763 lines
115 KiB
XML
763 lines
115 KiB
XML
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<mods:title id="53B26D495EECE631F917052538767FFE">Taxonomy of freshwater gobies of the genus Rhinogobius (Oxudercidae, Gobiiformes) from central Vietnam, with descriptions of two new species</mods:title>
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<mods:namePart id="38AFA5FB768E6F684E39C967FDA5BC08">Maeda, Ken</mods:namePart>
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<mods:affiliation id="C6325082B3B39D762B3B97AE6ABFA549">Marine Eco-Evo-Devo Unit, Okinawa Institute of Science and Technology Graduate University, 1919 - 1 Tancha, Onna, Okinawa 904 -</mods:affiliation>
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<mods:namePart id="53FB063A741233B830171C64B23C0F06">Kobayashi, Hirozumi</mods:namePart>
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<mods:affiliation id="0FD142EC129AF7D8FBC96FC07F571BA3">Tropical Biosphere Research Center, University of the Ryukyus, 1 Senbaru, Nishihara, Okinawa 903 - 0213, Japan. & Department of Zoology, National Museum of Nature and Science, 4 - 1 - 1 Amakubo, Tsukuba, Ibaraki 305 - 0005, Japan.</mods:affiliation>
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<mods:namePart id="61CF32C1AC4E9527BDCBFC2A67338B2E">Iida, Midori</mods:namePart>
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<mods:affiliation id="4EBB07E28D4C29574821B7F08D294F79">Marine Biological Station, Sado Island Center for Ecological Sustainability, Niigata University, 87 Tassha, Sado, Niigata 952 - 2135,</mods:affiliation>
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<mods:namePart id="09006AB94F10645E8E3634B37FA34C60">Tran, Hau Duc</mods:namePart>
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<mods:affiliation id="52ABC05843FB56660014EC087D277D1E">Institute of Natural Science, Hanoi National University of Education, 136 Xuan Thuy, Cau Giay, Hanoi, Vietnam.</mods:affiliation>
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<mods:date id="35365AC95A002948C2F053E2FFEEC412">2024</mods:date>
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<treatment id="7632823FFFE37D52A0C8FEACFBBAF8D8" ID-DOI="http://doi.org/10.5281/zenodo.13659792" ID-GBIF-Taxon="240259050" ID-Zenodo-Dep="13659792" LSID="urn:lsid:zoobank.org:act:FABEECA0-07E7-4CBE-886C-419EBB88F625" httpUri="http://treatment.plazi.org/id/7632823FFFE37D52A0C8FEACFBBAF8D8" lastPageId="14" lastPageNumber="521" pageId="11" pageNumber="518">
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<taxonomicName id="399B48AAFFE37D57A0C8FEACFE97FEA4" authorityName="Maeda & Kobayashi & Iida & Tran" authorityYear="2024" box="[151,375,294,321]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="518" phylum="Chordata" rank="species" species="nami" status="sp. nov.">
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<emphasis id="CCEFEF3BFFE37D57A0C8FEACFE97FEA4" bold="true" box="[151,375,294,321]" italics="true" pageId="11" pageNumber="518">Rhinogobius nami</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="D7DC5240FFE37D57A1D9FEADFE04FEA4" box="[390,484,295,321]" pageId="11" pageNumber="518" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="B68160A2FFE37D57A0C8FEC1FDDCFE80" box="[151,572,331,357]" pageId="11" pageNumber="518" type="description">
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<paragraph id="FE243329FFE37D57A0C8FEC1FDDCFE80" blockId="11.[151,572,294,357]" box="[151,572,331,357]" pageId="11" pageNumber="518">
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(
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<figureCitation id="66A02FACFFE37D57A0C0FEC1FF0FFE80" box="[159,239,331,357]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1899]" captionTargetBox="[158,1429,188,1844]" captionTargetId="figure-17@3.[151,1436,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Proportions of head length, upper-jaw length, first dorsal-, second dorsal-, anal-, and caudal-fin lengths to standard length (SL) of Rhinogobius rong (red circles), R. nami (light-blue circles), R. taenigena (deep-blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) collected from Da Nang City and Thua Thien Hue Province. Solid symbols represent males, open symbols represent females, and symbols filled with grey represent specimens of unknown sex (for R. similis). Symbols with an asterisk are holotypes." figureDoi="http://doi.org/10.5281/zenodo.13330565" httpUri="https://zenodo.org/record/13330565/files/figure.png" pageId="11" pageNumber="518">Figs. 2</figureCitation>
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,
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<figureCitation id="66A02FACFFE37D57A0A4FEC1FED0FE80" box="[251,304,331,357]" captionStart="FIGURE 3" captionStartId="4.[151,243,1970,1994]" captionTargetBox="[160,1429,786,1939]" captionTargetId="figure-313@4.[151,1436,777,1946]" captionTargetPageId="4" captionText="FIGURE 3. Arrangement of cephalic sensory pores and cutaneous sensory papillae in Rhinogobius rong (a–c, NSMT-P 147886) and Rhinogobius nami (d–f, NSMT-P 147890). a, d, lateral view; b, e, dorsal view; c, f, ventral view.AN, anterior naris; PN, posterior naris." figureDoi="http://doi.org/10.5281/zenodo.13330569" httpUri="https://zenodo.org/record/13330569/files/figure.png" pageId="11" pageNumber="518">3d–f</figureCitation>
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,
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<figureCitation id="66A02FACFFE37D57A163FEC1FE95FE80" box="[316,373,331,357]" captionStart="FIGURE 6" captionStartId="10.[151,250,1224,1248]" captionTargetBox="[157,1427,189,1194]" captionTargetId="figure-18@10.[151,1436,181,1200]" captionTargetPageId="10" captionText="FIGURE 6. Preserved specimens of Rhinogobius rong (a, b), R. nami (c, d), R. taenigena (e, f), R. leavelli (g, h), and R. similis (i, j) collected from Da Nang City and Thua Thien Hue Province, Vietnam. Specimens on left side are males, right are females. a, NSMT-P 147886, holotype, 45.6 mm SL; b, OISTICH 254, paratype, 52.7 mm SL; c, NSMT-P 147890, holotype, 24.9 mm SL; d, OISTICH 290, paratype, 27.0 mm SL; e, OISTICH 299, 23.6 mm SL; f, OISTICH 305, 34.1mm SL; g, NSMT-P 147888, 43.8 mm SL; h, NSMT-P 147887, 41.1 mm SL; i, OISTICH 240, 56.3 mm SL; j, OISTICH 241, 45.2 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13330583" httpUri="https://zenodo.org/record/13330583/files/figure.png" pageId="11" pageNumber="518">6c, d</figureCitation>
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,
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<figureCitation id="66A02FACFFE37D57A1DEFEC1FE71FE80" box="[385,401,331,357]" captionStart="FIGURE 7" captionStartId="13.[151,250,942,966]" captionTargetBox="[151,1435,181,918]" captionTargetId="figure-17@13.[151,1435,181,918]" captionTargetPageId="13" captionText="FIGURE 7. Rhinogobius nami immediately after fixation. a, NSMT-P 147890, holotype, male, 24.9 mm SL; b, OISTICH 282, paratype, female, 26.6 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13330585" httpUri="https://zenodo.org/record/13330585/files/figure.png" pageId="11" pageNumber="518">7</figureCitation>
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,
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<figureCitation id="66A02FACFFE37D57A1C2FEC1FE4DFE80" box="[413,429,331,357]" captionStart="FIGURE 8" captionStartId="13.[151,250,1907,1931]" captionTargetBox="[151,1435,1027,1883]" captionTargetId="figure-43@13.[151,1435,1027,1883]" captionTargetPageId="13" captionText="FIGURE 8. Live images of Rhinogobius nami in aquarium (a) and in situ underwater footage from Da Nang City, Vietnam (b–d). a, male, 12 January 2013; b, male, 13 January 2013; c, d, females, 13 January 2013 (photo by K. Maeda)." figureDoi="http://doi.org/10.5281/zenodo.13330587" httpUri="https://zenodo.org/record/13330587/files/figure.png" pageId="11" pageNumber="518">8</figureCitation>
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;
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<tableCitation id="B3190692FFE37D57A1E6FEC1FDD3FE80" box="[441,563,331,357]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStart-3="TABLE 4" captionStart-4="TABLE 5" captionStart-5="TABLE 6" captionStart-6="TABLE 7" captionStartId-0="5.[151,238,476,500]" captionStartId-1="5.[151,239,1153,1177]" captionStartId-2="6.[151,239,440,464]" captionStartId-3="7.[151,237,151,175]" captionStartId-4="7.[151,234,868,892]" captionStartId-5="8.[151,236,151,175]" captionStartId-6="8.[151,237,868,892]" captionText-0="TABLE 1. Number of vertebrae in various species of Rhinogobius examined in the present study. * Includes the holotype." captionText-1="TABLE 2. Number of soft rays on the second dorsal fin and anal fin of various species of Rhinogobius examined in the present study. * Includes the holotype." captionText-2="TABLE 3. Morphometrics of Rhinogobius rong and R. nami, expressed as percentages of standard length. D 1, first dorsal fin; D 2, second dorsal fin; A, anal fin; C, caudal fin; P 2, pelvic fin." captionText-3="TABLE 4. Pectoral-fin ray counts of various species of Rhinogobius examined in the present study. * Includes the holotype." captionText-4="TABLE 5. Longitudinal scale counts of various species of Rhinogobius examined in the present study. * Includes the holotype." captionText-5="TABLE 6. Transverse scale counts of various species of Rhinogobius examined in the present study. *Includes the holotype." captionText-6="TABLE 7. Predorsal scale counts of various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="11" pageNumber="518">Tables 1–7</tableCitation>
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)
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<typeStatus id="21208D8BFFE37D57A0C8FE19FEE4FE48" box="[151,260,403,429]" pageId="11" pageNumber="518" type="holotype">Holotype</typeStatus>
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</emphasis>
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.
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, male (
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SL),
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, a tributary of
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<collectingDate id="9A61EC01FFE37D57A262FE3DFD37FE34" box="[573,727,439,465]" pageId="11" pageNumber="518" value="2019-06-17">17 June 2019</collectingDate>
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|
||
<typeStatus id="21208D8BFFE37D57A098FE51FEDFFE10" box="[199,319,475,501]" pageId="11" pageNumber="518" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
.
|
||
<specimenCode id="AE3D9B52FFE37D57A111FE51FDC1FE10" box="[334,545,474,501]" collectionCode="HNUE-F" pageId="11" pageNumber="518">HNUE-F 351–352</specimenCode>
|
||
,
|
||
<specimenCount id="E89DF8A0FFE37D57A271FE51FD9AFE10" box="[558,634,475,501]" count="1" pageId="11" pageNumber="518" type="male">1 male</specimenCount>
|
||
(27.2) and
|
||
<specimenCount id="E89DF8A0FFE37D57A2A1FE51FC80FE10" box="[766,864,475,501]" count="1" pageId="11" pageNumber="518" type="female">1 female</specimenCount>
|
||
(
|
||
<quantity id="39639ECCFFE37D57A32FFE51FC37FE10" box="[880,983,475,501]" metricMagnitude="-2" metricUnit="m" metricValue="2.48" pageId="11" pageNumber="518" unit="mm" value="24.8">24.8 mm</quantity>
|
||
SL),
|
||
<collectorName id="536E56FFFFE37D57A448FE51FB9DFE10" box="[1047,1149,475,501]" pageId="11" pageNumber="518">Suoi Mo</collectorName>
|
||
, a tributary of
|
||
<location id="FB4465F2FFE37D57A57AFE51FA78FE10" LSID="urn:lsid:plazi:treatment:7632823FFFE37D52A0C8FEACFBBAF8D8:FB4465F2FFE37D57A57AFE51FA78FE10" box="[1317,1432,475,501]" latitude="15.999473" longLatPrecision="1" longitude="108.07019" municipality="Hoa Vang District" name="Song Han" pageId="11" pageNumber="518">Song Han</location>
|
||
,
|
||
<collectingMunicipality id="1E40A953FFE37D57A0C8FE75FEBFFDFC" box="[151,351,511,537]" pageId="11" pageNumber="518">Hoa Vang District</collectingMunicipality>
|
||
(
|
||
<geoCoordinate id="9BAF55EEFFE37D57A131FE74FDEDFDFC" box="[366,525,510,537]" degrees="15" direction="north" minutes="59" orientation="latitude" pageId="11" pageNumber="518" precision="1" seconds="58.1" value="15.999473">15°59’58.1”N</geoCoordinate>
|
||
<geoCoordinate id="9BAF55EEFFE37D57A24CFE75FD5FFDFC" box="[531,703,511,537]" degrees="108" direction="east" minutes="04" orientation="longitude" pageId="11" pageNumber="518" precision="1" seconds="12.7" value="108.07019">108°04’12.7”E</geoCoordinate>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<date id="8A2515E9FFE37D57A28DFE75FC6CFDFC" box="[722,908,511,537]" pageId="11" pageNumber="518" value="2018-01-27">27 January 2018</date>
|
||
;
|
||
<materialsCitation id="4EF33974FFE37D57A3C8FE75FDA5FDD8" ID-GBIF-Occurrence="4928480326" collectingDate="2019-06-17" county="Suoi Mo" latitude="16.00036" location="Hoa Vang District" longLatPrecision="1" longitude="108.05923" municipality="Song Han" pageId="11" pageNumber="518" specimenCode="HNUE-F 353-354" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Da Nang" typeStatus="paratype">
|
||
<specimenCode id="AE3D9B52FFE37D57A3C8FE75FB88FDFC" box="[919,1128,510,537]" collectionCode="HNUE-F" pageId="11" pageNumber="518">HNUE-F 353–354</specimenCode>
|
||
,
|
||
<specimenCount id="E89DF8A0FFE37D57A42CFE75FB5DFDFC" box="[1139,1213,511,537]" count="1" pageId="11" pageNumber="518" type="male">1 male</specimenCount>
|
||
(25.1) and
|
||
<specimenCount id="E89DF8A0FFE37D57A563FE75FA7CFDFC" box="[1340,1436,511,537]" count="1" pageId="11" pageNumber="518" type="female">1 female</specimenCount>
|
||
(
|
||
<quantity id="39639ECCFFE37D57A0C0FDA9FEE6FDD8" box="[159,262,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="2.87" pageId="11" pageNumber="518" unit="mm" value="28.7">28.7 mm</quantity>
|
||
SL), same data as holotype
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="4EF33974FFE37D57A20EFDA9FAC2FDD8" ID-GBIF-Occurrence="4928480340" box="[593,1314,547,573]" collectingDate="2019-06-17" county="Suoi Mo" latitude="16.00036" location="Hoa Vang District" longLatPrecision="1" longitude="108.05923" municipality="Song Han" pageId="11" pageNumber="518" specimenCode="NSMT-P 147889" specimenCount="1" stateProvince="Da Nang" typeStatus="paratype">
|
||
<specimenCode id="AE3D9B52FFE37D57A20EFDA9FCF3FDD8" box="[593,787,547,573]" collectionCode="NSMT-P" pageId="11" pageNumber="518">NSMT-P 147889</specimenCode>
|
||
, female (
|
||
<quantity id="39639ECCFFE37D57A323FDA9FC02FDD8" box="[892,994,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="3.2399999999999998" pageId="11" pageNumber="518" unit="mm" value="32.4">32.4 mm</quantity>
|
||
SL), same data as holotype
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="4EF33974FFE37D57A571FDA9FE1BFD60" ID-GBIF-Occurrence="4928480357" collectorName="Suoi Mo" latitude="15.999361" location="Song Han" longLatPrecision="1" longitude="108.06944" municipality="Hoa Vang District" pageId="11" pageNumber="518" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Da Nang" typeStatus="paratype">
|
||
OISTICH 272–273,
|
||
<specimenCount id="E89DF8A0FFE37D57A157FDCDFEB3FD84" box="[264,339,583,609]" count="1" pageId="11" pageNumber="518" type="male">1 male</specimenCount>
|
||
(
|
||
<quantity id="39639ECCFFE37D57A13CFDCDFE29FD84" box="[355,457,583,609]" metricMagnitude="-2" metricUnit="m" metricValue="2.69" pageId="11" pageNumber="518" unit="mm" value="26.9">26.9 mm</quantity>
|
||
SL) and
|
||
<specimenCount id="E89DF8A0FFE37D57A26DFDCDFD73FD84" box="[562,659,583,609]" count="1" pageId="11" pageNumber="518" type="female">1 female</specimenCount>
|
||
(
|
||
<quantity id="39639ECCFFE37D57A2FCFDCCFCE9FD84" box="[675,777,582,609]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" pageId="11" pageNumber="518" unit="mm" value="27.5">27.5 mm</quantity>
|
||
SL),
|
||
<collectorName id="536E56FFFFE37D57A317FDCDFC4DFD84" box="[840,941,583,609]" pageId="11" pageNumber="518">Suoi Mo</collectorName>
|
||
, a tributary of
|
||
<location id="FB4465F2FFE37D57A40CFDCDFB25FD84" LSID="urn:lsid:plazi:treatment:7632823FFFE37D52A0C8FEACFBBAF8D8:FB4465F2FFE37D57A40CFDCDFB25FD84" box="[1107,1221,583,609]" latitude="15.999361" longLatPrecision="1" longitude="108.06944" municipality="Hoa Vang District" name="Song Han" pageId="11" pageNumber="518" stateProvince="Da Nang">Song Han</location>
|
||
,
|
||
<collectingMunicipality id="1E40A953FFE37D57A48DFDCDFA7CFD84" box="[1234,1436,583,609]" pageId="11" pageNumber="518">Hoa Vang District</collectingMunicipality>
|
||
(
|
||
<geoCoordinate id="9BAF55EEFFE37D57A0FFFDE0FEDFFD60" box="[160,319,618,645]" degrees="15" direction="north" minutes="59" orientation="latitude" pageId="11" pageNumber="518" precision="1" seconds="57.7" value="15.999361">15°59’57.7”N</geoCoordinate>
|
||
<geoCoordinate id="9BAF55EEFFE37D57A119FDE1FE12FD60" box="[326,498,619,645]" degrees="108" direction="east" minutes="04" orientation="longitude" pageId="11" pageNumber="518" precision="1" seconds="10.0" value="108.06944">108°04’10.0”E</geoCoordinate>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="4EF33974FFE37D57A258FDE1FB9CFD4C" ID-GBIF-Occurrence="4928480308" collectingDate="2016-03-01" county="Hoa Vang District" latitude="15.955611" location="Suoi Hoa" longLatPrecision="1" longitude="107.97122" municipality="Song Han" pageId="11" pageNumber="518" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Da Nang" typeStatus="paratype">
|
||
<date id="8A2515E9FFE37D57A258FDE1FD49FD60" box="[519,681,619,645]" pageId="11" pageNumber="518" value="2016-03-01">
|
||
<collectingDate id="9A61EC01FFE37D57A258FDE1FD49FD60" box="[519,681,619,645]" pageId="11" pageNumber="518" value="2016-03-01">1 March 2016</collectingDate>
|
||
</date>
|
||
; OISTICH 274–275,
|
||
<specimenCount id="E89DF8A0FFE37D57A3C4FDE1FC07FD60" box="[923,999,619,645]" count="1" pageId="11" pageNumber="518" type="male">1 male</specimenCount>
|
||
(
|
||
<quantity id="39639ECCFFE37D57A3A8FDE0FBBDFD60" box="[1015,1117,618,645]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" pageId="11" pageNumber="518" unit="mm" value="27.5">27.5 mm</quantity>
|
||
SL) and
|
||
<specimenCount id="E89DF8A0FFE37D57A49AFDE1FAC7FD60" box="[1221,1319,619,645]" count="1" pageId="11" pageNumber="518" type="female">1 female</specimenCount>
|
||
(
|
||
<quantity id="39639ECCFFE37D57A569FDE1FA7DFD60" box="[1334,1437,619,645]" metricMagnitude="-2" metricUnit="m" metricValue="3.06" pageId="11" pageNumber="518" unit="mm" value="30.6">30.6 mm</quantity>
|
||
SL),
|
||
<location id="FB4465F2FFE37D57A091FD05FED7FD4C" LSID="urn:lsid:plazi:treatment:7632823FFFE37D52A0C8FEACFBBAF8D8:FB4465F2FFE37D57A091FD05FED7FD4C" box="[206,311,655,681]" county="Hoa Vang District" latitude="15.955611" longLatPrecision="1" longitude="107.97122" municipality="Song Han" name="Suoi Hoa" pageId="11" pageNumber="518" stateProvince="Da Nang">Suoi Hoa</location>
|
||
, a tributary of
|
||
<collectingMunicipality id="1E40A953FFE37D57A189FD05FDA5FD4C" box="[470,581,655,681]" pageId="11" pageNumber="518">Song Han</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="17454BA5FFE37D57A210FD05FCF6FD4C" box="[591,790,655,681]" pageId="11" pageNumber="518">Hoa Vang District</collectingCounty>
|
||
(
|
||
<geoCoordinate id="9BAF55EEFFE37D57A37BFD04FC23FD4C" box="[804,963,654,681]" degrees="15" direction="north" minutes="57" orientation="latitude" pageId="11" pageNumber="518" precision="1" seconds="20.2" value="15.955611">15°57’20.2”N</geoCoordinate>
|
||
<geoCoordinate id="9BAF55EEFFE37D57A397FD04FB94FD4C" box="[968,1140,654,681]" degrees="107" direction="east" minutes="58" orientation="longitude" pageId="11" pageNumber="518" precision="1" seconds="16.4" value="107.97122">107°58’16.4”E</geoCoordinate>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="4EF33974FFE37D57A4D8FD05FEACFD14" ID-GBIF-Occurrence="4928480352" collectingDate="2016-03-02" collectorName="Suoi Mo" latitude="15.999473" location="Song Han" longLatPrecision="1" longitude="108.07019" municipality="Hoa Vang District" pageId="11" pageNumber="518" specimenCount="3" specimenCount-female="3" stateProvince="Da Nang" typeStatus="paratype">
|
||
<date id="8A2515E9FFE37D57A4D8FD05FAC5FD4C" box="[1159,1317,655,681]" pageId="11" pageNumber="518" value="2016-03-02">
|
||
<collectingDate id="9A61EC01FFE37D57A4D8FD05FAC5FD4C" box="[1159,1317,655,681]" pageId="11" pageNumber="518" value="2016-03-02">2 March 2016</collectingDate>
|
||
</date>
|
||
; OISTICH 276–278,
|
||
<specimenCount id="E89DF8A0FFE37D57A154FD39FE9AFD28" box="[267,378,691,717]" count="3" pageId="11" pageNumber="518" type="female">3 females</specimenCount>
|
||
(28.0–
|
||
<quantity id="39639ECCFFE37D57A192FD39FDD5FD28" box="[461,565,691,717]" metricMagnitude="-2" metricUnit="m" metricValue="2.93" pageId="11" pageNumber="518" unit="mm" value="29.3">29.3 mm</quantity>
|
||
SL),
|
||
<collectorName id="536E56FFFFE37D57A224FD39FD03FD28" box="[635,739,691,717]" pageId="11" pageNumber="518">Suoi Mo</collectorName>
|
||
, a tributary of
|
||
<location id="FB4465F2FFE37D57A3CAFD39FBEAFD28" LSID="urn:lsid:plazi:treatment:7632823FFFE37D52A0C8FEACFBBAF8D8:FB4465F2FFE37D57A3CAFD39FBEAFD28" box="[917,1034,691,717]" latitude="15.999473" longLatPrecision="1" longitude="108.07019" municipality="Hoa Vang District" name="Song Han" pageId="11" pageNumber="518" stateProvince="Da Nang">Song Han</location>
|
||
,
|
||
<collectingMunicipality id="1E40A953FFE37D57A446FD39FB0AFD28" box="[1049,1258,691,717]" pageId="11" pageNumber="518">Hoa Vang District</collectingMunicipality>
|
||
(
|
||
<geoCoordinate id="9BAF55EEFFE37D57A4A2FD38FA7CFD28" box="[1277,1436,690,717]" degrees="15" direction="north" minutes="59" orientation="latitude" pageId="11" pageNumber="518" precision="1" seconds="58.1" value="15.999473">15°59’58.1”N</geoCoordinate>
|
||
<geoCoordinate id="9BAF55EEFFE37D57A0C8FD5DFEA3FD14" box="[151,323,727,753]" degrees="108" direction="east" minutes="04" orientation="longitude" pageId="11" pageNumber="518" precision="1" seconds="12.7" value="108.07019">108°04’12.7”E</geoCoordinate>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="4EF33974FFE37D57A106FD5DFD7AFCF0" ID-GBIF-Occurrence="4928480329" collectingDate="2018-01-27" collectorName="Suoi Hoa" latitude="15.955611" location="Song Han" longLatPrecision="1" longitude="107.97122" municipality="Hoa Vang District" pageId="11" pageNumber="518" specimenCount="1" stateProvince="Da Nang" typeStatus="paratype">
|
||
<date id="8A2515E9FFE37D57A106FD5DFDF7FD14" box="[345,535,727,753]" pageId="11" pageNumber="518" value="2018-01-27">
|
||
<collectingDate id="9A61EC01FFE37D57A106FD5DFDF7FD14" box="[345,535,727,753]" pageId="11" pageNumber="518" value="2018-01-27">27 January 2018</collectingDate>
|
||
</date>
|
||
; OISTICH 279, female (
|
||
<quantity id="39639ECCFFE37D57A36FFD5DFC77FD14" box="[816,919,727,753]" metricMagnitude="-2" metricUnit="m" metricValue="2.68" pageId="11" pageNumber="518" unit="mm" value="26.8">26.8 mm</quantity>
|
||
SL),
|
||
<collectorName id="536E56FFFFE37D57A387FD5DFBA4FD14" box="[984,1092,727,753]" pageId="11" pageNumber="518">Suoi Hoa</collectorName>
|
||
, a tributary of
|
||
<location id="FB4465F2FFE37D57A4B1FD5DFA81FD14" LSID="urn:lsid:plazi:treatment:7632823FFFE37D52A0C8FEACFBBAF8D8:FB4465F2FFE37D57A4B1FD5DFA81FD14" box="[1262,1377,727,753]" latitude="15.955611" longLatPrecision="1" longitude="107.97122" municipality="Hoa Vang District" name="Song Han" pageId="11" pageNumber="518" stateProvince="Da Nang">Song Han</location>
|
||
,
|
||
<collectingMunicipality id="1E40A953FFE37D57A531FD5DFECDFCF0" pageId="11" pageNumber="518">Hoa Vang District</collectingMunicipality>
|
||
(
|
||
<geoCoordinate id="9BAF55EEFFE37D57A161FD70FE3EFCF0" box="[318,478,762,789]" degrees="15" direction="north" minutes="57" orientation="latitude" pageId="11" pageNumber="518" precision="1" seconds="20.2" value="15.955611">15°57’20.2”N</geoCoordinate>
|
||
<geoCoordinate id="9BAF55EEFFE37D57A1B9FD70FD72FCF0" box="[486,658,762,789]" degrees="107" direction="east" minutes="58" orientation="longitude" pageId="11" pageNumber="518" precision="1" seconds="16.4" value="107.97122">107°58’16.4”E</geoCoordinate>
|
||
)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="4EF33974FFE37D57A2F8FD71FCFFFCDC" ID-GBIF-Occurrence="4928480316" collectingDate="2018-01-27" county="Suoi Mo" latitude="16.00036" location="Hoa Vang District" longLatPrecision="1" longitude="108.05923" municipality="Song Han" pageId="11" pageNumber="518" specimenCount="11" specimenCount-female="8" specimenCount-male="3" stateProvince="Da Nang" typeStatus="paratype">
|
||
<date id="8A2515E9FFE37D57A2F8FD71FC86FCF0" box="[679,870,763,789]" pageId="11" pageNumber="518" value="2018-01-27">
|
||
<collectingDate id="9A61EC01FFE37D57A2F8FD71FC86FCF0" box="[679,870,763,789]" pageId="11" pageNumber="518" value="2018-01-27">27 January 2018</collectingDate>
|
||
</date>
|
||
; OISTICH 280–290,
|
||
<specimenCount id="E89DF8A0FFE37D57A403FD71FB54FCF0" box="[1116,1204,763,789]" count="3" pageId="11" pageNumber="518" type="male">3 males</specimenCount>
|
||
(
|
||
<quantity id="39639ECCFFE37D57A49BFD70FA8AFCF0" box="[1220,1386,762,789]" metricMagnitude="-2" metricUnit="m" metricValue="2.68" metricValueMax="2.83" metricValueMin="2.53" pageId="11" pageNumber="518" unit="mm" value="26.8" valueMax="28.3" valueMin="25.3">25.3–28.3 mm</quantity>
|
||
SL) and
|
||
<specimenCount id="E89DF8A0FFE37D57A098FC95FED3FCDC" box="[199,307,799,825]" count="8" pageId="11" pageNumber="518" type="female">8 females</specimenCount>
|
||
(27.0–
|
||
<quantity id="39639ECCFFE37D57A1DDFC95FE06FCDC" box="[386,486,799,825]" metricMagnitude="-2" metricUnit="m" metricValue="3.06" pageId="11" pageNumber="518" unit="mm" value="30.6">30.6 mm</quantity>
|
||
SL), same data as holotype
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B68160A2FFE37D57A098FCC8FCC4FB98" pageId="11" pageNumber="518" type="diagnosis">
|
||
<paragraph id="FE243329FFE37D57A098FCC8FCC4FB98" blockId="11.[151,1437,403,2013]" pageId="11" pageNumber="518">
|
||
<emphasis id="CCEFEF3BFFE37D57A098FCC8FEA2FCB9" bold="true" box="[199,322,834,860]" pageId="11" pageNumber="518">Diagnosis.</emphasis>
|
||
Pectoral fin with 14–16 rays. Second dorsal and anal fins usually with one spine and eight soft rays. Number of vertebrae usually 26. Nape and posterior part of occipital region covered by cycloid scales; scaled area extending anteriorly to about halfway between posterior margins of preopercle and opercle. Longitudinal scales 28–31, transverse scales eight or nine, and predorsal scales 6–11. Cephalic sensory pore system usually with B´, C, D(S), E, F, H´, K´, L´, M´, N, and O´. Five, longitudinal, reddish-brown or dusky-brown stripes (dotted lines) running from trunk to caudal peduncle both in male and female; fourth (from dorsal) one slightly below lateral midline most distinct and others sometimes unclear. Male having three parallel, diagonally downward, reddish-brown or dusky-brown stripes on ventral part of cheek. Opercular membrane, interopercle, and lower part of subopercle of live male specimen light bluish-grey with many red or orange spots.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B68160A2FFE37D50A098FB0CFD01F894" lastPageId="12" lastPageNumber="519" pageId="11" pageNumber="518" type="description">
|
||
<paragraph id="FE243329FFE37D57A098FB0CFA89F990" blockId="11.[151,1437,403,2013]" pageId="11" pageNumber="518">
|
||
<emphasis id="CCEFEF3BFFE37D57A098FB0CFEB9FB45" bold="true" box="[199,345,1158,1184]" pageId="11" pageNumber="518">Description.</emphasis>
|
||
Body nearly cylindrical anteriorly and somewhat compressed posteriorly. Anterior tip of head pointed in female but rounded in male. Head larger in male than female (head length 31.3–34.5 vs. 28.4–30.9% of SL;
|
||
<figureCitation id="66A02FACFFE37D57A099FB45FEE9FB0C" box="[198,265,1231,1257]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1899]" captionTargetBox="[158,1429,188,1844]" captionTargetId="figure-17@3.[151,1436,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Proportions of head length, upper-jaw length, first dorsal-, second dorsal-, anal-, and caudal-fin lengths to standard length (SL) of Rhinogobius rong (red circles), R. nami (light-blue circles), R. taenigena (deep-blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) collected from Da Nang City and Thua Thien Hue Province. Solid symbols represent males, open symbols represent females, and symbols filled with grey represent specimens of unknown sex (for R. similis). Symbols with an asterisk are holotypes." figureDoi="http://doi.org/10.5281/zenodo.13330565" httpUri="https://zenodo.org/record/13330565/files/figure.png" pageId="11" pageNumber="518">Fig. 2</figureCitation>
|
||
,
|
||
<tableCitation id="B3190692FFE37D57A14CFB45FE87FB0C" box="[275,359,1231,1257]" captionStart="TABLE 3" captionStartId="6.[151,239,440,464]" captionText="TABLE 3. Morphometrics of Rhinogobius rong and R. nami, expressed as percentages of standard length. D 1, first dorsal fin; D 2, second dorsal fin; A, anal fin; C, caudal fin; P 2, pelvic fin." pageId="11" pageNumber="518">Table 3</tableCitation>
|
||
). Eyes located dorsolaterally. Mouth terminal and oblique with thick upper and lower lips. When mouth closed, anterior tips of upper and lower jaws almost even or lower jaw protruding slightly beyond upper jaw. Upper jaw extending posteriorly to center of eye in male, but ending below anterior part of eye (between anterior margin of eye and anterior margin of pupil) in female. Mouth larger in male than female (upper-jaw length 11.6– 13.8 vs. 8.4–9.6% of SL;
|
||
<figureCitation id="66A02FACFFE37D57A1E2FAD5FDE2FA9C" box="[445,514,1375,1401]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1899]" captionTargetBox="[158,1429,188,1844]" captionTargetId="figure-17@3.[151,1436,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Proportions of head length, upper-jaw length, first dorsal-, second dorsal-, anal-, and caudal-fin lengths to standard length (SL) of Rhinogobius rong (red circles), R. nami (light-blue circles), R. taenigena (deep-blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) collected from Da Nang City and Thua Thien Hue Province. Solid symbols represent males, open symbols represent females, and symbols filled with grey represent specimens of unknown sex (for R. similis). Symbols with an asterisk are holotypes." figureDoi="http://doi.org/10.5281/zenodo.13330565" httpUri="https://zenodo.org/record/13330565/files/figure.png" pageId="11" pageNumber="518">Fig. 2</figureCitation>
|
||
,
|
||
<tableCitation id="B3190692FFE37D57A250FAD5FD85FA9C" box="[527,613,1375,1401]" captionStart="TABLE 3" captionStartId="6.[151,239,440,464]" captionText="TABLE 3. Morphometrics of Rhinogobius rong and R. nami, expressed as percentages of standard length. D 1, first dorsal fin; D 2, second dorsal fin; A, anal fin; C, caudal fin; P 2, pelvic fin." pageId="11" pageNumber="518">Table 3</tableCitation>
|
||
). Upper and lower jaws usually with 2–4 rows of conical teeth; those in outermost row enlarged and those in inner rows irregularly aligned. Mental flap on chin usually round or rectangular, often with a small central projection. Anterior nostril short tubular, posterior nostril a pore. Cephalic sensory pore system with B´, C, D(S), E, F, H´, K´, and L´ in oculoscapular canal and M´, N, and O´ in preopercular canal of both left and right sides in 20 of
|
||
<specimenCount id="E89DF8A0FFE37D57A19BFA64FDBAF9EC" box="[452,602,1518,1545]" count="25" pageId="11" pageNumber="518" type="generic">25 specimens</specimenCount>
|
||
examined (including
|
||
<typeStatus id="21208D8BFFE37D57A314FA65FC51F9EC" box="[843,945,1519,1545]" pageId="11" pageNumber="518" type="holotype">holotype</typeStatus>
|
||
;
|
||
<figureCitation id="66A02FACFFE37D57A3E4FA65FBC5F9EC" box="[955,1061,1519,1545]" captionStart="FIGURE 3" captionStartId="4.[151,243,1970,1994]" captionTargetBox="[160,1429,786,1939]" captionTargetId="figure-313@4.[151,1436,777,1946]" captionTargetPageId="4" captionText="FIGURE 3. Arrangement of cephalic sensory pores and cutaneous sensory papillae in Rhinogobius rong (a–c, NSMT-P 147886) and Rhinogobius nami (d–f, NSMT-P 147890). a, d, lateral view; b, e, dorsal view; c, f, ventral view.AN, anterior naris; PN, posterior naris." figureDoi="http://doi.org/10.5281/zenodo.13330569" httpUri="https://zenodo.org/record/13330569/files/figure.png" pageId="11" pageNumber="518">Fig. 3d–f</figureCitation>
|
||
), but
|
||
<specimenCount id="E89DF8A0FFE37D57A43FFA65FAF3F9EC" box="[1120,1299,1519,1545]" count="3" pageId="11" pageNumber="518" type="generic">three specimens</specimenCount>
|
||
having pore G in one or both side(s), and
|
||
<specimenCount id="E89DF8A0FFE37D57A1BDF999FD69F9C8" box="[482,649,1555,1581]" count="2" pageId="11" pageNumber="518" type="generic">two specimens</specimenCount>
|
||
having an additional pore behind pore F (not G, but near F) in one or both side(s). Arrangement of cutaneous sensory papillae of head shown in
|
||
<figureCitation id="66A02FACFFE37D57A3B8F9BDFBB3F9B4" box="[999,1107,1591,1617]" captionStart="FIGURE 3" captionStartId="4.[151,243,1970,1994]" captionTargetBox="[160,1429,786,1939]" captionTargetId="figure-313@4.[151,1436,777,1946]" captionTargetPageId="4" captionText="FIGURE 3. Arrangement of cephalic sensory pores and cutaneous sensory papillae in Rhinogobius rong (a–c, NSMT-P 147886) and Rhinogobius nami (d–f, NSMT-P 147890). a, d, lateral view; b, e, dorsal view; c, f, ventral view.AN, anterior naris; PN, posterior naris." figureDoi="http://doi.org/10.5281/zenodo.13330569" httpUri="https://zenodo.org/record/13330569/files/figure.png" pageId="11" pageNumber="518">Fig. 3d–f</figureCitation>
|
||
. Cheek having no significant transverse row of papillae. Vertebrae 10+15=25 (n=1) or 10+16=26 (n=13;
|
||
<tableCitation id="B3190692FFE37D57A3BCF9D1FBD5F990" box="[995,1077,1627,1653]" captionStart="TABLE 1" captionStartId="5.[151,238,476,500]" captionText="TABLE 1. Number of vertebrae in various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="11" pageNumber="518">Table 1</tableCitation>
|
||
), P-V 3/II II I I 0/9 (n=14).
|
||
</paragraph>
|
||
<paragraph id="FE243329FFE37D50A098F9F5FEABFF30" blockId="11.[151,1437,403,2013]" lastBlockId="12.[151,1437,151,1977]" lastPageId="12" lastPageNumber="519" pageId="11" pageNumber="518">
|
||
First dorsal fin usually with six spines, but
|
||
<specimenCount id="E89DF8A0FFE37D57A2FEF9F5FCD9F97C" box="[673,825,1663,1689]" count="1" pageId="11" pageNumber="518" type="generic">one specimen</specimenCount>
|
||
with five. Second dorsal fin usually with one spine and eight soft rays, but
|
||
<specimenCount id="E89DF8A0FFE37D57A12EF929FDC5F958" box="[369,549,1699,1725]" count="3" pageId="11" pageNumber="518" type="generic">three specimens</specimenCount>
|
||
with one spine and seven soft rays (
|
||
<tableCitation id="B3190692FFE37D57A3E1F929FBF6F958" box="[958,1046,1699,1725]" captionStart="TABLE 2" captionStartId="5.[151,239,1153,1177]" captionText="TABLE 2. Number of soft rays on the second dorsal fin and anal fin of various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="11" pageNumber="518">Table 2</tableCitation>
|
||
). First and second dorsal-fin bases separated by a small interval (2.8–5.8% of SL;
|
||
<tableCitation id="B3190692FFE37D57A2EDF94DFCE8F904" box="[690,776,1735,1761]" captionStart="TABLE 3" captionStartId="6.[151,239,440,464]" captionText="TABLE 3. Morphometrics of Rhinogobius rong and R. nami, expressed as percentages of standard length. D 1, first dorsal fin; D 2, second dorsal fin; A, anal fin; C, caudal fin; P 2, pelvic fin." pageId="11" pageNumber="518">Table 3</tableCitation>
|
||
). In female, first dorsal fin rounded, almost semi-circular; posterior tip of fin (tips of fourth to sixth spines) not reaching second dorsal-fin origin. In male, first dorsal fin in parallelogram shape; posterior tip of fin (usually fourth spine) extending to, or exceeding base of second dorsal-fin spine, but never reaching base of second soft ray of second dorsal fin. Anal fin with one spine and seven (n=4) or eight (n=21) soft rays (
|
||
<tableCitation id="B3190692FFE37D57A1E5F8DDFDF3F894" box="[442,531,1879,1905]" captionStart="TABLE 2" captionStartId="5.[151,239,1153,1177]" captionText="TABLE 2. Number of soft rays on the second dorsal fin and anal fin of various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="11" pageNumber="518">Table 2</tableCitation>
|
||
). Caudal fin with 17 segmented rays including 12 (n=15), 13 (n=9), or 14 (n=1) branched rays; posterior margin rounded. Male having larger second dorsal, anal, and caudal fins than female (second dorsal-fin length 31.2–37.5 vs. 25.0–28.3% of SL, anal-fin length 27.6–30.1 vs. 23.5–27.2% of SL, and caudal-fin length 26.5–29.6 vs. 21.8–26.6% of SL;
|
||
<figureCitation id="66A02FACFFE37D57A2BCF849FCC8F838" box="[739,808,1987,2013]" captionStart="FIGURE 2" captionStartId="3.[151,250,1875,1899]" captionTargetBox="[158,1429,188,1844]" captionTargetId="figure-17@3.[151,1436,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Proportions of head length, upper-jaw length, first dorsal-, second dorsal-, anal-, and caudal-fin lengths to standard length (SL) of Rhinogobius rong (red circles), R. nami (light-blue circles), R. taenigena (deep-blue diamonds), R. leavelli (orange triangles), and R. similis (green squares) collected from Da Nang City and Thua Thien Hue Province. Solid symbols represent males, open symbols represent females, and symbols filled with grey represent specimens of unknown sex (for R. similis). Symbols with an asterisk are holotypes." figureDoi="http://doi.org/10.5281/zenodo.13330565" httpUri="https://zenodo.org/record/13330565/files/figure.png" pageId="11" pageNumber="518">Fig. 2</figureCitation>
|
||
,
|
||
<tableCitation id="B3190692FFE37D57A36AF849FC6BF838" box="[821,907,1987,2013]" captionStart="TABLE 3" captionStartId="6.[151,239,440,464]" captionText="TABLE 3. Morphometrics of Rhinogobius rong and R. nami, expressed as percentages of standard length. D 1, first dorsal fin; D 2, second dorsal fin; A, anal fin; C, caudal fin; P 2, pelvic fin." pageId="11" pageNumber="518">Table 3</tableCitation>
|
||
). Pectoral fin with 14–16 rays (mode 15 rays) (
|
||
<tableCitation id="B3190692FFE47D50A0C0FF1DFF16FF54" box="[159,246,151,177]" captionStart="TABLE 4" captionStartId="7.[151,237,151,175]" captionText="TABLE 4. Pectoral-fin ray counts of various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="12" pageNumber="519">Table 4</tableCitation>
|
||
). Pelvic fin with one spine and five soft rays; pelvic fins joined together to form a cuplike disc with fleshy bilobed frenum.
