171 lines
20 KiB
XML
171 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.93.87797" ID-Pensoft-Pub="1314-4049-93-57" ID-Pensoft-UUID="BB43F52F24035E65B1B2CFA0ECED9321" ModsDocID="1314-4049-93-57" checkinTime="1664560496946" checkinUser="pensoft" docAuthor="Bao, Dan-Feng, Bhat, Darbhe J., Boonmee, Saranyaphat, Hyde, Kevin D., Luo, Zong-Long & Nalumpang, Sarunya" docDate="2022" docId="A61481703B365B969677CAD862AE897E" docLanguage="en" docName="MycoKeys 93: 57-79" docOrigin="MycoKeys 93" docPubDate="2022-09-29" docSource="http://dx.doi.org/10.3897/mycokeys.93.87797" docTitle="Dematipyriforma nigrospora D. F. Bao, K. D. Hyde & Z. L. Luo 2022, comb. nov." docType="treatment" docVersion="1" id="BB43F52F24035E65B1B2CFA0ECED9321" lastPageNumber="57" masterDocId="BB43F52F24035E65B1B2CFA0ECED9321" masterDocTitle="Lignicolous freshwater ascomycetes from Thailand: Introducing Dematipyriforma muriformis sp. nov., one new combination and two new records in Pleurotheciaceae" masterLastPageNumber="79" masterPageNumber="57" pageNumber="57" updateTime="1664560496946" updateUser="pensoft">
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<mods:titleInfo>
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<mods:title>Lignicolous freshwater ascomycetes from Thailand: Introducing Dematipyriforma muriformis sp. nov., one new combination and two new records in Pleurotheciaceae</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Bao, Dan-Feng</mods:namePart>
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<mods:affiliation>College of Agronomy and Biosciences, Dali University, Dali 671003, Yunnan, China & Center of Excellence in Fungal Research, Mae Fah Luang University, Chiang Rai 57100, Thailand & Department of Entomology & Plant Pathology, Faculty of Agriculture, Chiang Mai University, Chiang Mai 50200, Thailand</mods:affiliation>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Bhat, Darbhe J.</mods:namePart>
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<mods:affiliation>128 / 1 - J, Azad Housing Society, Curca, Goa Velha- 403108, India</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Boonmee, Saranyaphat</mods:namePart>
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<mods:affiliation>Center of Excellence in Fungal Research, Mae Fah Luang University, Chiang Rai 57100, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hyde, Kevin D.</mods:namePart>
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<mods:affiliation>Center of Excellence in Fungal Research, Mae Fah Luang University, Chiang Rai 57100, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Luo, Zong-Long</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7307-4885</mods:nameIdentifier>
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<mods:affiliation>College of Agronomy and Biosciences, Dali University, Dali 671003, Yunnan, China</mods:affiliation>
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<mods:nameIdentifier type="email">514992672@qq.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Nalumpang, Sarunya</mods:namePart>
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<mods:affiliation>Department of Entomology & Plant Pathology, Faculty of Agriculture, Chiang Mai University, Chiang Mai 50200, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">sarunya.v@cmu.ac.th</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>MycoKeys</mods:title>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-09-29</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>93</mods:number>
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<mods:extent unit="page">
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<mods:start>57</mods:start>
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<mods:end>79</mods:end>
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<mods:url>http://dx.doi.org/10.3897/mycokeys.93.87797</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.93.87797</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-4049-93-57</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:A61481703B365B969677CAD862AE897E" httpUri="http://treatment.plazi.org/id/A61481703B365B969677CAD862AE897E" lastPageNumber="57" pageId="0" pageNumber="57">
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<subSubSection pageId="0" pageNumber="57" type="nomenclature">
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<paragraph pageId="0" pageNumber="57">
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<taxonomicName LSID="A6148170-3B36-5B96-9677-CAD862AE897E" authority="(Boonmee, D. F. Bao & K. D. Hyde) D. F. Bao, K. D. Hyde & Z. L. Luo" authorityName="D. F. Bao, K. D. Hyde & Z. L. Luo" authorityYear="2022" baseAuthorityName="Bao & Bhat & Boonmee & Hyde & Luo & Nalumpang" baseAuthorityYear="2022" class="Sordariomycetes" family="Pleurotheciaceae" genus="Dematipyriforma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Dematipyriforma nigrospora" order="Pleurotheciales" pageId="0" pageNumber="57" phylum="Ascomycota" rank="species" species="nigrospora" status="comb. nov.">Dematipyriforma nigrospora (Boonmee, D.F. Bao & K.D. Hyde) D.F. Bao, K.D. Hyde & Z.L. Luo</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="57">comb. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="57" type="reference_group">
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<paragraph pageId="0" pageNumber="57">
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<taxonomicName authorityName="Bao & Bhat & Boonmee & Hyde & Luo & Nalumpang" authorityYear="2022" baseAuthorityName="Bao & Bhat & Boonmee & Hyde & Luo & Nalumpang" baseAuthorityYear="2022" class="Sordariomycetes" family="Pleurotheciaceae" genus="Rhexoacrodictys" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhexoacrodictys nigrospora" order="Pleurotheciales" pageId="0" pageNumber="57" phylum="Ascomycota" rank="species" species="nigrospora" status="comb. nov.">Rhexoacrodictys nigrospora</taxonomicName>
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≡
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<taxonomicName authorityName="Bao & Bhat & Boonmee & Hyde & Luo & Nalumpang" authorityYear="2022" baseAuthorityName="Bao & Bhat & Boonmee & Hyde & Luo & Nalumpang" baseAuthorityYear="2022" class="Sordariomycetes" family="Pleurotheciaceae" genus="Rhexoacrodictys" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhexoacrodictys nigrospora" order="Pleurotheciales" pageId="0" pageNumber="57" phylum="Ascomycota" rank="species" species="nigrospora" status="comb. nov.">Rhexoacrodictys nigrospora</taxonomicName>
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Boonmee, D.F. Bao & K.D. Hyde, in Boonmee et al., Fungal Diversity 111: 200 (2021).
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</paragraph>
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</subSubSection>
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||
<subSubSection pageId="0" pageNumber="57" type="holotype">
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<paragraph pageId="0" pageNumber="57">Holotype.</paragraph>
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<paragraph pageId="0" pageNumber="57">Thailand, Phetchabun Province, on decaying bark, 25 July 2019, S. Boonmee, LSP03 (MFLU 21-0073).</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="57" type="description">
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<paragraph pageId="0" pageNumber="57">Descriptions and illustrations.</paragraph>
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<paragraph pageId="0" pageNumber="57">
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See
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<bibRefCitation DOI="https://doi.org/10.1007/s13225-021-00489-3" author="Boonmee, S" journalOrPublisher="Fungal Diversity" pageId="0" pageNumber="57" pagination="1 - 335" refId="B3" refString="Boonmee, S, Wanasinghe, DN, Calabon, MS, Huanraluek, N, Chandrasiri, SKU, Jones, EBG, Rossi, W, Leonardi, M, Singh, SK, Rana, S, Singh, PN, Maurya, DK, Lagashetti, AC, Choudhary, D, Dai, YC, Zhao, CL, Mu, YH, Yuan, HS, He, SH, Phookamsak, R, Jiang, HB, Martin, MP, Duenas, M, Telleria, MT, Kalucka, IL, Jagodzinski, AM, Liimatainen, K, Pereira, DS, Phillips, AJL, Suwannarach, N, Kumla, J, Khuna, S, Lumyong, S, Potter, TB, Shivas, RG, Sparks, AH, Vaghefi, N, Abdel-Wahab, MA, Abdel-Aziz, FA, Li, GJ, Lin, WF, Singh, U, Bhatt, RP, Lee, HB, Nguyen, TTT, Kirk, PM, Dutta, AK, Acharya, K, Sarma, VV, Niranjan, M, Rajeshkumar, KC, Ashtekar, N, Lad, S, Wijayawardene, NN, Bhat, DJ, Xu, RJ, Wijesinghe, SN, Shen, HW, Luo, ZL, Zhang, JY, Sysouphanthong, P, Thongklang, N, Bao, DF, Aluthmuhandiram, JVS, Abdollahzadeh, J, Javadi, A, Dovana, F, Usman, M, Khalid, AN, Dissanayake, AJ, Telagathoti, A, Probst, M, Peintner, U, Garrido-Benavent, I, Bona, L, Merenyi, Z, Boros, L, Zoltan, B, Stielow, JB, Jiang, N, Tian, CM, Shams, E, Dehghanizadeh, F, Pordel, A, Javan-Nikkhah, M, Denchev, TT, Denchev, CM, Kemler, M, Begerow, D, Deng, CY, Harrower, E, Bozorov, T, Kholmuradova, T, Gafforov, Y, Abdurazakov, A, Xu, JC, Mortimer, PE, Ren, GC, Jeewon, R, Maharachchikumbura, SSN, Phukhamsakda, C, Mapook, A, Hyde, KD, 2021. Fungal diversity notes 1387-1511: Taxonomic and phylogenetic contributions on genera and species of fungal taxa. Fungal Diversity 111 (1): 1 - 335, DOI: https://doi.org/10.1007/s13225-021-00489-3" title="Fungal diversity notes 1387 - 1511: Taxonomic and phylogenetic contributions on genera and species of fungal taxa." url="https://doi.org/10.1007/s13225-021-00489-3" volume="111" year="2021">Boonmee et al. (2021)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="57" type="notes">
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<paragraph pageId="0" pageNumber="57">Notes.</paragraph>
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<paragraph pageId="0" pageNumber="57">
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<taxonomicName authorityName="Bao & Bhat & Boonmee & Hyde & Luo & Nalumpang" authorityYear="2022" baseAuthorityName="Bao & Bhat & Boonmee & Hyde & Luo & Nalumpang" baseAuthorityYear="2022" class="Sordariomycetes" family="Pleurotheciaceae" genus="Rhexoacrodictys" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhexoacrodictys nigrospora" order="Pleurotheciales" pageId="0" pageNumber="57" phylum="Ascomycota" rank="species" species="nigrospora">
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<emphasis italics="true" pageId="0" pageNumber="57">Rhexoacrodictys nigrospora</emphasis>
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</taxonomicName>
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was introduced by
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<bibRefCitation DOI="https://doi.org/10.1007/s13225-021-00489-3" author="Boonmee, S" journalOrPublisher="Fungal Diversity" pageId="0" pageNumber="57" pagination="1 - 335" refId="B3" refString="Boonmee, S, Wanasinghe, DN, Calabon, MS, Huanraluek, N, Chandrasiri, SKU, Jones, EBG, Rossi, W, Leonardi, M, Singh, SK, Rana, S, Singh, PN, Maurya, DK, Lagashetti, AC, Choudhary, D, Dai, YC, Zhao, CL, Mu, YH, Yuan, HS, He, SH, Phookamsak, R, Jiang, HB, Martin, MP, Duenas, M, Telleria, MT, Kalucka, IL, Jagodzinski, AM, Liimatainen, K, Pereira, DS, Phillips, AJL, Suwannarach, N, Kumla, J, Khuna, S, Lumyong, S, Potter, TB, Shivas, RG, Sparks, AH, Vaghefi, N, Abdel-Wahab, MA, Abdel-Aziz, FA, Li, GJ, Lin, WF, Singh, U, Bhatt, RP, Lee, HB, Nguyen, TTT, Kirk, PM, Dutta, AK, Acharya, K, Sarma, VV, Niranjan, M, Rajeshkumar, KC, Ashtekar, N, Lad, S, Wijayawardene, NN, Bhat, DJ, Xu, RJ, Wijesinghe, SN, Shen, HW, Luo, ZL, Zhang, JY, Sysouphanthong, P, Thongklang, N, Bao, DF, Aluthmuhandiram, JVS, Abdollahzadeh, J, Javadi, A, Dovana, F, Usman, M, Khalid, AN, Dissanayake, AJ, Telagathoti, A, Probst, M, Peintner, U, Garrido-Benavent, I, Bona, L, Merenyi, Z, Boros, L, Zoltan, B, Stielow, JB, Jiang, N, Tian, CM, Shams, E, Dehghanizadeh, F, Pordel, A, Javan-Nikkhah, M, Denchev, TT, Denchev, CM, Kemler, M, Begerow, D, Deng, CY, Harrower, E, Bozorov, T, Kholmuradova, T, Gafforov, Y, Abdurazakov, A, Xu, JC, Mortimer, PE, Ren, GC, Jeewon, R, Maharachchikumbura, SSN, Phukhamsakda, C, Mapook, A, Hyde, KD, 2021. Fungal diversity notes 1387-1511: Taxonomic and phylogenetic contributions on genera and species of fungal taxa. Fungal Diversity 111 (1): 1 - 335, DOI: https://doi.org/10.1007/s13225-021-00489-3" title="Fungal diversity notes 1387 - 1511: Taxonomic and phylogenetic contributions on genera and species of fungal taxa." url="https://doi.org/10.1007/s13225-021-00489-3" volume="111" year="2021">Boonmee et al. (2021)</bibRefCitation>
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based on morphological characters and phylogenetic analysis. In our phylogenetic analysis,
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<taxonomicName class="Sordariomycetes" family="Pleurotheciaceae" kingdom="Fungi" lsidName="R. nigrospora" order="Pleurotheciales" pageId="0" pageNumber="57" phylum="Ascomycota" rank="species" species="nigrospora">
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<emphasis italics="true" pageId="0" pageNumber="57">R. nigrospora</emphasis>
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</taxonomicName>
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clustered with two
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<taxonomicName class="Sordariomycetes" family="Pleurotheciaceae" genus="Dematipyriforma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Dematipyriforma" order="Pleurotheciales" pageId="0" pageNumber="57" phylum="Ascomycota" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="57">Dematipyriforma</emphasis>
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</taxonomicName>
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species (
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<emphasis italics="true" pageId="0" pageNumber="57">
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<taxonomicName class="Sordariomycetes" family="Pleurotheciaceae" kingdom="Fungi" lsidName="D. aquilaria" order="Pleurotheciales" pageId="0" pageNumber="57" phylum="Ascomycota" rank="species" species="aquilaria">D. aquilaria</taxonomicName>
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and
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<taxonomicName class="Sordariomycetes" family="Pleurotheciaceae" kingdom="Fungi" lsidName="D. muriformis" order="Pleurotheciales" pageId="0" pageNumber="57" phylum="Ascomycota" rank="species" species="muriformis">D. muriformis</taxonomicName>
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</emphasis>
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) in a distinct clade within
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<taxonomicName class="Sordariomycetes" family="Pleurotheciaceae" kingdom="Fungi" lsidName="" order="Pleurotheciales" pageId="0" pageNumber="57" phylum="Ascomycota" rank="family">Pleurotheciaceae</taxonomicName>
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(Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phylogram based on a combined ITS, LSU SSU, RPB 2 and TEF 1 - α sequence data of selected members of four orders of the Savoryellomycetidae. Bootstrap support values for maximum likelihood (ML) greater than 70 % and Bayesian posterior probabilities (PP) greater than 0.95 are given as ML / PP above the nodes. Newly obtained sequences are indicated in red and ex-type strains are in bold." figureDoi="10.3897/mycokeys.93.87797.figure1" httpUri="https://binary.pensoft.net/fig/750598" pageId="0" pageNumber="57">1</figureCitation>
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). Therefore, we transfer
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<taxonomicName authorityName="Bao & Bhat & Boonmee & Hyde & Luo & Nalumpang" authorityYear="2022" baseAuthorityName="Bao & Bhat & Boonmee & Hyde & Luo & Nalumpang" baseAuthorityYear="2022" class="Sordariomycetes" family="Pleurotheciaceae" genus="Rhexoacrodictys" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhexoacrodictys nigrospora" order="Pleurotheciales" pageId="0" pageNumber="57" phylum="Ascomycota" rank="species" species="nigrospora">
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<emphasis italics="true" pageId="0" pageNumber="57">Rhexoacrodictys nigrospora</emphasis>
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</taxonomicName>
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to
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<taxonomicName class="Sordariomycetes" family="Pleurotheciaceae" genus="Dematipyriforma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Dematipyriforma" order="Pleurotheciales" pageId="0" pageNumber="57" phylum="Ascomycota" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="57">Dematipyriforma</emphasis>
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</taxonomicName>
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, as
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<taxonomicName authorityName="D. F. Bao, K. D. Hyde & Z. L. Luo" authorityYear="2022" baseAuthorityName="Bao & Bhat & Boonmee & Hyde & Luo & Nalumpang" baseAuthorityYear="2022" class="Sordariomycetes" family="Pleurotheciaceae" genus="Dematipyriforma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Dematipyriforma nigrospora" order="Pleurotheciales" pageId="0" pageNumber="57" phylum="Ascomycota" rank="species" species="nigrospora">
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<emphasis italics="true" pageId="0" pageNumber="57">Dematipyriforma nigrospora</emphasis>
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</taxonomicName>
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comb. nov.
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</paragraph>
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<paragraph pageId="0" pageNumber="57">
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<taxonomicName authorityName="D. F. Bao, K. D. Hyde & Z. L. Luo" authorityYear="2022" baseAuthorityName="Bao & Bhat & Boonmee & Hyde & Luo & Nalumpang" baseAuthorityYear="2022" class="Sordariomycetes" family="Pleurotheciaceae" genus="Dematipyriforma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Dematipyriforma nigrospora" order="Pleurotheciales" pageId="0" pageNumber="57" phylum="Ascomycota" rank="species" species="nigrospora">
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<emphasis italics="true" pageId="0" pageNumber="57">Dematipyriforma nigrospora</emphasis>
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</taxonomicName>
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resembles
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<taxonomicName class="Sordariomycetes" family="Pleurotheciaceae" kingdom="Fungi" lsidName="D. muriformis" order="Pleurotheciales" pageId="0" pageNumber="57" phylum="Ascomycota" rank="species" species="muriformis">
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<emphasis italics="true" pageId="0" pageNumber="57">D. muriformis</emphasis>
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</taxonomicName>
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in having micronematous or semi-macronematous, mononematous conidiophores and monoblastic, polyblastic, integrated, terminal conidiogenous cells. However,
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<taxonomicName class="Sordariomycetes" family="Pleurotheciaceae" kingdom="Fungi" lsidName="D. nigrospora" order="Pleurotheciales" pageId="0" pageNumber="57" phylum="Ascomycota" rank="species" species="nigrospora">
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<emphasis italics="true" pageId="0" pageNumber="57">D. nigrospora</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName class="Sordariomycetes" family="Pleurotheciaceae" kingdom="Fungi" lsidName="D. muriformis" order="Pleurotheciales" pageId="0" pageNumber="57" phylum="Ascomycota" rank="species" species="muriformis">
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<emphasis italics="true" pageId="0" pageNumber="57">D. muriformis</emphasis>
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</taxonomicName>
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in having brown to dark brown conidiophores and globose to subglobose, dark brown to black conidia (
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<bibRefCitation DOI="https://doi.org/10.1007/s13225-021-00489-3" author="Boonmee, S" journalOrPublisher="Fungal Diversity" pageId="0" pageNumber="57" pagination="1 - 335" refId="B3" refString="Boonmee, S, Wanasinghe, DN, Calabon, MS, Huanraluek, N, Chandrasiri, SKU, Jones, EBG, Rossi, W, Leonardi, M, Singh, SK, Rana, S, Singh, PN, Maurya, DK, Lagashetti, AC, Choudhary, D, Dai, YC, Zhao, CL, Mu, YH, Yuan, HS, He, SH, Phookamsak, R, Jiang, HB, Martin, MP, Duenas, M, Telleria, MT, Kalucka, IL, Jagodzinski, AM, Liimatainen, K, Pereira, DS, Phillips, AJL, Suwannarach, N, Kumla, J, Khuna, S, Lumyong, S, Potter, TB, Shivas, RG, Sparks, AH, Vaghefi, N, Abdel-Wahab, MA, Abdel-Aziz, FA, Li, GJ, Lin, WF, Singh, U, Bhatt, RP, Lee, HB, Nguyen, TTT, Kirk, PM, Dutta, AK, Acharya, K, Sarma, VV, Niranjan, M, Rajeshkumar, KC, Ashtekar, N, Lad, S, Wijayawardene, NN, Bhat, DJ, Xu, RJ, Wijesinghe, SN, Shen, HW, Luo, ZL, Zhang, JY, Sysouphanthong, P, Thongklang, N, Bao, DF, Aluthmuhandiram, JVS, Abdollahzadeh, J, Javadi, A, Dovana, F, Usman, M, Khalid, AN, Dissanayake, AJ, Telagathoti, A, Probst, M, Peintner, U, Garrido-Benavent, I, Bona, L, Merenyi, Z, Boros, L, Zoltan, B, Stielow, JB, Jiang, N, Tian, CM, Shams, E, Dehghanizadeh, F, Pordel, A, Javan-Nikkhah, M, Denchev, TT, Denchev, CM, Kemler, M, Begerow, D, Deng, CY, Harrower, E, Bozorov, T, Kholmuradova, T, Gafforov, Y, Abdurazakov, A, Xu, JC, Mortimer, PE, Ren, GC, Jeewon, R, Maharachchikumbura, SSN, Phukhamsakda, C, Mapook, A, Hyde, KD, 2021. Fungal diversity notes 1387-1511: Taxonomic and phylogenetic contributions on genera and species of fungal taxa. Fungal Diversity 111 (1): 1 - 335, DOI: https://doi.org/10.1007/s13225-021-00489-3" title="Fungal diversity notes 1387 - 1511: Taxonomic and phylogenetic contributions on genera and species of fungal taxa." url="https://doi.org/10.1007/s13225-021-00489-3" volume="111" year="2021">Boonmee et al. 2021</bibRefCitation>
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). Conidiophores of
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<taxonomicName class="Sordariomycetes" family="Pleurotheciaceae" kingdom="Fungi" lsidName="D. muriformis" order="Pleurotheciales" pageId="0" pageNumber="57" phylum="Ascomycota" rank="species" species="muriformis">
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<emphasis italics="true" pageId="0" pageNumber="57">D. muriformis</emphasis>
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</taxonomicName>
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are hyaline and conidia are ellipsoidal to obovoid, muriform, and subhyaline to pale olivaceous when young, olive to dark brown at maturity.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |