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<document id="1535B7F70C4F01E3BC3F02052ABF8D50" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629734358953" checkinUser="felipe" docAuthor="Nielson, M. W." docDate="2011" docId="27115005FFFE78466FEE1E4FFB71789C" docLanguage="en" docName="zt02953p085.pdf" docOrigin="Zootaxa 2953" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Boliviela caverna Nielson 2011, sp. nov." docType="treatment" docVersion="3" lastPageNumber="13" masterDocId="DB28287DFFF5784A6F791B38FFC97968" masterDocTitle="2953" masterLastPageNumber="85" masterPageNumber="1" pageNumber="12" updateTime="1699103988635" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:namePart id="4E91A6A2026C11414D8AD605514AEEB1">Nielson, M. W.</mods:namePart>
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<mods:date id="8BDF933BE7739201A323C8733AA2DA01">2011</mods:date>
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<mods:number id="D557E48F0E45BC1584165B999674416B">2011-07-08</mods:number>
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<treatment id="27115005FFFE78466FEE1E4FFB71789C" ID-DOI="http://doi.org/10.5281/zenodo.5288224" ID-Zenodo-Dep="5288224" LSID="urn:lsid:plazi:treatment:27115005FFFE78466FEE1E4FFB71789C" httpUri="http://treatment.plazi.org/id/27115005FFFE78466FEE1E4FFB71789C" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<subSubSection id="E7A2B298FFFE78416FEE1E4FFE0E7CF8" box="[151,455,1399,1424]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="AF07E113FFFE78416FEE1E4FFE0E7CF8" blockId="11.[151,455,1399,1460]" box="[151,455,1399,1424]" pageId="11" pageNumber="12">
<heading id="F44F567FFFFE78416FEE1E4FFE0E7CF8" bold="true" box="[151,455,1399,1424]" fontSize="10" level="2" pageId="11" pageNumber="12" reason="2">
<emphasis id="9DCC3D01FFFE78416FEE1E4FFE0E7CF8" bold="true" box="[151,455,1399,1424]" pageId="11" pageNumber="12">
<taxonomicName id="68B89A90FFFE78416FEE1E4FFEA97CE7" ID-CoL="efb92a1b-4fd8-45ec-977c-95d67d136773" authority="Nielson, 2011" authorityName="Nielson" authorityYear="2011" box="[151,352,1399,1424]" class="Insecta" family="Cicadellidae" genus="Boliviela" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="caverna" status="sp. nov.">
<emphasis id="9DCC3D01FFFE78416FEE1E4FFEA97CE7" bold="true" box="[151,352,1399,1424]" italics="true" pageId="11" pageNumber="12">Boliviela caverna</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="86FF807AFFFE78416E171E40FE0E7CF8" box="[366,455,1400,1424]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E7A2B298FFFE78466FEE1EA4FC1879BC" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="description">
<paragraph id="AF07E113FFFE78416FEE1EA4FE527CDC" blockId="11.[151,455,1399,1460]" box="[151,411,1436,1460]" pageId="11" pageNumber="12">
(
<figureCitation id="3783FD96FFFE78416FE71EA4FEC87CDC" box="[158,257,1436,1460]" captionStart="PLATE 1" captionStartId="41.[151,229,1959,1980]" captionTargetBox="[171,1414,151,1944]" captionTargetId="figure-73@41.[167,1432,136,1955]" captionTargetPageId="41" captionText="PLATE 1. AH. General habitus. (A) Dicodia recurvata, sp. nov.; (B) Boliviela cucurbita, sp. nov.; (C) Boliviela caverna, sp. nov.; (D) Boliviela biungulata, sp. nov.; (E) Collasuyusana recurvata, sp. nov.; (F) Collasuyusana collaris, sp. nov.; (G) Collasuyusana diversa, sp. nov.; (H) Collasuyusana spenceri, sp. nov." pageId="11" pageNumber="12">Plate 1C</figureCitation>
,
<figureCitation id="3783FD96FFFE78416E771EA4FE5A7CDC" box="[270,403,1436,1460]" captionStart="FIGURES 1724" captionStartId="48.[151,258,1947,1968]" captionTargetBox="[204,1374,168,1893]" captionTargetId="figure-76@48.[196,1380,159,1893]" captionTargetPageId="48" captionText="FIGURES 1724. Boliviela caverna, sp. nov. (17) pygofer, lateral view; (18) right subgenital plate, ventral view; (19) right style, lateral view; (20) right style, dorsal view; (21) aedeagus and dorsal connective, lateral view; (22) aedeagus, dorsal view; (23) connective, caudal view; (24) dorsal connective, attached to base of aedeagal socle, dorsal view." pageId="11" pageNumber="12">Figs. 1724</figureCitation>
)
</paragraph>
<paragraph id="AF07E113FFFE78416FEE1EDCFDB47C94" blockId="11.[151,1436,1508,2036]" box="[151,637,1508,1532]" pageId="11" pageNumber="12">
<emphasis id="9DCC3D01FFFE78416FEE1EDCFF3A7C94" bold="true" box="[151,243,1508,1532]" inLineHeading="true" pageId="11" pageNumber="12" reason="1">Length.</emphasis>
Male 8.50 mm., female unknown.
</paragraph>
<paragraph id="AF07E113FFFE78416FBE1D30FE287E74" blockId="11.[151,1436,1508,2036]" pageId="11" pageNumber="12">
<emphasis id="9DCC3D01FFFE78416FBE1D30FE057F48" bold="true" box="[199,460,1544,1568]" inLineHeading="true" pageId="11" pageNumber="12" reason="1">External morphology.</emphasis>
Moderately large, slender species. General color black throughout with numerous very small yellow spots on veins and one large yellow marking subapically on forewings; eyes grey; face dark brown, speckled with small yellow spots (Plate IC). Head significantly narrower than pronotum, anterior margin acutely angled; crown narrower than width of eyes, produced distally about ¼ distance of entire median length, lateral margins parallel, slightly carinate, disk depressed; pronotum with median length less than length of crown, surface sparsely bullated; mesonotum about equal in length to pronotum; forewings typical; clypeus elongate, broad in basal half, narrowed in apical half; clypellus short, about ¼ as long as clypeus, base very broad, inflated, lateral margins constricted medially.
</paragraph>
<paragraph id="AF07E113FFFE78466FBE1C10FC1879BC" blockId="11.[151,1436,1508,2036]" lastBlockId="12.[151,1436,151,500]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<emphasis id="9DCC3D01FFFE78416FBE1C10FEB37E28" bold="true" box="[199,378,1831,1856]" inLineHeading="true" pageId="11" pageNumber="12" reason="1">Male genitalia.</emphasis>
Pygofer narrow with 2 distinct caudal processes, caudodorsal process long, narrow, curved near middle, caudoventral process very long, broad in basal 1/3, attenuated in apical 2/3, apex sharply pointed (
<figureCitation id="3783FD96FFFE78416A171C74FF7F7EE0" captionStart="FIGURES 1724" captionStartId="48.[151,258,1947,1968]" captionTargetBox="[204,1374,168,1893]" captionTargetId="figure-76@48.[196,1380,159,1893]" captionTargetPageId="48" captionText="FIGURES 1724. Boliviela caverna, sp. nov. (17) pygofer, lateral view; (18) right subgenital plate, ventral view; (19) right style, lateral view; (20) right style, dorsal view; (21) aedeagus and dorsal connective, lateral view; (22) aedeagus, dorsal view; (23) connective, caudal view; (24) dorsal connective, attached to base of aedeagal socle, dorsal view." pageId="11" pageNumber="12">Fig. 17</figureCitation>
); right subgenital plate long, narrow, outer lateral margin expanded in distal ½, sparsely pilose (
<figureCitation id="3783FD96FFFE78416B841C48FA997EE0" box="[1277,1360,1904,1928]" captionStart="FIGURES 1724" captionStartId="48.[151,258,1947,1968]" captionTargetBox="[204,1374,168,1893]" captionTargetId="figure-76@48.[196,1380,159,1893]" captionTargetPageId="48" captionText="FIGURES 1724. Boliviela caverna, sp. nov. (17) pygofer, lateral view; (18) right subgenital plate, ventral view; (19) right style, lateral view; (20) right style, dorsal view; (21) aedeagus and dorsal connective, lateral view; (22) aedeagus, dorsal view; (23) connective, caudal view; (24) dorsal connective, attached to base of aedeagal socle, dorsal view." pageId="11" pageNumber="12">Fig. 18</figureCitation>
); right style long, narrow in middle 2/3, apex expanded in distal ¼ (
<figureCitation id="3783FD96FFFE78416C2D1CACFC287EC4" box="[852,993,1940,1964]" captionStart="FIGURES 1724" captionStartId="48.[151,258,1947,1968]" captionTargetBox="[204,1374,168,1893]" captionTargetId="figure-76@48.[196,1380,159,1893]" captionTargetPageId="48" captionText="FIGURES 1724. Boliviela caverna, sp. nov. (17) pygofer, lateral view; (18) right subgenital plate, ventral view; (19) right style, lateral view; (20) right style, dorsal view; (21) aedeagus and dorsal connective, lateral view; (22) aedeagus, dorsal view; (23) connective, caudal view; (24) dorsal connective, attached to base of aedeagal socle, dorsal view." pageId="11" pageNumber="12">Figs. 19, 20</figureCitation>
); aedeagus long, inflated along middle half in dorsal and lateral views, cavernate at middle with distal flange in lateral view, tubular, curved in apical 2/6, apical 1/6 digitate, sinuate and directed caudodorsally in lateral view, gonopore subapical (
<figureCitation id="3783FD96FFFE78416BFE1CE4FAC67E9C" box="[1159,1295,2012,2036]" captionStart="FIGURES 1724" captionStartId="48.[151,258,1947,1968]" captionTargetBox="[204,1374,168,1893]" captionTargetId="figure-76@48.[196,1380,159,1893]" captionTargetPageId="48" captionText="FIGURES 1724. Boliviela caverna, sp. nov. (17) pygofer, lateral view; (18) right subgenital plate, ventral view; (19) right style, lateral view; (20) right style, dorsal view; (21) aedeagus and dorsal connective, lateral view; (22) aedeagus, dorsal view; (23) connective, caudal view; (24) dorsal connective, attached to base of aedeagal socle, dorsal view." pageId="11" pageNumber="12">Figs. 21, 22</figureCitation>
); connective small, Y-shaped, stem very short (
<figureCitation id="3783FD96FFF978466D5B1BAFFDB179C7" box="[546,632,151,175]" captionStart="FIGURES 1724" captionStartId="48.[151,258,1947,1968]" captionTargetBox="[204,1374,168,1893]" captionTargetId="figure-76@48.[196,1380,159,1893]" captionTargetPageId="48" captionText="FIGURES 1724. Boliviela caverna, sp. nov. (17) pygofer, lateral view; (18) right subgenital plate, ventral view; (19) right style, lateral view; (20) right style, dorsal view; (21) aedeagus and dorsal connective, lateral view; (22) aedeagus, dorsal view; (23) connective, caudal view; (24) dorsal connective, attached to base of aedeagal socle, dorsal view." pageId="12" pageNumber="13">Fig. 23</figureCitation>
); dorsal connective long, narrow, bifurcate basally (
<figureCitation id="3783FD96FFF978466BA81BAFFAEE79C7" box="[1233,1319,151,175]" captionStart="FIGURES 1724" captionStartId="48.[151,258,1947,1968]" captionTargetBox="[204,1374,168,1893]" captionTargetId="figure-76@48.[196,1380,159,1893]" captionTargetPageId="48" captionText="FIGURES 1724. Boliviela caverna, sp. nov. (17) pygofer, lateral view; (18) right subgenital plate, ventral view; (19) right style, lateral view; (20) right style, dorsal view; (21) aedeagus and dorsal connective, lateral view; (22) aedeagus, dorsal view; (23) connective, caudal view; (24) dorsal connective, attached to base of aedeagal socle, dorsal view." pageId="12" pageNumber="13">Fig. 24</figureCitation>
), attached basally to dorso-subbasal shaft of aedeagus, not encircling shaft (
<figureCitation id="3783FD96FFF978466C081B84FC0D79BC" box="[881,964,188,212]" captionStart="FIGURES 1724" captionStartId="48.[151,258,1947,1968]" captionTargetBox="[204,1374,168,1893]" captionTargetId="figure-76@48.[196,1380,159,1893]" captionTargetPageId="48" captionText="FIGURES 1724. Boliviela caverna, sp. nov. (17) pygofer, lateral view; (18) right subgenital plate, ventral view; (19) right style, lateral view; (20) right style, dorsal view; (21) aedeagus and dorsal connective, lateral view; (22) aedeagus, dorsal view; (23) connective, caudal view; (24) dorsal connective, attached to base of aedeagal socle, dorsal view." pageId="12" pageNumber="13">Fig. 21</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E7A2B298FFF978466FBE1BE7FDF1780C" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="AF07E113FFF978466FBE1BE7FDF1780C" blockId="12.[151,1436,151,500]" pageId="12" pageNumber="13">
<emphasis id="9DCC3D01FFF978466FBE1BE7FE7D7990" bold="true" box="[199,436,223,248]" inLineHeading="true" pageId="12" pageNumber="13" reason="1">Material examined.</emphasis>
<materialsCitation id="1FD0EB4EFFF978466EB51BE7FACC7874" collectionCode="IAHC" collectorName="Nov &amp; Dec &amp; Malaise, D" country="Colombia" county="Cueva" elevation="2020" latitude="1.6166667" location="Borde de Rio Suaza" longLatPrecision="1308" longitude="-76.1" municipality="Cuacharos" pageId="12" pageNumber="13" specimenCount="1" stateProvince="Huila" typeStatus="holotype">
<typeStatus id="70035FB1FFF978466EB51BE7FDFD799F" box="[460,564,223,247]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
male.
<collectingCountry id="D7AFA183FFF978466DED1BE7FCE7799F" box="[660,814,223,247]" name="Colombia" pageId="12" pageNumber="13">COLOMBIA</collectingCountry>
:
<collectingRegion id="6D7C2FF1FFF978466C451BE7FCB4799F" box="[828,893,223,247]" country="Colombia" name="Huila" pageId="12" pageNumber="13">Huila</collectingRegion>
, PNN
<collectingCounty id="4666999FFFF978466CA91BE7FBDE799F" box="[976,1047,223,247]" pageId="12" pageNumber="13">Cueva</collectingCounty>
de los
<collectingMunicipality id="4F637B69FFF978466B0C1BE7FB27799F" box="[1141,1262,223,247]" pageId="12" pageNumber="13">Cuacharos</collectingMunicipality>
,
<location id="AA67B7C8FFF978466B871BE7FF147874" LSID="urn:lsid:plazi:treatment:27115005FFFE78466FEE1E4FFB71789C:AA67B7C8FFF978466B871BE7FF147874" country="Colombia" county="Cueva" latitude="1.6166667" longLatPrecision="1308" longitude="-76.1" municipality="Cuacharos" name="Borde de Rio Suaza" pageId="12" pageNumber="13" stateProvince="Huila">Borde de Rio Suaza</location>
,
<geoCoordinate id="CA8C87D4FFF978466F931A3CFE8D7874" box="[234,324,260,284]" degrees="1" direction="north" minutes="37" orientation="latitude" pageId="12" pageNumber="13" precision="925" value="1.6166667">1º 37N</geoCoordinate>
.
<geoCoordinate id="CA8C87D4FFF978466E211A3CFE7A7874" box="[344,435,260,284]" degrees="76" direction="west" minutes="6" orientation="longitude" pageId="12" pageNumber="13" precision="925" value="-76.1">76º6W</geoCoordinate>
.,
<quantity id="68404CF6FFF978466EBD1A3CFDE87873" box="[452,545,260,284]" metricMagnitude="3" metricUnit="m" metricValue="2.02" pageId="12" pageNumber="13" unit="m" value="2020.0">
<elevation id="24950620FFF978466EBD1A3CFDE87873" box="[452,545,260,284]" metricMagnitude="3" metricUnit="m" metricValue="2.02" pageId="12" pageNumber="13" unit="m" value="2020.0">2020 m</elevation>
</quantity>
., 28
<collectorName id="024D84C5FFF978466D201A3CFD407873" box="[601,649,260,283]" pageId="12" pageNumber="13">Nov</collectorName>
0111
<collectorName id="024D84C5FFF978466D981A3CFCC77873" box="[737,782,260,283]" pageId="12" pageNumber="13">Dec</collectorName>
01, M. 2542,
<collectorName id="024D84C5FFF978466CCE1A3CFBFE7873" box="[951,1079,260,284]" pageId="12" pageNumber="13">Malaise, D</collectorName>
. Campos (
<collectionCode id="C9A979D6FFF978466BCB1A3CFB347874" box="[1202,1277,260,284]" pageId="12" pageNumber="13">IAHC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="1FD0EB4EFFF978466A621A3CFDFD780C" collectionCode="MLBM" collectorName="Nov &amp; Feb &amp; Malaise, D" country="Colombia" county="Cueva" elevation="1950" latitude="1.6166667" location="Cabana Cedros" longLatPrecision="1308" longitude="-76.1" municipality="Cuachoros" pageId="12" pageNumber="13" specimenCount="1" specimenCount-male="1" stateProvince="Huila" typeStatus="paratype">
<typeStatus id="70035FB1FFF978466A621A3CFA497874" box="[1307,1408,260,284]" pageId="12" pageNumber="13" type="paratype">Paratype</typeStatus>
,
<specimenCount id="B9BE2A9AFFF978466AF71A3CFF197857" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
,
<collectingRegion id="6D7C2FF1FFF978466FA51A1FFED47857" box="[220,285,295,319]" country="Colombia" name="Huila" pageId="12" pageNumber="13">Huila</collectingRegion>
, PNN
<collectingCounty id="4666999FFFF978466E111A1FFE797857" box="[360,432,295,319]" pageId="12" pageNumber="13">Cueva</collectingCounty>
de los
<collectingMunicipality id="4F637B69FFF978466D791A1FFDB37857" box="[512,634,295,319]" pageId="12" pageNumber="13">Cuachoros</collectingMunicipality>
,
<location id="AA67B7C8FFF978466DFF1A1FFCFD7857" LSID="urn:lsid:plazi:treatment:27115005FFFE78466FEE1E4FFB71789C:AA67B7C8FFF978466DFF1A1FFCFD7857" box="[646,820,295,319]" country="Colombia" county="Cueva" latitude="1.6166667" longLatPrecision="1308" longitude="-76.1" municipality="Cuachoros" name="Cabana Cedros" pageId="12" pageNumber="13" stateProvince="Huila">Cabana Cedros</location>
,
<geoCoordinate id="CA8C87D4FFF978466C461A1FFC447857" box="[831,909,295,319]" degrees="1" direction="north" minutes="37" orientation="latitude" pageId="12" pageNumber="13" precision="925" value="1.6166667">1º37N</geoCoordinate>
.
<geoCoordinate id="CA8C87D4FFF978466CDB1A1FFC377857" box="[930,1022,295,319]" degrees="76" direction="west" minutes="6" orientation="longitude" pageId="12" pageNumber="13" precision="925" value="-76.1">76º6W</geoCoordinate>
.,
<quantity id="68404CF6FFF978466B771A1FFBA17857" box="[1038,1128,295,319]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="12" pageNumber="13" unit="m" value="1950.0">
<elevation id="24950620FFF978466B771A1FFBA17857" box="[1038,1128,295,319]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="12" pageNumber="13" unit="m" value="1950.0">1950 m</elevation>
</quantity>
., 22
<collectorName id="024D84C5FFF978466BE51A10FB057857" box="[1180,1228,296,319]" pageId="12" pageNumber="13">Nov</collectorName>
0112
<collectorName id="024D84C5FFF978466A581A1FFA827857" box="[1313,1355,295,319]" pageId="12" pageNumber="13">Feb</collectorName>
01, M. 2526,
<collectorName id="024D84C5FFF978466FA41A74FE92780B" box="[221,347,332,356]" pageId="12" pageNumber="13">Malaise, D</collectorName>
. Campos (
<collectionCode id="C9A979D6FFF978466EA91A74FDE5780C" box="[464,556,332,356]" pageId="12" pageNumber="13">MLBM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E7A2B298FFF978466FBE1A48FF1C78C4" pageId="12" pageNumber="13" type="etymology">
<paragraph id="AF07E113FFF978466FBE1A48FF1C78C4" blockId="12.[151,1436,151,500]" pageId="12" pageNumber="13">
<emphasis id="9DCC3D01FFF978466FBE1A48FE8778E0" bold="true" box="[199,334,368,392]" inLineHeading="true" pageId="12" pageNumber="13" reason="1">Etymology.</emphasis>
The name is descriptive for a large excavation in the middle of the inflated portion of the aedeagal shaft.
</paragraph>
</subSubSection>
<subSubSection id="E7A2B298FFF978466FBE1A80FB71789C" pageId="12" pageNumber="13" type="discussion">
<paragraph id="AF07E113FFF978466FBE1A80FB71789C" blockId="12.[151,1436,151,500]" pageId="12" pageNumber="13">
<emphasis id="9DCC3D01FFF978466FBE1A80FEF278B8" bold="true" box="[199,315,440,464]" inLineHeading="true" pageId="12" pageNumber="13" reason="1">Remarks.</emphasis>
This species is similar in male genitalia features to
<taxonomicName id="68B89A90FFF978466CF61A81FB9278B8" authority="DeLong" authorityName="DeLong" authorityYear="1969" box="[911,1115,440,464]" class="Insecta" family="Cicadellidae" genus="Boliviela" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="inflata">
<emphasis id="9DCC3D01FFF978466CF61A81FC6978B8" box="[911,928,441,464]" italics="true" pageId="12" pageNumber="13">B</emphasis>
.
<emphasis id="9DCC3D01FFF978466CD61A80FC3178B8" box="[943,1016,440,464]" italics="true" pageId="12" pageNumber="13">inflata</emphasis>
DeLong
</taxonomicName>
and can be separated by the larger size, the excavated aedeagal shaft and the short basal bifurcation of the dorsal apodeme.
</paragraph>
</subSubSection>
</treatment>
</document>