|
||
</paragraph>
|
||
<paragraph id="FE243329FFE47D50A098FF55FAC0FE10" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="519">
|
||
Ctenoid scales covering lateral, dorsal, and ventral surfaces of body posterior to flank except belly. Nape and posterior part of occipital region covered by cycloid scales; scaled area extending anteriorly around middle between posterior margin of preopercle and posterior margin of opercle both in male and female. Other regions of head naked. Belly and flank covered with cycloid scales. First and second dorsal, and anal-fin bases covered with ctenoid scales and a few small cycloid scales mixed. Proximal part of caudal fin covered with cycloid scales. Scales on breast (pre-pelvic area) difficult to detect, but cycloid scales observed in a few individuals. Pectoral-fin base probably naked (at least invisible without staining with alizarin red; see
|
||
<bibRefCitation id="9A0A4ED8FFE47D50A350FE3DFC01FE34" author="Suzuki, T. & Shibukawa, K. & Senou, H. & Chen, I. - S." box="[783,993,439,465]" pageId="12" pageNumber="519" pagination="227 - 238" refId="ref24564" refString="Suzuki, T., Shibukawa, K., Senou, H. & Chen, I. - S. (2016) Redescription of Rhinogobius similis Gill 1859 (Gobiidae: Gobionellinae), the type species of the genus Rhinogobius Gill 1859, with designation of the neotype. Ichthyological Research, 63, 227 - 238. https: // doi. org / 10.1007 / s 10228 - 015 - 0494 - 3" type="journal article" year="2016">
|
||
Suzuki
|
||
<emphasis id="CCEFEF3BFFE47D50A33BFE3DFC7DFE34" box="[868,925,439,465]" italics="true" pageId="12" pageNumber="519">et al.</emphasis>
|
||
2016
|
||
</bibRefCitation>
|
||
). Longitudinal scales 28–31, transverse scales eight or nine, transverse scales in caudal peduncle seven, and predorsal scales 6–11 (
|
||
<tableCitation id="B3190692FFE47D50A4CCFE51FAF3FE10" box="[1171,1299,474,501]" captionStart-0="TABLE 5" captionStart-1="TABLE 6" captionStart-2="TABLE 7" captionStartId-0="7.[151,234,868,892]" captionStartId-1="8.[151,236,151,175]" captionStartId-2="8.[151,237,868,892]" captionText-0="TABLE 5. Longitudinal scale counts of various species of Rhinogobius examined in the present study. * Includes the holotype." captionText-1="TABLE 6. Transverse scale counts of various species of Rhinogobius examined in the present study. *Includes the holotype." captionText-2="TABLE 7. Predorsal scale counts of various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="12" pageNumber="519">Tables 5–7</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FE243329FFE47D50A098FE75FB75FC64" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="519">
|
||
Color in life and fresh (
|
||
<figureCitation id="66A02FACFFE47D50A18BFE75FDC2FDFC" box="[468,546,511,537]" captionStart="FIGURE 7" captionStartId="13.[151,250,942,966]" captionTargetBox="[151,1435,181,918]" captionTargetId="figure-17@13.[151,1435,181,918]" captionTargetPageId="13" captionText="FIGURE 7. Rhinogobius nami immediately after fixation. a, NSMT-P 147890, holotype, male, 24.9 mm SL; b, OISTICH 282, paratype, female, 26.6 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13330585" httpUri="https://zenodo.org/record/13330585/files/figure.png" pageId="12" pageNumber="519">Figs. 7</figureCitation>
|
||
and
|
||
<figureCitation id="66A02FACFFE47D50A204FE75FD8BFDFC" box="[603,619,511,537]" captionStart="FIGURE 8" captionStartId="13.[151,250,1907,1931]" captionTargetBox="[151,1435,1027,1883]" captionTargetId="figure-43@13.[151,1435,1027,1883]" captionTargetPageId="13" captionText="FIGURE 8. Live images of Rhinogobius nami in aquarium (a) and in situ underwater footage from Da Nang City, Vietnam (b–d). a, male, 12 January 2013; b, male, 13 January 2013; c, d, females, 13 January 2013 (photo by K. Maeda)." figureDoi="http://doi.org/10.5281/zenodo.13330587" httpUri="https://zenodo.org/record/13330587/files/figure.png" pageId="12" pageNumber="519">8</figureCitation>
|
||
): in male, background of head and body cream or yellowish-brown, but whitish on ventral side. A clear, reddish-brown stripe across center of upper lip; stripe splitting to left and right sides at snout and extending to anterior margin of eyes and continuing posteriorly from posterior margin of eye to region above gill opening, but posterior half indistinct. Dorsal side of head above this stripe, grey or brown. Lower half of cheek and opercular region darker than upper half, sometimes strongly contrasting. Reddish-brown or dusky-brown spot at posterior part of upper lip. Three parallel, diagonally downward, reddish-brown or dusky-brown stripes on ventral part of cheek. Opercular membrane, interopercle, and lower part of subopercle light bluish-grey with many red or orange spots. Posterior part of lower lip also often light bluish-grey. Five longitudinal, reddish-brown or dusky-brown stripes (formed by dotted lines) running on lateral and dorsal sides from trunk to caudal peduncle; fourth one (counted from dorsal) slightly below lateral midline being most distinct and others sometimes unclear. Three to six indefinite dusky transverse bands sometimes appearing on lateral side of body.
|
||
</paragraph>
|
||
<paragraph id="FE243329FFE47D50A098FC01FC87FA78" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="519">
|
||
First dorsal fin with series of black or dusky-brown markings on middle parts of membranes between first to fifth spines; marking between first and second spines largest and clearest, and posterior ones sometimes unclear. Distal part of membrane light blue or white between first and second spines, and orange or red between second and fifth spines. Distal part of spines black or grey. Reddish-brown spots on proximal part of first to fourth spines; spines and membranes white between proximal reddish-brown spots and middle black or dusky-brown markings. Two or three indefinite reddish-brown stripes on membranes posterior to fifth spine and white between stripes (in most specimens, as in
|
||
<figureCitation id="66A02FACFFE47D50A10DFBE9FE41FB98" box="[338,417,1123,1149]" captionStart="FIGURE 8" captionStartId="13.[151,250,1907,1931]" captionTargetBox="[151,1435,1027,1883]" captionTargetId="figure-43@13.[151,1435,1027,1883]" captionTargetPageId="13" captionText="FIGURE 8. Live images of Rhinogobius nami in aquarium (a) and in situ underwater footage from Da Nang City, Vietnam (b–d). a, male, 12 January 2013; b, male, 13 January 2013; c, d, females, 13 January 2013 (photo by K. Maeda)." figureDoi="http://doi.org/10.5281/zenodo.13330587" httpUri="https://zenodo.org/record/13330587/files/figure.png" pageId="12" pageNumber="519">Fig. 8a</figureCitation>
|
||
, but stripes sometimes disordered, as in
|
||
<figureCitation id="66A02FACFFE47D50A301FBE9FC4DFB98" box="[862,941,1123,1149]" captionStart="FIGURE 7" captionStartId="13.[151,250,942,966]" captionTargetBox="[151,1435,181,918]" captionTargetId="figure-17@13.[151,1435,181,918]" captionTargetPageId="13" captionText="FIGURE 7. Rhinogobius nami immediately after fixation. a, NSMT-P 147890, holotype, male, 24.9 mm SL; b, OISTICH 282, paratype, female, 26.6 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13330585" httpUri="https://zenodo.org/record/13330585/files/figure.png" pageId="12" pageNumber="519">Fig. 7a</figureCitation>
|
||
). Second dorsal fin with three to six reddish-brown spots aligned along each spine and soft rays; membrane around these spots also reddish-brown and forming stripes across fin. Proximal and middle parts of spine, soft rays, and membranes white between reddish-brown spots. Distal margin of second dorsal fin white or light blue. Black, vertical rectangular or elliptical spot surrounded by cream and orange at middle of proximal part of caudal fin. Remainder of caudal fin white or translucent with many reddish-brown spots forming five to eight vertical bands, but ventral part orange or dark brown. Anal fin orange, but dusky brown distally with bluish-white margin. Pectoral fin greyish translucent or grey, but its proximal part light yellowish-brown or light grey with large black or reddish-brown blotch at middle of pectoral-fin base. Pelvic disc greyish translucent or dark grey with white or light blue margin.
|
||
</paragraph>
|
||
<paragraph id="FE243329FFE47D50A098FA2DFDC2F958" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="519">In female, markings and background color of head and body similar to those of male, but lacking three parallel, diagonally downward stripes from ventral part of cheek. Opercular membrane, interopercle, lower part of subopercle, and posterior part of lower lip yellowish-brown or dark grey without red spots. First dorsal fin translucent with one to five indefinite reddish-brown spots along each spine; proximal parts of spines white between reddish-brown spots; distal part of fin yellow. Markings and background of second dorsal, caudal, and pectoral fins similar to those of male, but white margin of second dorsal fin thinner and often indistinct, and ventral part of caudal fin translucent instead of orange. Anal fin translucent, but reddish or dark brown distally with thin, bluish-white margin. Pelvic disc translucent with white margin.</paragraph>
|
||
<paragraph id="FE243329FFE47D50A098F94DFD01F894" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="519">
|
||
Color in preservative (
|
||
<figureCitation id="66A02FACFFE47D50A193F94DFDDFF904" box="[460,575,1735,1761]" captionStart="FIGURE 6" captionStartId="10.[151,250,1224,1248]" captionTargetBox="[157,1427,189,1194]" captionTargetId="figure-18@10.[151,1436,181,1200]" captionTargetPageId="10" captionText="FIGURE 6. Preserved specimens of Rhinogobius rong (a, b), R. nami (c, d), R. taenigena (e, f), R. leavelli (g, h), and R. similis (i, j) collected from Da Nang City and Thua Thien Hue Province, Vietnam. Specimens on left side are males, right are females. a, NSMT-P 147886, holotype, 45.6 mm SL; b, OISTICH 254, paratype, 52.7 mm SL; c, NSMT-P 147890, holotype, 24.9 mm SL; d, OISTICH 290, paratype, 27.0 mm SL; e, OISTICH 299, 23.6 mm SL; f, OISTICH 305, 34.1mm SL; g, NSMT-P 147888, 43.8 mm SL; h, NSMT-P 147887, 41.1 mm SL; i, OISTICH 240, 56.3 mm SL; j, OISTICH 241, 45.2 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13330583" httpUri="https://zenodo.org/record/13330583/files/figure.png" pageId="12" pageNumber="519">Fig. 6c, d</figureCitation>
|
||
): background of head and body cream and that of fins translucent or light brown. Reddish-brown markings observed in life becoming dark-brown, and dusky-brown markings becoming black. Light bluish-grey color on opercular membrane, interopercle, and lower part of subopercle of male becoming dark brown, and red and orange spots becoming light brown (dark and pale pattern opposite from that in life). Red, orange, white, and bluish colors of fins disappeared.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B68160A2FFE47D52A098F8F1FBBAF8D8" lastPageId="14" lastPageNumber="521" pageId="12" pageNumber="519" type="etymology">
|
||
<paragraph id="FE243329FFE47D50A098F8F1FD94F85C" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="519">
|
||
<emphasis id="CCEFEF3BFFE47D50A098F8F1FEAEF870" bold="true" box="[199,334,1915,1941]" pageId="12" pageNumber="519">Etymology.</emphasis>
|
||
The specific name of the new species is dedicated to the late Chu Hoang Nam, who helped our surveys in
|
||
<collectingCountry id="868C73B9FFE47D50A14FF815FE8FF85C" box="[272,367,1951,1977]" name="Vietnam" pageId="12" pageNumber="519">Vietnam</collectingCountry>
|
||
during 2018 and 2019.
|
||
</paragraph>
|
||
<caption id="AAE463A1FFE57D51A0C8FC24FE32FC0F" ID-DOI="http://doi.org/10.5281/zenodo.13330585" ID-Zenodo-Dep="13330585" httpUri="https://zenodo.org/record/13330585/files/figure.png" pageId="13" pageNumber="520" startId="13.[151,250,942,966]" targetBox="[151,1435,181,918]" targetPageId="13" targetType="figure">
|
||
<paragraph id="FE243329FFE57D51A0C8FC24FE32FC0F" blockId="13.[151,1436,942,1003]" pageId="13" pageNumber="520">
|
||
<emphasis id="CCEFEF3BFFE57D51A0C8FC24FEF3FC23" bold="true" box="[151,275,942,966]" pageId="13" pageNumber="520">FIGURE 7.</emphasis>
|
||
<taxonomicName id="399B48AAFFE57D51A146FC24FE33FC23" authorityName="Maeda & Kobayashi & Iida & Tran" authorityYear="2024" box="[281,467,942,966]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="520" phylum="Chordata" rank="species" species="nami">
|
||
<emphasis id="CCEFEF3BFFE57D51A146FC24FE33FC23" box="[281,467,942,966]" italics="true" pageId="13" pageNumber="520">Rhinogobius nami</emphasis>
|
||
</taxonomicName>
|
||
immediately after fixation. a, NSMT-P 147890, holotype, male, 24.9 mm SL; b, OISTICH 282, paratype, female, 26.6 mm SL.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="AAE463A1FFE57D51A0C8F8F9FAE8F84A" ID-DOI="http://doi.org/10.5281/zenodo.13330587" ID-Zenodo-Dep="13330587" httpUri="https://zenodo.org/record/13330587/files/figure.png" pageId="13" pageNumber="520" startId="13.[151,250,1907,1931]" targetBox="[151,1435,1027,1883]" targetPageId="13" targetType="figure">
|
||
<paragraph id="FE243329FFE57D51A0C8F8F9FAE8F84A" blockId="13.[151,1437,1907,1967]" pageId="13" pageNumber="520">
|
||
<emphasis id="CCEFEF3BFFE57D51A0C8F8F9FEF5F86E" bold="true" box="[151,277,1907,1931]" pageId="13" pageNumber="520">FIGURE 8.</emphasis>
|
||
Live images of
|
||
<taxonomicName id="399B48AAFFE57D51A19EF8F9FD9CF86E" authorityName="Maeda & Kobayashi & Iida & Tran" authorityYear="2024" box="[449,636,1907,1931]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="520" phylum="Chordata" rank="species" species="nami">
|
||
<emphasis id="CCEFEF3BFFE57D51A19EF8F9FD9CF86E" box="[449,636,1907,1931]" italics="true" pageId="13" pageNumber="520">Rhinogobius nami</emphasis>
|
||
</taxonomicName>
|
||
in aquarium (a) and
|
||
<emphasis id="CCEFEF3BFFE57D51A306F8FEFC7AF86E" box="[857,922,1908,1931]" italics="true" pageId="13" pageNumber="520">in situ</emphasis>
|
||
underwater footage from Da Nang City, Vietnam (b–d). a, male, 12 January 2013; b, male, 13 January 2013; c, d, females, 13 January 2013 (photo by K. Maeda).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FE243329FFE67D52A098FF1CFECDFE6D" blockId="14.[151,1437,150,644]" pageId="14" pageNumber="521">
|
||
<emphasis id="CCEFEF3BFFE67D52A098FF1CFE82FF55" bold="true" box="[199,354,150,176]" pageId="14" pageNumber="521">Comparison.</emphasis>
|
||
<taxonomicName id="399B48AAFFE67D52A134FF1CFDDAFF55" authorityName="Maeda & Kobayashi & Iida & Tran" authorityYear="2024" box="[363,570,150,176]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="nami">
|
||
<emphasis id="CCEFEF3BFFE67D52A134FF1CFDDAFF55" box="[363,570,150,176]" italics="true" pageId="14" pageNumber="521">Rhinogobius nami</emphasis>
|
||
</taxonomicName>
|
||
shares many morphological features with
|
||
<taxonomicName id="399B48AAFFE67D52A47AFF1DFF2FFF31" authority="Chen & Miller, 2014" authorityName="Chen & Miller" authorityYear="2014" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="sangenloensis">
|
||
<emphasis id="CCEFEF3BFFE67D52A47AFF1DFB03FF55" box="[1061,1251,150,176]" italics="true" pageId="14" pageNumber="521">R. sangenloensis</emphasis>
|
||
<bibRefCitation id="9A0A4ED8FFE67D52A4B2FF1CFF2FFF31" author="Chen, I. - S. & Miller, P. J." pageId="14" pageNumber="521" pagination="124 - 129" refId="ref23034" refString="Chen, I. - S. & Miller, P. J. (2014) A new freshwater goby of Rhinogobius (Teleostei: Gobiidae) from Hainan Island, southern China. Journal of Marine Science and Technology, 21 (Supplement), 124 - 129. https: // doi. org / 10.6119 / JMST- 013 - 1219 - 10" type="journal article" year="2014">Chen & Miller, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="3C5FFDCBFFE67D52A145FF30FE8EFF31" box="[282,366,186,212]" country="China" name="Hainan" pageId="14" pageNumber="521">Hainan</collectingRegion>
|
||
,
|
||
<collectingCountry id="868C73B9FFE67D52A123FF30FE23FF31" box="[380,451,186,212]" name="China" pageId="14" pageNumber="521">China</collectingCountry>
|
||
. The coloration is nearly identical, including the longitudinal stripes on body and fin markings, but the male of new species has three reddish-brown or dusky-brown stripes on cheek (absent in
|
||
<taxonomicName id="399B48AAFFE67D52A5DBFF55FED4FEF9" authorityName="Chen & Miller" authorityYear="2014" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="sangenloensis">
|
||
<emphasis id="CCEFEF3BFFE67D52A5DBFF55FED4FEF9" italics="true" pageId="14" pageNumber="521">R. sangenloensis</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="9A0A4ED8FFE67D52A11CFE88FDCBFEF9" author="Chen, I. - S. & Miller, P. J." box="[323,555,258,284]" pageId="14" pageNumber="521" pagination="124 - 129" refId="ref23034" refString="Chen, I. - S. & Miller, P. J. (2014) A new freshwater goby of Rhinogobius (Teleostei: Gobiidae) from Hainan Island, southern China. Journal of Marine Science and Technology, 21 (Supplement), 124 - 129. https: // doi. org / 10.6119 / JMST- 013 - 1219 - 10" type="journal article" year="2014">Chen & Miller 2014</bibRefCitation>
|
||
), and the mode count of pectoral-fin rays is also different (
|
||
<quantity id="39639ECCFFE67D52A4E4FE88FB15FEF9" box="[1211,1269,258,285]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="14" pageNumber="521" unit="in" value="15.0">15 in</quantity>
|
||
<taxonomicName id="399B48AAFFE67D52A4A4FE89FAB2FEF9" authorityName="Maeda & Kobayashi & Iida & Tran" authorityYear="2024" box="[1275,1362,259,284]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="nami">
|
||
<emphasis id="CCEFEF3BFFE67D52A4A4FE89FAB2FEF9" box="[1275,1362,259,284]" italics="true" pageId="14" pageNumber="521">R. nami</emphasis>
|
||
</taxonomicName>
|
||
vs.
|
||
<quantity id="39639ECCFFE67D52A5DFFE88FF4DFEA5" metricMagnitude="-1" metricUnit="m" metricValue="4.318" pageId="14" pageNumber="521" unit="in" value="17.0">17 in</quantity>
|
||
<taxonomicName id="399B48AAFFE67D52A0EBFEADFE90FEA5" authorityName="Chen & Miller" authorityYear="2014" box="[180,368,294,320]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="sangenloensis">
|
||
<emphasis id="CCEFEF3BFFE67D52A0EBFEADFE90FEA5" box="[180,368,294,320]" italics="true" pageId="14" pageNumber="521">R. sangenloensis</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="9A0A4ED8FFE67D52A120FEACFD88FEA5" author="Chen, I. - S. & Miller, P. J." box="[383,616,294,320]" pageId="14" pageNumber="521" pagination="124 - 129" refId="ref23034" refString="Chen, I. - S. & Miller, P. J. (2014) A new freshwater goby of Rhinogobius (Teleostei: Gobiidae) from Hainan Island, southern China. Journal of Marine Science and Technology, 21 (Supplement), 124 - 129. https: // doi. org / 10.6119 / JMST- 013 - 1219 - 10" type="journal article" year="2014">Chen & Miller 2014</bibRefCitation>
|
||
). Both males and females of
|
||
<taxonomicName id="399B48AAFFE67D52A3F2FEADFBE4FEA5" authorityName="Maeda & Kobayashi & Iida & Tran" authorityYear="2024" box="[941,1028,295,320]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="nami">
|
||
<emphasis id="CCEFEF3BFFE67D52A3F2FEADFBE4FEA5" box="[941,1028,295,320]" italics="true" pageId="14" pageNumber="521">R. nami</emphasis>
|
||
</taxonomicName>
|
||
have a black, vertical rectangular or elliptical spot at the middle of caudal-fin base, while
|
||
<taxonomicName id="399B48AAFFE67D52A2A7FEC1FC56FE81" authorityName="Chen & Miller" authorityYear="2014" box="[760,950,330,356]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="sangenloensis">
|
||
<emphasis id="CCEFEF3BFFE67D52A2A7FEC1FC56FE81" box="[760,950,330,356]" italics="true" pageId="14" pageNumber="521">R. sangenloensis</emphasis>
|
||
</taxonomicName>
|
||
female has a black horizontal bar (
|
||
<bibRefCitation id="9A0A4ED8FFE67D52A51EFEC0FEC1FE6D" author="Chen, I. - S. & Miller, P. J." pageId="14" pageNumber="521" pagination="124 - 129" refId="ref23034" refString="Chen, I. - S. & Miller, P. J. (2014) A new freshwater goby of Rhinogobius (Teleostei: Gobiidae) from Hainan Island, southern China. Journal of Marine Science and Technology, 21 (Supplement), 124 - 129. https: // doi. org / 10.6119 / JMST- 013 - 1219 - 10" type="journal article" year="2014">Chen & Miller 2014</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FE243329FFE67D52A098FE18FE6EFD61" blockId="14.[151,1437,150,644]" pageId="14" pageNumber="521">
|
||
The three oblique stripes on the cheek of male specimens of
|
||
<taxonomicName id="399B48AAFFE67D52A32BFE19FC2BFE49" authorityName="Maeda & Kobayashi & Iida & Tran" authorityYear="2024" box="[884,971,403,428]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="nami">
|
||
<emphasis id="CCEFEF3BFFE67D52A32BFE19FC2BFE49" box="[884,971,403,428]" italics="true" pageId="14" pageNumber="521">R. nami</emphasis>
|
||
</taxonomicName>
|
||
is one of the features of the new species. Several Chinese and Vietnamese species of the genus also have oblique stripes on the cheek. The new species however differs from most of these species having a thinner and/or more numerous stripes (in
|
||
<taxonomicName id="399B48AAFFE67D52A4BFFE51FE68FDFD" authority="Huang & Chen, 2007" authorityName="Huang & Chen" authorityYear="2007" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="changtinensis">
|
||
<emphasis id="CCEFEF3BFFE67D52A4BFFE51FA7CFE11" box="[1248,1436,474,500]" italics="true" pageId="14" pageNumber="521">R. changtinensis</emphasis>
|
||
<bibRefCitation id="9A0A4ED8FFE67D52A0C8FE75FE68FDFD" author="Huang, S. - P. & Chen, I. - S." box="[151,392,510,537]" pageId="14" pageNumber="521" pagination="101 - 110" refId="ref23822" refString="Huang, S. - P. & Chen, I. - S. (2007) Three new species of Rhinogobius Gill, 1859 (Teleeostei: Gobiidae) from the Hanjiang Basin, southern China. Raffles Bulletin of Zoology, Supplement 14, 101 - 110." type="journal article" year="2007">Huang & Chen, 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="399B48AAFFE67D52A1CDFE75FC08FDFD" authority="Chen, Wang, Chen & Shao, 2022" authorityName="Chen, Wang, Chen & Shao" authorityYear="2022" box="[402,1000,510,537]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="lingtongyanensis">
|
||
<emphasis id="CCEFEF3BFFE67D52A1CDFE75FD8EFDFD" box="[402,622,510,536]" italics="true" pageId="14" pageNumber="521">R. lingtongyanensis</emphasis>
|
||
<bibRefCitation id="9A0A4ED8FFE67D52A22BFE74FC08FDFD" author="Chen, I. - S. & Wang, S. - C. & Chen, K. - Y. & Shao, K. - T." box="[628,1000,510,537]" pageId="14" pageNumber="521" pagination="18 - 28" refId="ref23302" refString="Chen, I. - S., Wang, S. - C., Chen, K. - Y. & Shao, K. - T. (2022 a) A new freshwater goby of Rhinogobius lingtongyanensis (Teleostei, Gobiidae) from the Dongshi river basin, Fujian Province, southeastern China. Zootaxa, 5189 (1), 18 - 28. https: // doi. org / 10.11646 / zootaxa. 5189.1.5" type="journal article" year="2022">Chen, Wang, Chen & Shao, 2022</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="399B48AAFFE67D52A3ADFE75FF32FDD9" authority="Chen, Cheng & Shao, 2008" authorityName="Chen, Cheng & Shao" authorityYear="2008" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="longyanensis">
|
||
<emphasis id="CCEFEF3BFFE67D52A3ADFE75FB43FDFD" box="[1010,1187,510,536]" italics="true" pageId="14" pageNumber="521">R. longyanensis</emphasis>
|
||
<bibRefCitation id="9A0A4ED8FFE67D52A4F6FE74FF32FDD9" author="Chen, I. - S. & Cheng, Y. - H. & Shao, K. - T." pageId="14" pageNumber="521" pagination="335 - 343" refId="ref23230" refString="Chen, I. - S., Cheng, Y. - H. & Shao, K. - T. (2008) A new species of Rhinogobius (Teleostei: Gobiidae) from the Julongjiang basin in Fujian Province, China. Ichthyological Research, 55, 335 - 343. https: // doi. org / 10.1007 / s 10228 - 008 - 0045 - 2" type="journal article" year="2008">Chen, Cheng & Shao, 2008</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="399B48AAFFE67D52A083FDA9FD9AFDD9" authority="Huang & Chen, 2007" authorityName="Huang & Chen" authorityYear="2007" box="[220,634,546,573]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="lungwoensis">
|
||
<emphasis id="CCEFEF3BFFE67D52A083FDA9FE64FDD9" box="[220,388,546,572]" italics="true" pageId="14" pageNumber="521">R. lungwoensis</emphasis>
|
||
<bibRefCitation id="9A0A4ED8FFE67D52A1D4FDA9FD9AFDD9" author="Huang, S. - P. & Chen, I. - S." box="[395,634,546,573]" pageId="14" pageNumber="521" pagination="101 - 110" refId="ref23822" refString="Huang, S. - P. & Chen, I. - S. (2007) Three new species of Rhinogobius Gill, 1859 (Teleeostei: Gobiidae) from the Hanjiang Basin, southern China. Raffles Bulletin of Zoology, Supplement 14, 101 - 110." type="journal article" year="2007">Huang & Chen, 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="399B48AAFFE67D52A2DBFDA9FBC9FDD9" authority="Chen & Miller, 2008" authorityName="Chen & Miller" authorityYear="2008" box="[644,1065,546,572]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="rubrolineatus">
|
||
<emphasis id="CCEFEF3BFFE67D52A2DBFDA9FCD9FDD9" box="[644,825,546,572]" italics="true" pageId="14" pageNumber="521">R. rubrolineatus</emphasis>
|
||
<bibRefCitation id="9A0A4ED8FFE67D52A31FFDA8FBC9FDD9" author="Chen, I. - S. & Miller, P. J." box="[832,1065,546,572]" pageId="14" pageNumber="521" pagination="225 - 232" refId="ref22979" refString="Chen, I. - S. & Miller, P. J. (2008) Two new freshwater gobies of genus Rhinogobius (Teleostei: Gobiidae) in southern China, around the northern region of the South China Sea. Raffles Bulletin of Zoology, Supplement 19, 225 - 232." type="journal article" year="2008">Chen & Miller, 2008</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="399B48AAFFE67D52A46CFDA9FA78FDD9" authority="Chen & Miller, 2008" authorityName="Chen & Miller" authorityYear="2008" box="[1075,1432,546,572]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="sagittus">
|
||
<emphasis id="CCEFEF3BFFE67D52A46CFDA9FB48FDD9" box="[1075,1192,547,572]" italics="true" pageId="14" pageNumber="521">R. sagittus</emphasis>
|
||
<bibRefCitation id="9A0A4ED8FFE67D52A4F0FDA8FA78FDD9" author="Chen, I. - S. & Miller, P. J." box="[1199,1432,546,572]" pageId="14" pageNumber="521" pagination="225 - 232" refId="ref22979" refString="Chen, I. - S. & Miller, P. J. (2008) Two new freshwater gobies of genus Rhinogobius (Teleostei: Gobiidae) in southern China, around the northern region of the South China Sea. Raffles Bulletin of Zoology, Supplement 19, 225 - 232." type="journal article" year="2008">Chen & Miller, 2008</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="399B48AAFFE67D52A097FDCDFDF8FD85" authority="Chen & Fang, 2006" authorityName="Chen & Fang" authorityYear="2006" box="[200,536,582,609]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="wangi">
|
||
<emphasis id="CCEFEF3BFFE67D52A097FDCDFECCFD85" box="[200,300,583,608]" italics="true" pageId="14" pageNumber="521">R. wangi</emphasis>
|
||
<bibRefCitation id="9A0A4ED8FFE67D52A16AFDCCFDF8FD85" author="Chen, I. - S. & Fang, L. - S." box="[309,536,582,609]" pageId="14" pageNumber="521" pagination="247 - 253" refId="ref22819" refString="Chen, I. - S. & Fang, L. - S. (2006) A new species of Rhinogobius (Teleostei: Gobiidae) from the Hanjiang Basin in Guangdong Province, China. Ichthyological Research, 53, 247 - 253. https: // doi. org / 10.1007 / s 10228 - 006 - 0342 - 6" type="journal article" year="2006">Chen & Fang, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
) (
|
||
<bibRefCitation id="9A0A4ED8FFE67D52A270FDCCFCEFFD85" author="Chen, I. - S. & Fang, L. - S." box="[559,783,582,609]" pageId="14" pageNumber="521" pagination="247 - 253" refId="ref22819" refString="Chen, I. - S. & Fang, L. - S. (2006) A new species of Rhinogobius (Teleostei: Gobiidae) from the Hanjiang Basin in Guangdong Province, China. Ichthyological Research, 53, 247 - 253. https: // doi. org / 10.1007 / s 10228 - 006 - 0342 - 6" type="journal article" year="2006">Chen & Fang 2006</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="9A0A4ED8FFE67D52A344FDCCFBE6FD85" author="Chen, I. - S. & Miller, P. J." box="[795,1030,582,608]" pageId="14" pageNumber="521" pagination="225 - 232" refId="ref22979" refString="Chen, I. - S. & Miller, P. J. (2008) Two new freshwater gobies of genus Rhinogobius (Teleostei: Gobiidae) in southern China, around the northern region of the South China Sea. Raffles Bulletin of Zoology, Supplement 19, 225 - 232." type="journal article" year="2008">Chen & Miller 2008</bibRefCitation>
|
||
; Chen
|
||
<emphasis id="CCEFEF3BFFE67D52A409FDCDFB70FD85" box="[1110,1168,582,608]" italics="true" pageId="14" pageNumber="521">et al.</emphasis>
|
||
2008, 2022a;
|
||
<bibRefCitation id="9A0A4ED8FFE67D52A56CFDCDFEF3FD61" author="Huang, S. - P. & Chen, I. - S." pageId="14" pageNumber="521" pagination="101 - 110" refId="ref23822" refString="Huang, S. - P. & Chen, I. - S. (2007) Three new species of Rhinogobius Gill, 1859 (Teleeostei: Gobiidae) from the Hanjiang Basin, southern China. Raffles Bulletin of Zoology, Supplement 14, 101 - 110." type="journal article" year="2007">Huang & Chen 2007</bibRefCitation>
|
||
) (
|
||
<tableCitation id="B3190692FFE67D52A176FDE0FE60FD61" box="[297,384,618,644]" captionStart="TABLE 8" captionStartId="14.[151,239,692,716]" captionText="TABLE 8. Various species of Rhinogobius with oblique stripes on cheeks." pageId="14" pageNumber="521">Table 8</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="AAE463A1FFE67D52A0C8FD3EFC51FD29" box="[151,945,692,716]" pageId="14" pageNumber="521" startId="14.[151,239,692,716]" targetType="table">
|
||
<paragraph id="FE243329FFE67D52A0C8FD3EFC51FD29" blockId="14.[151,945,692,716]" box="[151,945,692,716]" pageId="14" pageNumber="521">
|
||
<emphasis id="CCEFEF3BFFE67D52A0C8FD3EFEEBFD29" bold="true" box="[151,267,692,716]" pageId="14" pageNumber="521">TABLE 8.</emphasis>
|
||
Various species of
|
||
<taxonomicName id="399B48AAFFE67D52A186FD3EFD80FD29" box="[473,608,692,716]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="genus">
|
||
<emphasis id="CCEFEF3BFFE67D52A186FD3EFD80FD29" box="[473,608,692,716]" italics="true" pageId="14" pageNumber="521">Rhinogobius</emphasis>
|
||
</taxonomicName>
|
||
with oblique stripes on cheeks.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FE243329FFE67D52A12AFD56FB6EFB32" pageId="14" pageNumber="521">
|
||
<table id="8C9BC189FFE682A3A0FCFD56FA6AFB3D" box="[163,1418,732,1240]" gridcols="5" gridrows="13" pageId="14" pageNumber="521">
|
||
<tr id="40AB316BFFE682A3A0FCFD56FA6AFD11" box="[163,1418,732,756]" gridrow="0" pageId="14" pageNumber="521" rowspan-0="1">
|
||
<th id="037A5817FFE682A3A12AFD56FDCBFD11" box="[373,555,732,756]" gridcol="1" gridrow="0" pageId="14" pageNumber="521">Number of stripes</th>
|
||
<th id="037A5817FFE682A3A217FD56FD09FD11" box="[584,745,732,756]" gridcol="2" gridrow="0" pageId="14" pageNumber="521">Stripe thickness</th>
|
||
<th id="037A5817FFE682A3A35DFD56FBBCFD11" box="[770,1116,732,756]" gridcol="3" gridrow="0" pageId="14" pageNumber="521">Position of stripe</th>
|
||
<th id="037A5817FFE682A3A42AFD56FA6AFD11" box="[1141,1418,732,756]" gridcol="4" gridrow="0" pageId="14" pageNumber="521">Number of vertebrae (mode)</th>
|
||
</tr>
|
||
<tr id="40AB316BFFE682A3A0FCFC8EFA6AFCF8" box="[163,1418,772,797]" gridrow="1" pageId="14" pageNumber="521">
|
||
<th id="037A5817FFE682A3A0FCFC8EFEBDFCF8" box="[163,349,772,797]" gridcol="0" gridrow="1" pageId="14" pageNumber="521">
|
||
<taxonomicName id="399B48AAFFE67D52A0FCFC8FFF12FCF9" authorityName="Maeda & Kobayashi & Iida & Tran" authorityYear="2024" box="[163,242,773,796]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="nami" status="n. sp.">
|
||
<emphasis id="CCEFEF3BFFE67D52A0FCFC8FFF12FCF9" box="[163,242,773,796]" italics="true" pageId="14" pageNumber="521">R. nami</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D7DC5240FFE67D52A0A7FC8FFECEFCF9" box="[248,302,773,796]" pageId="14" pageNumber="521" rank="species">n. sp.</taxonomicNameLabel>
|
||
</th>
|
||
<td id="037A5817FFE682A3A12AFC8EFDCBFCF8" box="[373,555,772,797]" gridcol="1" gridrow="1" pageId="14" pageNumber="521">3</td>
|
||
<td id="037A5817FFE682A3A217FC8EFD09FCF8" box="[584,745,772,797]" gridcol="2" gridrow="1" pageId="14" pageNumber="521">Broad</td>
|
||
<td id="037A5817FFE682A3A35DFC8EFBBCFCF8" box="[770,1116,772,797]" gridcol="3" gridrow="1" pageId="14" pageNumber="521">Limited to lower half of the cheek</td>
|
||
<td id="037A5817FFE682A3A42AFC8EFA6AFCF8" box="[1141,1418,772,797]" gridcol="4" gridrow="1" pageId="14" pageNumber="521">26</td>
|
||
</tr>
|
||
<tr id="40AB316BFFE682A3A0FCFCA7FA6AFCA0" box="[163,1418,813,837]" gridrow="2" pageId="14" pageNumber="521">
|
||
<th id="037A5817FFE682A3A0FCFCA7FEBDFCA0" box="[163,349,813,837]" gridcol="0" gridrow="2" pageId="14" pageNumber="521">
|
||
<taxonomicName id="399B48AAFFE67D52A0FCFCA7FEDCFCA0" authorityName="Huang & Chen" authorityYear="2007" box="[163,316,813,837]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="lungwoensis">
|
||
<emphasis id="CCEFEF3BFFE67D52A0FCFCA7FEDCFCA0" box="[163,316,813,837]" italics="true" pageId="14" pageNumber="521">R. lungwoensis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="037A5817FFE682A3A12AFCA7FDCBFCA0" box="[373,555,813,837]" gridcol="1" gridrow="2" pageId="14" pageNumber="521">1</td>
|
||
<td id="037A5817FFE682A3A217FCA7FD09FCA0" box="[584,745,813,837]" gridcol="2" gridrow="2" pageId="14" pageNumber="521">Slightly broad</td>
|
||
<td id="037A5817FFE682A3A35DFCA7FBBCFCA0" box="[770,1116,813,837]" gridcol="3" gridrow="2" pageId="14" pageNumber="521">Limited to lower half of the cheek</td>
|
||
<td id="037A5817FFE682A3A42AFCA7FA6AFCA0" box="[1141,1418,813,837]" gridcol="4" gridrow="2" pageId="14" pageNumber="521">28</td>
|
||
</tr>
|
||
<tr id="40AB316BFFE682A3A0FCFCDFFA6AFC88" box="[163,1418,853,877]" gridrow="3" pageId="14" pageNumber="521">
|
||
<th id="037A5817FFE682A3A0FCFCDFFEBDFC88" box="[163,349,853,877]" gridcol="0" gridrow="3" pageId="14" pageNumber="521">
|
||
<taxonomicName id="399B48AAFFE67D52A0FCFCDCFEA8FC88" authorityName="Chen & Miller" authorityYear="2008" box="[163,328,853,877]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="rubrolineatus">
|
||
<emphasis id="CCEFEF3BFFE67D52A0FCFCDCFEA8FC88" box="[163,328,853,877]" italics="true" pageId="14" pageNumber="521">R. rubrolineatus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="037A5817FFE682A3A12AFCDFFDCBFC88" box="[373,555,853,877]" gridcol="1" gridrow="3" pageId="14" pageNumber="521">2</td>
|
||
<td id="037A5817FFE682A3A217FCDFFD09FC88" box="[584,745,853,877]" gridcol="2" gridrow="3" pageId="14" pageNumber="521">Thin</td>
|
||
<td id="037A5817FFE682A3A35DFCDFFBBCFC88" box="[770,1116,853,877]" gridcol="3" gridrow="3" pageId="14" pageNumber="521">Reaching upper half of the cheek</td>
|
||
<td id="037A5817FFE682A3A42AFCDFFA6AFC88" box="[1141,1418,853,877]" gridcol="4" gridrow="3" pageId="14" pageNumber="521">26</td>
|
||
</tr>
|
||
<tr id="40AB316BFFE682A3A0FCFCF7FA6AFC70" box="[163,1418,893,917]" gridrow="4" pageId="14" pageNumber="521">
|
||
<th id="037A5817FFE682A3A0FCFCF7FEBDFC70" box="[163,349,893,917]" gridcol="0" gridrow="4" pageId="14" pageNumber="521">
|
||
<taxonomicName id="399B48AAFFE67D52A0FCFCF4FEAAFC70" authorityName="Huang & Chen" authorityYear="2007" box="[163,330,893,917]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="changtinensis">
|
||
<emphasis id="CCEFEF3BFFE67D52A0FCFCF4FEAAFC70" box="[163,330,893,917]" italics="true" pageId="14" pageNumber="521">R. changtinensis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="037A5817FFE682A3A12AFCF7FDCBFC70" box="[373,555,893,917]" gridcol="1" gridrow="4" pageId="14" pageNumber="521">3</td>
|
||
<td id="037A5817FFE682A3A217FCF7FD09FC70" box="[584,745,893,917]" gridcol="2" gridrow="4" pageId="14" pageNumber="521">Thin</td>
|
||
<td id="037A5817FFE682A3A35DFCF7FBBCFC70" box="[770,1116,893,917]" gridcol="3" gridrow="4" pageId="14" pageNumber="521">Reaching upper half of the cheek</td>
|
||
<td id="037A5817FFE682A3A42AFCF7FA6AFC70" box="[1141,1418,893,917]" gridcol="4" gridrow="4" pageId="14" pageNumber="521">27</td>
|
||
</tr>
|
||
<tr id="40AB316BFFE682A3A0FCFC2FFA6AFC5B" box="[163,1418,933,958]" gridrow="5" pageId="14" pageNumber="521">
|
||
<th id="037A5817FFE682A3A0FCFC2FFEBDFC5B" box="[163,349,933,958]" gridcol="0" gridrow="5" pageId="14" pageNumber="521">
|
||
<taxonomicName id="399B48AAFFE67D52A0FCFC2CFEA3FC5B" authorityName="Chen, Cheng & Shao" authorityYear="2008" box="[163,323,934,958]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="longyanensis">
|
||
<emphasis id="CCEFEF3BFFE67D52A0FCFC2CFEA3FC5B" box="[163,323,934,958]" italics="true" pageId="14" pageNumber="521">R. longyanensis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="037A5817FFE682A3A12AFC2FFDCBFC5B" box="[373,555,933,958]" gridcol="1" gridrow="5" pageId="14" pageNumber="521">3</td>
|
||
<td id="037A5817FFE682A3A217FC2FFD09FC5B" box="[584,745,933,958]" gridcol="2" gridrow="5" pageId="14" pageNumber="521">Thin</td>
|
||
<td id="037A5817FFE682A3A35DFC2FFBBCFC5B" box="[770,1116,933,958]" gridcol="3" gridrow="5" pageId="14" pageNumber="521">Limited to lower half of the cheek</td>
|
||
<td id="037A5817FFE682A3A42AFC2FFA6AFC5B" box="[1141,1418,933,958]" gridcol="4" gridrow="5" pageId="14" pageNumber="521">27</td>
|
||
</tr>
|
||
<tr id="40AB316BFFE682A3A0FCFC44FA6AFC03" box="[163,1418,974,998]" gridrow="6" pageId="14" pageNumber="521">
|
||
<th id="037A5817FFE682A3A0FCFC44FEBDFC03" box="[163,349,974,998]" gridcol="0" gridrow="6" pageId="14" pageNumber="521">
|
||
<taxonomicName id="399B48AAFFE67D52A0FCFC45FEBDFC03" authorityName="Chen, Wang, Chen & Shao" authorityYear="2022" box="[163,349,974,998]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="lingtongyanensis">
|
||
<emphasis id="CCEFEF3BFFE67D52A0FCFC45FEBDFC03" box="[163,349,974,998]" italics="true" pageId="14" pageNumber="521">R. lingtongyanensis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="037A5817FFE682A3A12AFC44FDCBFC03" box="[373,555,974,998]" gridcol="1" gridrow="6" pageId="14" pageNumber="521">4</td>
|
||
<td id="037A5817FFE682A3A217FC44FD09FC03" box="[584,745,974,998]" gridcol="2" gridrow="6" pageId="14" pageNumber="521">Thin</td>
|
||
<td id="037A5817FFE682A3A35DFC44FBBCFC03" box="[770,1116,974,998]" gridcol="3" gridrow="6" pageId="14" pageNumber="521">Limited to lower half of the cheek</td>
|
||
<td id="037A5817FFE682A3A42AFC44FA6AFC03" box="[1141,1418,974,998]" gridcol="4" gridrow="6" pageId="14" pageNumber="521">26</td>
|
||
</tr>
|
||
<tr id="40AB316BFFE682A3A0FCFC7CFA6AFBEB" box="[163,1418,1014,1038]" gridrow="7" pageId="14" pageNumber="521">
|
||
<th id="037A5817FFE682A3A0FCFC7CFEBDFBEB" box="[163,349,1014,1038]" gridcol="0" gridrow="7" pageId="14" pageNumber="521">
|
||
<taxonomicName id="399B48AAFFE67D52A0FCFC7DFEEEFBEB" authorityName="Chen & Miller" authorityYear="2008" box="[163,270,1015,1038]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="sagittus">
|
||
<emphasis id="CCEFEF3BFFE67D52A0FCFC7DFEEEFBEB" box="[163,270,1015,1038]" italics="true" pageId="14" pageNumber="521">R. sagittus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="037A5817FFE682A3A12AFC7CFDCBFBEB" box="[373,555,1014,1038]" gridcol="1" gridrow="7" pageId="14" pageNumber="521">4–5</td>
|
||
<td id="037A5817FFE682A3A217FC7CFD09FBEB" box="[584,745,1014,1038]" gridcol="2" gridrow="7" pageId="14" pageNumber="521">Thin</td>
|
||
<td id="037A5817FFE682A3A35DFC7CFBBCFBEB" box="[770,1116,1014,1038]" gridcol="3" gridrow="7" pageId="14" pageNumber="521">Reaching upper half of the cheek</td>
|
||
<td id="037A5817FFE682A3A42AFC7CFA6AFBEB" box="[1141,1418,1014,1038]" gridcol="4" gridrow="7" pageId="14" pageNumber="521">26</td>
|
||
</tr>
|
||
<tr id="40AB316BFFE682A3A0FCFB94FA6AFBD3" box="[163,1418,1054,1078]" gridrow="8" pageId="14" pageNumber="521">
|
||
<th id="037A5817FFE682A3A0FCFB94FEBDFBD3" box="[163,349,1054,1078]" gridcol="0" gridrow="8" pageId="14" pageNumber="521">
|
||
<taxonomicName id="399B48AAFFE67D52A0FCFB95FF1DFBD3" authorityName="Chen & Fang" authorityYear="2006" box="[163,253,1055,1078]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="wangi">
|
||
<emphasis id="CCEFEF3BFFE67D52A0FCFB95FF1DFBD3" box="[163,253,1055,1078]" italics="true" pageId="14" pageNumber="521">R. wangi</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="037A5817FFE682A3A12AFB94FDCBFBD3" box="[373,555,1054,1078]" gridcol="1" gridrow="8" pageId="14" pageNumber="521">modally 7</td>
|
||
<td id="037A5817FFE682A3A217FB94FD09FBD3" box="[584,745,1054,1078]" gridcol="2" gridrow="8" pageId="14" pageNumber="521">Thin</td>
|
||
<td id="037A5817FFE682A3A35DFB94FBBCFBD3" box="[770,1116,1054,1078]" gridcol="3" gridrow="8" pageId="14" pageNumber="521">Reaching upper half of the cheek</td>
|
||
<td id="037A5817FFE682A3A42AFB94FA6AFBD3" box="[1141,1418,1054,1078]" gridcol="4" gridrow="8" pageId="14" pageNumber="521">27</td>
|
||
</tr>
|
||
<tr id="40AB316BFFE682A3A0FCFBCDFA6AFBBA" box="[163,1418,1095,1119]" gridrow="9" pageId="14" pageNumber="521">
|
||
<th id="037A5817FFE682A3A0FCFBCDFEBDFBBA" box="[163,349,1095,1119]" gridcol="0" gridrow="9" pageId="14" pageNumber="521">
|
||
<taxonomicName id="399B48AAFFE67D52A0FCFBCDFEF2FBBA" authorityName="Chen & Kottelat" authorityYear="2005" box="[163,274,1095,1119]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="sulcatus">
|
||
<emphasis id="CCEFEF3BFFE67D52A0FCFBCDFEF2FBBA" box="[163,274,1095,1119]" italics="true" pageId="14" pageNumber="521">R. sulcatus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="037A5817FFE682A3A12AFBCDFDCBFBBA" box="[373,555,1095,1119]" gridcol="1" gridrow="9" pageId="14" pageNumber="521">3–4</td>
|
||
<td id="037A5817FFE682A3A217FBCDFD09FBBA" box="[584,745,1095,1119]" gridcol="2" gridrow="9" pageId="14" pageNumber="521">Broad</td>
|
||
<td id="037A5817FFE682A3A35DFBCDFBBCFBBA" box="[770,1116,1095,1119]" gridcol="3" gridrow="9" pageId="14" pageNumber="521">Reaching upper half of the cheek</td>
|
||
<td id="037A5817FFE682A3A42AFBCDFA6AFBBA" box="[1141,1418,1095,1119]" gridcol="4" gridrow="9" pageId="14" pageNumber="521">27</td>
|
||
</tr>
|
||
<tr id="40AB316BFFE682A3A0FCFBE5FA6AFB62" box="[163,1418,1135,1159]" gridrow="10" pageId="14" pageNumber="521">
|
||
<th id="037A5817FFE682A3A0FCFBE5FEBDFB62" box="[163,349,1135,1159]" gridcol="0" gridrow="10" pageId="14" pageNumber="521">
|
||
<taxonomicName id="399B48AAFFE67D52A0FCFBFAFEC0FB62" baseAuthorityName="Herre" baseAuthorityYear="1935" box="[163,288,1135,1159]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="duospilus">
|
||
<emphasis id="CCEFEF3BFFE67D52A0FCFBFAFEC0FB62" box="[163,288,1135,1159]" italics="true" pageId="14" pageNumber="521">R. duospilus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="037A5817FFE682A3A12AFBE5FDCBFB62" box="[373,555,1135,1159]" gridcol="1" gridrow="10" pageId="14" pageNumber="521">1–3</td>
|
||
<td id="037A5817FFE682A3A217FBE5FD09FB62" box="[584,745,1135,1159]" gridcol="2" gridrow="10" pageId="14" pageNumber="521">Thin or broad</td>
|
||
<td id="037A5817FFE682A3A35DFBE5FBBCFB62" box="[770,1116,1135,1159]" gridcol="3" gridrow="10" pageId="14" pageNumber="521">Reaching upper half of the cheek</td>
|
||
<td id="037A5817FFE682A3A42AFBE5FA6AFB62" box="[1141,1418,1135,1159]" gridcol="4" gridrow="10" pageId="14" pageNumber="521">27</td>
|
||
</tr>
|
||
<tr id="40AB316BFFE682A3A0FCFB1DFA6AFB4A" box="[163,1418,1175,1199]" gridrow="11" pageId="14" pageNumber="521">
|
||
<th id="037A5817FFE682A3A0FCFB1DFEBDFB4A" box="[163,349,1175,1199]" gridcol="0" gridrow="11" pageId="14" pageNumber="521">
|
||
<taxonomicName id="399B48AAFFE67D52A0FCFB12FEDCFB4A" authorityName="Huang & Chen" authorityYear="2007" box="[163,316,1175,1199]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="ponkouensis">
|
||
<emphasis id="CCEFEF3BFFE67D52A0FCFB12FEDCFB4A" box="[163,316,1175,1199]" italics="true" pageId="14" pageNumber="521">R. ponkouensis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="037A5817FFE682A3A12AFB1DFDCBFB4A" box="[373,555,1175,1199]" gridcol="1" gridrow="11" pageId="14" pageNumber="521">2</td>
|
||
<td id="037A5817FFE682A3A217FB1DFD09FB4A" box="[584,745,1175,1199]" gridcol="2" gridrow="11" pageId="14" pageNumber="521">Broad</td>
|
||
<td id="037A5817FFE682A3A35DFB1DFBBCFB4A" box="[770,1116,1175,1199]" gridcol="3" gridrow="11" pageId="14" pageNumber="521">Reaching upper half of the cheek</td>
|
||
<td id="037A5817FFE682A3A42AFB1DFA6AFB4A" box="[1141,1418,1175,1199]" gridcol="4" gridrow="11" pageId="14" pageNumber="521">28</td>
|
||
</tr>
|
||
<tr id="40AB316BFFE682A3A0FCFB35FA6AFB3D" box="[163,1418,1215,1240]" gridrow="12" pageId="14" pageNumber="521">
|
||
<th id="037A5817FFE682A3A0FCFB35FEBDFB3D" box="[163,349,1215,1240]" gridcol="0" gridrow="12" pageId="14" pageNumber="521">
|
||
<taxonomicName id="399B48AAFFE67D52A0FCFB4AFEFFFB32" authorityName="Chen & Kottelat" authorityYear="2005" box="[163,287,1216,1239]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="virgigena">
|
||
<emphasis id="CCEFEF3BFFE67D52A0FCFB4AFEFFFB32" box="[163,287,1216,1239]" italics="true" pageId="14" pageNumber="521">R. virgigena</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="037A5817FFE682A3A12AFB35FDCBFB3D" box="[373,555,1215,1240]" gridcol="1" gridrow="12" pageId="14" pageNumber="521">1–3</td>
|
||
<td id="037A5817FFE682A3A217FB35FD09FB3D" box="[584,745,1215,1240]" gridcol="2" gridrow="12" pageId="14" pageNumber="521">Thin or broad</td>
|
||
<td id="037A5817FFE682A3A35DFB35FBBCFB3D" box="[770,1116,1215,1240]" gridcol="3" gridrow="12" pageId="14" pageNumber="521">Limited to lower half of the cheek</td>
|
||
<td id="037A5817FFE682A3A42AFB35FA6AFB3D" box="[1141,1418,1215,1240]" gridcol="4" gridrow="12" pageId="14" pageNumber="521">26</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="FE243329FFE67D52A098FA8DFE24FA30" blockId="14.[151,1437,1287,1854]" pageId="14" pageNumber="521">
|
||
Male of
|
||
<taxonomicName id="399B48AAFFE67D52A177FA82FD4CFAC7" authority="Chen & Kottelat, 2005" authorityName="Chen & Kottelat" authorityYear="2005" box="[296,684,1287,1314]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="sulcatus">
|
||
<emphasis id="CCEFEF3BFFE67D52A177FA82FE43FAC4" box="[296,419,1287,1313]" italics="true" pageId="14" pageNumber="521">R. sulcatus</emphasis>
|
||
<bibRefCitation id="9A0A4ED8FFE67D52A1F4FA8DFD4CFAC7" author="Chen, I. - S. & Kottelat, M." box="[427,684,1287,1314]" pageId="14" pageNumber="521" pagination="1407 - 1429" refId="ref22926" refString="Chen, I. - S. & Kottelat, M. (2005) Four new freshwater gobies of the genus Rhinogobius (Teleostei: Gobiidae) from northern Vietnam. Journal of Natural History, 39, 1407 - 1429. https: // doi. org / 10.1080 / 00222930400008736" type="journal article" year="2005">Chen & Kottelat, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
has three broad stripes similar to those of the new species, but the stripes are longer (often reaching upper half of cheek) (
|
||
<figureCitation id="66A02FACFFE67D52A35FFAA1FCB1FAA0" box="[768,849,1323,1349]" captionStart="FIGURE 9" captionStartId="16.[151,250,207,231]" captionTargetBox="[147,1441,181,1982]" captionTargetId="figure-17@15.[147,1441,181,1954]" captionTargetPageId="15" captionText="FIGURE 9. Comparative material of the genus Rhinogobius from northern Vietnam (a–l) and Japan (m–p) observed in the present study. Photographs were taken immediately after fixation. Specimens on left side are males, right are females. a, b, R. sulcatus (a, OISTICH 184, 30.4 mm SL; b, OISTICH 190, 32.4 mm SL); c, d, R. boa (c, OISTICH 161, 25.4 mm SL; d, OISTICH 160, 28.0 mm SL); e, f, R. virgigena (e, NSMT-P 147872, 36.1 mm SL; f, OISTICH 154, 33.7 mm SL); g, h, R. variolatus (g, OISTICH 169, 31.4 mm SL; h, OISTICH 170, 28.8 mm SL); i, j, R. leavelli (i, OISTICH 209, 38.1 mm SL; j, OISTICH 210, 35.4 mm SL); k, l, R. similis (k, OISTICH 222, 54.4 mm SL; l, NSMT-P 147882, 44.7 mm SL); m, n, R. similis from Okinawa Island (m, URM-P 49149, 48.6 mm SL; n, OISTICH 2, 63.6 mm SL); o, p, R. similis, from Izu Peninsula (o, OISTICH 330, 47.6 mm SL; p, OISTICH 332, 51.0 mm SL)." figureDoi="http://doi.org/10.5281/zenodo.13688016" httpUri="https://zenodo.org/record/13688016/files/figure.png" pageId="14" pageNumber="521">Fig. 9a</figureCitation>
|
||
), and often branched and connected with each other, forming a reticulate pattern.
|
||
<taxonomicName id="399B48AAFFE67D52A18DFAC5FD22FA8C" authorityName="Chen & Kottelat" authorityYear="2005" box="[466,706,1359,1385]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="sulcatus">
|
||
<emphasis id="CCEFEF3BFFE67D52A18DFAC5FD22FA8C" box="[466,706,1359,1385]" italics="true" pageId="14" pageNumber="521">Rhinogobius sulcatus</emphasis>
|
||
</taxonomicName>
|
||
also differs from the new species in lacking longitudinal, reddish-brown or dusky-brown stripes on sides of the body (
|
||
<figureCitation id="66A02FACFFE67D52A2BFFAF9FCABFA68" box="[736,843,1395,1421]" captionStart="FIGURE 9" captionStartId="16.[151,250,207,231]" captionTargetBox="[147,1441,181,1982]" captionTargetId="figure-17@15.[147,1441,181,1954]" captionTargetPageId="15" captionText="FIGURE 9. Comparative material of the genus Rhinogobius from northern Vietnam (a–l) and Japan (m–p) observed in the present study. Photographs were taken immediately after fixation. Specimens on left side are males, right are females. a, b, R. sulcatus (a, OISTICH 184, 30.4 mm SL; b, OISTICH 190, 32.4 mm SL); c, d, R. boa (c, OISTICH 161, 25.4 mm SL; d, OISTICH 160, 28.0 mm SL); e, f, R. virgigena (e, NSMT-P 147872, 36.1 mm SL; f, OISTICH 154, 33.7 mm SL); g, h, R. variolatus (g, OISTICH 169, 31.4 mm SL; h, OISTICH 170, 28.8 mm SL); i, j, R. leavelli (i, OISTICH 209, 38.1 mm SL; j, OISTICH 210, 35.4 mm SL); k, l, R. similis (k, OISTICH 222, 54.4 mm SL; l, NSMT-P 147882, 44.7 mm SL); m, n, R. similis from Okinawa Island (m, URM-P 49149, 48.6 mm SL; n, OISTICH 2, 63.6 mm SL); o, p, R. similis, from Izu Peninsula (o, OISTICH 330, 47.6 mm SL; p, OISTICH 332, 51.0 mm SL)." figureDoi="http://doi.org/10.5281/zenodo.13688016" httpUri="https://zenodo.org/record/13688016/files/figure.png" pageId="14" pageNumber="521">Fig. 9a, b</figureCitation>
|
||
) (five stripes in male and female of
|
||
<taxonomicName id="399B48AAFFE67D52A4BDFAFEFAD9FA68" authorityName="Maeda & Kobayashi & Iida & Tran" authorityYear="2024" box="[1250,1337,1396,1421]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="nami">
|
||
<emphasis id="CCEFEF3BFFE67D52A4BDFAFEFAD9FA68" box="[1250,1337,1396,1421]" italics="true" pageId="14" pageNumber="521">R. nami</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="66A02FACFFE67D52A518FAF9FF47FA54" captionStart="FIGURE 7" captionStartId="13.[151,250,942,966]" captionTargetBox="[151,1435,181,918]" captionTargetId="figure-17@13.[151,1435,181,918]" captionTargetPageId="13" captionText="FIGURE 7. Rhinogobius nami immediately after fixation. a, NSMT-P 147890, holotype, male, 24.9 mm SL; b, OISTICH 282, paratype, female, 26.6 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13330585" httpUri="https://zenodo.org/record/13330585/files/figure.png" pageId="14" pageNumber="521">Fig. 7a, b</figureCitation>
|
||
), seven soft rays on anal fin (vs. usually eight in
|
||
<taxonomicName id="399B48AAFFE67D52A28CFA12FCCBFA54" authorityName="Maeda & Kobayashi & Iida & Tran" authorityYear="2024" box="[723,811,1432,1457]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="nami">
|
||
<emphasis id="CCEFEF3BFFE67D52A28CFA12FCCBFA54" box="[723,811,1432,1457]" italics="true" pageId="14" pageNumber="521">R. nami</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<tableCitation id="B3190692FFE67D52A365FA1DFC6FFA54" box="[826,911,1431,1457]" captionStart="TABLE 2" captionStartId="5.[151,239,1153,1177]" captionText="TABLE 2. Number of soft rays on the second dorsal fin and anal fin of various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="14" pageNumber="521">Table 2</tableCitation>
|
||
), 16 pectoral-fin rays (vs. 15;
|
||
<tableCitation id="B3190692FFE67D52A4BAFA1DFADAFA54" box="[1253,1338,1431,1457]" captionStart="TABLE 4" captionStartId="7.[151,237,151,175]" captionText="TABLE 4. Pectoral-fin ray counts of various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="14" pageNumber="521">Table 4</tableCitation>
|
||
), and 27 vertebrae (vs. 26;
|
||
<tableCitation id="B3190692FFE67D52A13EFA31FE53FA30" box="[353,435,1467,1493]" captionStart="TABLE 1" captionStartId="5.[151,238,476,500]" captionText="TABLE 1. Number of vertebrae in various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="14" pageNumber="521">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FE243329FFE67D52A098FA55FAF0F96C" blockId="14.[151,1437,1287,1854]" pageId="14" pageNumber="521">
|
||
Males of
|
||
<taxonomicName id="399B48AAFFE67D52A16DFA6AFDBFFA1F" authority="(Herre, 1935)" baseAuthorityName="Herre" baseAuthorityYear="1935" box="[306,607,1503,1530]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="duospilus">
|
||
<emphasis id="CCEFEF3BFFE67D52A16DFA6AFE5CFA1C" box="[306,444,1503,1529]" italics="true" pageId="14" pageNumber="521">R. duospilus</emphasis>
|
||
(Herre, 1935)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="399B48AAFFE67D52A234FA6AFBEFFA1C" authority="Huang & Chen, 2007" authorityName="Huang & Chen" authorityYear="2007" box="[619,1039,1503,1530]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="ponkouensis">
|
||
<emphasis id="CCEFEF3BFFE67D52A234FA6AFCF4FA1C" box="[619,788,1503,1529]" italics="true" pageId="14" pageNumber="521">R. ponkouensis</emphasis>
|
||
<bibRefCitation id="9A0A4ED8FFE67D52A343FA6AFBEFFA1C" author="Huang, S. - P. & Chen, I. - S." box="[796,1039,1503,1530]" pageId="14" pageNumber="521" pagination="101 - 110" refId="ref23822" refString="Huang, S. - P. & Chen, I. - S. (2007) Three new species of Rhinogobius Gill, 1859 (Teleeostei: Gobiidae) from the Hanjiang Basin, southern China. Raffles Bulletin of Zoology, Supplement 14, 101 - 110." type="journal article" year="2007">Huang & Chen, 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="399B48AAFFE67D52A415FA6AFF32F9FB" authority="Chen & Kottelat, 2005" authorityName="Chen & Kottelat" authorityYear="2005" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="virgigena">
|
||
<emphasis id="CCEFEF3BFFE67D52A415FA6AFB34FA1C" box="[1098,1236,1504,1529]" italics="true" pageId="14" pageNumber="521">R. virgigena</emphasis>
|
||
<bibRefCitation id="9A0A4ED8FFE67D52A483FA55FF32F9FB" author="Chen, I. - S. & Kottelat, M." pageId="14" pageNumber="521" pagination="1407 - 1429" refId="ref22926" refString="Chen, I. - S. & Kottelat, M. (2005) Four new freshwater gobies of the genus Rhinogobius (Teleostei: Gobiidae) from northern Vietnam. Journal of Natural History, 39, 1407 - 1429. https: // doi. org / 10.1080 / 00222930400008736" type="journal article" year="2005">Chen & Kottelat, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
, have two or three stripes on cheek, but lack the longitudinal stripes along sides of the body (
|
||
<bibRefCitation id="9A0A4ED8FFE67D52A484F989FA8AF9FB" author="Herre, A. W. C. T." box="[1243,1386,1539,1566]" pageId="14" pageNumber="521" pagination="285 - 293" refId="ref23697" refString="Herre, A. W. C. T. (1935 a) Notes on fishes in the Zoological Museum of Stanford University. VI. New and rare Hong Kong fishes obtained in 1934. Hong Kong Naturalist, 6, 285 - 293." type="journal article" year="1935">Herre 1935a</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="9A0A4ED8FFE67D52A52AF98EFEF1F9A4" author="Wu, H. - L. & Ni, Y." pageId="14" pageNumber="521" pagination="259 - 314" refId="ref24977" refString="Wu, H. - L. & Ni, Y. (1986) Gobioidei. In: Chinese Academy of Fishery Sciences (Ed.), The freshwater and estuaries fishes of Hainan Island. Guangdong Science and Technology Press, Guangzhou, pp. 259 - 314. [in Chinese]" type="book chapter" year="1986">Wu & Ni 1986</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="9A0A4ED8FFE67D52A144F9A2FE64F9A4" author="Wu, H. - L." box="[283,388,1575,1602]" pageId="14" pageNumber="521" pagination="422 - 504" refId="ref24900" refString="Wu, H. - L. (1991) Gobioidei. In: Pan, J. - H., Zhong, L., Zheng, C. - Y., Wu, H. - L. & Liu, J. (Eds.), The freshwater fishes of Guangdong Province. Guangdong Science and Technology Press, Guangzhou, pp. 422 - 504. [in Chinese]" type="book chapter" year="1991">Wu 1991</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="9A0A4ED8FFE67D52A1D0F9ADFD69F9A7" author="Chen, I. - S. & Kottelat, M." box="[399,649,1575,1602]" pageId="14" pageNumber="521" pagination="1407 - 1429" refId="ref22926" refString="Chen, I. - S. & Kottelat, M. (2005) Four new freshwater gobies of the genus Rhinogobius (Teleostei: Gobiidae) from northern Vietnam. Journal of Natural History, 39, 1407 - 1429. https: // doi. org / 10.1080 / 00222930400008736" type="journal article" year="2005">Chen & Kottelat 2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="9A0A4ED8FFE67D52A2CBF9A2FC9EF9A4" author="Huang, S. - P. & Chen, I. - S." box="[660,894,1575,1602]" pageId="14" pageNumber="521" pagination="101 - 110" refId="ref23822" refString="Huang, S. - P. & Chen, I. - S. (2007) Three new species of Rhinogobius Gill, 1859 (Teleeostei: Gobiidae) from the Hanjiang Basin, southern China. Raffles Bulletin of Zoology, Supplement 14, 101 - 110." type="journal article" year="2007">Huang & Chen 2007</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="9A0A4ED8FFE67D52A3D7F9A2FBBDF9A4" author="Wu, H. - L. & Zhong, J. - S." box="[904,1117,1575,1602]" pageId="14" pageNumber="521" refId="ref25033" refString="Wu, H. - L. & Zhong, J. - S. (2008) Fauna Sinica, Osteichthyes, Perciformes (V), Gobioidei. Science Press, Beijing, 951 pp., 16 pls. [in Chinese]" type="book" year="2008">Wu & Zhong 2008</bibRefCitation>
|
||
; observation on
|
||
<taxonomicName id="399B48AAFFE67D52A54CF9A2FA7CF9A4" authorityName="Chen & Kottelat" authorityYear="2005" box="[1299,1436,1576,1601]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="virgigena">
|
||
<emphasis id="CCEFEF3BFFE67D52A54CF9A2FA7CF9A4" box="[1299,1436,1576,1601]" italics="true" pageId="14" pageNumber="521">R. virgigena</emphasis>
|
||
</taxonomicName>
|
||
in the present study;
|
||
<figureCitation id="66A02FACFFE67D52A1DBF9C1FE0DF980" box="[388,493,1611,1637]" captionStart="FIGURE 9" captionStartId="16.[151,250,207,231]" captionTargetBox="[147,1441,181,1982]" captionTargetId="figure-17@15.[147,1441,181,1954]" captionTargetPageId="15" captionText="FIGURE 9. Comparative material of the genus Rhinogobius from northern Vietnam (a–l) and Japan (m–p) observed in the present study. Photographs were taken immediately after fixation. Specimens on left side are males, right are females. a, b, R. sulcatus (a, OISTICH 184, 30.4 mm SL; b, OISTICH 190, 32.4 mm SL); c, d, R. boa (c, OISTICH 161, 25.4 mm SL; d, OISTICH 160, 28.0 mm SL); e, f, R. virgigena (e, NSMT-P 147872, 36.1 mm SL; f, OISTICH 154, 33.7 mm SL); g, h, R. variolatus (g, OISTICH 169, 31.4 mm SL; h, OISTICH 170, 28.8 mm SL); i, j, R. leavelli (i, OISTICH 209, 38.1 mm SL; j, OISTICH 210, 35.4 mm SL); k, l, R. similis (k, OISTICH 222, 54.4 mm SL; l, NSMT-P 147882, 44.7 mm SL); m, n, R. similis from Okinawa Island (m, URM-P 49149, 48.6 mm SL; n, OISTICH 2, 63.6 mm SL); o, p, R. similis, from Izu Peninsula (o, OISTICH 330, 47.6 mm SL; p, OISTICH 332, 51.0 mm SL)." figureDoi="http://doi.org/10.5281/zenodo.13688016" httpUri="https://zenodo.org/record/13688016/files/figure.png" pageId="14" pageNumber="521">Fig. 9e, f</figureCitation>
|
||
) (vs. five stripes in
|
||
<taxonomicName id="399B48AAFFE67D52A290F9C6FCC7F980" authorityName="Maeda & Kobayashi & Iida & Tran" authorityYear="2024" box="[719,807,1612,1637]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="nami">
|
||
<emphasis id="CCEFEF3BFFE67D52A290F9C6FCC7F980" box="[719,807,1612,1637]" italics="true" pageId="14" pageNumber="521">R. nami</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="66A02FACFFE67D52A368F9C1FC45F980" box="[823,933,1611,1638]" captionStart="FIGURE 7" captionStartId="13.[151,250,942,966]" captionTargetBox="[151,1435,181,918]" captionTargetId="figure-17@13.[151,1435,181,918]" captionTargetPageId="13" captionText="FIGURE 7. Rhinogobius nami immediately after fixation. a, NSMT-P 147890, holotype, male, 24.9 mm SL; b, OISTICH 282, paratype, female, 26.6 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13330585" httpUri="https://zenodo.org/record/13330585/files/figure.png" pageId="14" pageNumber="521">Fig. 7a, b</figureCitation>
|
||
), 27 and 28 vertebrae in
|
||
<taxonomicName id="399B48AAFFE67D52A49EF9C6FAACF980" baseAuthorityName="Herre" baseAuthorityYear="1935" box="[1217,1356,1611,1637]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="duospilus">
|
||
<emphasis id="CCEFEF3BFFE67D52A49EF9C6FAACF980" box="[1217,1356,1611,1637]" italics="true" pageId="14" pageNumber="521">R. duospilus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="399B48AAFFE67D52A5DBF9C6FEC1F96C" authorityName="Huang & Chen" authorityYear="2007" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="ponkouensis">
|
||
<emphasis id="CCEFEF3BFFE67D52A5DBF9C6FEC1F96C" italics="true" pageId="14" pageNumber="521">R. ponkouensis</emphasis>
|
||
</taxonomicName>
|
||
, respectively (
|
||
<bibRefCitation id="9A0A4ED8FFE67D52A199F9E5FD52F96C" author="Huang, S. - P. & Chen, I. - S." box="[454,690,1647,1673]" pageId="14" pageNumber="521" pagination="101 - 110" refId="ref23822" refString="Huang, S. - P. & Chen, I. - S. (2007) Three new species of Rhinogobius Gill, 1859 (Teleeostei: Gobiidae) from the Hanjiang Basin, southern China. Raffles Bulletin of Zoology, Supplement 14, 101 - 110." type="journal article" year="2007">Huang & Chen 2007</bibRefCitation>
|
||
) (vs. usually
|
||
<quantity id="39639ECCFFE67D52A317F9E5FC61F96C" box="[840,897,1647,1673]" metricMagnitude="-1" metricUnit="m" metricValue="6.604" pageId="14" pageNumber="521" unit="in" value="26.0">26 in</quantity>
|
||
<taxonomicName id="399B48AAFFE67D52A3D7F9FAFBF1F96C" authorityName="Chen & Kottelat" authorityYear="2005" box="[904,1041,1648,1673]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="virgigena">
|
||
<emphasis id="CCEFEF3BFFE67D52A3D7F9FAFBF1F96C" box="[904,1041,1648,1673]" italics="true" pageId="14" pageNumber="521">R. virgigena</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="399B48AAFFE67D52A417F9FAFB7FF96C" authorityName="Maeda & Kobayashi & Iida & Tran" authorityYear="2024" box="[1096,1183,1648,1673]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="nami">
|
||
<emphasis id="CCEFEF3BFFE67D52A417F9FAFB7FF96C" box="[1096,1183,1648,1673]" italics="true" pageId="14" pageNumber="521">R. nami</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<tableCitation id="B3190692FFE67D52A4F2F9E5FB1FF96C" box="[1197,1279,1647,1673]" captionStart="TABLE 1" captionStartId="5.[151,238,476,500]" captionText="TABLE 1. Number of vertebrae in various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="14" pageNumber="521">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FE243329FFE67D52A098F919FBBAF8D8" blockId="14.[151,1437,1287,1854]" pageId="14" pageNumber="521">
|
||
Among species from central
|
||
<collectingCountry id="868C73B9FFE67D52A24FF919FD8FF948" box="[528,623,1683,1709]" name="Vietnam" pageId="14" pageNumber="521">Vietnam</collectingCountry>
|
||
described in the present study,
|
||
<taxonomicName id="399B48AAFFE67D52A388F91EFBD0F948" authorityName="Maeda & Kobayashi & Iida & Tran" authorityYear="2024" box="[983,1072,1684,1709]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="nami">
|
||
<emphasis id="CCEFEF3BFFE67D52A388F91EFBD0F948" box="[983,1072,1684,1709]" italics="true" pageId="14" pageNumber="521">R. nami</emphasis>
|
||
</taxonomicName>
|
||
is most similar to
|
||
<taxonomicName id="399B48AAFFE67D52A559F91EFA75F948" authorityName="Chen, Kottelat & Miller" authorityYear="1999" box="[1286,1429,1684,1709]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="taenigena">
|
||
<emphasis id="CCEFEF3BFFE67D52A559F91EFA75F948" box="[1286,1429,1684,1709]" italics="true" pageId="14" pageNumber="521">R. taenigena</emphasis>
|
||
</taxonomicName>
|
||
, sharing the same markings on fins, and the red and light bluish-grey patterns on opercular membrane, interopercle, and lower part of subopercle of the male (
|
||
<figureCitation id="66A02FACFFE67D52A2DFF951FD3DF913" box="[640,733,1755,1782]" captionStart="FIGURE 7" captionStartId="13.[151,250,942,966]" captionTargetBox="[151,1435,181,918]" captionTargetId="figure-17@13.[151,1435,181,918]" captionTargetPageId="13" captionText="FIGURE 7. Rhinogobius nami immediately after fixation. a, NSMT-P 147890, holotype, male, 24.9 mm SL; b, OISTICH 282, paratype, female, 26.6 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13330585" httpUri="https://zenodo.org/record/13330585/files/figure.png" pageId="14" pageNumber="521">Figs. 7a</figureCitation>
|
||
and
|
||
<figureCitation id="66A02FACFFE67D52A347F951FC81F910" box="[792,865,1755,1781]" captionStart="FIGURE 11" captionStartId="19.[151,250,1968,1992]" captionTargetBox="[151,1435,337,1952]" captionTargetId="figure-108@19.[151,1435,337,1952]" captionTargetPageId="19" captionText="FIGURE 11. Rhinogobius taenigena immediately after fixation. a, OISTICH 303, male, 30.8 mm SL; b, OISTICH 312, male, 28.8 mm SL; c, OISTICH 304, female, 30.5 mm SL; d, OISTICH 311, female, 33.4 mm SL." figureDoi="http://doi.org/10.5281/zenodo.13330591" httpUri="https://zenodo.org/record/13330591/files/figure.png" pageId="14" pageNumber="521">11a, b</figureCitation>
|
||
). However,
|
||
<taxonomicName id="399B48AAFFE67D52A3B1F956FBA7F910" authorityName="Maeda & Kobayashi & Iida & Tran" authorityYear="2024" box="[1006,1095,1756,1781]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="nami">
|
||
<emphasis id="CCEFEF3BFFE67D52A3B1F956FBA7F910" box="[1006,1095,1756,1781]" italics="true" pageId="14" pageNumber="521">R. nami</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="399B48AAFFE67D52A5DBF956FEE5F8FC" authorityName="Chen, Kottelat & Miller" authorityYear="1999" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="taenigena">
|
||
<emphasis id="CCEFEF3BFFE67D52A5DBF956FEE5F8FC" italics="true" pageId="14" pageNumber="521">R. taenigena</emphasis>
|
||
</taxonomicName>
|
||
in the presence of five longitudinal stripes on sides of the body (absent in
|
||
<taxonomicName id="399B48AAFFE67D52A43FF88AFB0EF8FC" authorityName="Chen, Kottelat & Miller" authorityYear="1999" box="[1120,1262,1792,1817]" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="521" phylum="Chordata" rank="species" species="taenigena">
|
||
<emphasis id="CCEFEF3BFFE67D52A43FF88AFB0EF8FC" box="[1120,1262,1792,1817]" italics="true" pageId="14" pageNumber="521">R. taenigena</emphasis>
|
||
</taxonomicName>
|
||
), three oblique stripes on the male’s cheek (vs. absent) and presence of 26 vertebrae (vs. 27;
|
||
<tableCitation id="B3190692FFE67D52A3A8F8A9FBA9F8D8" box="[1015,1097,1827,1853]" captionStart="TABLE 1" captionStartId="5.[151,238,476,500]" captionText="TABLE 1. Number of vertebrae in various species of Rhinogobius examined in the present study. * Includes the holotype." pageId="14" pageNumber="521">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |