa; 64, propodeal triangle." figureDoi="http://doi.org/10.5281/zenodo.12712088" httpUri="https://zenodo.org/record/12712088/files/figure.png" pageId="20" pageNumber="471">Figs. 60, 62, 63</figureCitation>
;
<taxonomicName id="48DC4D68B85A643FFE47DBA7EB03FF32" authorityName="melittoides Friese" authorityYear="1899" box="[444,601,223,249]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="melittoides">
<emphasis id="BDA8EAF9B85A643FFE47DBA7EB03FF32" box="[444,601,223,249]" italics="true" pageId="20" pageNumber="471">A. melittoides</emphasis>
</taxonomicName>
has abundant reddish-yellow markings on the discs of T1–2, with tergal margins of T3–4 only faintly lightened hyaline-whitish,
<figureCitation id="17E72A6EB85A643FFCE1DA7BEA34FED6" box="[794,878,258,285]" captionStart="FIGURES 65–73" captionStartId="22.[151,264,1903,1927]" captionTargetBox="[165,1421,181,1878]" captionTargetId="figure-33@22.[165,1421,181,1878]" captionTargetPageId="22" captionText="FIGURES 65–73. Andrena (Melittoides) melittoides Friese, female (65–68) and male (69–73): 65, 69, habitus; 66, 70, head; 67, flocculus and scopa; 68, forewing venation; 71, genitalia; 72, eighth sternum; 73, seventh sternum." figureDoi="http://doi.org/10.5281/zenodo.12712090" httpUri="https://zenodo.org/record/12712090/files/figure.png" pageId="20" pageNumber="471">Fig. 65</figureCitation>
), and by the smaller body size of
<quantity id="48249B0EB85A643FFB0CDA7BEC28FED6" box="[1271,1394,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="20" pageNumber="471" unit="mm" value="12.5" valueMax="13.0" valueMin="12.0">12–13 mm</quantity>
(
<taxonomicName id="48DC4D68B85A643FFA7FDA7BE848FE8A" authorityName="melittoides Friese" authorityYear="1899" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="melittoides">
<emphasis id="BDA8EAF9B85A643FFA7FDA7BE848FE8A" italics="true" pageId="20" pageNumber="471">A. melittoides</emphasis>
</taxonomicName>
has a body length of
<quantity id="48249B0EB85A643FFDF8DA5FEBDAFE8A" box="[515,640,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="20" pageNumber="471" unit="mm" value="16.5" valueMax="17.0" valueMin="16.0">16–17 mm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="C7C66560B85A643FFF3CDA32E8F5FE42" pageId="20" pageNumber="471" type="distribution">
<paragraph id="8F6336EBB85A643FFF3CDA32EB35FEAE" blockId="20.[151,1437,151,1005]" box="[199,623,330,357]" pageId="20" pageNumber="471">
<emphasis id="BDA8EAF9B85A643FFF3CDA32E839FEAF" bold="true" box="[199,355,330,356]" pageId="20" pageNumber="471">Distribution:</emphasis>
Central
<collectingCountry id="F7CB767BB85A643FFE38DA33EB5CFEAE" box="[451,518,331,357]" name="China" pageId="20" pageNumber="471">China</collectingCountry>
(
<collectingRegion id="4D18F809B85A643FFDEEDA33EB39FEAE" box="[533,611,331,357]" country="China" name="Gansu" pageId="20" pageNumber="471">Gansu</collectingRegion>
).
</paragraph>
<paragraph id="8F6336EBB85A643FFF3CDA16E8F5FE42" blockId="20.[151,1437,151,1005]" box="[199,431,366,393]" pageId="20" pageNumber="471">
<emphasis id="BDA8EAF9B85A643FFF3CDA16E837FE43" bold="true" box="[199,365,366,392]" pageId="20" pageNumber="471">Flight period:</emphasis>
June.
</paragraph>
</subSubSection>
<subSubSection id="C7C66560B85A643FFF3CDAEBED90FE3E" pageId="20" pageNumber="471" type="description">
<paragraph id="8F6336EBB85A643FFF3CDAEBE88AFE66" blockId="20.[151,1437,151,1005]" box="[199,464,403,429]" pageId="20" pageNumber="471">
<emphasis id="BDA8EAF9B85A643FFF3CDAEBE8DFFE66" bold="true" box="[199,389,403,429]" pageId="20" pageNumber="471">Flower records:</emphasis>
None.
</paragraph>
<paragraph id="8F6336EBB85A643FFF3CDACFEB73FE1A" blockId="20.[151,1437,151,1005]" box="[199,553,438,465]" pageId="20" pageNumber="471">
<emphasis id="BDA8EAF9B85A643FFF3CDACFE8F6FE1B" bold="true" box="[199,428,438,465]" pageId="20" pageNumber="471">Pollen preferences:</emphasis>
Unknown.
</paragraph>
<paragraph id="8F6336EBB85A643FFF3CDAA2ED90FE3E" blockId="20.[151,1437,151,1005]" box="[199,1226,474,501]" pageId="20" pageNumber="471">
<emphasis id="BDA8EAF9B85A643FFF3CDAA2EB46FE3F" bold="true" box="[199,540,474,501]" pageId="20" pageNumber="471">Genetic sequences available:</emphasis>
UCE data will be made public following future publications.
</paragraph>
</subSubSection>
<subSubSection id="C7C66560B85A643FFF3CDA86EC17FDF6" pageId="20" pageNumber="471" type="materials_examined">
<paragraph id="8F6336EBB85A643FFF3CDA86EC17FDF6" blockId="20.[151,1437,151,1005]" pageId="20" pageNumber="471">
<materialsCitation id="3FB43CB6B85A643FFF3CDA86EA10FDF6" ID-GBIF-Occurrence="4904901348" collectingDate="1998-06-01" collectingDateMax="1998-06-15" collectingDateMin="1998-06-01" collectionCode="OLML" country="China" elevation="3500" latitude="35.1984" location="Xiahe Labrang" longLatPrecision="6" longitude="102.5098" pageId="20" pageNumber="471" specimenCount="1" specimenCount-female="1" stateProvince="Gansu" typeStatus="holotype">
<emphasis id="BDA8EAF9B85A643FFF3CDA86E888FDD3" bold="true" box="[199,466,510,536]" pageId="20" pageNumber="471">
<typeStatus id="50678849B85A643FFF3CDA86E830FDD3" box="[199,362,510,536]" pageId="20" pageNumber="471" type="holotype">HOLOTYPE</typeStatus>
:
<collectingCountry id="F7CB767BB85A643FFE8EDA86E888FDD3" box="[373,466,510,536]" name="China" pageId="20" pageNumber="471">CHINA</collectingCountry>
</emphasis>
•
<specimenCount id="99DAFD62B85A643FFE1CDA87E8A7FDD2" box="[487,509,511,537]" count="1" pageId="20" pageNumber="471" type="female">♀</specimenCount>
;
<collectingRegion id="4D18F809B85A643FFDF1DA87EB08FDD2" box="[522,594,511,537]" country="China" name="Gansu" pageId="20" pageNumber="471">Kansu</collectingRegion>
mer. [
<collectingRegion id="4D18F809B85A643FFD6EDA87EBBEFDD2" box="[661,740,511,537]" country="China" name="Gansu" pageId="20" pageNumber="471">Gansu</collectingRegion>
],
<location id="8A036030B85A643FFD0EDA87EAC0FDD2" LSID="urn:lsid:plazi:treatment:077587FDB85F643FFF6CDDC5EC53FC26:8A036030B85A643FFD0EDA87EAC0FDD2" box="[757,922,511,537]" country="China" latitude="35.1984" longLatPrecision="6" longitude="102.5098" name="Xiahe Labrang" pageId="20" pageNumber="471" stateProvince="Gansu">Xiahe Labrang</location>
[Labrang Monastery];
<geoCoordinate id="EAE8502CB85A643FFB60DA86EC44FDD2" box="[1179,1310,510,537]" degrees="35.1984" direction="north" orientation="latitude" pageId="20" pageNumber="471" precision="5" value="35.1984">35.1984° N</geoCoordinate>
,
<geoCoordinate id="EAE8502CB85A643FFAD3DA86E9F1FDF6" degrees="102.5098" direction="east" orientation="longitude" pageId="20" pageNumber="471" precision="5" value="102.5098">102.5098° E</geoCoordinate>
;
<elevation id="04F1D1D8B85A643FFF4CD95BE8D5FDF6" box="[183,399,547,573]" metricMagnitude="3" metricUnit="m" metricValue="3.5" metricValueMax="3.7" metricValueMin="3.3" pageId="20" pageNumber="471" unit="m" value="3500.0" valueMax="3700.0" valueMin="3300.0">
<quantity id="48249B0EB85A643FFF4CD95BE808FDF6" box="[183,338,547,573]" metricMagnitude="3" metricUnit="m" metricValue="3.5" metricValueMax="3.7" metricValueMin="3.3" pageId="20" pageNumber="471" unit="m" value="3500.0" valueMax="3700.0" valueMin="3300.0">3300–3700 m</quantity>
a.s.l.
</elevation>
;
<date id="FB62102BB85A643FFE67D95AEB11FDF6" box="[412,587,546,573]" pageId="20" pageNumber="471" value="1998-06-01" valueMax="1998-06-15" valueMin="1998-06-01">
<collectingDate id="EB26E9C3B85A643FFE67D95AEB11FDF6" box="[412,587,546,573]" pageId="20" pageNumber="471" value="1998-06-01" valueMax="1998-06-15" valueMin="1998-06-01">1–15 Jun. 1998</collectingDate>
</date>
;
<taxonomicName id="48DC4D68B85A643FFDADD95BEBE2FDF6" authorityName="Puton" authorityYear="1879" box="[598,696,547,573]" class="Magnoliopsida" family="Fabaceae" genus="Vicia" kingdom="Plantae" order="Fabales" pageId="20" pageNumber="471" phylum="Tracheophyta" rank="species" species="major">V. Major</taxonomicName>
leg.;
<collectionCode id="E9CDAE2EB85A643FFD03D95BEA10FDF6" box="[760,842,547,573]" country="Austria" httpUri="http://grbio.org/cool/pp0t-48s3" name="Oberösterreichisches Landesmuseum" pageId="20" pageNumber="471" type="Museum">OLML</collectionCode>
</materialsCitation>
.—
<materialsCitation id="3FB43CB6B85A643FFC91D95AEC13FDF6" ID-GBIF-Occurrence="4904901375" box="[874,1353,546,573]" collectingDate="1998-06-01" collectingDateMax="1998-06-15" collectingDateMin="1998-06-01" collectionCode="OLML" country="China" elevation="3500" latitude="35.1984" location="Xiahe Labrang" longLatPrecision="6" longitude="102.5098" pageId="20" pageNumber="471" specimenCount="1" specimenCount-female="1" stateProvince="Gansu" typeStatus="paratype">
<emphasis id="BDA8EAF9B85A643FFC91D95AED31FDF7" bold="true" box="[874,1131,546,572]" pageId="20" pageNumber="471">
<typeStatus id="50678849B85A643FFC91D95AED59FDF7" box="[874,1027,546,572]" pageId="20" pageNumber="471" type="paratype">PARATYPE</typeStatus>
:
<collectingCountry id="F7CB767BB85A643FFBF4D95AED31FDF7" box="[1039,1131,546,572]" name="China" pageId="20" pageNumber="471">CHINA</collectingCountry>
</emphasis>
•
<specimenCount id="99DAFD62B85A643FFB79D95BEDFDFDF6" box="[1154,1191,547,573]" count="1" pageId="20" pageNumber="471" type="female">1♀</specimenCount>
; ibid.;
<collectionCode id="E9CDAE2EB85A643FFB0DD95BEC13FDF6" box="[1270,1353,547,573]" country="Austria" httpUri="http://grbio.org/cool/pp0t-48s3" name="Oberösterreichisches Landesmuseum" pageId="20" pageNumber="471" type="Museum">OLML</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C7C66560B85A643FFF3CD93FEC48FD62" pageId="20" pageNumber="471" type="etymology">
<paragraph id="8F6336EBB85A643FFF3CD93FEC48FD62" blockId="20.[151,1437,151,1005]" pageId="20" pageNumber="471">
<emphasis id="BDA8EAF9B85A643FFF3CD93FE814FDAA" bold="true" box="[199,334,583,609]" pageId="20" pageNumber="471">Etymology.</emphasis>
Feminine singular nominative form of the Latin adjective
<emphasis id="BDA8EAF9B85A643FFC17D93FED1FFDAB" box="[1004,1093,583,608]" italics="true" pageId="20" pageNumber="471">remotus</emphasis>
meaning distant or remote, in reference to the strong separation between the true members of the subgenus
<taxonomicName id="48DC4D68B85A643FFC06D913EDF4FD4E" box="[1021,1198,619,645]" pageId="20" pageNumber="471" rank="subGenus" subGenus="Warnckandrena">
<emphasis id="BDA8EAF9B85A643FFC06D913EDF4FD4E" box="[1021,1198,619,645]" italics="true" pageId="20" pageNumber="471">Warnckandrena</emphasis>
</taxonomicName>
with an eastern limit in northern
<collectingCountry id="F7CB767BB85A643FFEE1D9F7E810FD62" box="[282,330,655,681]" name="Iran" pageId="20" pageNumber="471">Iran</collectingCountry>
, and
<taxonomicName id="48DC4D68B85A643FFE7FD9F7E8ABFD63" authorityName="Pisanty &amp; Wood" authorityYear="2024" box="[388,497,655,681]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="remota">
<emphasis id="BDA8EAF9B85A643FFE7FD9F7E8ABFD63" box="[388,497,655,681]" italics="true" pageId="20" pageNumber="471">A. remota</emphasis>
</taxonomicName>
which is currently known from
<quantity id="48249B0EB85A643FFCA0D9F7EA9FFD62" box="[859,965,655,681]" metricMagnitude="6" metricUnit="m" metricValue="4.0" pageId="20" pageNumber="471" unit="km" value="4000.0">4,000 km</quantity>
further east in Central
<collectingCountry id="F7CB767BB85A643FFB3CD9F7EC54FD62" box="[1223,1294,655,681]" name="China" pageId="20" pageNumber="471">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C7C66560B85A643FFF3CD9CBEC53FC26" pageId="20" pageNumber="471" type="discussion">
<paragraph id="8F6336EBB85A643FFF3CD9CBEC53FC26" blockId="20.[151,1437,151,1005]" pageId="20" pageNumber="471">
<emphasis id="BDA8EAF9B85A643FFF3CD9CBE861FD06" bold="true" box="[199,315,691,717]" pageId="20" pageNumber="471">Remarks.</emphasis>
In a preliminary molecular phylogenetic analysis of ultraconserved element (UCE)data,
<taxonomicName id="48DC4D68B85A643FFAF9D9CBEC36FD07" authorityName="Pisanty &amp; Wood" authorityYear="2024" box="[1282,1388,691,717]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="remota">
<emphasis id="BDA8EAF9B85A643FFAF9D9CBEC36FD07" box="[1282,1388,691,717]" italics="true" pageId="20" pageNumber="471">A.remota</emphasis>
</taxonomicName>
was placed within the most basal clade of
<taxonomicName id="48DC4D68B85A643FFDCCD9AFEBCCFD3A" authorityName="Confalonierii Guiglia" authorityYear="1929" box="[567,662,727,753]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="genus">
<emphasis id="BDA8EAF9B85A643FFDCCD9AFEBCCFD3A" box="[567,662,727,753]" italics="true" pageId="20" pageNumber="471">Andrena</emphasis>
</taxonomicName>
, as sister to
<taxonomicName id="48DC4D68B85A643FFCE4D9AFEACBFD3B" baseAuthorityName="Morawitz" baseAuthorityYear="1877" box="[799,913,727,753]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="curiosa">
<emphasis id="BDA8EAF9B85A643FFCE4D9AFEACBFD3B" box="[799,913,727,753]" italics="true" pageId="20" pageNumber="471">A. curiosa</emphasis>
</taxonomicName>
+
<taxonomicName id="48DC4D68B85A643FFC56D9AFED16FD3A" authorityName="Perez" authorityYear="1903" box="[941,1100,727,753]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="ramlehiana">
<emphasis id="BDA8EAF9B85A643FFC56D9AFED16FD3A" box="[941,1100,727,753]" italics="true" pageId="20" pageNumber="471">A. ramlehiana</emphasis>
</taxonomicName>
, i.e. the
<taxonomicName id="48DC4D68B85A643FFB51D9AFEC01FD3A" box="[1194,1371,727,753]" pageId="20" pageNumber="471" rank="subGenus" subGenus="Warnckandrena">
<emphasis id="BDA8EAF9B85A643FFB51D9AFEC01FD3A" box="[1194,1371,727,753]" italics="true" pageId="20" pageNumber="471">Warnckandrena</emphasis>
</taxonomicName>
clade (corresponding to clade 2 of
<taxonomicName id="48DC4D68B85A643FFE2CD983EB6CFCDE" authorityName="Confalonierii Guiglia" authorityYear="1929" box="[471,566,763,789]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="genus">
<emphasis id="BDA8EAF9B85A643FFE2CD983EB6CFCDE" box="[471,566,763,789]" italics="true" pageId="20" pageNumber="471">Andrena</emphasis>
</taxonomicName>
in
<bibRefCitation id="EB4D4B1AB85A643FFDA2D983EA76FCDE" author="Pisanty, G. &amp; Richter, R. &amp; Martin, T. &amp; Dettman, J. &amp; Cardinal, S." box="[601,812,763,789]" pageId="20" pageNumber="471" refId="ref29908" refString="Pisanty, G., Richter, R., Martin, T., Dettman, J. &amp; Cardinal, S. (2022) Molecular phylogeny, historical biogeography and revised classifcation of andrenine bees (Hymenoptera: Andrenidae). Molecular Phylogenetics and Evolution, 170, 107151. https: // doi. org / 10.1016 / j. ympev. 2021.107151" type="journal volume" year="2022">
<taxonomicName id="48DC4D68B85A643FFDA2D983EBF1FCDE" authorityName="Pisanty &amp; Wood" authorityYear="2024" box="[601,683,763,789]" pageId="20" pageNumber="477" rank="subGenus" status="subgen. nov." subGenus="Pisanty">Pisanty</taxonomicName>
<emphasis id="BDA8EAF9B85A643FFD49D983EBB0FCDE" box="[690,746,763,789]" italics="true" pageId="20" pageNumber="471">et al.</emphasis>
2022
</bibRefCitation>
; S. Bossert, M. Branstetter and the authors, unpublished results). However, though it is clearly related to
<taxonomicName id="48DC4D68B85A643FFD59D867EA09FCF2" box="[674,851,799,825]" pageId="20" pageNumber="471" rank="subGenus" subGenus="Warnckandrena">
<emphasis id="BDA8EAF9B85A643FFD59D867EA09FCF2" box="[674,851,799,825]" italics="true" pageId="20" pageNumber="471">Warnckandrena</emphasis>
</taxonomicName>
,
<taxonomicName id="48DC4D68B85A643FFCA5D867EA93FCF3" authorityName="Pisanty &amp; Wood" authorityYear="2024" box="[862,969,799,825]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="remota">
<emphasis id="BDA8EAF9B85A643FFCA5D867EA93FCF3" box="[862,969,799,825]" italics="true" pageId="20" pageNumber="471">A. remota</emphasis>
</taxonomicName>
displays several morphological characters that differ markedly from typical members of the subgenus, and it also occurs
<quantity id="48249B0EB85A643FFBF8D83BED37FC96" box="[1027,1133,835,861]" metricMagnitude="6" metricUnit="m" metricValue="4.0" pageId="20" pageNumber="471" unit="km" value="4000.0">4,000 km</quantity>
outside of its geographical range, deep in the Eastern Palaearctic realm. Further phylogenetic analysis, as well as more sampling in Central
<collectingCountry id="F7CB767BB85A643FFF6CD8F3E980FC6E" box="[151,218,907,933]" name="China" pageId="20" pageNumber="471">China</collectingCountry>
(including the collection of males and possible related species) are needed to assess the subgeneric position of this taxon. If
<taxonomicName id="48DC4D68B85A643FFEB2D8D7E8ECFC03" authorityName="Pisanty &amp; Wood" authorityYear="2024" box="[329,438,943,969]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="remota">
<emphasis id="BDA8EAF9B85A643FFEB2D8D7E8ECFC03" box="[329,438,943,969]" italics="true" pageId="20" pageNumber="471">A. remota</emphasis>
</taxonomicName>
is indeed confirmed as the sister group to
<taxonomicName id="48DC4D68B85A643FFC75D8D7ED65FC02" box="[910,1087,943,969]" pageId="20" pageNumber="471" rank="subGenus" subGenus="Warnckandrena">
<emphasis id="BDA8EAF9B85A643FFC75D8D7ED65FC02" box="[910,1087,943,969]" italics="true" pageId="20" pageNumber="471">Warnckandrena</emphasis>
</taxonomicName>
in the strict sense, it may merit erection of a separate subgenus. For now, we leave
<taxonomicName id="48DC4D68B85A643FFD2CD8ABEA1EFC27" authorityName="Pisanty &amp; Wood" authorityYear="2024" box="[727,836,979,1005]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="471" phylum="Arthropoda" rank="species" species="remota">
<emphasis id="BDA8EAF9B85A643FFD2CD8ABEA1EFC27" box="[727,836,979,1005]" italics="true" pageId="20" pageNumber="471">A. remota</emphasis>
</taxonomicName>
as
<emphasis id="BDA8EAF9B85A643FFC91D8ABED5FFC26" box="[874,1029,979,1005]" italics="true" pageId="20" pageNumber="471">incertae sedis</emphasis>
, pending further study.
</paragraph>
</subSubSection>
</treatment>
</document>
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