treatments-xml/data/6D/0E/87/6D0E87E4EA0F616F44F8F0E0FEDAFE69.xml

<document id="4E674C3189DB52EA666146E07F7F314A" ID-DOI="10.5281/zenodo.5276563" ID-ISSN="0253-116X" ID-Zenodo-Dep="5276563" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630012980284" checkinUser="carolina" docAuthor="Assing, V." docDate="2009" docId="6D0E87E4EA0F616F44F8F0E0FEDAFE69" docLanguage="en" docName="LinzerbiolBeitr.2009.41.2.1191-1246.pdf" docOrigin="Linzer biologische Beiträge 41 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Geostiba (Tropogastrosipalia) sarica Assing 2009, nov.sp." docType="treatment" docVersion="3" lastPageNumber="1208" masterDocId="9137FF9CEA01617E444AF536FFEAFFD1" masterDocTitle="A revision of Geostiba of the Western Palaearctic region. XIX. New species from Turkey and Iran and additional records, with an updated key and a catalogue of the species of the Eastern Mediterranean, the Caucasus, and adjacent regions (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="1246" masterPageNumber="1191" pageNumber="1205" updateTime="1699136907799" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="0BEEF9A28013CBB09DE17EA7C13F5F74">A revision of Geostiba of the Western Palaearctic region. XIX. New species from Turkey and Iran and additional records, with an updated key and a catalogue of the species of the Eastern Mediterranean, the Caucasus, and adjacent regions (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="3F5F23FD8647835B53D7AA9519D4DDE4">Assing, V.</mods:namePart>
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<mods:title id="D1AB780B4959145AD7256FDF5A2EB622">Linzer biologische Beiträge</mods:title>
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<mods:date id="181A04E052A177820B1A2087B7A1A3F5">2009</mods:date>
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<subSubSection id="ADBD6579EA0F617044F8F0E0FCEEFA3D" box="[178,772,1494,1516]" pageId="14" pageNumber="1205" type="nomenclature">
<paragraph id="E51836F2EA0F617044F8F0E0FCEEFA3D" blockId="14.[178,772,1494,1516]" box="[178,772,1494,1516]" pageId="14" pageNumber="1205">
<heading id="BE50819EEA0F617044F8F0E0FCEEFA3D" box="[178,772,1494,1516]" fontSize="8" level="2" pageId="14" pageNumber="1205" reason="2">
<emphasis id="D7D3EAE0EA0F617044F8F0E0FD6BFA3D" bold="true" box="[178,641,1494,1516]" pageId="14" pageNumber="1205">
<taxonomicName id="22A74D71EA0F617044F8F0E0FDC4FA3D" ID-CoL="6KCVQ" authority="Assing, 2009" authorityName="Assing" authorityYear="2009" box="[178,558,1494,1516]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1205" phylum="Arthropoda" rank="species" species="sarica" status="sp. nov." subGenus="Tropogastrosipalia">
<emphasis id="D7D3EAE0EA0F617044F8F0E0FEE7FA3D" bold="true" box="[178,269,1494,1516]" italics="true" pageId="14" pageNumber="1205">Geostiba</emphasis>
(
<emphasis id="D7D3EAE0EA0F61704556F0E0FE0BFA3D" bold="true" box="[284,481,1494,1516]" italics="true" pageId="14" pageNumber="1205">Tropogastrosipalia</emphasis>
) 
<emphasis id="D7D3EAE0EA0F617045A5F0E1FDC4FA3D" bold="true" box="[495,558,1495,1516]" italics="true" pageId="14" pageNumber="1205">sarica</emphasis>
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<taxonomicNameLabel id="CCE0579BEA0F6170467EF0E0FD6BFA3D" box="[564,641,1494,1516]" pageId="14" pageNumber="1205" rank="species">nov.sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="7D9C2A77EA0F617046C4F0E0FD14FA3D" box="[654,766,1494,1516]" captionStart-0="Figs 32-39" captionStart-1="Figs 40-47" captionStartId-0="14.[178,218,1365,1385]" captionStartId-1="16.[178,218,1255,1275]" captionTargetBox-0="[183,1070,219,1338]" captionTargetBox-1="[182,1056,219,1229]" captionTargetId-0="figure-0@14.[178,1084,219,1357]" captionTargetId-1="figure-203@16.[178,1085,219,1244]" captionTargetPageId-0="14" captionTargetPageId-1="16" captionText-0="Figs 32-39: Geostiba sarica nov.sp. from the environs of Sangdeh (type locality): (32) male habitus; (33) male forebody; (34) male abominal segments VI-VIII in lateral view; (35) male abominal segments VII-VIII in dorsal view; (36-38) median lobe of aedeagus in lateral view; (39) spermatheca. Scale bars: 32: 1.0 mm; 33-35: 0.5 mm; 36-39: 0.1 mm." captionText-1="Figs 40-47: Geostiba sarica nov.sp. from the environs of Qolqol: (40) male habitus; (41) male forebody; (42) male abdominal segments V-VII in lateral view; (43) male abdominal segments VI- VII in dorsal view; (44) male tergite VIII; (45) male sternite VIII; (46) median lobe of aedeagus in lateral view; (47) spermatheca. Scale bars: 40: 1.0 mm; 41-42: 0.5 mm; 43-45: 0.2 mm; 46-47: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5276579" figureDoi-1="http://doi.org/10.5281/zenodo.5276581" httpUri-0="https://zenodo.org/record/5276579/files/figure.png" httpUri-1="https://zenodo.org/record/5276581/files/figure.png" pageId="14" pageNumber="1205">Figs 32-47</figureCitation>
)
</heading>
</paragraph>
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<paragraph id="E51836F2EA0F617144F8F334FDA1FEFE" blockId="14.[178,1085,1537,1604]" lastBlockId="15.[178,1085,214,449]" lastPageId="15" lastPageNumber="1206" pageId="14" pageNumber="1205">
<materialsCitation id="55CF3CAFEA0F617144F8F334FDA1FEFE" collectingDate="2008-05-29" collectionCode="A" collectorName="Mohammadabad &amp; Mts. &amp; Slope &amp; NE Sangdeh &amp; Putz &amp; Prov. &amp; E Qolqol" country="Iran" county="Sari County" determinerName="V. Assing" elevation="1533" lastPageId="15" lastPageNumber="1206" latitude="36.171116" location="Sari County" longLatPrecision="1" longitude="53.271534" municipality="Fagus forest" pageId="14" pageNumber="1205" specimenCount="8" specimenCount-female="8" stateProvince="Mazandaran" typeStatus="holotype">
<typeStatus id="3A1C8850EA0F617044F8F334FEEDF9C7" box="[178,263,1538,1558]" pageId="14" pageNumber="1205" type="holotype">Holotype</typeStatus>
Ƌ: &quot; 
<collectingCountry id="9DB07662EA0F6170457DF334FEB5F9C7" box="[311,351,1538,1558]" name="Iran" pageId="14" pageNumber="1205">Iran</collectingCountry>
, Prov. 
<collectingRegion id="2763F810EA0F617045EEF334FDFEF9C7" box="[420,532,1538,1558]" country="Iran" name="Mazandaran" pageId="14" pageNumber="1205">Mazandaran</collectingRegion>
[IR08-01], 
<collectingCounty id="0C794E7EEA0F617046CFF334FD1CF9C7" box="[645,758,1538,1558]" pageId="14" pageNumber="1205">Sari County</collectingCounty>
, 
<collectorName id="48525324EA0F61704748F334FC77F9C7" box="[770,925,1538,1558]" pageId="14" pageNumber="1205">Mohammadabad</collectorName>
, 
<collectingRegion id="2763F810EA0F617047E3F334FC0CF9C7" box="[937,998,1538,1558]" country="Iran" name="Alborz" pageId="14" pageNumber="1205">Elburz</collectingRegion>
<collectorName id="48525324EA0F617047A4F334FBF0F9C7" box="[1006,1050,1538,1558]" pageId="14" pageNumber="1205">Mts.</collectorName>
, N- 
<collectorName id="48525324EA0F617044F8F32FFF02F9FC" box="[178,232,1561,1581]" pageId="14" pageNumber="1205">Slope</collectorName>
, 
<collectorName id="48525324EA0F617044BDF32FFE98F9FC" box="[247,370,1561,1581]" pageId="14" pageNumber="1205">NE Sangdeh</collectorName>
, 
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<elevation id="6E8AD1C1EA0F617045CAF32FFE24F9FC" box="[384,462,1561,1581]" metricMagnitude="3" metricUnit="m" metricValue="1.533" pageId="14" pageNumber="1205" unit="m" value="1533.0">1533 m</elevation>
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, 
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, 
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[recte: 
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, 53°09.578&quot;E], 
<collectingMunicipality id="057CAC88EA0F617044F8F306FECCF995" box="[178,294,1584,1604]" pageId="14" pageNumber="1205">Fagus forest</collectingMunicipality>
, leaves debris, sifted, 
<date id="91191032EA0F617045B2F306FDB0F995" box="[504,602,1584,1604]" pageId="14" pageNumber="1205" value="2008-05-29">
<collectingDate id="815DE9DAEA0F617045B2F306FDB0F995" box="[504,602,1584,1604]" pageId="14" pageNumber="1205" value="2008-05-29">29.V.2008</collectingDate>
</date>
, leg. 
<collectionCode id="83B6AE37EA0F617046C5F306FD48F995" box="[655,674,1584,1604]" pageId="14" pageNumber="1205">A</collectionCode>
. 
<collectorName id="48525324EA0F617046E7F306FD3FF995" box="[685,725,1584,1604]" pageId="14" pageNumber="1205">Pütz</collectorName>
/ 
<typeStatus id="3A1C8850EA0F617046A6F306FCA1F995" box="[748,843,1584,1604]" pageId="14" pageNumber="1205" type="holotype">Holotypus</typeStatus>
Ƌ 
<taxonomicName id="22A74D71EA0F61704724F306FBEBF995" authority="Assing, 2009" authorityName="Assing" authorityYear="2009" box="[878,1025,1584,1604]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1205" phylum="Arthropoda" rank="species" species="sarica" status="sp. nov.">
<emphasis id="D7D3EAE0EA0F61704724F306FBEBF995" box="[878,1025,1584,1604]" italics="true" pageId="14" pageNumber="1205">Geostiba sarica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCE0579BEA0F61704043F306FBD7F995" box="[1033,1085,1584,1604]" pageId="14" pageNumber="1205" rank="species">sp. n.</taxonomicNameLabel>
det. 
<determinerName id="10C4DA36EA0E61714491F5E1FED3FF3A" box="[219,313,215,235]" pageId="15" pageNumber="1206">V. Assing</determinerName>
2009&quot; (cPüt). 
<typeStatus id="3A1C8850EA0E6171458CF5E1FDC8FF3A" box="[454,546,215,235]" pageId="15" pageNumber="1206" type="paratype">Paratypes</typeStatus>
: 5ƋƋ, 
<specimenCount id="F3A1FD7BEA0E61714627F5E1FD76FF21" box="[621,668,215,240]" pageId="15" pageNumber="1206" type="female">6♀♀♀</specimenCount>
: same data as holotype (cPüt, cAss); 1Ƌ, 
<specimenCount id="F3A1FD7BEA0E617144F8F5D8FF3FFED6" box="[178,213,238,263]" pageId="15" pageNumber="1206" type="female">2♀♀</specimenCount>
: &quot; 
<collectingCountry id="9DB07662EA0E617144BDF5D8FEF5FED3" box="[247,287,238,258]" name="Iran" pageId="15" pageNumber="1206">Iran</collectingCountry>
, 
<collectorName id="48525324EA0E61714560F5D8FEB1FED3" box="[298,347,238,258]" pageId="15" pageNumber="1206">Prov.</collectorName>
<collectingRegion id="2763F810EA0E6171452BF5D8FE3BFED3" box="[353,465,238,258]" country="Iran" name="Mazandaran" pageId="15" pageNumber="1206">Mazandaran</collectingRegion>
[IR08-03], 
<location id="E0786029EA0E61714674F5D8FD44FED3" LSID="urn:lsid:plazi:treatment:6D0E87E4EA0F616F44F8F0E0FEDAFE69:E0786029EA0E61714674F5D8FD44FED3" box="[574,686,238,258]" country="Iran" county="Sari County" latitude="36.171116" longLatPrecision="1" longitude="53.271534" municipality="Fagus forest" name="Sari County" pageId="15" pageNumber="1206" stateProvince="Mazandaran">Sari County</location>
, 
<collectorName id="48525324EA0E617146F3F5D8FCBEFED3" box="[697,852,238,258]" pageId="15" pageNumber="1206">Mohammadabad</collectorName>
, 
<collectingRegion id="2763F810EA0E61714714F5D8FC70FED3" box="[862,922,238,258]" country="Iran" name="Alborz" pageId="15" pageNumber="1206">Elburz</collectingRegion>
<collectorName id="48525324EA0E617147EBF5D8FC27FED3" box="[929,973,238,258]" pageId="15" pageNumber="1206">Mts.</collectorName>
, N-Slope, 
<collectorName id="48525324EA0E61714065F5D8FF19FEC9" pageId="15" pageNumber="1206">E Qolqol</collectorName>
, 
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, 
<geoCoordinate id="80935035EA0E617145DAF432FDE0FEC9" box="[400,522,260,280]" degrees="53" direction="east" minutes="16" orientation="longitude" pageId="15" pageNumber="1206" precision="1" seconds="29.2" value="53.274776">53°16'29.2&quot;E</geoCoordinate>
[recte: 
<geoCoordinate id="80935035EA0E61714611F432FD3BFEC9" box="[603,721,260,280]" degrees="36" direction="north" minutes="10.267" orientation="latitude" pageId="15" pageNumber="1206" precision="1" value="36.171116">36°10.267'N</geoCoordinate>
, 
<geoCoordinate id="80935035EA0E617146AAF432FCB9FEC9" box="[736,851,260,280]" degrees="53" direction="east" minutes="16.292" orientation="longitude" pageId="15" pageNumber="1206" precision="1" value="53.271534">53°16.292'E</geoCoordinate>
], 
<quantity id="225F9B17EA0E61714723F432FC41FEC9" box="[873,939,260,280]" metricMagnitude="2" metricUnit="m" metricValue="9.16" pageId="15" pageNumber="1206" unit="m" value="916.0">
<elevation id="6E8AD1C1EA0E61714723F432FC41FEC9" box="[873,939,260,280]" metricMagnitude="2" metricUnit="m" metricValue="9.16" pageId="15" pageNumber="1206" unit="m" value="916.0">916 m</elevation>
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, leaves debris, sifted, 
<date id="91191032EA0E617144BAF42DFEB8FEFE" box="[240,338,283,303]" pageId="15" pageNumber="1206" value="2008-05-29">
<collectingDate id="815DE9DAEA0E617144BAF42DFEB8FEFE" box="[240,338,283,303]" pageId="15" pageNumber="1206" value="2008-05-29">29.V.2008</collectingDate>
</date>
, leg. 
<collectionCode id="83B6AE37EA0E617145C8F42DFE7FFEFE" box="[386,405,283,303]" pageId="15" pageNumber="1206">A</collectionCode>
. Pütz&quot; (cPüt, cAss).
</materialsCitation>
</paragraph>
<paragraph id="E51836F2EA0E617144F8F400FF13FE10" blockId="15.[178,1085,214,449]" pageId="15" pageNumber="1206">
D e s c r i p t i o n: Body length 
<quantity id="225F9B17EA0E6171466EF400FD4AFE9A" box="[548,672,310,332]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="3.2" metricValueMin="2.1" pageId="15" pageNumber="1206" unit="mm" value="2.65" valueMax="3.2" valueMin="2.1">2.1-3.2 mm</quantity>
. Habitus as in 
<figureCitation id="7D9C2A77EA0E6171470FF400FC72FE9D" box="[837,920,310,332]" captionStart="Figs 32-39" captionStartId="14.[178,218,1365,1385]" captionTargetBox="[183,1070,219,1338]" captionTargetId="figure-0@14.[178,1084,219,1357]" captionTargetPageId="14" captionText="Figs 32-39: Geostiba sarica nov.sp. from the environs of Sangdeh (type locality): (32) male habitus; (33) male forebody; (34) male abominal segments VI-VIII in lateral view; (35) male abominal segments VII-VIII in dorsal view; (36-38) median lobe of aedeagus in lateral view; (39) spermatheca. Scale bars: 32: 1.0 mm; 33-35: 0.5 mm; 36-39: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276579" httpUri="https://zenodo.org/record/5276579/files/figure.png" pageId="15" pageNumber="1206">Figs 32</figureCitation>
, 
<figureCitation id="7D9C2A77EA0E617147ECF400FC28FE9D" box="[934,962,310,332]" captionStart="Figs 40-47" captionStartId="16.[178,218,1255,1275]" captionTargetBox="[182,1056,219,1229]" captionTargetId="figure-203@16.[178,1085,219,1244]" captionTargetPageId="16" captionText="Figs 40-47: Geostiba sarica nov.sp. from the environs of Qolqol: (40) male habitus; (41) male forebody; (42) male abdominal segments V-VII in lateral view; (43) male abdominal segments VI- VII in dorsal view; (44) male tergite VIII; (45) male sternite VIII; (46) median lobe of aedeagus in lateral view; (47) spermatheca. Scale bars: 40: 1.0 mm; 41-42: 0.5 mm; 43-45: 0.2 mm; 46-47: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276581" httpUri="https://zenodo.org/record/5276581/files/figure.png" pageId="15" pageNumber="1206">40</figureCitation>
. Coloration variable: body uniformly reddish to more or less distinctly bicoloured, with the head and the abdomen blackish to blackish-brown and the pronotum, elytra, and the abdominal apex paler reddish to brown; legs yellowish to yellowish-brown; antennae reddish to brown.
</paragraph>
<paragraph id="E51836F2EA0E617144F8F4E6FC04FDF1" blockId="15.[178,1085,464,544]" pageId="15" pageNumber="1206">
Head 1.0-1.1 times as long as wide (
<figureCitation id="7D9C2A77EA0E6171460FF4E6FD72FE37" box="[581,664,464,486]" captionStart="Figs 32-39" captionStartId="14.[178,218,1365,1385]" captionTargetBox="[183,1070,219,1338]" captionTargetId="figure-0@14.[178,1084,219,1357]" captionTargetPageId="14" captionText="Figs 32-39: Geostiba sarica nov.sp. from the environs of Sangdeh (type locality): (32) male habitus; (33) male forebody; (34) male abominal segments VI-VIII in lateral view; (35) male abominal segments VII-VIII in dorsal view; (36-38) median lobe of aedeagus in lateral view; (39) spermatheca. Scale bars: 32: 1.0 mm; 33-35: 0.5 mm; 36-39: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276579" httpUri="https://zenodo.org/record/5276579/files/figure.png" pageId="15" pageNumber="1206">Figs 33</figureCitation>
, 
<figureCitation id="7D9C2A77EA0E617146E3F4E6FD2FFE37" box="[681,709,464,486]" captionStart="Figs 40-47" captionStartId="16.[178,218,1255,1275]" captionTargetBox="[182,1056,219,1229]" captionTargetId="figure-203@16.[178,1085,219,1244]" captionTargetPageId="16" captionText="Figs 40-47: Geostiba sarica nov.sp. from the environs of Qolqol: (40) male habitus; (41) male forebody; (42) male abdominal segments V-VII in lateral view; (43) male abdominal segments VI- VII in dorsal view; (44) male tergite VIII; (45) male sternite VIII; (46) median lobe of aedeagus in lateral view; (47) spermatheca. Scale bars: 40: 1.0 mm; 41-42: 0.5 mm; 43-45: 0.2 mm; 46-47: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276581" httpUri="https://zenodo.org/record/5276581/files/figure.png" pageId="15" pageNumber="1206">41</figureCitation>
); punctation extremely fine, barely noticeable; surface with shallow, but distinct microreticulation. Eyes weakly convex and relatively small, approximately 1/3 the length of postocular region in dorsal view.
</paragraph>
<paragraph id="E51836F2EA0E617144F8F706FC3DFD51" blockId="15.[178,1085,560,640]" pageId="15" pageNumber="1206">
Pronotum with moderately pronounced sexual dimorphism (
<figureCitation id="7D9C2A77EA0E61714750F706FC83FD97" box="[794,873,560,582]" captionStart="Figs 32-39" captionStartId="14.[178,218,1365,1385]" captionTargetBox="[183,1070,219,1338]" captionTargetId="figure-0@14.[178,1084,219,1357]" captionTargetPageId="14" captionText="Figs 32-39: Geostiba sarica nov.sp. from the environs of Sangdeh (type locality): (32) male habitus; (33) male forebody; (34) male abominal segments VI-VIII in lateral view; (35) male abominal segments VII-VIII in dorsal view; (36-38) median lobe of aedeagus in lateral view; (39) spermatheca. Scale bars: 32: 1.0 mm; 33-35: 0.5 mm; 36-39: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276579" httpUri="https://zenodo.org/record/5276579/files/figure.png" pageId="15" pageNumber="1206">Figs 33</figureCitation>
, 
<figureCitation id="7D9C2A77EA0E6171473CF706FC78FD97" box="[886,914,560,582]" captionStart="Figs 40-47" captionStartId="16.[178,218,1255,1275]" captionTargetBox="[182,1056,219,1229]" captionTargetId="figure-203@16.[178,1085,219,1244]" captionTargetPageId="16" captionText="Figs 40-47: Geostiba sarica nov.sp. from the environs of Qolqol: (40) male habitus; (41) male forebody; (42) male abdominal segments V-VII in lateral view; (43) male abdominal segments VI- VII in dorsal view; (44) male tergite VIII; (45) male sternite VIII; (46) median lobe of aedeagus in lateral view; (47) spermatheca. Scale bars: 40: 1.0 mm; 41-42: 0.5 mm; 43-45: 0.2 mm; 46-47: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276581" httpUri="https://zenodo.org/record/5276581/files/figure.png" pageId="15" pageNumber="1206">41</figureCitation>
), weakly oblong (Ƌ) to weakly transverse (
<specimenCount id="F3A1FD7BEA0E6171458FF767FE39FDB9" box="[453,467,593,616]" pageId="15" pageNumber="1206" type="female">♀</specimenCount>
), and approximately 1.2 times as wide as head; punctation as fine as that of head; microreticulation clearly more distinct than that of head.
</paragraph>
<paragraph id="E51836F2EA0E617144F8F7A6FE49FD2C" blockId="15.[178,1085,656,765]" pageId="15" pageNumber="1206">
Elytra with moderately pronounced sexual dimorphism, very short, 0.45-0.55 times as long as pronotum; in small specimens approximately 1.2 times, in large specimens almost 1.4 times as wide as pronotum; punctation much more distinct than that of head and pronotum (
<figureCitation id="7D9C2A77EA0E6171456BF7D1FE85FD2C" box="[289,367,743,765]" captionStart="Figs 32-39" captionStartId="14.[178,218,1365,1385]" captionTargetBox="[183,1070,219,1338]" captionTargetId="figure-0@14.[178,1084,219,1357]" captionTargetPageId="14" captionText="Figs 32-39: Geostiba sarica nov.sp. from the environs of Sangdeh (type locality): (32) male habitus; (33) male forebody; (34) male abominal segments VI-VIII in lateral view; (35) male abominal segments VII-VIII in dorsal view; (36-38) median lobe of aedeagus in lateral view; (39) spermatheca. Scale bars: 32: 1.0 mm; 33-35: 0.5 mm; 36-39: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276579" httpUri="https://zenodo.org/record/5276579/files/figure.png" pageId="15" pageNumber="1206">Figs 33</figureCitation>
, 
<figureCitation id="7D9C2A77EA0E61714531F7D1FE7DFD2C" box="[379,407,743,765]" captionStart="Figs 40-47" captionStartId="16.[178,218,1255,1275]" captionTargetBox="[182,1056,219,1229]" captionTargetId="figure-203@16.[178,1085,219,1244]" captionTargetPageId="16" captionText="Figs 40-47: Geostiba sarica nov.sp. from the environs of Qolqol: (40) male habitus; (41) male forebody; (42) male abdominal segments V-VII in lateral view; (43) male abdominal segments VI- VII in dorsal view; (44) male tergite VIII; (45) male sternite VIII; (46) median lobe of aedeagus in lateral view; (47) spermatheca. Scale bars: 40: 1.0 mm; 41-42: 0.5 mm; 43-45: 0.2 mm; 46-47: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276581" httpUri="https://zenodo.org/record/5276581/files/figure.png" pageId="15" pageNumber="1206">41</figureCitation>
).
</paragraph>
<paragraph id="E51836F2EA0E617144F8F63BFC94FCAA" blockId="15.[178,1085,781,891]" pageId="15" pageNumber="1206">
Abdomen at least slightly wider than elytra; anterior tergites without sexual dimorphism; punctation moderately fine and moderately sparse on anterior tergites, very fine and very sparse on posterior tergites; posterior margin of tergite VII without palisade fringe; posterior margin of tergite VIII weakly convex in both sexes (
<figureCitation id="7D9C2A77EA0E6171476FF653FC98FCAA" box="[805,882,869,891]" captionStart="Figs 40-47" captionStartId="16.[178,218,1255,1275]" captionTargetBox="[182,1056,219,1229]" captionTargetId="figure-203@16.[178,1085,219,1244]" captionTargetPageId="16" captionText="Figs 40-47: Geostiba sarica nov.sp. from the environs of Qolqol: (40) male habitus; (41) male forebody; (42) male abdominal segments V-VII in lateral view; (43) male abdominal segments VI- VII in dorsal view; (44) male tergite VIII; (45) male sternite VIII; (46) median lobe of aedeagus in lateral view; (47) spermatheca. Scale bars: 40: 1.0 mm; 41-42: 0.5 mm; 43-45: 0.2 mm; 46-47: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276581" httpUri="https://zenodo.org/record/5276581/files/figure.png" pageId="15" pageNumber="1206">Fig. 44</figureCitation>
).
</paragraph>
<paragraph id="E51836F2EA0E617144F8F6BEFBD3FB81" blockId="15.[178,1085,904,1104]" pageId="15" pageNumber="1206">
Ƌ (with fully developed secondary sexual characters): pronotum approximately as long as wide or weakly oblong, posterior margin in the middle distinctly pointed, posterior half of midline usually indistinctly keeled (
<figureCitation id="7D9C2A77EA0E6171462CF6F2FD5EFC0B" box="[614,692,964,986]" captionStart="Figs 32-39" captionStartId="14.[178,218,1365,1385]" captionTargetBox="[183,1070,219,1338]" captionTargetId="figure-0@14.[178,1084,219,1357]" captionTargetPageId="14" captionText="Figs 32-39: Geostiba sarica nov.sp. from the environs of Sangdeh (type locality): (32) male habitus; (33) male forebody; (34) male abominal segments VI-VIII in lateral view; (35) male abominal segments VII-VIII in dorsal view; (36-38) median lobe of aedeagus in lateral view; (39) spermatheca. Scale bars: 32: 1.0 mm; 33-35: 0.5 mm; 36-39: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276579" httpUri="https://zenodo.org/record/5276579/files/figure.png" pageId="15" pageNumber="1206">Figs 33</figureCitation>
, 
<figureCitation id="7D9C2A77EA0E617146F5F6F2FD31FC0B" box="[703,731,964,986]" captionStart="Figs 40-47" captionStartId="16.[178,218,1255,1275]" captionTargetBox="[182,1056,219,1229]" captionTargetId="figure-203@16.[178,1085,219,1244]" captionTargetPageId="16" captionText="Figs 40-47: Geostiba sarica nov.sp. from the environs of Qolqol: (40) male habitus; (41) male forebody; (42) male abdominal segments V-VII in lateral view; (43) male abdominal segments VI- VII in dorsal view; (44) male tergite VIII; (45) male sternite VIII; (46) median lobe of aedeagus in lateral view; (47) spermatheca. Scale bars: 40: 1.0 mm; 41-42: 0.5 mm; 43-45: 0.2 mm; 46-47: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276581" httpUri="https://zenodo.org/record/5276581/files/figure.png" pageId="15" pageNumber="1206">41</figureCitation>
); elytra with granulose punctation, at anterior half of suture with not very pronounced elevation composed of dense granula, surface without distinct impressions; tergite VII with short and apically acute median process at posterior margin (
<figureCitation id="7D9C2A77EA0E61714599F12AFDA9FBE3" box="[467,579,1052,1074]" captionStart="Figs 32-39" captionStartId="14.[178,218,1365,1385]" captionTargetBox="[183,1070,219,1338]" captionTargetId="figure-0@14.[178,1084,219,1357]" captionTargetPageId="14" captionText="Figs 32-39: Geostiba sarica nov.sp. from the environs of Sangdeh (type locality): (32) male habitus; (33) male forebody; (34) male abominal segments VI-VIII in lateral view; (35) male abominal segments VII-VIII in dorsal view; (36-38) median lobe of aedeagus in lateral view; (39) spermatheca. Scale bars: 32: 1.0 mm; 33-35: 0.5 mm; 36-39: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276579" httpUri="https://zenodo.org/record/5276579/files/figure.png" pageId="15" pageNumber="1206">Figs 34-35</figureCitation>
, 
<figureCitation id="7D9C2A77EA0E61714605F12AFD67FBE3" box="[591,653,1052,1074]" captionStart="Figs 40-47" captionStartId="16.[178,218,1255,1275]" captionTargetBox="[182,1056,219,1229]" captionTargetId="figure-203@16.[178,1085,219,1244]" captionTargetPageId="16" captionText="Figs 40-47: Geostiba sarica nov.sp. from the environs of Qolqol: (40) male habitus; (41) male forebody; (42) male abdominal segments V-VII in lateral view; (43) male abdominal segments VI- VII in dorsal view; (44) male tergite VIII; (45) male sternite VIII; (46) median lobe of aedeagus in lateral view; (47) spermatheca. Scale bars: 40: 1.0 mm; 41-42: 0.5 mm; 43-45: 0.2 mm; 46-47: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276581" httpUri="https://zenodo.org/record/5276581/files/figure.png" pageId="15" pageNumber="1206">42-43</figureCitation>
); posterior margin of sternite VIII broadly convex (
<figureCitation id="7D9C2A77EA0E61714542F10CFEBFFB81" box="[264,341,1082,1104]" captionStart="Figs 40-47" captionStartId="16.[178,218,1255,1275]" captionTargetBox="[182,1056,219,1229]" captionTargetId="figure-203@16.[178,1085,219,1244]" captionTargetPageId="16" captionText="Figs 40-47: Geostiba sarica nov.sp. from the environs of Qolqol: (40) male habitus; (41) male forebody; (42) male abdominal segments V-VII in lateral view; (43) male abdominal segments VI- VII in dorsal view; (44) male tergite VIII; (45) male sternite VIII; (46) median lobe of aedeagus in lateral view; (47) spermatheca. Scale bars: 40: 1.0 mm; 41-42: 0.5 mm; 43-45: 0.2 mm; 46-47: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276581" httpUri="https://zenodo.org/record/5276581/files/figure.png" pageId="15" pageNumber="1206">Fig. 45</figureCitation>
); aedeagus with cristal process of highly variable shape (
<figureCitation id="7D9C2A77EA0E617147DFF10CFBECFB81" box="[917,1030,1082,1104]" captionStart="Figs 32-39" captionStartId="14.[178,218,1365,1385]" captionTargetBox="[183,1070,219,1338]" captionTargetId="figure-0@14.[178,1084,219,1357]" captionTargetPageId="14" captionText="Figs 32-39: Geostiba sarica nov.sp. from the environs of Sangdeh (type locality): (32) male habitus; (33) male forebody; (34) male abominal segments VI-VIII in lateral view; (35) male abominal segments VII-VIII in dorsal view; (36-38) median lobe of aedeagus in lateral view; (39) spermatheca. Scale bars: 32: 1.0 mm; 33-35: 0.5 mm; 36-39: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276579" httpUri="https://zenodo.org/record/5276579/files/figure.png" pageId="15" pageNumber="1206">Figs 36-38</figureCitation>
, 
<figureCitation id="7D9C2A77EA0E6171405BF10CFBC4FB81" box="[1041,1070,1082,1104]" captionStart="Figs 40-47" captionStartId="16.[178,218,1255,1275]" captionTargetBox="[182,1056,219,1229]" captionTargetId="figure-203@16.[178,1085,219,1244]" captionTargetPageId="16" captionText="Figs 40-47: Geostiba sarica nov.sp. from the environs of Qolqol: (40) male habitus; (41) male forebody; (42) male abdominal segments V-VII in lateral view; (43) male abdominal segments VI- VII in dorsal view; (44) male tergite VIII; (45) male sternite VIII; (46) median lobe of aedeagus in lateral view; (47) spermatheca. Scale bars: 40: 1.0 mm; 41-42: 0.5 mm; 43-45: 0.2 mm; 46-47: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276581" httpUri="https://zenodo.org/record/5276581/files/figure.png" pageId="15" pageNumber="1206">46</figureCitation>
).
</paragraph>
<paragraph id="E51836F2EA0E617144F8F155FE50FB7E" blockId="15.[178,1085,1119,1199]" pageId="15" pageNumber="1206">
<specimenCount id="F3A1FD7BEA0E617144F8F155FF2AFBAB" box="[178,192,1123,1146]" pageId="15" pageNumber="1206" type="female">♀</specimenCount>
: pronotum weakly transverse, approximately 1.05 times as wide as long, posterior margin weakly convex, in the middle truncate; punctation of elytra not granulose; spermatheca as in 
<figureCitation id="7D9C2A77EA0E6171450BF1AFFE64FB7E" box="[321,398,1177,1199]" captionStart="Figs 32-39" captionStartId="14.[178,218,1365,1385]" captionTargetBox="[183,1070,219,1338]" captionTargetId="figure-0@14.[178,1084,219,1357]" captionTargetPageId="14" captionText="Figs 32-39: Geostiba sarica nov.sp. from the environs of Sangdeh (type locality): (32) male habitus; (33) male forebody; (34) male abominal segments VI-VIII in lateral view; (35) male abominal segments VII-VIII in dorsal view; (36-38) median lobe of aedeagus in lateral view; (39) spermatheca. Scale bars: 32: 1.0 mm; 33-35: 0.5 mm; 36-39: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276579" httpUri="https://zenodo.org/record/5276579/files/figure.png" pageId="15" pageNumber="1206">Figs 39</figureCitation>
, 
<figureCitation id="7D9C2A77EA0E617145D3F1AFFE50FB7E" box="[409,442,1177,1199]" captionStart="Figs 40-47" captionStartId="16.[178,218,1255,1275]" captionTargetBox="[182,1056,219,1229]" captionTargetId="figure-203@16.[178,1085,219,1244]" captionTargetPageId="16" captionText="Figs 40-47: Geostiba sarica nov.sp. from the environs of Qolqol: (40) male habitus; (41) male forebody; (42) male abdominal segments V-VII in lateral view; (43) male abdominal segments VI- VII in dorsal view; (44) male tergite VIII; (45) male sternite VIII; (46) median lobe of aedeagus in lateral view; (47) spermatheca. Scale bars: 40: 1.0 mm; 41-42: 0.5 mm; 43-45: 0.2 mm; 46-47: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276581" httpUri="https://zenodo.org/record/5276581/files/figure.png" pageId="15" pageNumber="1206">47.</figureCitation>
</paragraph>
<paragraph id="E51836F2EA0E617144F8F1F6FDACFB23" blockId="15.[178,1085,1215,1266]" pageId="15" pageNumber="1206">E t y m o l o g y: The specific epithet (adjective) is derived from the name of the county where the species was collected.</paragraph>
<paragraph id="E51836F2EA0E617144F8F034FC75F9EF" blockId="15.[178,1085,1282,1598]" pageId="15" pageNumber="1206">
I n t r a s p e c i f i c v a r i a t i o n: Coloration, size, and proportions are subject to pronounced variation; the body tends to be more slender in smaller than in larger specimens. Also, as is usual with species of the subgenus 
<taxonomicName id="22A74D71EA0E61714687F00BFC65FA82" box="[717,911,1341,1363]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1206" phylum="Arthropoda" rank="subGenus" subGenus="Tropogastrosipalia">
<emphasis id="D7D3EAE0EA0E61714687F00BFC65FA82" box="[717,911,1341,1363]" italics="true" pageId="15" pageNumber="1206">Tropogastrosipalia</emphasis>
</taxonomicName>
, the male secondary sexual characters are almost or completely obsolete in small males. Remarkably, the cristal process of the aedeagus is highly variable, too, even in material from the same locality (
<figureCitation id="7D9C2A77EA0E61714546F0A3FE95FA7A" box="[268,383,1429,1451]" captionStart="Figs 32-39" captionStartId="14.[178,218,1365,1385]" captionTargetBox="[183,1070,219,1338]" captionTargetId="figure-0@14.[178,1084,219,1357]" captionTargetPageId="14" captionText="Figs 32-39: Geostiba sarica nov.sp. from the environs of Sangdeh (type locality): (32) male habitus; (33) male forebody; (34) male abominal segments VI-VIII in lateral view; (35) male abominal segments VII-VIII in dorsal view; (36-38) median lobe of aedeagus in lateral view; (39) spermatheca. Scale bars: 32: 1.0 mm; 33-35: 0.5 mm; 36-39: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276579" httpUri="https://zenodo.org/record/5276579/files/figure.png" pageId="15" pageNumber="1206">Figs 36-38</figureCitation>
). The male from the environs of Qolqol differs from the males from the 
<typeStatus id="3A1C8850EA0E61714493F085FEEFFA19" box="[217,261,1459,1480]" pageId="15" pageNumber="1206">type</typeStatus>
locality not only by the much larger and broader cristal process (
<figureCitation id="7D9C2A77EA0E617147F3F084FBEDFA19" box="[953,1031,1458,1480]" captionStart="Figs 40-47" captionStartId="16.[178,218,1255,1275]" captionTargetBox="[182,1056,219,1229]" captionTargetId="figure-203@16.[178,1085,219,1244]" captionTargetPageId="16" captionText="Figs 40-47: Geostiba sarica nov.sp. from the environs of Qolqol: (40) male habitus; (41) male forebody; (42) male abdominal segments V-VII in lateral view; (43) male abdominal segments VI- VII in dorsal view; (44) male tergite VIII; (45) male sternite VIII; (46) median lobe of aedeagus in lateral view; (47) spermatheca. Scale bars: 40: 1.0 mm; 41-42: 0.5 mm; 43-45: 0.2 mm; 46-47: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276581" httpUri="https://zenodo.org/record/5276581/files/figure.png" pageId="15" pageNumber="1206">Fig. 46</figureCitation>
), but also by the slightly denser and more distinctly granulose punctation of the elytra (
<figureCitation id="7D9C2A77EA0E61714059F0E6FF26F9D2" captionStart="Figs 40-47" captionStartId="16.[178,218,1255,1275]" captionTargetBox="[182,1056,219,1229]" captionTargetId="figure-203@16.[178,1085,219,1244]" captionTargetPageId="16" captionText="Figs 40-47: Geostiba sarica nov.sp. from the environs of Qolqol: (40) male habitus; (41) male forebody; (42) male abdominal segments V-VII in lateral view; (43) male abdominal segments VI- VII in dorsal view; (44) male tergite VIII; (45) male sternite VIII; (46) median lobe of aedeagus in lateral view; (47) spermatheca. Scale bars: 40: 1.0 mm; 41-42: 0.5 mm; 43-45: 0.2 mm; 46-47: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276581" httpUri="https://zenodo.org/record/5276581/files/figure.png" pageId="15" pageNumber="1206">Fig. 41</figureCitation>
) and by the dorso-ventrally slightly more compressed process of the abdominal tergite VII (
<figureCitation id="7D9C2A77EA0E617144AEF33DFED9F9F0" box="[228,307,1547,1569]" captionStart="Figs 40-47" captionStartId="16.[178,218,1255,1275]" captionTargetBox="[182,1056,219,1229]" captionTargetId="figure-203@16.[178,1085,219,1244]" captionTargetPageId="16" captionText="Figs 40-47: Geostiba sarica nov.sp. from the environs of Qolqol: (40) male habitus; (41) male forebody; (42) male abdominal segments V-VII in lateral view; (43) male abdominal segments VI- VII in dorsal view; (44) male tergite VIII; (45) male sternite VIII; (46) median lobe of aedeagus in lateral view; (47) spermatheca. Scale bars: 40: 1.0 mm; 41-42: 0.5 mm; 43-45: 0.2 mm; 46-47: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5276581" httpUri="https://zenodo.org/record/5276581/files/figure.png" pageId="15" pageNumber="1206">Fig. 42</figureCitation>
). In view of the variability observed in the material from the 
<typeStatus id="3A1C8850EA0E617147FCF33DFC08F9F1" box="[950,994,1547,1568]" pageId="15" pageNumber="1206">type</typeStatus>
locality, these differences are attributed to intra- rather than interspecific variation.
</paragraph>
<caption id="B1D8667AEA11616E44F8F1D1FF36FA87" ID-DOI="http://doi.org/10.5281/zenodo.5276581" ID-Zenodo-Dep="5276581" httpUri="https://zenodo.org/record/5276581/files/figure.png" pageId="16" pageNumber="1207" startId="16.[178,218,1255,1275]" targetBox="[182,1056,219,1229]" targetPageId="16">
<paragraph id="E51836F2EA11616E44F8F1D1FF36FA87" blockId="16.[178,1085,1255,1366]" pageId="16" pageNumber="1207">
<emphasis id="D7D3EAE0EA11616E44F8F1D1FEF3FB2A" bold="true" box="[178,281,1255,1275]" pageId="16" pageNumber="1207">Figs 40-47</emphasis>
: 
<taxonomicName id="22A74D71EA11616E4562F1D1FE56FB2A" authority="Assing, 2009" authorityName="Assing" authorityYear="2009" box="[296,444,1255,1275]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1207" phylum="Arthropoda" rank="species" species="sarica" status="sp. nov.">
<emphasis id="D7D3EAE0EA11616E4562F1D1FE56FB2A" box="[296,444,1255,1275]" italics="true" pageId="16" pageNumber="1207">Geostiba sarica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCE0579BEA11616E458FF1D1FDEDFB2A" box="[453,519,1255,1275]" pageId="16" pageNumber="1207" rank="species">nov.sp.</taxonomicNameLabel>
from the environs of Qolqol: (
<emphasis id="D7D3EAE0EA11616E477EF1D1FCA1FB2A" bold="true" box="[820,843,1255,1275]" pageId="16" pageNumber="1207">40</emphasis>
) male habitus; (
<emphasis id="D7D3EAE0EA11616E47A3F1D1FBEAFB2A" bold="true" box="[1001,1024,1255,1275]" pageId="16" pageNumber="1207">41</emphasis>
) male forebody; (
<emphasis id="D7D3EAE0EA11616E4552F1C8FEC5FAC3" bold="true" box="[280,303,1278,1298]" pageId="16" pageNumber="1207">42</emphasis>
) male abdominal segments V-VII in lateral view; (
<emphasis id="D7D3EAE0EA11616E474FF1C8FCF6FAC3" bold="true" box="[773,796,1278,1298]" pageId="16" pageNumber="1207">43</emphasis>
) male abdominal segments VI- VII in dorsal view; (
<emphasis id="D7D3EAE0EA11616E4524F023FE6FFAF8" bold="true" box="[366,389,1301,1321]" pageId="16" pageNumber="1207">44</emphasis>
) male tergite VIII; (
<emphasis id="D7D3EAE0EA11616E460AF023FDBDFAF8" bold="true" box="[576,599,1301,1321]" pageId="16" pageNumber="1207">45</emphasis>
) male sternite VIII; (
<emphasis id="D7D3EAE0EA11616E4756F023FCD9FAF8" bold="true" box="[796,819,1301,1321]" pageId="16" pageNumber="1207">46</emphasis>
) median lobe of aedeagus in lateral view; (
<emphasis id="D7D3EAE0EA11616E457BF01DFEA2FAEE" bold="true" box="[305,328,1323,1343]" pageId="16" pageNumber="1207">47</emphasis>
) spermatheca. Scale bars: 40: 1.0 mm; 41-42: 0.5 mm; 43-45: 0.2 mm; 46-47: 0.1 mm.
</paragraph>
</caption>
<paragraph id="E51836F2EA11616F44F8F052FEAFFEFA" blockId="16.[178,1085,1380,1607]" lastBlockId="17.[178,1084,219,299]" lastPageId="17" lastPageNumber="1208" pageId="16" pageNumber="1207">
C o m p a r a t i v e n o t e s: Previously, only two species of the subgenus 
<taxonomicName id="22A74D71EA11616E44F8F0B4FE9FFA49" box="[178,373,1410,1432]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1207" phylum="Arthropoda" rank="subGenus" subGenus="Tropogastrosipalia">
<emphasis id="D7D3EAE0EA11616E44F8F0B4FE9FFA49" box="[178,373,1410,1432]" italics="true" pageId="16" pageNumber="1207">Tropogastrosipalia</emphasis>
</taxonomicName>
were known from 
<collectingCountry id="9DB07662EA11616E4603F0B4FD9EFA46" box="[585,628,1410,1431]" name="Iran" pageId="16" pageNumber="1207">Iran</collectingCountry>
, 
<taxonomicName id="22A74D71EA11616E46CEF0B4FC9CFA49" authority="PACE 1983" authorityName="PACE" authorityYear="1983" box="[644,886,1410,1432]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1207" phylum="Arthropoda" rank="species" species="sengleti">
<emphasis id="D7D3EAE0EA11616E46CEF0B4FD1DFA49" box="[644,759,1410,1432]" italics="true" pageId="16" pageNumber="1207">G. sengleti</emphasis>
PACE 1983
</taxonomicName>
(
<collectingRegion id="2763F810EA11616E47C2F0B4FF01FA64" country="Iran" name="Mazandaran" pageId="16" pageNumber="1207">Mazandaran province</collectingRegion>
, 
<geoCoordinate id="80935035EA11616E44B2F0A9FEA7FA64" box="[248,333,1439,1461]" degrees="37" direction="north" minutes="20" orientation="latitude" pageId="16" pageNumber="1207" precision="925" value="37.333332">37°20'N</geoCoordinate>
, 
<geoCoordinate id="80935035EA11616E4510F0A9FE47FA64" box="[346,429,1439,1461]" degrees="55" direction="east" minutes="44" orientation="longitude" pageId="16" pageNumber="1207" precision="925" value="55.733334">55°44'E</geoCoordinate>
] and 
<taxonomicName id="22A74D71EA11616E45A0F0A9FD23FA64" authority="PACE 1983" authorityName="PACE" authorityYear="1983" box="[490,713,1439,1461]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1207" phylum="Arthropoda" rank="species" species="huberi">
<emphasis id="D7D3EAE0EA11616E45A0F0A9FDA4FA64" box="[490,590,1439,1461]" italics="true" pageId="16" pageNumber="1207">G. huberi</emphasis>
PACE 1983
</taxonomicName>
(between Bonjurd and Shapasand). The latter is a species of doubtful status, since its description is based on a single female; 
<taxonomicName id="22A74D71EA11616E44F8F0ECFE9FFA21" box="[178,373,1498,1520]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1235" phylum="Arthropoda" rank="subGenus" subGenus="Tropogastrosipalia">
<emphasis id="D7D3EAE0EA11616E44F8F0ECFE9FFA21" box="[178,373,1498,1520]" italics="true" pageId="16" pageNumber="1207">Tropogastrosipalia</emphasis>
</taxonomicName>
species can be distinguished only based on the male sexual characters. The new species differs from 
<taxonomicName id="22A74D71EA11616E4651F0C1FD61F9DC" authorityName="PACE" authorityYear="1983" box="[539,651,1527,1549]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1207" phylum="Arthropoda" rank="species" species="sengleti">
<emphasis id="D7D3EAE0EA11616E4651F0C1FD61F9DC" box="[539,651,1527,1549]" italics="true" pageId="16" pageNumber="1207">G. sengleti</emphasis>
</taxonomicName>
by the posteriorly more distinctly pointed male pronotum, the absence of distinct sutural carinae and the presence of distinctly granulose punctation on the male elytra, the shape of the cristal process of the aedeagus, and the completely different shape of the spermatheca. For figures of 
<taxonomicName id="22A74D71EA10616F47EAF5EDFBF8FF20" authorityName="PACE" authorityYear="1983" box="[928,1042,219,241]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1208" phylum="Arthropoda" rank="species" species="sengleti">
<emphasis id="D7D3EAE0EA10616F47EAF5EDFBF8FF20" box="[928,1042,219,241]" italics="true" pageId="17" pageNumber="1208">G. sengleti</emphasis>
</taxonomicName>
see 
<bibRefCitation id="81364B03EA10616F44F8F5CFFED6FEDF" author="PACE R" box="[178,316,248,270]" pageId="17" pageNumber="1208" pagination="129 - 139" refId="ref43768" refString="PACE R. (1983 a): Nuove specie europee ed asiatiche del genere Geostiba THOMSON (Coleoptera Staphylinidae). - Giornale italiano di Entomologia 1: 129 - 139." type="journal article" year="1983">PACE (1983a)</bibRefCitation>
and 
<bibRefCitation id="81364B03EA10616F4524F5CEFDF8FEDF" author="ASSING V" box="[366,530,248,270]" pageId="17" pageNumber="1208" pagination="903 - 1006" refId="ref43550" refString="ASSING V. (2005 a): A revision of the species of Geostiba THOMSON and Tropimenelytron PACE of the Eastern Mediterranean, the Caucasus, and adjacent regions (Coleoptera: Staphylinidae, Aleocharinae). - Linzer biologische Beitrage 37 (2): 903 - 1006." type="journal article" year="2005">ASSING (2005a)</bibRefCitation>
, for an illustration of the spermatheca of 
<taxonomicName id="22A74D71EA10616F47FFF5CEFBFCFEDF" authorityName="PACE" authorityYear="1983" box="[949,1046,248,270]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1208" phylum="Arthropoda" rank="species" species="huberi">
<emphasis id="D7D3EAE0EA10616F47FFF5CEFBFCFEDF" box="[949,1046,248,270]" italics="true" pageId="17" pageNumber="1208">G. huberi</emphasis>
</taxonomicName>
see 
<bibRefCitation id="81364B03EA10616F44F8F420FEABFEFA" author="PACE R" box="[178,321,277,299]" pageId="17" pageNumber="1208" pagination="3 - 46" refId="ref43801" refString="PACE R. (1983 b): Specie del genere Geostiba THOMSON raccolte dal Dr. C. Besuchet e collaboratori in Marocco, nella Penisola Iberica e Balcanica, e nel Medio Oriente (Coleoptera, Staphylinidae). - Revue suisse de Zoologie 90: 3 - 46." type="journal article" year="1983">PACE (1983b)</bibRefCitation>
.
</paragraph>
<paragraph id="E51836F2EA10616F44F8F47DFEDAFE69" blockId="17.[178,1084,330,440]" pageId="17" pageNumber="1208">
D i s t r i b u t i o n a n d b i o n o m i c s: 
<taxonomicName id="22A74D71EA10616F46F9F47CFCB0FEB1" authorityName="Assing" authorityYear="2009" box="[691,858,330,352]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1208" phylum="Arthropoda" rank="species" species="sarica">
<emphasis id="D7D3EAE0EA10616F46F9F47CFCB0FEB1" box="[691,858,330,352]" italics="true" pageId="17" pageNumber="1208">Geostiba sarica</emphasis>
</taxonomicName>
is known from two localities in Sari county, 
<collectingRegion id="2763F810EA10616F45FFF45EFD7FFEAF" box="[437,661,360,382]" country="Iran" name="Mazandaran" pageId="17" pageNumber="1208">Mazandaran province</collectingRegion>
, northern 
<collectingCountry id="9DB07662EA10616F46B5F45EFCC1FEAC" box="[767,811,360,381]" name="Iran" pageId="17" pageNumber="1208">Iran</collectingCountry>
. The 
<typeStatus id="3A1C8850EA10616F472CF45EFC78FEAC" box="[870,914,360,381]" pageId="17" pageNumber="1208">type</typeStatus>
specimens were sifted from leaf litter, at least partly in a beech forest, at altitudes of approximately 920 and 
<quantity id="225F9B17EA10616F4496F494FEC1FE69" box="[220,299,418,440]" metricMagnitude="3" metricUnit="m" metricValue="1.53" pageId="17" pageNumber="1208" unit="m" value="1530.0">1530 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>