plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/07/75/87/077587FDB84C642FFF6CD944ECC7FB11.xml

643 lines
103 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="E7E507C7BFBF10BFC2E1A49DD30B7C77" ID-CLB-Dataset="299666" ID-DOI="10.11646/zootaxa.5474.5.1" ID-GBIF-Dataset="f6a45a12-d3b6-44a7-997e-75eb5ebe9901" ID-ISSN="1175-5326" ID-Zenodo-Dep="12712066" ID-ZooBank="3DDAA7F0-27D0-43E8-AAFB-E56F0E24FB3F" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1720373654954" checkinUser="plazi" docAuthor="Pisanty, Gideon &amp; Wood, Thomas James" docDate="2024" docId="077587FDB84C642FFF6CD944ECC7FB11" docLanguage="en" docName="zootaxa.5474.5.1.pdf" docOrigin="Zootaxa 5474 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5474.5.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Warnckandrena (Warnckandrena) Pisanty, subgen. nov." docType="treatment" docVersion="4" lastPageNumber="455" masterDocId="FB4CFF85B84E642BFFFBDB78E95AFFCB" masterDocTitle="The early-diverging subgenera of the bee genus Andrena (Hymenoptera: Andrenidae) in the Old World" masterLastPageNumber="488" masterPageNumber="451" pageNumber="453" updateTime="1720652268470" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="5E17D937F312E718104AC9CD6BF192DB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6D5E4BD32E418C0BE1B77CBF14E7375F">
<mods:title id="539C25AF843922F7EAF2FAC03F49D5D0">The early-diverging subgenera of the bee genus Andrena (Hymenoptera: Andrenidae) in the Old World</mods:title>
</mods:titleInfo>
<mods:name id="8D78A7D3989DB35418874E3D891D1B59" type="personal">
<mods:role id="E05240E476AAE5DE444C60E65EEAE799">
<mods:roleTerm id="8F772123EE9BB4FDD0AB01C7707D17DD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F08E91A4EA7BB93F8BF5951347DAD212">Pisanty, Gideon</mods:namePart>
<mods:affiliation id="F17A4426A1E47FE520453D9B6213A09D">The Steinhardt Museum of Natural History, Tel Aviv University, Tel Aviv 6997801, Israel.</mods:affiliation>
</mods:name>
<mods:name id="DA6411A65340344BE8B0B694EE2A642D" type="personal">
<mods:role id="DEAF4479CBAE9AA776C7673AA3382F61">
<mods:roleTerm id="0EB494BC4993178D0B6115003AB5D767">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="50CDEFA36076C6A6A3DA3EE78D0E87B3">Wood, Thomas James</mods:namePart>
<mods:affiliation id="A70903C035E9BDAB3EDBCA720F5C632B">Naturalis Biodiversity Center, Darwinweg 2, 2333 CR, Leiden, the Netherlands.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="6E2E1A6A3352E9C20F60F968B59F45F8">text</mods:typeOfResource>
<mods:relatedItem id="ED9E1A196B8137AEB77AC4C02807D946" type="host">
<mods:titleInfo id="B28B4E8D6814CADD73E674F3F59410F6">
<mods:title id="B4CC1F0D359C010F826D9333DFADC905">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="DCAF4818B14A77EAFB41D93F64725F66">
<mods:date id="2ADBFBE85531A54ACC5C6D990D73EEA3">2024</mods:date>
<mods:detail id="9FD82E3B153A0BD182AD0030A762D075" type="pubDate">
<mods:number id="A950740BD5051510BFF6E1CDEADA0B4B">2024-07-02</mods:number>
</mods:detail>
<mods:detail id="1FC0249C3C1342754D8D60D85C363A48" type="volume">
<mods:number id="F0EB558E64E7354A39DBB51204A9A9AE">5474</mods:number>
</mods:detail>
<mods:detail id="5067888BE0259D3BC067F26276E546CD" type="issue">
<mods:number id="B6558B49271A92B7F489B3447CC68C1B">5</mods:number>
</mods:detail>
<mods:extent id="583DA63E2AA2FD76FEBBC0975276D4E4" unit="page">
<mods:start id="E39349E2AF74DB4F38E5A99EEC215BDB">451</mods:start>
<mods:end id="034797D718129488A35B9B1C9E4C2E53">488</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="8C2C4F95A0A57ED8D01DA0741BF857B7">
<mods:url id="FDE026FDAABEF3DA407298D44EB827BC">http://dx.doi.org/10.11646/zootaxa.5474.5.1</mods:url>
</mods:location>
<mods:classification id="E31ACD4B1DF463CCD2E97CDA691A90DD">journal article</mods:classification>
<mods:identifier id="25A6C6807A00D5B5BAC806760412D1E4" type="CLB-Dataset">299666</mods:identifier>
<mods:identifier id="400930FA836DB2B085B7C0E8BF0F43C2" type="DOI">10.11646/zootaxa.5474.5.1</mods:identifier>
<mods:identifier id="24AB3D0F633D09A4C393E403BF0247FE" type="GBIF-Dataset">f6a45a12-d3b6-44a7-997e-75eb5ebe9901</mods:identifier>
<mods:identifier id="914FE3B84F4BB2E299BAD23EB557DF53" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B7960CCC5350CEE518FA5869B0FE827C" type="Zenodo-Dep">12712066</mods:identifier>
<mods:identifier id="EAAB12D6D1611580F853007153A2A5A2" type="ZooBank">3DDAA7F0-27D0-43E8-AAFB-E56F0E24FB3F</mods:identifier>
</mods:mods>
<treatment id="077587FDB84C642FFF6CD944ECC7FB11" ID-DOI="http://doi.org/10.5281/zenodo.12710333" ID-GBIF-Taxon="235032106" ID-Zenodo-Dep="12710333" LSID="urn:lsid:plazi:treatment:077587FDB84C642FFF6CD944ECC7FB11" httpUri="http://treatment.plazi.org/id/077587FDB84C642FFF6CD944ECC7FB11" lastPageId="4" lastPageNumber="455" pageId="2" pageNumber="453">
<subSubSection id="C7C66560B84C6429FF6CD944EBBAFD9D" box="[151,736,571,598]" pageId="2" pageNumber="453" type="nomenclature">
<paragraph id="8F6336EBB84C6429FF6CD944EBBAFD9D" blockId="2.[151,736,571,598]" box="[151,736,571,598]" pageId="2" pageNumber="453">
<heading id="D42B8187B84C6429FF6CD944EBBAFD9D" bold="true" box="[151,736,571,598]" fontSize="11" level="1" pageId="2" pageNumber="453" reason="1">
<emphasis id="BDA8EAF9B84C6429FF6CD944EBBAFD9D" bold="true" box="[151,736,571,598]" pageId="2" pageNumber="453">
Subgenus
<taxonomicName id="48DC4D68B84C6429FEECD943EB65FD9D" authority="Pisanty" authorityName="Pisanty" box="[279,575,571,598]" class="Insecta" family="Andrenidae" genus="Warnckandrena" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="genus">
<emphasis id="BDA8EAF9B84C6429FEECD943E880FD9D" bold="true" box="[279,474,571,598]" italics="true" pageId="2" pageNumber="453">Warnckandrena</emphasis>
Pisanty
</taxonomicName>
<taxonomicNameLabel id="A69B5782B84C6429FDBDD944EBBAFD9D" box="[582,736,572,598]" pageId="2" pageNumber="453" rank="subGenus">subgen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C7C66560B84C6429FF6CD9FCEA3CFD55" box="[151,870,644,670]" pageId="2" pageNumber="453" type="type_taxon">
<paragraph id="8F6336EBB84C6429FF6CD9FCEA3CFD55" blockId="2.[151,1437,644,2039]" box="[151,870,644,670]" pageId="2" pageNumber="453">
<emphasis id="BDA8EAF9B84C6429FF6CD9FCE86FFD55" bold="true" box="[151,309,644,670]" pageId="2" pageNumber="453">
<typeStatus id="50678849B84C6429FF6CD9FCE988FD55" box="[151,210,644,670]" pageId="2" pageNumber="453">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="48DC4D68B84C6429FEC7D9FCEBFEFD55" authority="Morawitz, 1877" authorityName="Morawitz" authorityYear="1877" box="[316,676,644,670]" class="Insecta" family="Melittidae" genus="Melitta" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="curiosa">
<emphasis id="BDA8EAF9B84C6429FEC7D9FCE8BDFD55" box="[316,487,644,670]" italics="true" pageId="2" pageNumber="453">Melitta curiosa</emphasis>
<bibRefCitation id="EB4D4B1AB84C6429FE15D9FCEBFEFD55" author="Morawitz, F." box="[494,676,644,670]" pageId="2" pageNumber="453" pagination="3 - 112" refId="ref29486" refString="Morawitz, F. (1877) Nachtrag zur Bienenfauna Caucasiens. Horae Societatis Entomologicae Rossicae, 14 (1), 3 - 112." type="journal article" year="1877">Morawitz, 1877</bibRefCitation>
</taxonomicName>
, designated here.
</paragraph>
</subSubSection>
<subSubSection id="C7C66560B84C6429FF3CD9D0EB70F83D" pageId="2" pageNumber="453" type="description">
<paragraph id="8F6336EBB84C6429FF3CD9D0EB70F83D" blockId="2.[151,1437,644,2039]" pageId="2" pageNumber="453">
<emphasis id="BDA8EAF9B84C6429FF3CD9D0E803FD09" bold="true" box="[199,345,680,706]" pageId="2" pageNumber="453">Description.</emphasis>
Medium to large-sized (
<quantity id="48249B0EB84C6429FD8AD9D0EBAAFD09" box="[625,752,680,706]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.8" metricValueMin="1.0" pageId="2" pageNumber="453" unit="mm" value="14.0" valueMax="18.0" valueMin="10.0">10–18 mm</quantity>
), usually robust bees (
<figureCitation id="17E72A6EB84C6429FC16D9D0ED67FD09" box="[1005,1085,680,706]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1944,1968]" captionTargetBox="[253,1335,179,1918]" captionTargetId="figure-17@5.[253,1335,179,1918]" captionTargetPageId="5" captionText="FIGURES 1–9. Andrena (Warnckandrena) curiosa (Morawitz), female: 1, flagellum; 2, vertex and mesosoma; 3, metasoma; 4, profile of propodeum; 5, propodeal triangle; 6, flocculus and scopa; 7, forewing venation; 8, habitus; 9, head." figureDoi="http://doi.org/10.5281/zenodo.12712068" httpUri="https://zenodo.org/record/12712068/files/figure.png" pageId="2" pageNumber="453">Figs. 8</figureCitation>
,
<figureCitation id="17E72A6EB84C6429FBB0D9D0EDB1FD09" box="[1099,1259,680,706]" captionStart="FIGURES 10–17" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[263,1335,181,1920]" captionTargetId="figure-33@6.[252,1335,181,1920]" captionTargetPageId="6" captionText="FIGURES 10–17. Andrena (Warnckandrena) female habitus (10, 12, 14, 16) and heads (11, 13, 15, 17): 10–11, A. innesi Gribodo; 12–13, A. papio sp. nov.; 14–15, A. ramlehiana Pérez; 16–17, A. otoolei sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12712072" httpUri="https://zenodo.org/record/12712072/files/figure.png" pageId="2" pageNumber="453">10, 12, 14, 16</figureCitation>
,
<figureCitation id="17E72A6EB84C6429FB03D9D0ECCDFD09" box="[1272,1431,680,706]" captionStart="FIGURES 18–29" captionStartId="7.[151,264,1846,1870]" captionTargetBox="[158,1426,181,1822]" captionTargetId="figure-17@7.[151,1436,181,1822]" captionTargetPageId="7" captionText="FIGURES 18–29. Andrena (Warnckandrena) male habitus (18, 21, 24, 27), heads (19, 22, 25, 28) and genitalia (20, 23, 26, 29): 18–20, A. coromanda Warncke; 21–23, A. papio sp. nov.; 24–26, A. otoolei sp. nov.; 27–29, A. curiosa (Morawitz)." figureDoi="http://doi.org/10.5281/zenodo.12712076" httpUri="https://zenodo.org/record/12712076/files/figure.png" pageId="2" pageNumber="453">18, 21, 24, 27</figureCitation>
, 30, 33). Integument dark, male clypeus yellow, male paraocular area dark. Head 1.0–1.2 times broader than long (
<figureCitation id="17E72A6EB84C6429FF64D988E9ABFCC1" box="[159,241,752,778]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1944,1968]" captionTargetBox="[253,1335,179,1918]" captionTargetId="figure-17@5.[253,1335,179,1918]" captionTargetPageId="5" captionText="FIGURES 1–9. Andrena (Warnckandrena) curiosa (Morawitz), female: 1, flagellum; 2, vertex and mesosoma; 3, metasoma; 4, profile of propodeum; 5, propodeal triangle; 6, flocculus and scopa; 7, forewing venation; 8, habitus; 9, head." figureDoi="http://doi.org/10.5281/zenodo.12712068" httpUri="https://zenodo.org/record/12712068/files/figure.png" pageId="2" pageNumber="453">Figs. 9</figureCitation>
,
<figureCitation id="17E72A6EB84C6429FF04D988E8FBFCC1" box="[255,417,752,779]" captionStart="FIGURES 10–17" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[263,1335,181,1920]" captionTargetId="figure-33@6.[252,1335,181,1920]" captionTargetPageId="6" captionText="FIGURES 10–17. Andrena (Warnckandrena) female habitus (10, 12, 14, 16) and heads (11, 13, 15, 17): 10–11, A. innesi Gribodo; 12–13, A. papio sp. nov.; 14–15, A. ramlehiana Pérez; 16–17, A. otoolei sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12712072" httpUri="https://zenodo.org/record/12712072/files/figure.png" pageId="2" pageNumber="453">11, 13, 15, 17</figureCitation>
,
<figureCitation id="17E72A6EB84C6429FE4BD988EB09FCC1" box="[432,595,752,779]" captionStart="FIGURES 18–29" captionStartId="7.[151,264,1846,1870]" captionTargetBox="[158,1426,181,1822]" captionTargetId="figure-17@7.[151,1436,181,1822]" captionTargetPageId="7" captionText="FIGURES 18–29. Andrena (Warnckandrena) male habitus (18, 21, 24, 27), heads (19, 22, 25, 28) and genitalia (20, 23, 26, 29): 18–20, A. coromanda Warncke; 21–23, A. papio sp. nov.; 24–26, A. otoolei sp. nov.; 27–29, A. curiosa (Morawitz)." figureDoi="http://doi.org/10.5281/zenodo.12712076" httpUri="https://zenodo.org/record/12712076/files/figure.png" pageId="2" pageNumber="453">19, 22, 25, 28</figureCitation>
, 31, 34). Mandible normally developed in both sexes, strongly bidentate. Condylar lamella of female mandible well-developed. Galea finely shagreened, matt to shiny, impunctate or very finely punctured, apex more or less rounded. Maxillary palpus extending as far as galea or beyond, six-segmented, segment 2 longest, 1.2–1.7 times longer than 1. Shape of glossa highly variable, 3–9 times longer than broad. Labial palpus shorter than or subequal to glossa, four-segmented, segment 1 longest. Subgenal coronet well-developed along hind and inner margin of paramandibular process. Labral process broader than long, trapezoidal, rectangular or arched, unnotched. Clypeus weakly to strongly protuberant, arched or medially flattened, 1.2–1.8 times broader than long. Malar space weakly to strongly developed, 0.2 to 0.6 times as long as width of mandible base. Inner margins of compound eyes parallel-sided to convergent above (
<figureCitation id="17E72A6EB84C6429FCA3DF68EAFDFBE1" box="[856,935,1040,1066]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1944,1968]" captionTargetBox="[253,1335,179,1918]" captionTargetId="figure-17@5.[253,1335,179,1918]" captionTargetPageId="5" captionText="FIGURES 1–9. Andrena (Warnckandrena) curiosa (Morawitz), female: 1, flagellum; 2, vertex and mesosoma; 3, metasoma; 4, profile of propodeum; 5, propodeal triangle; 6, flocculus and scopa; 7, forewing venation; 8, habitus; 9, head." figureDoi="http://doi.org/10.5281/zenodo.12712068" httpUri="https://zenodo.org/record/12712068/files/figure.png" pageId="2" pageNumber="453">Figs. 9</figureCitation>
,
<figureCitation id="17E72A6EB84C6429FC49DF68ED11FBE1" box="[946,1099,1040,1067]" captionStart="FIGURES 10–17" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[263,1335,181,1920]" captionTargetId="figure-33@6.[252,1335,181,1920]" captionTargetPageId="6" captionText="FIGURES 10–17. Andrena (Warnckandrena) female habitus (10, 12, 14, 16) and heads (11, 13, 15, 17): 10–11, A. innesi Gribodo; 12–13, A. papio sp. nov.; 14–15, A. ramlehiana Pérez; 16–17, A. otoolei sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12712072" httpUri="https://zenodo.org/record/12712072/files/figure.png" pageId="2" pageNumber="453">11, 13, 15, 17</figureCitation>
,
<figureCitation id="17E72A6EB84C6429FBADDF68EDABFBE1" box="[1110,1265,1040,1067]" captionStart="FIGURES 18–29" captionStartId="7.[151,264,1846,1870]" captionTargetBox="[158,1426,181,1822]" captionTargetId="figure-17@7.[151,1436,181,1822]" captionTargetPageId="7" captionText="FIGURES 18–29. Andrena (Warnckandrena) male habitus (18, 21, 24, 27), heads (19, 22, 25, 28) and genitalia (20, 23, 26, 29): 18–20, A. coromanda Warncke; 21–23, A. papio sp. nov.; 24–26, A. otoolei sp. nov.; 27–29, A. curiosa (Morawitz)." figureDoi="http://doi.org/10.5281/zenodo.12712076" httpUri="https://zenodo.org/record/12712076/files/figure.png" pageId="2" pageNumber="453">19, 22, 25, 28</figureCitation>
, 31, 34). Facial foveae broad and long, sometimes narrowing at mid-length, sometimes separated from compound eye by raised linear area, fovea width equals 0.5–0.7 times antennocular distance. Distance of fovea from lateral ocellus less than 1 ocellar diameter (
<figureCitation id="17E72A6EB84C6429FE51DF04E8AAFB5D" box="[426,496,1148,1174]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1944,1968]" captionTargetBox="[253,1335,179,1918]" captionTargetId="figure-17@5.[253,1335,179,1918]" captionTargetPageId="5" captionText="FIGURES 1–9. Andrena (Warnckandrena) curiosa (Morawitz), female: 1, flagellum; 2, vertex and mesosoma; 3, metasoma; 4, profile of propodeum; 5, propodeal triangle; 6, flocculus and scopa; 7, forewing venation; 8, habitus; 9, head." figureDoi="http://doi.org/10.5281/zenodo.12712068" httpUri="https://zenodo.org/record/12712068/files/figure.png" pageId="2" pageNumber="453">Fig. 2</figureCitation>
). Flagellomere
<quantity id="48249B0EB84C6429FD5BDF04EB96FB5D" box="[672,716,1148,1174]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="2" pageNumber="453" unit="in" value="1.0">1 in</quantity>
females 0.9–1.1 times as long as 2+3+4 (
<figureCitation id="17E72A6EB84C6429FB59DF04EDBEFB5D" box="[1186,1252,1148,1174]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1944,1968]" captionTargetBox="[253,1335,179,1918]" captionTargetId="figure-17@5.[253,1335,179,1918]" captionTargetPageId="5" captionText="FIGURES 1–9. Andrena (Warnckandrena) curiosa (Morawitz), female: 1, flagellum; 2, vertex and mesosoma; 3, metasoma; 4, profile of propodeum; 5, propodeal triangle; 6, flocculus and scopa; 7, forewing venation; 8, habitus; 9, head." figureDoi="http://doi.org/10.5281/zenodo.12712068" httpUri="https://zenodo.org/record/12712068/files/figure.png" pageId="2" pageNumber="453">Fig. 1</figureCitation>
), in males more variable, but always longer than 2 and shorter than 2+3+4. Vertex and gena more or less normally developed, genal area 0.7–1.3 times as broad as compound eye, ocelloccipital distance 1–2 times the diameter of the lateral ocellus (
<figureCitation id="17E72A6EB84C6429FF64DF90E9B2FAC9" box="[159,232,1256,1282]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1944,1968]" captionTargetBox="[253,1335,179,1918]" captionTargetId="figure-17@5.[253,1335,179,1918]" captionTargetPageId="5" captionText="FIGURES 1–9. Andrena (Warnckandrena) curiosa (Morawitz), female: 1, flagellum; 2, vertex and mesosoma; 3, metasoma; 4, profile of propodeum; 5, propodeal triangle; 6, flocculus and scopa; 7, forewing venation; 8, habitus; 9, head." figureDoi="http://doi.org/10.5281/zenodo.12712068" httpUri="https://zenodo.org/record/12712068/files/figure.png" pageId="2" pageNumber="453">Fig. 2</figureCitation>
). Preoccipital ridge dorsally weakly to moderately carinate, laterally rounded in females, slightly angled in males. Dorsolateral angle of pronotum slightly to moderately elevated, sometimes creating a transverse dorsal pronotal ridge that does not descend much laterally. Mesepisternum, propodeum and dorsal mesosomal surfaces finely shagreened, punctation very fine, shallow and often inconspicuous (
<figureCitation id="17E72A6EB84C6429FC02DE2CEDD1FAA4" box="[1017,1163,1364,1391]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1944,1968]" captionTargetBox="[253,1335,179,1918]" captionTargetId="figure-17@5.[253,1335,179,1918]" captionTargetPageId="5" captionText="FIGURES 1–9. Andrena (Warnckandrena) curiosa (Morawitz), female: 1, flagellum; 2, vertex and mesosoma; 3, metasoma; 4, profile of propodeum; 5, propodeal triangle; 6, flocculus and scopa; 7, forewing venation; 8, habitus; 9, head." figureDoi="http://doi.org/10.5281/zenodo.12712068" httpUri="https://zenodo.org/record/12712068/files/figure.png" pageId="2" pageNumber="453">Figs. 2, 4, 5</figureCitation>
). Profile of propodeum weakly separated into narrow, curved basal region versus broad, vertical apical region, basal region about 0.9 times as wide as metanotum (
<figureCitation id="17E72A6EB84C6429FE1EDEE4EB71FA7D" box="[485,555,1436,1462]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1944,1968]" captionTargetBox="[253,1335,179,1918]" captionTargetId="figure-17@5.[253,1335,179,1918]" captionTargetPageId="5" captionText="FIGURES 1–9. Andrena (Warnckandrena) curiosa (Morawitz), female: 1, flagellum; 2, vertex and mesosoma; 3, metasoma; 4, profile of propodeum; 5, propodeal triangle; 6, flocculus and scopa; 7, forewing venation; 8, habitus; 9, head." figureDoi="http://doi.org/10.5281/zenodo.12712068" httpUri="https://zenodo.org/record/12712068/files/figure.png" pageId="2" pageNumber="453">Fig. 4</figureCitation>
). Propodeal triangle very finely shagreened, without rugae (
<figureCitation id="17E72A6EB84C6429FB2BDEE4EC4FFA7C" box="[1232,1301,1436,1463]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1944,1968]" captionTargetBox="[253,1335,179,1918]" captionTargetId="figure-17@5.[253,1335,179,1918]" captionTargetPageId="5" captionText="FIGURES 1–9. Andrena (Warnckandrena) curiosa (Morawitz), female: 1, flagellum; 2, vertex and mesosoma; 3, metasoma; 4, profile of propodeum; 5, propodeal triangle; 6, flocculus and scopa; 7, forewing venation; 8, habitus; 9, head." figureDoi="http://doi.org/10.5281/zenodo.12712068" httpUri="https://zenodo.org/record/12712068/files/figure.png" pageId="2" pageNumber="453">Fig. 5</figureCitation>
). Propodeal corbicula incomplete, corbicular fringe thick, limited to dorsoposterior margin of corbicula, composed of long, minutely to moderately plumose hairs (
<figureCitation id="17E72A6EB84C6429FDACDE9CEBC4FA35" box="[599,670,1508,1534]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1944,1968]" captionTargetBox="[253,1335,179,1918]" captionTargetId="figure-17@5.[253,1335,179,1918]" captionTargetPageId="5" captionText="FIGURES 1–9. Andrena (Warnckandrena) curiosa (Morawitz), female: 1, flagellum; 2, vertex and mesosoma; 3, metasoma; 4, profile of propodeum; 5, propodeal triangle; 6, flocculus and scopa; 7, forewing venation; 8, habitus; 9, head." figureDoi="http://doi.org/10.5281/zenodo.12712068" httpUri="https://zenodo.org/record/12712068/files/figure.png" pageId="2" pageNumber="453">Fig. 4</figureCitation>
). Inner side of hind femur rounded. Inner hind tibial spur normally developed, not broadened or curved. Femoral and tibial scopae composed of simple hairs. Flocculus present, more or less complete (
<figureCitation id="17E72A6EB84C6429FEC5DD54E8D8F98D" box="[318,386,1580,1606]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1944,1968]" captionTargetBox="[253,1335,179,1918]" captionTargetId="figure-17@5.[253,1335,179,1918]" captionTargetPageId="5" captionText="FIGURES 1–9. Andrena (Warnckandrena) curiosa (Morawitz), female: 1, flagellum; 2, vertex and mesosoma; 3, metasoma; 4, profile of propodeum; 5, propodeal triangle; 6, flocculus and scopa; 7, forewing venation; 8, habitus; 9, head." figureDoi="http://doi.org/10.5281/zenodo.12712068" httpUri="https://zenodo.org/record/12712068/files/figure.png" pageId="2" pageNumber="453">Fig. 6</figureCitation>
). Hind pretarsal claw with distinct inner tooth. Forewing with three submarginal cells. Stigma slender, 4–5 times longer than broad, about as broad as prestigma. Submarginal crossvein 1 enters marginal cell 5–6 vein widths from stigma. Recurrent vein 1 enters submarginal cell 2 at its middle or proximal to it. Recurrent vein 2 enters submarginal cell 3 at 0.6 its length. Nervulus antefurcal to interstitial (
<figureCitation id="17E72A6EB84C6429FBC7DDE0EDDAF979" box="[1084,1152,1688,1714]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1944,1968]" captionTargetBox="[253,1335,179,1918]" captionTargetId="figure-17@5.[253,1335,179,1918]" captionTargetPageId="5" captionText="FIGURES 1–9. Andrena (Warnckandrena) curiosa (Morawitz), female: 1, flagellum; 2, vertex and mesosoma; 3, metasoma; 4, profile of propodeum; 5, propodeal triangle; 6, flocculus and scopa; 7, forewing venation; 8, habitus; 9, head." figureDoi="http://doi.org/10.5281/zenodo.12712068" httpUri="https://zenodo.org/record/12712068/files/figure.png" pageId="2" pageNumber="453">Fig. 7</figureCitation>
). Jugal lobe of hind wing 0.7–0.8 times shorter than vannal lobe. Profile of tergum 1 clearly separated into declivous basal region versus horizontal posterior region, basal region with medial longitudinal rim. Tergal discs shagreened, densely and finely punctured, discs 1–3 often with conspicuous hair cover. Tergal apical hair bands moderately developed (
<figureCitation id="17E72A6EB84C6429FABCDC7CECD4F8D5" box="[1351,1422,1796,1822]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1944,1968]" captionTargetBox="[253,1335,179,1918]" captionTargetId="figure-17@5.[253,1335,179,1918]" captionTargetPageId="5" captionText="FIGURES 1–9. Andrena (Warnckandrena) curiosa (Morawitz), female: 1, flagellum; 2, vertex and mesosoma; 3, metasoma; 4, profile of propodeum; 5, propodeal triangle; 6, flocculus and scopa; 7, forewing venation; 8, habitus; 9, head." figureDoi="http://doi.org/10.5281/zenodo.12712068" httpUri="https://zenodo.org/record/12712068/files/figure.png" pageId="2" pageNumber="453">Fig. 3</figureCitation>
). Female pygidial plate normally developed, without distinct elevated medial area; male pygidial plate undeveloped. Male sternum 7 normally developed, with two medioapical lobes which are fully separated to partly fused medially (
<figureCitation id="17E72A6EB84C6429FF64DC08E871F841" box="[159,299,1904,1930]" captionStart="FIGURES 30–43" captionStartId="8.[151,264,1657,1681]" captionTargetBox="[165,1421,181,1633]" captionTargetId="figure-33@8.[165,1421,181,1633]" captionTargetPageId="8" captionText="FIGURES 30–43. Andrena (Warnckandrena) male habitus (30, 33), heads (31, 34) genitalia (32, 35), eighth sterna (36–41) and seventh sterna (42–43): 30–32, 38, A. innesi Gribodo; 33–35, 41, A. ramlehiana Pérez; 36, A. coromanda Warncke; 37, 42, A. curiosa (Morawitz); 39, A. otoolei sp. nov.; 40, 43, A. papio sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12712078" httpUri="https://zenodo.org/record/12712078/files/figure.png" pageId="2" pageNumber="453">Figs. 42–43</figureCitation>
). Male sternum 8 more or less normally developed, columnar, without lateral extensions, pilosity of ventral side more or less uniform, apical process often broadened (
<figureCitation id="17E72A6EB84C6429FC67DCECED72F865" box="[924,1064,1940,1966]" captionStart="FIGURES 30–43" captionStartId="8.[151,264,1657,1681]" captionTargetBox="[165,1421,181,1633]" captionTargetId="figure-33@8.[165,1421,181,1633]" captionTargetPageId="8" captionText="FIGURES 30–43. Andrena (Warnckandrena) male habitus (30, 33), heads (31, 34) genitalia (32, 35), eighth sterna (36–41) and seventh sterna (42–43): 30–32, 38, A. innesi Gribodo; 33–35, 41, A. ramlehiana Pérez; 36, A. coromanda Warncke; 37, 42, A. curiosa (Morawitz); 39, A. otoolei sp. nov.; 40, 43, A. papio sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12712078" httpUri="https://zenodo.org/record/12712078/files/figure.png" pageId="2" pageNumber="453">Figs. 36–41</figureCitation>
). Genital capsule variable, often highly modified. Dorsal gonocoxite lobe absent or developed, gonostylus blade broadened, penis valves elongate and narrow. Volsella often enlarged.
</paragraph>
</subSubSection>
<subSubSection id="C7C66560B84D6428FF3CDBEEE8F8FC76" pageId="3" pageNumber="454" type="diagnosis">
<paragraph id="8F6336EBB84D6428FF3CDBEEE8A4FEA9" blockId="3.[151,1437,150,1454]" pageId="3" pageNumber="454">
<emphasis id="BDA8EAF9B84D6428FF3CDBEEE818FF7B" bold="true" box="[199,322,150,176]" pageId="3" pageNumber="454">Diagnosis.</emphasis>
<taxonomicName id="48DC4D68B84D6428FEB1DBEFE8A1FF7A" box="[330,507,151,177]" pageId="3" pageNumber="471" rank="subGenus" subGenus="Warnckandrena">
<emphasis id="BDA8EAF9B84D6428FEB1DBEFE8A1FF7A" box="[330,507,151,177]" italics="true" pageId="3" pageNumber="454">Warnckandrena</emphasis>
</taxonomicName>
does not possess any unique single character which is fully diagnostic. However, the subgenus can be characterised by the combination of robust body shape, large and broad facial foveae, very long flagellomere 1 especially in females, finely sculptured mesosomal and metasomal surfaces, weakly differentiated and strongly sloping profile of propodeum, and rather modified genitalia. Other important characters, such as mouthparts, clypeal protrusion and malar space, are too variable among the included species to provide any meaningful diagnostic criterion.
</paragraph>
<paragraph id="8F6336EBB84D6428FF3CDA13EB9DFDFC" blockId="3.[151,1437,150,1454]" pageId="3" pageNumber="454">
<taxonomicName id="48DC4D68B84D6428FF3CDA13E8F1FE4E" authorityName="melittoides Friese" authorityYear="1899" box="[199,427,363,389]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="melittoides">
<emphasis id="BDA8EAF9B84D6428FF3CDA13E8F1FE4E" box="[199,427,363,389]" italics="true" pageId="3" pageNumber="454">Andrena melittoides</emphasis>
</taxonomicName>
, the sole representative of subgenus
<taxonomicName id="48DC4D68B84D6428FCA3DA13EA8CFE4E" box="[856,982,363,389]" pageId="3" pageNumber="454" rank="subGenus" subGenus="Melittoides">
<emphasis id="BDA8EAF9B84D6428FCA3DA13EA8CFE4E" box="[856,982,363,389]" italics="true" pageId="3" pageNumber="454">Melittoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A69B5782B84D6428FC1BDA14ED4FFE4D" box="[992,1045,364,390]" pageId="3" pageNumber="454" sensu="stricto">s.str.</taxonomicNameLabel>
, is very easily distinguished from members of
<taxonomicName id="48DC4D68B84D6428FED5DAF7E885FE62" box="[302,479,399,425]" pageId="3" pageNumber="471" rank="subGenus" subGenus="Warnckandrena">
<emphasis id="BDA8EAF9B84D6428FED5DAF7E885FE62" box="[302,479,399,425]" italics="true" pageId="3" pageNumber="454">Warnckandrena</emphasis>
</taxonomicName>
by the basally orange-coloured terga 1–3 (
<figureCitation id="17E72A6EB84D6428FC19DAF7ED2FFE62" box="[994,1141,399,426]" captionStart="FIGURES 65–73" captionStartId="22.[151,264,1903,1927]" captionTargetBox="[165,1421,181,1878]" captionTargetId="figure-33@22.[165,1421,181,1878]" captionTargetPageId="22" captionText="FIGURES 65–73. Andrena (Melittoides) melittoides Friese, female (65–68) and male (69–73): 65, 69, habitus; 66, 70, head; 67, flocculus and scopa; 68, forewing venation; 71, genitalia; 72, eighth sternum; 73, seventh sternum." figureDoi="http://doi.org/10.5281/zenodo.12712090" httpUri="https://zenodo.org/record/12712090/files/figure.png" pageId="3" pageNumber="454">Figs. 65, 69</figureCitation>
), and the male’s yellowmarked paraocular areas (
<figureCitation id="17E72A6EB84D6428FE3ADACAEB4FFE06" box="[449,533,434,461]" captionStart="FIGURES 65–73" captionStartId="22.[151,264,1903,1927]" captionTargetBox="[165,1421,181,1878]" captionTargetId="figure-33@22.[165,1421,181,1878]" captionTargetPageId="22" captionText="FIGURES 65–73. Andrena (Melittoides) melittoides Friese, female (65–68) and male (69–73): 65, 69, habitus; 66, 70, head; 67, flocculus and scopa; 68, forewing venation; 71, genitalia; 72, eighth sternum; 73, seventh sternum." figureDoi="http://doi.org/10.5281/zenodo.12712090" httpUri="https://zenodo.org/record/12712090/files/figure.png" pageId="3" pageNumber="454">Fig. 70</figureCitation>
) and highly unusual genitalia (
<figureCitation id="17E72A6EB84D6428FC84DACAEA8CFE06" box="[895,982,434,461]" captionStart="FIGURES 65–73" captionStartId="22.[151,264,1903,1927]" captionTargetBox="[165,1421,181,1878]" captionTargetId="figure-33@22.[165,1421,181,1878]" captionTargetPageId="22" captionText="FIGURES 65–73. Andrena (Melittoides) melittoides Friese, female (65–68) and male (69–73): 65, 69, habitus; 66, 70, head; 67, flocculus and scopa; 68, forewing venation; 71, genitalia; 72, eighth sternum; 73, seventh sternum." figureDoi="http://doi.org/10.5281/zenodo.12712090" httpUri="https://zenodo.org/record/12712090/files/figure.png" pageId="3" pageNumber="454">Fig. 71</figureCitation>
). Other important diagnostic characters include the very short glossa (
<figureCitation id="17E72A6EB84D6428FE08DAAEEBDAFE3B" box="[499,640,470,496]" captionStart="FIGURES 65–73" captionStartId="22.[151,264,1903,1927]" captionTargetBox="[165,1421,181,1878]" captionTargetId="figure-33@22.[165,1421,181,1878]" captionTargetPageId="22" captionText="FIGURES 65–73. Andrena (Melittoides) melittoides Friese, female (65–68) and male (69–73): 65, 69, habitus; 66, 70, head; 67, flocculus and scopa; 68, forewing venation; 71, genitalia; 72, eighth sternum; 73, seventh sternum." figureDoi="http://doi.org/10.5281/zenodo.12712090" httpUri="https://zenodo.org/record/12712090/files/figure.png" pageId="3" pageNumber="454">Figs. 66, 70</figureCitation>
; this trait highly variable in
<taxonomicName id="48DC4D68B84D6428FC30DAAEED26FE3B" box="[971,1148,470,496]" pageId="3" pageNumber="471" rank="subGenus" subGenus="Warnckandrena">
<emphasis id="BDA8EAF9B84D6428FC30DAAEED26FE3B" box="[971,1148,470,496]" italics="true" pageId="3" pageNumber="454">Warnckandrena</emphasis>
</taxonomicName>
), more strongly plumose mesosomal hair (especially on the propodeum), very strongly postfurcal nervulus (
<figureCitation id="17E72A6EB84D6428FBD4DA81EDDAFDD8" box="[1071,1152,505,531]" captionStart="FIGURES 65–73" captionStartId="22.[151,264,1903,1927]" captionTargetBox="[165,1421,181,1878]" captionTargetId="figure-33@22.[165,1421,181,1878]" captionTargetPageId="22" captionText="FIGURES 65–73. Andrena (Melittoides) melittoides Friese, female (65–68) and male (69–73): 65, 69, habitus; 66, 70, head; 67, flocculus and scopa; 68, forewing venation; 71, genitalia; 72, eighth sternum; 73, seventh sternum." figureDoi="http://doi.org/10.5281/zenodo.12712090" httpUri="https://zenodo.org/record/12712090/files/figure.png" pageId="3" pageNumber="454">Fig. 68</figureCitation>
) (antefurcal to interstitial in
<taxonomicName id="48DC4D68B84D6428FF4FD965E83FFDFC" box="[180,357,541,567]" pageId="3" pageNumber="471" rank="subGenus" subGenus="Warnckandrena">
<emphasis id="BDA8EAF9B84D6428FF4FD965E83FFDFC" box="[180,357,541,567]" italics="true" pageId="3" pageNumber="454">Warnckandrena</emphasis>
</taxonomicName>
), and narrower scopa (
<figureCitation id="17E72A6EB84D6428FD9CD965EBE1FDFC" box="[615,699,541,567]" captionStart="FIGURES 65–73" captionStartId="22.[151,264,1903,1927]" captionTargetBox="[165,1421,181,1878]" captionTargetId="figure-33@22.[165,1421,181,1878]" captionTargetPageId="22" captionText="FIGURES 65–73. Andrena (Melittoides) melittoides Friese, female (65–68) and male (69–73): 65, 69, habitus; 66, 70, head; 67, flocculus and scopa; 68, forewing venation; 71, genitalia; 72, eighth sternum; 73, seventh sternum." figureDoi="http://doi.org/10.5281/zenodo.12712090" httpUri="https://zenodo.org/record/12712090/files/figure.png" pageId="3" pageNumber="454">Fig. 67</figureCitation>
).
</paragraph>
<paragraph id="8F6336EBB84D6428FF3CD938E8F8FC76" blockId="3.[151,1437,150,1454]" pageId="3" pageNumber="454">
Aside from
<taxonomicName id="48DC4D68B84D6428FEACD938E88FFD91" authorityName="Friese" authorityYear="1921" box="[343,469,576,602]" class="Insecta" family="Andrenidae" genus="Melittoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="BDA8EAF9B84D6428FEACD938E88FFD91" box="[343,469,576,602]" italics="true" pageId="3" pageNumber="454">Melittoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A69B5782B84D6428FE1AD939EB4DFD90" box="[481,535,577,603]" pageId="3" pageNumber="454" sensu="stricto">s.str.</taxonomicNameLabel>
,
<taxonomicName id="48DC4D68B84D6428FDD3D938EB83FD91" box="[552,729,576,602]" pageId="3" pageNumber="471" rank="subGenus" subGenus="Warnckandrena">
<emphasis id="BDA8EAF9B84D6428FDD3D938EB83FD91" box="[552,729,576,602]" italics="true" pageId="3" pageNumber="454">Warnckandrena</emphasis>
</taxonomicName>
are most easily confused with members of the species-rich subgenus
<taxonomicName id="48DC4D68B84D6428FEF2D91CE8FDFDB5" box="[265,423,612,638]" pageId="3" pageNumber="454" rank="subGenus" subGenus="Truncandrena">
<emphasis id="BDA8EAF9B84D6428FEF2D91CE8FDFDB5" box="[265,423,612,638]" italics="true" pageId="3" pageNumber="454">Truncandrena</emphasis>
</taxonomicName>
. The best way to distinguish these two subgenera is by the sculpture of the mesosoma. In
<taxonomicName id="48DC4D68B84D6428FF40D9FFE836FD6A" box="[187,364,647,673]" pageId="3" pageNumber="471" rank="subGenus" subGenus="Warnckandrena">
<emphasis id="BDA8EAF9B84D6428FF40D9FFE836FD6A" box="[187,364,647,673]" italics="true" pageId="3" pageNumber="454">Warnckandrena</emphasis>
</taxonomicName>
, all dorsal and lateral mesosomal surfaces (mesepisternum, entire propodeum, mesonotum and scutellum) are very finely shagreened or alveolated and without strong punctation or any striations, and the propodeum in profile is strongly sloping almost in its entirety, without a clearly differentiated horizontal basal region (
<figureCitation id="17E72A6EB84D6428FF16D98AE82BFCC7" box="[237,369,753,780]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1944,1968]" captionTargetBox="[253,1335,179,1918]" captionTargetId="figure-17@5.[253,1335,179,1918]" captionTargetPageId="5" captionText="FIGURES 1–9. Andrena (Warnckandrena) curiosa (Morawitz), female: 1, flagellum; 2, vertex and mesosoma; 3, metasoma; 4, profile of propodeum; 5, propodeal triangle; 6, flocculus and scopa; 7, forewing venation; 8, habitus; 9, head." figureDoi="http://doi.org/10.5281/zenodo.12712068" httpUri="https://zenodo.org/record/12712068/files/figure.png" pageId="3" pageNumber="454">Figs. 2, 4, 5</figureCitation>
). By comparison, the mesosomal sculpture of
<taxonomicName id="48DC4D68B84D6428FC89D98AED4AFCC7" box="[882,1040,754,780]" pageId="3" pageNumber="478" rank="subGenus" subGenus="Truncandrena">
<emphasis id="BDA8EAF9B84D6428FC89D98AED4AFCC7" box="[882,1040,754,780]" italics="true" pageId="3" pageNumber="454">Truncandrena</emphasis>
</taxonomicName>
species is highly variable, but all of them have distinct punctation or striation on some mesosomal surfaces, and/or a strongly differentiated basal region of the propodeum. Similar characters can be used to distinguish the subgenus
<taxonomicName id="48DC4D68B84D6428FBDFD840EDF5FC99" authorityName="Warncke" authorityYear="1968" box="[1060,1199,824,850]" class="Insecta" family="Andrenidae" genus="Nobandrena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="BDA8EAF9B84D6428FBDFD840EDF5FC99" box="[1060,1199,824,850]" italics="true" pageId="3" pageNumber="454">Nobandrena</emphasis>
</taxonomicName>
. In members of this subgenus, the propodeum is very strongly divided into basal versus apical regions with an almost 90 degree angle between them, and the basal part is very broad and almost horizontal. In addition, the males have yellow markings on the paraocular areas.
</paragraph>
</subSubSection>
<subSubSection id="C7C66560B84D6428FF3CD8BEEDB7FB80" pageId="3" pageNumber="454" type="biology_ecology">
<paragraph id="8F6336EBB84D6428FF3CD8BEEDB7FB80" blockId="3.[151,1437,150,1454]" pageId="3" pageNumber="454">
<emphasis id="BDA8EAF9B84D6428FF3CD8BEE87DFC2B" bold="true" box="[199,295,966,992]" pageId="3" pageNumber="454">Biology.</emphasis>
<taxonomicName id="48DC4D68B84D6428FED0D8BEE886FC2B" box="[299,476,966,992]" pageId="3" pageNumber="471" rank="subGenus" subGenus="Warnckandrena">
<emphasis id="BDA8EAF9B84D6428FED0D8BEE886FC2B" box="[299,476,966,992]" italics="true" pageId="3" pageNumber="454">Warnckandrena</emphasis>
</taxonomicName>
are active in springtime on diverse plant families. Pollen collection data are available for four species, all of which appear to be pollen specialists. Three plant families are known pollen hosts (
<taxonomicName id="48DC4D68B84D6428FAF8D892ECC2FBCF" authorityName="G.T.Burnett" authorityYear="1835" box="[1283,1432,1002,1028]" class="Magnoliopsida" family="Brassicaceae" kingdom="Plantae" order="Brassicales" pageId="3" pageNumber="454" phylum="Tracheophyta" rank="family">Brassicaceae</taxonomicName>
,
<taxonomicName id="48DC4D68B84D6428FF6CDF75E858FBEC" box="[151,258,1037,1063]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="3" pageNumber="454" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
,
<taxonomicName id="48DC4D68B84D6428FEF6DF75E8F0FBEC" box="[269,426,1037,1063]" class="Magnoliopsida" family="Papaveraceae" kingdom="Plantae" order="Ranunculales" pageId="3" pageNumber="454" phylum="Tracheophyta" rank="family">Papaveraceae</taxonomicName>
), and additional families are exploited as sources of nectar. The use of diverse flower hosts can explain the great variation in glossal length and head morphology among species of this group.
</paragraph>
</subSubSection>
<subSubSection id="C7C66560B84D6428FF3CDF2CEA49FB59" pageId="3" pageNumber="454" type="distribution">
<paragraph id="8F6336EBB84D6428FF3CDF2CEA49FB59" blockId="3.[151,1437,150,1454]" pageId="3" pageNumber="454">
<emphasis id="BDA8EAF9B84D6428FF3CDF2CE839FBA5" bold="true" box="[199,355,1108,1134]" pageId="3" pageNumber="454">Distribution:</emphasis>
North Africa, Aegean Islands and
<collectingCountry id="F7CB767BB84D6428FD00DF2CEA16FBA5" box="[763,844,1108,1134]" name="Turkey" pageId="3" pageNumber="454">Turkey</collectingCountry>
, the Levant, the Caucasus, and
<collectingCountry id="F7CB767BB84D6428FB39DF2DEDA9FBA4" box="[1218,1267,1109,1135]" name="Iran" pageId="3" pageNumber="454">Iran</collectingCountry>
. The centre of diversity is in
<collectingCountry id="F7CB767BB84D6428FECDDF00E8DCFB59" box="[310,390,1144,1170]" name="Turkey" pageId="3" pageNumber="454">Turkey</collectingCountry>
with five of the six known species.
</paragraph>
</subSubSection>
<subSubSection id="C7C66560B84D6428FF3CDFE3E9BAFA65" pageId="3" pageNumber="454" type="etymology">
<paragraph id="8F6336EBB84D6428FF3CDFE3E846FB37" blockId="3.[151,1437,150,1454]" pageId="3" pageNumber="454">
<emphasis id="BDA8EAF9B84D6428FF3CDFE3E814FB7E" bold="true" box="[199,334,1179,1205]" pageId="3" pageNumber="454">Etymology.</emphasis>
The subgenus is named in honour of Klaus Warncke (1937–1993), the renowned German expert on
<taxonomicName id="48DC4D68B84D6428FF41DFC7E843FB12" box="[186,281,1215,1241]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="BDA8EAF9B84D6428FF41DFC7E843FB12" box="[186,281,1215,1241]" italics="true" pageId="3" pageNumber="454">Andrena</emphasis>
</taxonomicName>
(as well as many other bee groups), in combination with the latter genus name. The gender of the name is feminine.
</paragraph>
<paragraph id="8F6336EBB84D6428FF3CDE7EE9BAFA65" blockId="3.[151,1437,150,1454]" pageId="3" pageNumber="454">
<emphasis id="BDA8EAF9B84D6428FF3CDE7EE8CAFAD4" bold="true" box="[199,400,1285,1312]" pageId="3" pageNumber="454">Included species.</emphasis>
<taxonomicName id="48DC4D68B84D6428FE6DDE7EEA7CFAEB" authority="Warncke, 1975" authorityName="Warncke" authorityYear="1975" box="[406,806,1285,1312]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="coromanda">
<emphasis id="BDA8EAF9B84D6428FE6DDE7EEB20FAEB" box="[406,634,1286,1312]" italics="true" pageId="3" pageNumber="454">Andrena coromanda</emphasis>
<bibRefCitation id="EB4D4B1AB84D6428FD84DE7EEA7CFAEB" author="Warncke, K." box="[639,806,1285,1312]" pageId="3" pageNumber="454" pagination="29 - 102" refId="ref30277" refString="Warncke, K. (1975) Die Sandbienen der Turkei (Hymenoptera, Apoidea, Andrena), Teil B. Beschreibungen der neuen Andrena Arten bzw. Unterarten. Mitteilungen der Munchner Entomologischen Gesellschaft, 65, 29 - 102." type="journal article" year="1975">Warncke, 1975</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F7CB767BB84D6428FCC8DE7EEADFFAEB" box="[819,901,1286,1312]" name="Turkey" pageId="3" pageNumber="454">Turkey</collectingCountry>
and
<collectingCountry id="F7CB767BB84D6428FC42DE7EEAB0FAEB" box="[953,1002,1286,1312]" name="Iran" pageId="3" pageNumber="454">Iran</collectingCountry>
),
<taxonomicName id="48DC4D68B84D6428FC07DE7EEC6DFAEB" authority="(Morawitz, 1877)" baseAuthorityName="Morawitz" baseAuthorityYear="1877" box="[1020,1335,1286,1312]" class="Insecta" family="Melittidae" genus="Melitta" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="curiosa">
<emphasis id="BDA8EAF9B84D6428FC07DE7EED37FAD4" box="[1020,1133,1286,1312]" italics="true" pageId="3" pageNumber="454">A. curiosa</emphasis>
(
<bibRefCitation id="EB4D4B1AB84D6428FB80DE7EEC75FAEB" author="Morawitz, F." box="[1147,1327,1286,1312]" pageId="3" pageNumber="454" pagination="3 - 112" refId="ref29486" refString="Morawitz, F. (1877) Nachtrag zur Bienenfauna Caucasiens. Horae Societatis Entomologicae Rossicae, 14 (1), 3 - 112." type="journal article" year="1877">Morawitz, 1877</bibRefCitation>
)
</taxonomicName>
(
<collectingCountry id="F7CB767BB84D6428FABFDE7EECC2FAEB" box="[1348,1432,1286,1312]" name="Greece" pageId="3" pageNumber="454">Greece</collectingCountry>
, Levant,
<collectingCountry id="F7CB767BB84D6428FF0EDE51E81CFA88" box="[245,326,1321,1347]" name="Turkey" pageId="3" pageNumber="454">Turkey</collectingCountry>
, the Caucasus, and
<collectingCountry id="F7CB767BB84D6428FDD2DE51EB00FA88" box="[553,602,1321,1347]" name="Iran" pageId="3" pageNumber="454">Iran</collectingCountry>
),
<taxonomicName id="48DC4D68B84D6428FD8BDE51EAD8FA88" authority="Gribodo, 1894" authorityName="Gribodo" authorityYear="1894" box="[624,898,1321,1347]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="innesi">
<emphasis id="BDA8EAF9B84D6428FD8BDE51EB8EFA88" box="[624,724,1321,1347]" italics="true" pageId="3" pageNumber="454">A. innesi</emphasis>
<bibRefCitation id="EB4D4B1AB84D6428FD26DE51EAD8FA88" author="Gribodo, G." box="[733,898,1321,1347]" pageId="3" pageNumber="454" pagination="262 - 314" refId="ref28719" refString="Gribodo, G. (1894) Note Imenotterologiche. Nota 2. Nuove generi e n. specie di Imenotteri antofili ed osservazioni sopra alcune specie gia conosciute. Bollettino della Societa entomologica italiana, 26, 76 - 136 + 262 - 314." type="book chapter" year="1894">Gribodo, 1894</bibRefCitation>
</taxonomicName>
(North Africa, Levant, and
<collectingCountry id="F7CB767BB84D6428FB3FDE51EC42FA88" box="[1220,1304,1321,1347]" name="Turkey" pageId="3" pageNumber="454">Turkey</collectingCountry>
),
<taxonomicName id="48DC4D68B84D6428FAD6DE51ECC6FA88" authorityName="Pisanty &amp; Wood" authorityYear="2024" box="[1325,1436,1321,1347]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="otoolei">
<emphasis id="BDA8EAF9B84D6428FAD6DE51ECC6FA88" box="[1325,1436,1321,1347]" italics="true" pageId="3" pageNumber="454">A. otoolei</emphasis>
</taxonomicName>
Pisanty
<emphasis id="BDA8EAF9B84D6428FF0FDE35E808FAAC" bold="true" box="[244,338,1357,1383]" pageId="3" pageNumber="454">
<taxonomicNameLabel id="A69B5782B84D6428FF0FDE35E808FAAC" box="[244,338,1357,1383]" pageId="3" pageNumber="454" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Levant and
<collectingCountry id="F7CB767BB84D6428FE0ADE34EB1FFAAD" box="[497,581,1356,1382]" name="Turkey" pageId="3" pageNumber="454">Turkey</collectingCountry>
),
<taxonomicName id="48DC4D68B84D6428FDA7DE35EBE5FAAD" authorityName="Pisanty &amp; Wood" authorityYear="2024" box="[604,703,1357,1383]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="papio">
<emphasis id="BDA8EAF9B84D6428FDA7DE35EBE5FAAD" box="[604,703,1357,1383]" italics="true" pageId="3" pageNumber="454">A. papio</emphasis>
</taxonomicName>
Pisanty
<emphasis id="BDA8EAF9B84D6428FCDCDE35EADEFAAC" bold="true" box="[807,900,1357,1383]" pageId="3" pageNumber="454">
<taxonomicNameLabel id="A69B5782B84D6428FCDCDE35EADEFAAC" box="[807,900,1357,1383]" pageId="3" pageNumber="454" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<collectingCountry id="F7CB767BB84D6428FC6DDE34EAB6FAAD" box="[918,1004,1356,1382]" name="Turkey" pageId="3" pageNumber="454">Turkey</collectingCountry>
),
<taxonomicName id="48DC4D68B84D6428FBF8DE35EC60FAAC" authority="Perez, 1903" authorityName="Perez" authorityYear="1903" box="[1027,1338,1356,1383]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="ramlehiana">
<emphasis id="BDA8EAF9B84D6428FBF8DE35EDFDFAAD" box="[1027,1191,1356,1383]" italics="true" pageId="3" pageNumber="454">A. ramlehiana</emphasis>
<bibRefCitation id="EB4D4B1AB84D6428FB49DE35EC60FAAC" author="Perez, J." box="[1202,1338,1357,1383]" pageId="3" pageNumber="454" refId="ref29873" refString="Perez, J. (1903) Especes nouvelles de Melliferes (palearctiques). Proces-verbaux des seances de la Societe Linneenne de Bordeaux, 58, LXXVIII - XCIII + CCVIII - CCXXXVI." type="journal volume" year="1903">Pérez, 1903</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F7CB767BB84D6428FAB7DE34ECCAFAAD" box="[1356,1424,1356,1382]" name="Israel" pageId="3" pageNumber="454">Israel</collectingCountry>
). Species belonging to
<taxonomicName id="48DC4D68B84D6428FE71DE08EB61FA41" box="[394,571,1392,1418]" pageId="3" pageNumber="471" rank="subGenus" subGenus="Warnckandrena">
<emphasis id="BDA8EAF9B84D6428FE71DE08EB61FA41" box="[394,571,1392,1418]" italics="true" pageId="3" pageNumber="454">Warnckandrena</emphasis>
</taxonomicName>
were originally assigned to the subgenus
<taxonomicName id="48DC4D68B84D6428FBECDE08EC72FA41" authority="Friese, 1921" authorityName="Friese" authorityYear="1921" box="[1047,1320,1392,1418]" pageId="3" pageNumber="454" rank="subGenus" subGenus="Melittoides">
<emphasis id="BDA8EAF9B84D6428FBECDE08EDCFFA41" box="[1047,1173,1392,1418]" italics="true" pageId="3" pageNumber="454">Melittoides</emphasis>
<bibRefCitation id="EB4D4B1AB84D6428FB66DE08EC72FA41" author="Friese, H." box="[1181,1320,1392,1418]" pageId="3" pageNumber="454" pagination="177 - 180" refId="ref28662" refString="Friese, H. (1921) Eine neue Bienengattung Melittoides n. gen. (Hym.). In: Fahringer, J. &amp; Friese, H., Eine Hymenopteren- Ausbeute aus dem Amanusgebirge. Anhang. Archiv fur Naturgeschichte, 87 A (3), pp. 177 - 180." type="journal article" year="1921">Friese, 1921</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EB4D4B1AB84D6428FACCDE08E989FA65" author="Warncke, K." pageId="3" pageNumber="454" pagination="1 - 110" refId="ref30179" refString="Warncke, K. (1968) Die Untergattungen der westpalaarktischen Bienengattung Andrena F. Memorias e Estudos Mus´eu Zoologico da Universidade de Coimbra, 307, 1 - 110." type="journal article" year="1968">Warncke 1968</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C7C66560B84D642FFF6CDE84ECC6FE7E" lastPageId="4" lastPageNumber="455" pageId="3" pageNumber="454" type="key">
<paragraph id="8F6336EBB84D6428FF6CDE84EB3AF9DD" blockId="3.[151,608,1531,1558]" box="[151,608,1531,1558]" pageId="3" pageNumber="454">
<heading id="D42B8187B84D6428FF6CDE84EB3AF9DD" bold="true" box="[151,608,1531,1558]" fontSize="11" level="1" pageId="3" pageNumber="454" reason="1">
<emphasis id="BDA8EAF9B84D6428FF6CDE84EB3AF9DD" bold="true" box="[151,608,1531,1558]" pageId="3" pageNumber="454">
Key to the species of
<taxonomicName id="48DC4D68B84D6428FE66DE83EB3AF9DD" authority="Pisanty" authorityName="Pisanty" box="[413,608,1531,1558]" class="Insecta" family="Andrenidae" genus="Warnckandrena" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="471" phylum="Arthropoda" rank="subGenus" subGenus="Warnckandrena">
<emphasis id="BDA8EAF9B84D6428FE66DE83EB3AF9DD" bold="true" box="[413,608,1531,1558]" italics="true" pageId="3" pageNumber="454">Warnckandrena</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8F6336EBB84D6428FF6CDD3CE9A3F995" blockId="3.[151,1437,1604,2040]" box="[151,249,1604,1630]" pageId="3" pageNumber="454">
<emphasis id="BDA8EAF9B84D6428FF6CDD3CE9A3F995" bold="true" box="[151,249,1604,1630]" pageId="3" pageNumber="454">Females</emphasis>
</paragraph>
<keyStep id="34282E4EB84D6428FF6CDD1DECC6F924" pageId="3" pageNumber="454">
<paragraph id="8F6336EBB84D6428FF6CDD1DECC6F97E" blockId="3.[151,1437,1604,2040]" pageId="3" pageNumber="454">
<keyLead id="342D95DEB84D6428FF6CDD1DECC6F97E" pageId="3" pageNumber="454">
1- Mid and hind basitarsi orange. Disc of tergum 4 with black hair. Basal half of clypeus shagreened, apical half smooth. Malar area developed, about 0.4 times as long as broad. Glossa short, 3 times longer than broad. Body length
<quantity id="48249B0EB84D6428FB20DDFAEC19F953" box="[1243,1347,1666,1688]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="3" pageNumber="454" unit="mm" value="14.0" valueMax="15.0" valueMin="13.0">13–15 mm</quantity>
(
<collectingCountry id="F7CB767BB84D6428FAA9DDFAECC3F953" box="[1362,1433,1666,1688]" name="Turkey" pageId="3" pageNumber="454">Turkey</collectingCountry>
,
<collectingCountry id="F7CB767BB84D6428FF19DDD8E850F97E" box="[226,266,1696,1717]" name="Iran" pageId="3" pageNumber="454">Iran</collectingCountry>
)....................................................................................
<taxonomicName id="48DC4D68B84D6428FAEBDDD8ECC6F97E" ID-CoL="DRDY" authorityName="Warncke" authorityYear="1975" box="[1296,1436,1695,1717]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="coromanda">
<emphasis id="BDA8EAF9B84D6428FAEBDDD8ECC6F97E" bold="true" box="[1296,1436,1695,1717]" italics="true" pageId="3" pageNumber="454">A. coromanda</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8F6336EBB84D6428FF6CDDC5ECC6F924" blockId="3.[151,1437,1604,2040]" pageId="3" pageNumber="454">
<keyLead id="342D95DEB84D6428FF6CDDC5ECC6F924" pageId="3" pageNumber="454">
- All basitarsi dark (
<figureCitation id="17E72A6EB84D6428FE6EDDC5E883F919" box="[405,473,1725,1747]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1944,1968]" captionTargetBox="[253,1335,179,1918]" captionTargetId="figure-17@5.[253,1335,179,1918]" captionTargetPageId="5" captionText="FIGURES 1–9. Andrena (Warnckandrena) curiosa (Morawitz), female: 1, flagellum; 2, vertex and mesosoma; 3, metasoma; 4, profile of propodeum; 5, propodeal triangle; 6, flocculus and scopa; 7, forewing venation; 8, habitus; 9, head." figureDoi="http://doi.org/10.5281/zenodo.12712068" httpUri="https://zenodo.org/record/12712068/files/figure.png" pageId="3" pageNumber="454">Figs. 8</figureCitation>
,
<figureCitation id="17E72A6EB84D6428FE1FDDC5EB36F918" box="[484,620,1725,1747]" captionStart="FIGURES 10–17" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[263,1335,181,1920]" captionTargetId="figure-33@6.[252,1335,181,1920]" captionTargetPageId="6" captionText="FIGURES 10–17. Andrena (Warnckandrena) female habitus (10, 12, 14, 16) and heads (11, 13, 15, 17): 10–11, A. innesi Gribodo; 12–13, A. papio sp. nov.; 14–15, A. ramlehiana Pérez; 16–17, A. otoolei sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12712072" httpUri="https://zenodo.org/record/12712072/files/figure.png" pageId="3" pageNumber="454">10, 12, 14, 16</figureCitation>
). Disc of tergum 4 with mostly light- to reddish-coloured hair (
<figureCitation id="17E72A6EB84D6428FB36DDC5EC72F919" box="[1229,1320,1725,1747]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1944,1968]" captionTargetBox="[253,1335,179,1918]" captionTargetId="figure-17@5.[253,1335,179,1918]" captionTargetPageId="5" captionText="FIGURES 1–9. Andrena (Warnckandrena) curiosa (Morawitz), female: 1, flagellum; 2, vertex and mesosoma; 3, metasoma; 4, profile of propodeum; 5, propodeal triangle; 6, flocculus and scopa; 7, forewing venation; 8, habitus; 9, head." figureDoi="http://doi.org/10.5281/zenodo.12712068" httpUri="https://zenodo.org/record/12712068/files/figure.png" pageId="3" pageNumber="454">Figs. 3, 8</figureCitation>
,
<figureCitation id="17E72A6EB84D6428FACFDDC5ECCFF918" box="[1332,1429,1725,1747]" captionStart="FIGURES 10–17" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[263,1335,181,1920]" captionTargetId="figure-33@6.[252,1335,181,1920]" captionTargetPageId="6" captionText="FIGURES 10–17. Andrena (Warnckandrena) female habitus (10, 12, 14, 16) and heads (11, 13, 15, 17): 10–11, A. innesi Gribodo; 12–13, A. papio sp. nov.; 14–15, A. ramlehiana Pérez; 16–17, A. otoolei sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12712072" httpUri="https://zenodo.org/record/12712072/files/figure.png" pageId="3" pageNumber="454">10, 12, 16</figureCitation>
) and/or clypeus entirely shagreened (
<figureCitation id="17E72A6EB84D6428FDC6DDA2EBDEF93B" box="[573,644,1754,1776]" captionStart="FIGURES 10–17" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[263,1335,181,1920]" captionTargetId="figure-33@6.[252,1335,181,1920]" captionTargetPageId="6" captionText="FIGURES 10–17. Andrena (Warnckandrena) female habitus (10, 12, 14, 16) and heads (11, 13, 15, 17): 10–11, A. innesi Gribodo; 12–13, A. papio sp. nov.; 14–15, A. ramlehiana Pérez; 16–17, A. otoolei sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12712072" httpUri="https://zenodo.org/record/12712072/files/figure.png" pageId="3" pageNumber="454">Fig. 15</figureCitation>
). Malar area, glossa and body length variable............................ 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="34282E4EB84D6428FF6CDD80ECC6F8AE" pageId="3" pageNumber="454">
<paragraph id="8F6336EBB84D6428FF6CDD80ECC6F88C" blockId="3.[151,1437,1604,2040]" pageId="3" pageNumber="454">
<keyLead id="342D95DEB84D6428FF6CDD80ECC6F88C" pageId="3" pageNumber="454">
2- Clypeus largely shiny, smooth to very superficially shagreened, relatively sparsely punctured, distance between punctures 1–2 puncture diameters (
<figureCitation id="17E72A6EB84D6428FE5FDC6DE884F8E1" box="[420,478,1813,1835]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1944,1968]" captionTargetBox="[253,1335,179,1918]" captionTargetId="figure-17@5.[253,1335,179,1918]" captionTargetPageId="5" captionText="FIGURES 1–9. Andrena (Warnckandrena) curiosa (Morawitz), female: 1, flagellum; 2, vertex and mesosoma; 3, metasoma; 4, profile of propodeum; 5, propodeal triangle; 6, flocculus and scopa; 7, forewing venation; 8, habitus; 9, head." figureDoi="http://doi.org/10.5281/zenodo.12712068" httpUri="https://zenodo.org/record/12712068/files/figure.png" pageId="3" pageNumber="454">Fig. 9</figureCitation>
). Glossa 6 times longer than broad (
<figureCitation id="17E72A6EB84D6428FCC9DC6DEA30F8E1" box="[818,874,1813,1835]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1944,1968]" captionTargetBox="[253,1335,179,1918]" captionTargetId="figure-17@5.[253,1335,179,1918]" captionTargetPageId="5" captionText="FIGURES 1–9. Andrena (Warnckandrena) curiosa (Morawitz), female: 1, flagellum; 2, vertex and mesosoma; 3, metasoma; 4, profile of propodeum; 5, propodeal triangle; 6, flocculus and scopa; 7, forewing venation; 8, habitus; 9, head." figureDoi="http://doi.org/10.5281/zenodo.12712068" httpUri="https://zenodo.org/record/12712068/files/figure.png" pageId="3" pageNumber="454">Fig. 8</figureCitation>
). Body length
<quantity id="48249B0EB84D6428FC0DDC6DED06F8E1" box="[1014,1116,1813,1835]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="3" pageNumber="454" unit="mm" value="16.0" valueMax="17.0" valueMin="15.0">15–17 mm</quantity>
(Aegean Islands, Levant,
<collectingCountry id="F7CB767BB84D6428FAA8DC6DECC3F8E0" box="[1363,1433,1813,1835]" name="Turkey" pageId="3" pageNumber="454">Turkey</collectingCountry>
, Caucasus,
<collectingCountry id="F7CB767BB84D6428FEB3DC4AE82AF88C" box="[328,368,1842,1863]" name="Iran" pageId="3" pageNumber="454">Iran</collectingCountry>
)................................................................................
<taxonomicName id="48DC4D68B84D6428FACCDC4AECC6F88C" baseAuthorityName="Morawitz" baseAuthorityYear="1877" box="[1335,1436,1842,1863]" class="Insecta" family="Melittidae" genus="Melitta" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="curiosa">
<emphasis id="BDA8EAF9B84D6428FACCDC4AECC6F88C" bold="true" box="[1335,1436,1842,1863]" italics="true" pageId="3" pageNumber="454">A. curiosa</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8F6336EBB84D6428FF6CDC28ECC6F8AE" blockId="3.[151,1437,1604,2040]" box="[151,1436,1871,1894]" pageId="3" pageNumber="454">
<keyLead id="342D95DEB84D6428FF6CDC28ECC6F8AE" box="[151,1436,1871,1894]" pageId="3" pageNumber="454">
- Clypeus strongly shagreened at least in basal half (
<figureCitation id="17E72A6EB84D6428FD32DC28EADEF8AE" box="[713,900,1871,1894]" captionStart="FIGURES 10–17" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[263,1335,181,1920]" captionTargetId="figure-33@6.[252,1335,181,1920]" captionTargetPageId="6" captionText="FIGURES 10–17. Andrena (Warnckandrena) female habitus (10, 12, 14, 16) and heads (11, 13, 15, 17): 10–11, A. innesi Gribodo; 12–13, A. papio sp. nov.; 14–15, A. ramlehiana Pérez; 16–17, A. otoolei sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12712072" httpUri="https://zenodo.org/record/12712072/files/figure.png" pageId="3" pageNumber="454">Figs. 11, 13, 15, 17</figureCitation>
). Glossa and body length variable................ 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="34282E4EB84D642FFF6CDC15ECC6FF01" lastPageId="4" lastPageNumber="455" pageId="3" pageNumber="454">
<paragraph id="8F6336EBB84D6428FF6CDC15ECC6F877" blockId="3.[151,1437,1604,2040]" pageId="3" pageNumber="454">
<keyLead id="342D95DEB84D6428FF6CDC15ECC6F877" pageId="3" pageNumber="454">
3- Clypeus densely punctured except for shiny impunctate medioapical area (distance between punctures 0–1 puncture diameters), and usually with slightly elevated midline (
<figureCitation id="17E72A6EB84D6428FD7FDCF2EB91F854" box="[644,715,1930,1952]" captionStart="FIGURES 10–17" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[263,1335,181,1920]" captionTargetId="figure-33@6.[252,1335,181,1920]" captionTargetPageId="6" captionText="FIGURES 10–17. Andrena (Warnckandrena) female habitus (10, 12, 14, 16) and heads (11, 13, 15, 17): 10–11, A. innesi Gribodo; 12–13, A. papio sp. nov.; 14–15, A. ramlehiana Pérez; 16–17, A. otoolei sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12712072" httpUri="https://zenodo.org/record/12712072/files/figure.png" pageId="3" pageNumber="454">Fig. 11</figureCitation>
). Malar area linear, 0.2 times as long as broad (
<figureCitation id="17E72A6EB84D6428FB63DCF2EC57F854" box="[1176,1293,1930,1952]" captionStart="FIGURES 10–17" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[263,1335,181,1920]" captionTargetId="figure-33@6.[252,1335,181,1920]" captionTargetPageId="6" captionText="FIGURES 10–17. Andrena (Warnckandrena) female habitus (10, 12, 14, 16) and heads (11, 13, 15, 17): 10–11, A. innesi Gribodo; 12–13, A. papio sp. nov.; 14–15, A. ramlehiana Pérez; 16–17, A. otoolei sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12712072" httpUri="https://zenodo.org/record/12712072/files/figure.png" pageId="3" pageNumber="454">Figs. 10–11</figureCitation>
). Glossa short, 3.5 times longer than broad (
<figureCitation id="17E72A6EB84D6428FE0DDCD0EB64F876" box="[502,574,1960,1982]" captionStart="FIGURES 10–17" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[263,1335,181,1920]" captionTargetId="figure-33@6.[252,1335,181,1920]" captionTargetPageId="6" captionText="FIGURES 10–17. Andrena (Warnckandrena) female habitus (10, 12, 14, 16) and heads (11, 13, 15, 17): 10–11, A. innesi Gribodo; 12–13, A. papio sp. nov.; 14–15, A. ramlehiana Pérez; 16–17, A. otoolei sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12712072" httpUri="https://zenodo.org/record/12712072/files/figure.png" pageId="3" pageNumber="454">Fig. 10</figureCitation>
). Body length
<quantity id="48249B0EB84D6428FD30DCDFEA68F876" box="[715,818,1959,1981]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="3" pageNumber="454" unit="mm" value="14.0" valueMax="15.0" valueMin="13.0">13–15 mm</quantity>
(North Africa, Levant,
<collectingCountry id="F7CB767BB84D6428FBEEDCD0ED01F875" box="[1045,1115,1960,1982]" name="Turkey" pageId="3" pageNumber="454">Turkey</collectingCountry>
)...................
<taxonomicName id="48DC4D68B84D6428FABFDCD0ECC6F877" ID-CoL="DRWX" authorityName="Gribodo" authorityYear="1894" box="[1348,1436,1959,1981]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="innesi">
<emphasis id="BDA8EAF9B84D6428FABFDCD0ECC6F877" bold="true" box="[1348,1436,1959,1981]" italics="true" pageId="3" pageNumber="454">A. innesi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8F6336EBB84D642FFF6CDCBDECC6FF01" blockId="3.[151,1437,1604,2040]" lastBlockId="4.[151,1437,151,437]" lastPageId="4" lastPageNumber="455" pageId="3" pageNumber="454">
<keyLead id="342D95DEB84D642FFF6CDCBDECC6FF01" lastPageId="4" lastPageNumber="455" pageId="3" pageNumber="454">
- Clypeus uniformly shagreened and matt at least in basal ¾, more sparsely punctured (distance between punctures 0.5–2 puncture diameters), without shiny impunctate medioapical area, without elevated midline (
<figureCitation id="17E72A6EB84D6428FC02DC9AEDCBF83C" box="[1017,1169,2018,2040]" captionStart="FIGURES 10–17" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[263,1335,181,1920]" captionTargetId="figure-33@6.[252,1335,181,1920]" captionTargetPageId="6" captionText="FIGURES 10–17. Andrena (Warnckandrena) female habitus (10, 12, 14, 16) and heads (11, 13, 15, 17): 10–11, A. innesi Gribodo; 12–13, A. papio sp. nov.; 14–15, A. ramlehiana Pérez; 16–17, A. otoolei sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12712072" httpUri="https://zenodo.org/record/12712072/files/figure.png" pageId="3" pageNumber="454">Figs. 13, 15, 17</figureCitation>
). Malar area longer, at least 0.3 times as long as broad (
<figureCitation id="17E72A6EB84A642FFE13DBEFEB04FF66" box="[488,606,151,173]" captionStart="FIGURES 10–17" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[263,1335,181,1920]" captionTargetId="figure-33@6.[252,1335,181,1920]" captionTargetPageId="6" captionText="FIGURES 10–17. Andrena (Warnckandrena) female habitus (10, 12, 14, 16) and heads (11, 13, 15, 17): 10–11, A. innesi Gribodo; 12–13, A. papio sp. nov.; 14–15, A. ramlehiana Pérez; 16–17, A. otoolei sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12712072" httpUri="https://zenodo.org/record/12712072/files/figure.png" pageId="4" pageNumber="455">Figs. 12–17</figureCitation>
). Glossa longer, 6–9 times longer than broad (
<figureCitation id="17E72A6EB84A642FFBE2DBEFEDE8FF66" box="[1049,1202,151,173]" captionStart="FIGURES 10–17" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[263,1335,181,1920]" captionTargetId="figure-33@6.[252,1335,181,1920]" captionTargetPageId="6" captionText="FIGURES 10–17. Andrena (Warnckandrena) female habitus (10, 12, 14, 16) and heads (11, 13, 15, 17): 10–11, A. innesi Gribodo; 12–13, A. papio sp. nov.; 14–15, A. ramlehiana Pérez; 16–17, A. otoolei sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12712072" httpUri="https://zenodo.org/record/12712072/files/figure.png" pageId="4" pageNumber="455">Figs. 12, 14, 16</figureCitation>
). Body length variable. ................................................................................................... 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="34282E4EB84A642FFF6CDBAAECC6FE8B" pageId="4" pageNumber="455">
<paragraph id="8F6336EBB84A642FFF6CDBAAECC6FECE" blockId="4.[151,1437,151,437]" pageId="4" pageNumber="455">
<keyLead id="342D95DEB84A642FFF6CDBAAECC6FECE" pageId="4" pageNumber="455">
4- Malar area almost half as long as broad (
<figureCitation id="17E72A6EB84A642FFD97DBAAEBB9FF2C" box="[620,739,210,232]" captionStart="FIGURES 10–17" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[263,1335,181,1920]" captionTargetId="figure-33@6.[252,1335,181,1920]" captionTargetPageId="6" captionText="FIGURES 10–17. Andrena (Warnckandrena) female habitus (10, 12, 14, 16) and heads (11, 13, 15, 17): 10–11, A. innesi Gribodo; 12–13, A. papio sp. nov.; 14–15, A. ramlehiana Pérez; 16–17, A. otoolei sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12712072" httpUri="https://zenodo.org/record/12712072/files/figure.png" pageId="4" pageNumber="455">Figs. 12–13</figureCitation>
). Clypeus slightly flattened centrally. Tergum 4 with light-coloured hair. Glossa 8 times longer than broad. Body length
<quantity id="48249B0EB84A642FFD5DDB97EA57FECE" box="[678,781,239,261]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="4" pageNumber="455" unit="mm" value="17.0" valueMax="18.0" valueMin="16.0">16–18 mm</quantity>
(
<collectingCountry id="F7CB767BB84A642FFCE1DB97EA38FECE" box="[794,866,239,261]" name="Turkey" pageId="4" pageNumber="455">Turkey</collectingCountry>
)................................
<emphasis id="BDA8EAF9B84A642FFB0CDB97ECC6FECE" bold="true" box="[1271,1436,239,261]" pageId="4" pageNumber="455">
<taxonomicName id="48DC4D68B84A642FFB0CDB97EC10FECF" authorityName="Pisanty &amp; Wood" authorityYear="2024" box="[1271,1354,239,260]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="papio" status="sp. nov.">
<emphasis id="BDA8EAF9B84A642FFB0CDB97EC10FECF" bold="true" box="[1271,1354,239,260]" italics="true" pageId="4" pageNumber="455">A. papio</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A69B5782B84A642FFAABDB88ECC6FECE" box="[1360,1436,240,261]" pageId="4" pageNumber="455" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8F6336EBB84A642FFF6CDA75ECC6FE8B" blockId="4.[151,1437,151,437]" pageId="4" pageNumber="455">
<keyLead id="342D95DEB84A642FFF6CDA75ECC6FE8B" pageId="4" pageNumber="455">
- Malar area 0.3 times as long as broad (
<figureCitation id="17E72A6EB84A642FFDAADA75EB9CFEE9" box="[593,710,269,291]" captionStart="FIGURES 10–17" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[263,1335,181,1920]" captionTargetId="figure-33@6.[252,1335,181,1920]" captionTargetPageId="6" captionText="FIGURES 10–17. Andrena (Warnckandrena) female habitus (10, 12, 14, 16) and heads (11, 13, 15, 17): 10–11, A. innesi Gribodo; 12–13, A. papio sp. nov.; 14–15, A. ramlehiana Pérez; 16–17, A. otoolei sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12712072" httpUri="https://zenodo.org/record/12712072/files/figure.png" pageId="4" pageNumber="455">Figs. 14–17</figureCitation>
). Clypeus evenly domed. Tergum 4 with light- or dark-coloured hair. Glossa 6–9 times longer than broad. Body length
<quantity id="48249B0EB84A642FFD8CDA52EB87FE8B" box="[631,733,298,320]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.5" metricValueMin="1.1" pageId="4" pageNumber="455" unit="mm" value="13.0" valueMax="15.0" valueMin="11.0">11–15 mm</quantity>
......................................................... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="34282E4EB84A642FFF6CDA3FECC6FE7E" pageId="4" pageNumber="455">
<paragraph id="8F6336EBB84A642FFF6CDA3FECC7FEB1" blockId="4.[151,1437,151,437]" pageId="4" pageNumber="455">
<keyLead id="342D95DEB84A642FFF6CDA3FECC7FEB1" pageId="4" pageNumber="455">
5- Tergum 4 with dark brown hair, in sharp contrast to the light-coloured terga 1–3 (
<figureCitation id="17E72A6EB84A642FFC13DA3FED6AFE96" box="[1000,1072,327,349]" captionStart="FIGURES 10–17" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[263,1335,181,1920]" captionTargetId="figure-33@6.[252,1335,181,1920]" captionTargetPageId="6" captionText="FIGURES 10–17. Andrena (Warnckandrena) female habitus (10, 12, 14, 16) and heads (11, 13, 15, 17): 10–11, A. innesi Gribodo; 12–13, A. papio sp. nov.; 14–15, A. ramlehiana Pérez; 16–17, A. otoolei sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12712072" httpUri="https://zenodo.org/record/12712072/files/figure.png" pageId="4" pageNumber="455">Fig. 14</figureCitation>
). Clypeus fully shagreened up to apex (
<figureCitation id="17E72A6EB84A642FFF12DA1DE86AFEB0" box="[233,304,357,379]" captionStart="FIGURES 10–17" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[263,1335,181,1920]" captionTargetId="figure-33@6.[252,1335,181,1920]" captionTargetPageId="6" captionText="FIGURES 10–17. Andrena (Warnckandrena) female habitus (10, 12, 14, 16) and heads (11, 13, 15, 17): 10–11, A. innesi Gribodo; 12–13, A. papio sp. nov.; 14–15, A. ramlehiana Pérez; 16–17, A. otoolei sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12712072" httpUri="https://zenodo.org/record/12712072/files/figure.png" pageId="4" pageNumber="455">Fig. 15</figureCitation>
). Glossa 6 times longer than broad. Body length
<quantity id="48249B0EB84A642FFCFDDA1DEA37FEB1" box="[774,877,357,378]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="4" pageNumber="455" unit="mm" value="12.0" valueMax="13.0" valueMin="11.0">11–13 mm</quantity>
(
<collectingCountry id="F7CB767BB84A642FFC81DA1DEAE8FEB0" box="[890,946,357,379]" name="Israel" pageId="4" pageNumber="455">Israel</collectingCountry>
)............................
<taxonomicName id="48DC4D68B84A642FFAF5DA1DECC7FEB1" ID-CoL="DSRH" authorityName="Perez" authorityYear="1903" box="[1294,1437,356,378]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="ramlehiana">
<emphasis id="BDA8EAF9B84A642FFAF5DA1DECC7FEB1" bold="true" box="[1294,1437,356,378]" italics="true" pageId="4" pageNumber="455">A. ramlehiana</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8F6336EBB84A642FFF6CDAFBECC6FE7E" blockId="4.[151,1437,151,437]" pageId="4" pageNumber="455">
<keyLead id="342D95DEB84A642FFF6CDAFBECC6FE7E" pageId="4" pageNumber="455">
- Terga 1–4 all with white or yellowish hair (
<figureCitation id="17E72A6EB84A642FFD71DAFAEB89FE53" box="[650,723,386,408]" captionStart="FIGURES 10–17" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[263,1335,181,1920]" captionTargetId="figure-33@6.[252,1335,181,1920]" captionTargetPageId="6" captionText="FIGURES 10–17. Andrena (Warnckandrena) female habitus (10, 12, 14, 16) and heads (11, 13, 15, 17): 10–11, A. innesi Gribodo; 12–13, A. papio sp. nov.; 14–15, A. ramlehiana Pérez; 16–17, A. otoolei sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12712072" httpUri="https://zenodo.org/record/12712072/files/figure.png" pageId="4" pageNumber="455">Fig. 16</figureCitation>
). Apical margin of clypeus often smooth to weakly shagreened (
<figureCitation id="17E72A6EB84A642FFAB3DAFAECCAFE5C" box="[1352,1424,386,408]" captionStart="FIGURES 10–17" captionStartId="6.[151,264,1944,1968]" captionTargetBox="[263,1335,181,1920]" captionTargetId="figure-33@6.[252,1335,181,1920]" captionTargetPageId="6" captionText="FIGURES 10–17. Andrena (Warnckandrena) female habitus (10, 12, 14, 16) and heads (11, 13, 15, 17): 10–11, A. innesi Gribodo; 12–13, A. papio sp. nov.; 14–15, A. ramlehiana Pérez; 16–17, A. otoolei sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12712072" httpUri="https://zenodo.org/record/12712072/files/figure.png" pageId="4" pageNumber="455">Fig. 17</figureCitation>
). Glossa 9 times longer than broad. Body length
<quantity id="48249B0EB84A642FFD5DDAE7EA57FE7E" box="[678,781,415,437]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="4" pageNumber="455" unit="mm" value="14.0" valueMax="15.0" valueMin="13.0">13–15 mm</quantity>
(Levant and
<collectingCountry id="F7CB767BB84A642FFC76DAE7EA89FE7E" box="[909,979,415,437]" name="Turkey" pageId="4" pageNumber="455">Turkey</collectingCountry>
)......................
<emphasis id="BDA8EAF9B84A642FFB10DAE7ECC6FE7E" bold="true" box="[1259,1436,415,437]" pageId="4" pageNumber="455">
<taxonomicName id="48DC4D68B84A642FFB10DAE7EC10FE7E" authorityName="Pisanty &amp; Wood" authorityYear="2024" box="[1259,1354,415,437]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="otoolei" status="sp. nov.">
<emphasis id="BDA8EAF9B84A642FFB10DAE7EC10FE7E" bold="true" box="[1259,1354,415,437]" italics="true" pageId="4" pageNumber="455">A. otoolei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A69B5782B84A642FFAABDAD8ECC6FE7E" box="[1360,1436,416,437]" pageId="4" pageNumber="455" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
<subSubSection id="C7C66560B84A642FFF6CDA9CE985FE35" box="[151,223,484,510]" pageId="4" pageNumber="455" type="multiple">
<paragraph id="8F6336EBB84A642FFF6CDA9CE985FE35" blockId="4.[151,1437,484,1243]" box="[151,223,484,510]" pageId="4" pageNumber="455">
<emphasis id="BDA8EAF9B84A642FFF6CDA9CE985FE35" bold="true" box="[151,223,484,510]" pageId="4" pageNumber="455">Males</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C7C66560B84A642FFF6CD97DECC7FB11" pageId="4" pageNumber="455" type="key">
<keyStep id="34282E4EB84A642FFF6CD97DECC6FD44" pageId="4" pageNumber="455">
<paragraph id="8F6336EBB84A642FFF6CD97DECC6FD9E" blockId="4.[151,1437,484,1243]" pageId="4" pageNumber="455">
<keyLead id="342D95DEB84A642FFF6CD97DECC6FD9E" pageId="4" pageNumber="455">
1- Flagellomere 1 1.2 times as long as 2. Hind basitarsi and often also mid basitarsi orange (
<figureCitation id="17E72A6EB84A642FFBBDD97DEDD4FDD1" box="[1094,1166,517,539]" captionStart="FIGURES 18–29" captionStartId="7.[151,264,1846,1870]" captionTargetBox="[158,1426,181,1822]" captionTargetId="figure-17@7.[151,1436,181,1822]" captionTargetPageId="7" captionText="FIGURES 18–29. Andrena (Warnckandrena) male habitus (18, 21, 24, 27), heads (19, 22, 25, 28) and genitalia (20, 23, 26, 29): 18–20, A. coromanda Warncke; 21–23, A. papio sp. nov.; 24–26, A. otoolei sp. nov.; 27–29, A. curiosa (Morawitz)." figureDoi="http://doi.org/10.5281/zenodo.12712076" httpUri="https://zenodo.org/record/12712076/files/figure.png" pageId="4" pageNumber="455">Fig. 18</figureCitation>
). Dorsal lobe of gonocoxite large, extending more than half the distance to the gonostylus blade (
<figureCitation id="17E72A6EB84A642FFC8AD95AEAE2FDFC" box="[881,952,546,568]" captionStart="FIGURES 18–29" captionStartId="7.[151,264,1846,1870]" captionTargetBox="[158,1426,181,1822]" captionTargetId="figure-17@7.[151,1436,181,1822]" captionTargetPageId="7" captionText="FIGURES 18–29. Andrena (Warnckandrena) male habitus (18, 21, 24, 27), heads (19, 22, 25, 28) and genitalia (20, 23, 26, 29): 18–20, A. coromanda Warncke; 21–23, A. papio sp. nov.; 24–26, A. otoolei sp. nov.; 27–29, A. curiosa (Morawitz)." figureDoi="http://doi.org/10.5281/zenodo.12712076" httpUri="https://zenodo.org/record/12712076/files/figure.png" pageId="4" pageNumber="455">Fig. 20</figureCitation>
). Glossa short, 3 times longer than broad (
<figureCitation id="17E72A6EB84A642FFAB1D95AECCAFDFC" box="[1354,1424,546,568]" captionStart="FIGURES 18–29" captionStartId="7.[151,264,1846,1870]" captionTargetBox="[158,1426,181,1822]" captionTargetId="figure-17@7.[151,1436,181,1822]" captionTargetPageId="7" captionText="FIGURES 18–29. Andrena (Warnckandrena) male habitus (18, 21, 24, 27), heads (19, 22, 25, 28) and genitalia (20, 23, 26, 29): 18–20, A. coromanda Warncke; 21–23, A. papio sp. nov.; 24–26, A. otoolei sp. nov.; 27–29, A. curiosa (Morawitz)." figureDoi="http://doi.org/10.5281/zenodo.12712076" httpUri="https://zenodo.org/record/12712076/files/figure.png" pageId="4" pageNumber="455">Fig. 18</figureCitation>
). Body Length
<quantity id="48249B0EB84A642FFE9DD947E885FD9E" box="[358,479,575,597]" metricMagnitude="-2" metricUnit="m" metricValue="1.125" metricValueMax="1.2" metricValueMin="1.05" pageId="4" pageNumber="455" unit="mm" value="11.25" valueMax="12.0" valueMin="10.5">10.5–12 mm</quantity>
(
<collectingCountry id="F7CB767BB84A642FFE17D947EB69FD9E" box="[492,563,575,597]" name="Turkey" pageId="4" pageNumber="455">Turkey</collectingCountry>
,
<collectingCountry id="F7CB767BB84A642FFDC7D938EB3EFD9E" box="[572,612,576,597]" name="Iran" pageId="4" pageNumber="455">Iran</collectingCountry>
)........................................................
<taxonomicName id="48DC4D68B84A642FFAEBD947ECC6FD9E" authorityName="Warncke" authorityYear="1975" box="[1296,1436,575,597]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="coromanda">
<emphasis id="BDA8EAF9B84A642FFAEBD947ECC6FD9E" bold="true" box="[1296,1436,575,597]" italics="true" pageId="4" pageNumber="455">A. coromanda</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8F6336EBB84A642FFF6CD925ECC6FD44" blockId="4.[151,1437,484,1243]" pageId="4" pageNumber="455">
<keyLead id="342D95DEB84A642FFF6CD925ECC6FD44" pageId="4" pageNumber="455">
- Flagellomere 1 much longer than 2, at least 0.9 times as long as 2+3 (
<figureCitation id="17E72A6EB84A642FFC84D925EAAEFDB9" box="[895,1012,605,627]" captionStart="FIGURES 18–29" captionStartId="7.[151,264,1846,1870]" captionTargetBox="[158,1426,181,1822]" captionTargetId="figure-17@7.[151,1436,181,1822]" captionTargetPageId="7" captionText="FIGURES 18–29. Andrena (Warnckandrena) male habitus (18, 21, 24, 27), heads (19, 22, 25, 28) and genitalia (20, 23, 26, 29): 18–20, A. coromanda Warncke; 21–23, A. papio sp. nov.; 24–26, A. otoolei sp. nov.; 27–29, A. curiosa (Morawitz)." figureDoi="http://doi.org/10.5281/zenodo.12712076" httpUri="https://zenodo.org/record/12712076/files/figure.png" pageId="4" pageNumber="455">Figs. 25, 28</figureCitation>
, 31). All basitarsi dark (
<figureCitation id="17E72A6EB84A642FFB20D925EC2EFDB9" box="[1243,1396,605,627]" captionStart="FIGURES 18–29" captionStartId="7.[151,264,1846,1870]" captionTargetBox="[158,1426,181,1822]" captionTargetId="figure-17@7.[151,1436,181,1822]" captionTargetPageId="7" captionText="FIGURES 18–29. Andrena (Warnckandrena) male habitus (18, 21, 24, 27), heads (19, 22, 25, 28) and genitalia (20, 23, 26, 29): 18–20, A. coromanda Warncke; 21–23, A. papio sp. nov.; 24–26, A. otoolei sp. nov.; 27–29, A. curiosa (Morawitz)." figureDoi="http://doi.org/10.5281/zenodo.12712076" httpUri="https://zenodo.org/record/12712076/files/figure.png" pageId="4" pageNumber="455">Figs. 21, 24, 27</figureCitation>
, 30, 33). Dorsal lobe of gonocoxite absent or smaller (
<figureCitation id="17E72A6EB84A642FFD44D902EA0DFD44" box="[703,855,634,656]" captionStart="FIGURES 18–29" captionStartId="7.[151,264,1846,1870]" captionTargetBox="[158,1426,181,1822]" captionTargetId="figure-17@7.[151,1436,181,1822]" captionTargetPageId="7" captionText="FIGURES 18–29. Andrena (Warnckandrena) male habitus (18, 21, 24, 27), heads (19, 22, 25, 28) and genitalia (20, 23, 26, 29): 18–20, A. coromanda Warncke; 21–23, A. papio sp. nov.; 24–26, A. otoolei sp. nov.; 27–29, A. curiosa (Morawitz)." figureDoi="http://doi.org/10.5281/zenodo.12712076" httpUri="https://zenodo.org/record/12712076/files/figure.png" pageId="4" pageNumber="455">Figs. 23, 26, 29</figureCitation>
, 32, 35). Glossa and body length variable.............. 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="34282E4EB84A642FFF6CD9E0ECC6FCCE" pageId="4" pageNumber="455">
<paragraph id="8F6336EBB84A642FFF6CD9E0ECC6FD01" blockId="4.[151,1437,484,1243]" pageId="4" pageNumber="455">
<keyLead id="342D95DEB84A642FFF6CD9E0ECC6FD01" pageId="4" pageNumber="455">
2- Gonocoxites curved medioapically, without dorsal lobes (
<figureCitation id="17E72A6EB84A642FFCEDD9EFEAD5FD66" box="[790,911,663,685]" captionStart="FIGURES 18–29" captionStartId="7.[151,264,1846,1870]" captionTargetBox="[158,1426,181,1822]" captionTargetId="figure-17@7.[151,1436,181,1822]" captionTargetPageId="7" captionText="FIGURES 18–29. Andrena (Warnckandrena) male habitus (18, 21, 24, 27), heads (19, 22, 25, 28) and genitalia (20, 23, 26, 29): 18–20, A. coromanda Warncke; 21–23, A. papio sp. nov.; 24–26, A. otoolei sp. nov.; 27–29, A. curiosa (Morawitz)." figureDoi="http://doi.org/10.5281/zenodo.12712076" httpUri="https://zenodo.org/record/12712076/files/figure.png" pageId="4" pageNumber="455">Figs. 23, 26</figureCitation>
). Glossa long, 8–9 times longer than broad (
<figureCitation id="17E72A6EB84A642FFABCD9EFECCBFD66" box="[1351,1425,663,685]" captionStart="FIGURES 18–29" captionStartId="7.[151,264,1846,1870]" captionTargetBox="[158,1426,181,1822]" captionTargetId="figure-17@7.[151,1436,181,1822]" captionTargetPageId="7" captionText="FIGURES 18–29. Andrena (Warnckandrena) male habitus (18, 21, 24, 27), heads (19, 22, 25, 28) and genitalia (20, 23, 26, 29): 18–20, A. coromanda Warncke; 21–23, A. papio sp. nov.; 24–26, A. otoolei sp. nov.; 27–29, A. curiosa (Morawitz)." figureDoi="http://doi.org/10.5281/zenodo.12712076" httpUri="https://zenodo.org/record/12712076/files/figure.png" pageId="4" pageNumber="455">Fig. 24</figureCitation>
). Body length
<quantity id="48249B0EB84A642FFEA5D9CDE88DFD01" box="[350,471,693,715]" metricMagnitude="-2" metricUnit="m" metricValue="1.375" metricValueMax="1.45" metricValueMin="1.3" pageId="4" pageNumber="455" unit="mm" value="13.75" valueMax="14.5" valueMin="13.0">13–14.5 mm</quantity>
............................................................................... 3
</keyLead>
</paragraph>
<paragraph id="8F6336EBB84A642FFF6CD9ABECC6FCCE" blockId="4.[151,1437,484,1243]" pageId="4" pageNumber="455">
<keyLead id="342D95DEB84A642FFF6CD9ABECC6FCCE" pageId="4" pageNumber="455">
- Gonocoxites with distinct dorsal lobes (
<figureCitation id="17E72A6EB84A642FFD9AD9AAEBE8FD2C" box="[609,690,722,744]" captionStart="FIGURES 18–29" captionStartId="7.[151,264,1846,1870]" captionTargetBox="[158,1426,181,1822]" captionTargetId="figure-17@7.[151,1436,181,1822]" captionTargetPageId="7" captionText="FIGURES 18–29. Andrena (Warnckandrena) male habitus (18, 21, 24, 27), heads (19, 22, 25, 28) and genitalia (20, 23, 26, 29): 18–20, A. coromanda Warncke; 21–23, A. papio sp. nov.; 24–26, A. otoolei sp. nov.; 27–29, A. curiosa (Morawitz)." figureDoi="http://doi.org/10.5281/zenodo.12712076" httpUri="https://zenodo.org/record/12712076/files/figure.png" pageId="4" pageNumber="455">Figs. 29</figureCitation>
, 32, 35). Glossa shorter, 3.5–6.5 times longer than broad. Body length
<quantity id="48249B0EB84A642FFA9BD9AAE85DFCCE" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="4" pageNumber="455" unit="mm" value="12.5" valueMax="15.0" valueMin="10.0">10–15 mm</quantity>
................................................................................................ 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="34282E4EB84A642FFF6CD875ECC6FC5C" pageId="4" pageNumber="455">
<paragraph id="8F6336EBB84A642FFF6CD875ECC6FCF4" blockId="4.[151,1437,484,1243]" pageId="4" pageNumber="455">
<keyLead id="342D95DEB84A642FFF6CD875ECC6FCF4" pageId="4" pageNumber="455">
3- Malar area 0.5–0.6 times as long as broad (
<figureCitation id="17E72A6EB84A642FFD69D875EA50FCE9" box="[658,778,781,803]" captionStart="FIGURES 18–29" captionStartId="7.[151,264,1846,1870]" captionTargetBox="[158,1426,181,1822]" captionTargetId="figure-17@7.[151,1436,181,1822]" captionTargetPageId="7" captionText="FIGURES 18–29. Andrena (Warnckandrena) male habitus (18, 21, 24, 27), heads (19, 22, 25, 28) and genitalia (20, 23, 26, 29): 18–20, A. coromanda Warncke; 21–23, A. papio sp. nov.; 24–26, A. otoolei sp. nov.; 27–29, A. curiosa (Morawitz)." figureDoi="http://doi.org/10.5281/zenodo.12712076" httpUri="https://zenodo.org/record/12712076/files/figure.png" pageId="4" pageNumber="455">Figs. 21–22</figureCitation>
). Penis valves uniformly broad, digiti very large. Inner margins of gonocoxites strongly diverging apically (
<figureCitation id="17E72A6EB84A642FFD90D852EBEBFCF4" box="[619,689,810,832]" captionStart="FIGURES 18–29" captionStartId="7.[151,264,1846,1870]" captionTargetBox="[158,1426,181,1822]" captionTargetId="figure-17@7.[151,1436,181,1822]" captionTargetPageId="7" captionText="FIGURES 18–29. Andrena (Warnckandrena) male habitus (18, 21, 24, 27), heads (19, 22, 25, 28) and genitalia (20, 23, 26, 29): 18–20, A. coromanda Warncke; 21–23, A. papio sp. nov.; 24–26, A. otoolei sp. nov.; 27–29, A. curiosa (Morawitz)." figureDoi="http://doi.org/10.5281/zenodo.12712076" httpUri="https://zenodo.org/record/12712076/files/figure.png" pageId="4" pageNumber="455">Fig. 23</figureCitation>
) (
<collectingCountry id="F7CB767BB84A642FFD3FD852EA56FC8B" box="[708,780,810,832]" name="Turkey" pageId="4" pageNumber="455">Turkey</collectingCountry>
)........................................
<emphasis id="BDA8EAF9B84A642FFB0CD852ECC6FCF4" bold="true" box="[1271,1436,810,831]" pageId="4" pageNumber="455">
<taxonomicName id="48DC4D68B84A642FFB0CD852EC10FCF4" authorityName="Pisanty &amp; Wood" authorityYear="2024" box="[1271,1354,810,831]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="papio" status="sp. nov.">
<emphasis id="BDA8EAF9B84A642FFB0CD852EC10FCF4" bold="true" box="[1271,1354,810,831]" italics="true" pageId="4" pageNumber="455">A. papio</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A69B5782B84A642FFAABD852ECC6FCF4" box="[1360,1436,810,831]" pageId="4" pageNumber="455" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8F6336EBB84A642FFF6CD830ECC6FC5C" blockId="4.[151,1437,484,1243]" pageId="4" pageNumber="455">
<keyLead id="342D95DEB84A642FFF6CD830ECC6FC5C" pageId="4" pageNumber="455">
- Malar area 0.3–0.4 times as long as broad (
<figureCitation id="17E72A6EB84A642FFD70D83FEA59FC96" box="[651,771,839,861]" captionStart="FIGURES 18–29" captionStartId="7.[151,264,1846,1870]" captionTargetBox="[158,1426,181,1822]" captionTargetId="figure-17@7.[151,1436,181,1822]" captionTargetPageId="7" captionText="FIGURES 18–29. Andrena (Warnckandrena) male habitus (18, 21, 24, 27), heads (19, 22, 25, 28) and genitalia (20, 23, 26, 29): 18–20, A. coromanda Warncke; 21–23, A. papio sp. nov.; 24–26, A. otoolei sp. nov.; 27–29, A. curiosa (Morawitz)." figureDoi="http://doi.org/10.5281/zenodo.12712076" httpUri="https://zenodo.org/record/12712076/files/figure.png" pageId="4" pageNumber="455">Figs. 24–25</figureCitation>
). Penis valves slightly swollen medially, digiti small, hidden behind gonocoxite in dorsal view. Inner margins of gonocoxites parallel almost to apex (
<figureCitation id="17E72A6EB84A642FFC10D81DED6BFCB0" box="[1003,1073,869,891]" captionStart="FIGURES 18–29" captionStartId="7.[151,264,1846,1870]" captionTargetBox="[158,1426,181,1822]" captionTargetId="figure-17@7.[151,1436,181,1822]" captionTargetPageId="7" captionText="FIGURES 18–29. Andrena (Warnckandrena) male habitus (18, 21, 24, 27), heads (19, 22, 25, 28) and genitalia (20, 23, 26, 29): 18–20, A. coromanda Warncke; 21–23, A. papio sp. nov.; 24–26, A. otoolei sp. nov.; 27–29, A. curiosa (Morawitz)." figureDoi="http://doi.org/10.5281/zenodo.12712076" httpUri="https://zenodo.org/record/12712076/files/figure.png" pageId="4" pageNumber="455">Fig. 26</figureCitation>
) (Levant,
<collectingCountry id="F7CB767BB84A642FFB6ED81DED81FCB0" box="[1173,1243,869,891]" name="Turkey" pageId="4" pageNumber="455">Turkey</collectingCountry>
)....................................................................................................
<emphasis id="BDA8EAF9B84A642FFB10D8FAECC6FC5C" bold="true" box="[1259,1436,898,920]" pageId="4" pageNumber="455">
<taxonomicName id="48DC4D68B84A642FFB10D8FAEC10FC53" authorityName="Pisanty &amp; Wood" authorityYear="2024" box="[1259,1354,898,920]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="otoolei" status="sp. nov.">
<emphasis id="BDA8EAF9B84A642FFB10D8FAEC10FC53" bold="true" box="[1259,1354,898,920]" italics="true" pageId="4" pageNumber="455">A. otoolei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A69B5782B84A642FFAABD8FAECC6FC5C" box="[1360,1436,898,919]" pageId="4" pageNumber="455" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="34282E4EB84A642FFF6CD8D8ECC6FB83" pageId="4" pageNumber="455">
<paragraph id="8F6336EBB84A642FFF6CD8D8ECC6FC24" blockId="4.[151,1437,484,1243]" pageId="4" pageNumber="455">
<keyLead id="342D95DEB84A642FFF6CD8D8ECC6FC24" pageId="4" pageNumber="455">
4- Dorsal gonocoxite lobes well-developed, parallel-sided, relatively broad, rounded apically. Digiti short, hidden under gonocoxite in dorsal view. (
<figureCitation id="17E72A6EB84A642FFE80D8C5E899FC19" box="[379,451,957,979]" captionStart="FIGURES 18–29" captionStartId="7.[151,264,1846,1870]" captionTargetBox="[158,1426,181,1822]" captionTargetId="figure-17@7.[151,1436,181,1822]" captionTargetPageId="7" captionText="FIGURES 18–29. Andrena (Warnckandrena) male habitus (18, 21, 24, 27), heads (19, 22, 25, 28) and genitalia (20, 23, 26, 29): 18–20, A. coromanda Warncke; 21–23, A. papio sp. nov.; 24–26, A. otoolei sp. nov.; 27–29, A. curiosa (Morawitz)." figureDoi="http://doi.org/10.5281/zenodo.12712076" httpUri="https://zenodo.org/record/12712076/files/figure.png" pageId="4" pageNumber="455">Fig. 29</figureCitation>
). Head as long as broad. Clypeus smooth and shiny at least in apical ½ (
<figureCitation id="17E72A6EB84A642FFB82D8C5ED9BFC19" box="[1145,1217,957,979]" captionStart="FIGURES 18–29" captionStartId="7.[151,264,1846,1870]" captionTargetBox="[158,1426,181,1822]" captionTargetId="figure-17@7.[151,1436,181,1822]" captionTargetPageId="7" captionText="FIGURES 18–29. Andrena (Warnckandrena) male habitus (18, 21, 24, 27), heads (19, 22, 25, 28) and genitalia (20, 23, 26, 29): 18–20, A. coromanda Warncke; 21–23, A. papio sp. nov.; 24–26, A. otoolei sp. nov.; 27–29, A. curiosa (Morawitz)." figureDoi="http://doi.org/10.5281/zenodo.12712076" httpUri="https://zenodo.org/record/12712076/files/figure.png" pageId="4" pageNumber="455">Fig. 28</figureCitation>
). Body length
<quantity id="48249B0EB84A642FFAB5D8C5E85DFC3B" metricMagnitude="-2" metricUnit="m" metricValue="1.425" metricValueMax="1.5" metricValueMin="1.35" pageId="4" pageNumber="455" unit="mm" value="14.25" valueMax="15.0" valueMin="13.5">13.5–15 mm</quantity>
(Aegean Islands, Levant,
<collectingCountry id="F7CB767BB84A642FFDF8D8A2EB13FC3B" box="[515,585,986,1008]" name="Turkey" pageId="4" pageNumber="455">Turkey</collectingCountry>
, Caucasus,
<collectingCountry id="F7CB767BB84A642FFD43D8A2EBBAFC24" box="[696,736,986,1007]" name="Iran" pageId="4" pageNumber="455">Iran</collectingCountry>
).................................................
<taxonomicName id="48DC4D68B84A642FFACCD8A2ECC6FC24" baseAuthorityName="Morawitz" baseAuthorityYear="1877" box="[1335,1436,986,1007]" class="Insecta" family="Melittidae" genus="Melitta" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="curiosa">
<emphasis id="BDA8EAF9B84A642FFACCD8A2ECC6FC24" bold="true" box="[1335,1436,986,1007]" italics="true" pageId="4" pageNumber="455">A. curiosa</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8F6336EBB84A642FFF6CD880ECC6FB83" blockId="4.[151,1437,484,1243]" pageId="4" pageNumber="455">
<keyLead id="342D95DEB84A642FFF6CD880ECC6FB83" pageId="4" pageNumber="455">
- Dorsal gonocoxite lobes short, converging apically, narrow and pointed. Digiti well-developed, knob-shaped, clearly visible above gonocoxites (
<figureCitation id="17E72A6EB84A642FFE5ADF6DEB4EFBE0" box="[417,532,1045,1067]" captionStart="FIGURES 30–43" captionStartId="8.[151,264,1657,1681]" captionTargetBox="[165,1421,181,1633]" captionTargetId="figure-33@8.[165,1421,181,1633]" captionTargetPageId="8" captionText="FIGURES 30–43. Andrena (Warnckandrena) male habitus (30, 33), heads (31, 34) genitalia (32, 35), eighth sterna (36–41) and seventh sterna (42–43): 30–32, 38, A. innesi Gribodo; 33–35, 41, A. ramlehiana Pérez; 36, A. coromanda Warncke; 37, 42, A. curiosa (Morawitz); 39, A. otoolei sp. nov.; 40, 43, A. papio sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12712078" httpUri="https://zenodo.org/record/12712078/files/figure.png" pageId="4" pageNumber="455">Figs. 32, 35</figureCitation>
). Head 1.1–1.2 times broader than long. Clypeus fully shagreened to apically smooth (
<figureCitation id="17E72A6EB84A642FFAB2DF6DE9A7FB8C" captionStart="FIGURES 30–43" captionStartId="8.[151,264,1657,1681]" captionTargetBox="[165,1421,181,1633]" captionTargetId="figure-33@8.[165,1421,181,1633]" captionTargetPageId="8" captionText="FIGURES 30–43. Andrena (Warnckandrena) male habitus (30, 33), heads (31, 34) genitalia (32, 35), eighth sterna (36–41) and seventh sterna (42–43): 30–32, 38, A. innesi Gribodo; 33–35, 41, A. ramlehiana Pérez; 36, A. coromanda Warncke; 37, 42, A. curiosa (Morawitz); 39, A. otoolei sp. nov.; 40, 43, A. papio sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12712078" httpUri="https://zenodo.org/record/12712078/files/figure.png" pageId="4" pageNumber="455">Figs. 31, 34</figureCitation>
). Body length
<quantity id="48249B0EB84A642FFE71DF4AE8ABFB83" box="[394,497,1074,1096]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.4" metricValueMin="1.0" pageId="4" pageNumber="455" unit="mm" value="12.0" valueMax="14.0" valueMin="10.0">10–14 mm</quantity>
............................................................................. 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="34282E4EB84A642FFF6CDF37ECC7FB11" pageId="4" pageNumber="455">
<paragraph id="8F6336EBB84A642FFF6CDF37ECC6FB54" blockId="4.[151,1437,484,1243]" pageId="4" pageNumber="455">
<keyLead id="342D95DEB84A642FFF6CDF37ECC6FB54" pageId="4" pageNumber="455">
5- Gonostyli strongly curved towards each other, with strong inward bulge at mid length (
<figureCitation id="17E72A6EB84A642FFBD8DF37ED31FBAE" box="[1059,1131,1103,1125]" captionStart="FIGURES 30–43" captionStartId="8.[151,264,1657,1681]" captionTargetBox="[165,1421,181,1633]" captionTargetId="figure-33@8.[165,1421,181,1633]" captionTargetPageId="8" captionText="FIGURES 30–43. Andrena (Warnckandrena) male habitus (30, 33), heads (31, 34) genitalia (32, 35), eighth sterna (36–41) and seventh sterna (42–43): 30–32, 38, A. innesi Gribodo; 33–35, 41, A. ramlehiana Pérez; 36, A. coromanda Warncke; 37, 42, A. curiosa (Morawitz); 39, A. otoolei sp. nov.; 40, 43, A. papio sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12712078" httpUri="https://zenodo.org/record/12712078/files/figure.png" pageId="4" pageNumber="455">Fig. 32</figureCitation>
). Flagellomere 1 as long as 2+3 or slightly shorter. Glossa short, 3.5 times longer than broad. Apical ¼–½ of clypeus shiny, more or less smooth (
<figureCitation id="17E72A6EB84A642FFAF0DF15EC0AFB49" box="[1291,1360,1133,1155]" captionStart="FIGURES 30–43" captionStartId="8.[151,264,1657,1681]" captionTargetBox="[165,1421,181,1633]" captionTargetId="figure-33@8.[165,1421,181,1633]" captionTargetPageId="8" captionText="FIGURES 30–43. Andrena (Warnckandrena) male habitus (30, 33), heads (31, 34) genitalia (32, 35), eighth sterna (36–41) and seventh sterna (42–43): 30–32, 38, A. innesi Gribodo; 33–35, 41, A. ramlehiana Pérez; 36, A. coromanda Warncke; 37, 42, A. curiosa (Morawitz); 39, A. otoolei sp. nov.; 40, 43, A. papio sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12712078" httpUri="https://zenodo.org/record/12712078/files/figure.png" pageId="4" pageNumber="455">Fig. 31</figureCitation>
) (North Africa, Levant,
<collectingCountry id="F7CB767BB84A642FFE82DFF2E8E5FB6B" box="[377,447,1162,1184]" name="Turkey" pageId="4" pageNumber="455">Turkey</collectingCountry>
).........................................................................
<taxonomicName id="48DC4D68B84A642FFABFDFF2ECC6FB54" authorityName="Gribodo" authorityYear="1894" box="[1348,1436,1162,1183]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="innesi">
<emphasis id="BDA8EAF9B84A642FFABFDFF2ECC6FB54" bold="true" box="[1348,1436,1162,1183]" italics="true" pageId="4" pageNumber="455">A. innesi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8F6336EBB84A642FFF6CDFD0ECC7FB11" blockId="4.[151,1437,484,1243]" pageId="4" pageNumber="455">
<keyLead id="342D95DEB84A642FFF6CDFD0ECC7FB11" pageId="4" pageNumber="455">
- Gonostyli extended apically, with very weak inward bulge at mid length (
<figureCitation id="17E72A6EB84A642FFC42DFDFED59FB76" box="[953,1027,1191,1213]" captionStart="FIGURES 30–43" captionStartId="8.[151,264,1657,1681]" captionTargetBox="[165,1421,181,1633]" captionTargetId="figure-33@8.[165,1421,181,1633]" captionTargetPageId="8" captionText="FIGURES 30–43. Andrena (Warnckandrena) male habitus (30, 33), heads (31, 34) genitalia (32, 35), eighth sterna (36–41) and seventh sterna (42–43): 30–32, 38, A. innesi Gribodo; 33–35, 41, A. ramlehiana Pérez; 36, A. coromanda Warncke; 37, 42, A. curiosa (Morawitz); 39, A. otoolei sp. nov.; 40, 43, A. papio sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12712078" httpUri="https://zenodo.org/record/12712078/files/figure.png" pageId="4" pageNumber="455">Fig. 35</figureCitation>
). Flagellomere 1 longer than 2+3. Glossa longer, 6–6.5 times longer than broad. Clypeus fully shagreened (
<figureCitation id="17E72A6EB84A642FFCACDFBDEAC6FB11" box="[855,924,1221,1243]" captionStart="FIGURES 30–43" captionStartId="8.[151,264,1657,1681]" captionTargetBox="[165,1421,181,1633]" captionTargetId="figure-33@8.[165,1421,181,1633]" captionTargetPageId="8" captionText="FIGURES 30–43. Andrena (Warnckandrena) male habitus (30, 33), heads (31, 34) genitalia (32, 35), eighth sterna (36–41) and seventh sterna (42–43): 30–32, 38, A. innesi Gribodo; 33–35, 41, A. ramlehiana Pérez; 36, A. coromanda Warncke; 37, 42, A. curiosa (Morawitz); 39, A. otoolei sp. nov.; 40, 43, A. papio sp. nov." figureDoi="http://doi.org/10.5281/zenodo.12712078" httpUri="https://zenodo.org/record/12712078/files/figure.png" pageId="4" pageNumber="455">Fig. 34</figureCitation>
) (
<collectingCountry id="F7CB767BB84A642FFC4BDFBDEAB2FB10" box="[944,1000,1221,1243]" name="Israel" pageId="4" pageNumber="455">Israel</collectingCountry>
).......................
<taxonomicName id="48DC4D68B84A642FFAF5DFBDECC7FB11" authorityName="Perez" authorityYear="1903" box="[1294,1437,1220,1242]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="ramlehiana">
<emphasis id="BDA8EAF9B84A642FFAF5DFBDECC7FB11" bold="true" box="[1294,1437,1220,1242]" italics="true" pageId="4" pageNumber="455">A. ramlehiana</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink