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<document id="69AA8D0D5CCEF0D131DCA4918973D378" ID-DOI="10.11646/zootaxa.3846.4.2" ID-GBIF-Dataset="ea136b1c-dc96-4c41-b078-01ca553453a0" ID-ISSN="1175-5326" ID-Zenodo-Dep="252519" ID-ZooBank="71A80CC3-CDC1-4291-863B-FA7501D60AF5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460467899762" checkinUser="plazi" docAuthor="Câmara, J. T. &amp; Rafael, J. A." docDate="2014" docId="03F6943FE066FF8EC7C7FD5EAD3617AC" docLanguage="en" docName="zt03846p546.pdf" docOrigin="Zootaxa 3846 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Opeatocerata hadrophallus Câmara &amp; Rafael, 2014, sp. nov." docType="treatment" docVersion="11" lastPageNumber="521" masterDocId="FFCFEC47E075FF9DC750FFEAA8211077" masterDocTitle="Revision of Opeatocerata Melander, 1928 (Diptera: Empididae: Empidinae)" masterLastPageNumber="546" masterPageNumber="502" pageNumber="521" updateTime="1698305230595" updateUser="plazi">
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<mods:title id="9F62C64EE273A6FD4E4FEC7E1C137B62">Revision of Opeatocerata Melander, 1928 (Diptera: Empididae: Empidinae)</mods:title>
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<mods:namePart id="5361785B628E669FF2914500E16BE6D6">Câmara, J. T.</mods:namePart>
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<mods:namePart id="206EB45FF2D0BCE10657B64C231297EF">Rafael, J. A.</mods:namePart>
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<mods:date id="D113C8A1B09322124FF7A9F754BB19E1">2014</mods:date>
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<treatment id="03F6943FE066FF8EC7C7FD5EAD3617AC" ID-DOI="http://doi.org/10.5281/zenodo.5285239" ID-GBIF-Taxon="119514712" ID-Zenodo-Dep="5285239" LSID="urn:lsid:plazi:treatment:03F6943FE066FF8EC7C7FD5EAD3617AC" httpUri="http://treatment.plazi.org/id/03F6943FE066FF8EC7C7FD5EAD3617AC" lastPageNumber="521" pageId="19" pageNumber="521">
<subSubSection id="C34576A2E066FF8EC7C7FD5EA9061298" pageId="19" pageNumber="521" type="nomenclature">
<paragraph id="8BE02529E066FF8EC7C7FD5EAA6412B9" blockId="19.[151,581,692,751]" box="[151,581,692,718]" pageId="19" pageNumber="521">
<heading id="D0A89245E066FF8EC7C7FD5EAA6412B9" bold="true" box="[151,581,692,718]" fontSize="11" level="1" pageId="19" pageNumber="521" reason="1">
<emphasis id="B92BF93BE066FF8EC7C7FD5EAA6412B9" bold="true" box="[151,581,692,718]" pageId="19" pageNumber="521">
<taxonomicName id="4C5F5EAAE066FF8EC7C7FD5EA9C112B9" ID-CoL="8D3LK" box="[151,480,692,718]" class="Insecta" family="Empididae" genus="Opeatocerata" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="521" phylum="Arthropoda" rank="species" species="hadrophallus" status="sp. nov.">
<emphasis id="B92BF93BE066FF8EC7C7FD5EA9C112B9" bold="true" box="[151,480,692,718]" italics="true" pageId="19" pageNumber="521">Opeatocerata hadrophallus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2184440E066FF8EC6B7FD5EAA6412B9" box="[487,581,692,718]" pageId="19" pageNumber="521" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE02529E066FF8EC7C7FD3CA9061298" blockId="19.[151,581,692,751]" box="[151,295,726,751]" pageId="19" pageNumber="521">
(
<figureCitation id="136439ACE066FF8EC7CFFD3CA93E1298" box="[159,287,726,751]" captionStart="FIGURES 90 99" captionStartId="20.[151,264,1816,1839]" captionTargetBox="[164,1411,193,1785]" captionTargetId="figure@20.[164,1422,193,1796]" captionTargetPageId="20" captionText="FIGURES 90 99. Opeatocerata hadrophallus sp. nov., holotype ♂. 90. habitus, lateral view; 91. wing; 92. abdomen from tergite 6 to terminalia, lateral view; 93. tergites 4 8, dorsal view; 94. terminalia, lateral view; 95. lobes of cerci, dorsal view; 96. descendant plate of anterior cercus and hypoproct, posterior view; 97. subepandrial sclerite and bacilliform sclerite, dorsal view; 98. ejaculatory apodeme, phallus and hypandrium, lateral view; 99. hypandrium, ventral view." httpUri="https://zenodo.org/record/252531/files/figure.png" pageId="19" pageNumber="521">Figs 9099</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34576A2E066FF8EC7C7FCF4AC0E1308" pageId="19" pageNumber="521" type="diagnosis">
<paragraph id="8BE02529E066FF8EC7C7FCF4AC0E1308" blockId="19.[151,1437,798,2011]" pageId="19" pageNumber="521">
<emphasis id="B92BF93BE066FF8EC7C7FCF4A92B1340" bold="true" box="[151,266,798,823]" pageId="19" pageNumber="521">Diagnosis</emphasis>
. Tergites 36 with paramedian subtriangular black spots (
<figureCitation id="136439ACE066FF8EC4E8FCF4AC311340" box="[952,1040,798,823]" captionStart="FIGURES 90 99" captionStartId="20.[151,264,1816,1839]" captionTargetBox="[164,1411,193,1785]" captionTargetId="figure@20.[164,1422,193,1796]" captionTargetPageId="20" captionText="FIGURES 90 99. Opeatocerata hadrophallus sp. nov., holotype ♂. 90. habitus, lateral view; 91. wing; 92. abdomen from tergite 6 to terminalia, lateral view; 93. tergites 4 8, dorsal view; 94. terminalia, lateral view; 95. lobes of cerci, dorsal view; 96. descendant plate of anterior cercus and hypoproct, posterior view; 97. subepandrial sclerite and bacilliform sclerite, dorsal view; 98. ejaculatory apodeme, phallus and hypandrium, lateral view; 99. hypandrium, ventral view." httpUri="https://zenodo.org/record/252531/files/figure.png" pageId="19" pageNumber="521">Fig. 93</figureCitation>
); descendant plate of the anterior cercus subtrapezoidal with a small median constriction in posterior view (
<figureCitation id="136439ACE066FF8EC49FFCA8AC02132C" box="[975,1059,834,859]" captionStart="FIGURES 90 99" captionStartId="20.[151,264,1816,1839]" captionTargetBox="[164,1411,193,1785]" captionTargetId="figure@20.[164,1422,193,1796]" captionTargetPageId="20" captionText="FIGURES 90 99. Opeatocerata hadrophallus sp. nov., holotype ♂. 90. habitus, lateral view; 91. wing; 92. abdomen from tergite 6 to terminalia, lateral view; 93. tergites 4 8, dorsal view; 94. terminalia, lateral view; 95. lobes of cerci, dorsal view; 96. descendant plate of anterior cercus and hypoproct, posterior view; 97. subepandrial sclerite and bacilliform sclerite, dorsal view; 98. ejaculatory apodeme, phallus and hypandrium, lateral view; 99. hypandrium, ventral view." httpUri="https://zenodo.org/record/252531/files/figure.png" pageId="19" pageNumber="521">Fig. 96</figureCitation>
); phallus with narrow base, wider in mostly, pointed apically; short, not extending beyond the hypandrium (
<figureCitation id="136439ACE066FF8EC49EFC8CAC021308" box="[974,1059,870,895]" captionStart="FIGURES 90 99" captionStartId="20.[151,264,1816,1839]" captionTargetBox="[164,1411,193,1785]" captionTargetId="figure@20.[164,1422,193,1796]" captionTargetPageId="20" captionText="FIGURES 90 99. Opeatocerata hadrophallus sp. nov., holotype ♂. 90. habitus, lateral view; 91. wing; 92. abdomen from tergite 6 to terminalia, lateral view; 93. tergites 4 8, dorsal view; 94. terminalia, lateral view; 95. lobes of cerci, dorsal view; 96. descendant plate of anterior cercus and hypoproct, posterior view; 97. subepandrial sclerite and bacilliform sclerite, dorsal view; 98. ejaculatory apodeme, phallus and hypandrium, lateral view; 99. hypandrium, ventral view." httpUri="https://zenodo.org/record/252531/files/figure.png" pageId="19" pageNumber="521">Fig. 98</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34576A2E066FF8EC797FC60AB7C1670" pageId="19" pageNumber="521" type="description">
<paragraph id="8BE02529E066FF8EC797FC60AB861594" blockId="19.[151,1437,798,2011]" pageId="19" pageNumber="521">
<emphasis id="B92BF93BE066FF8EC797FC60A97013D4" bold="true" box="[199,337,906,931]" pageId="19" pageNumber="521">Description</emphasis>
.
<emphasis id="B92BF93BE066FF8EC630FC60A9BC13D4" bold="true" box="[352,413,906,931]" pageId="19" pageNumber="521">Male</emphasis>
(
<figureCitation id="136439ACE066FF8EC6FDFC60AA2313D4" box="[429,514,906,931]" captionStart="FIGURES 90 99" captionStartId="20.[151,264,1816,1839]" captionTargetBox="[164,1411,193,1785]" captionTargetId="figure@20.[164,1422,193,1796]" captionTargetPageId="20" captionText="FIGURES 90 99. Opeatocerata hadrophallus sp. nov., holotype ♂. 90. habitus, lateral view; 91. wing; 92. abdomen from tergite 6 to terminalia, lateral view; 93. tergites 4 8, dorsal view; 94. terminalia, lateral view; 95. lobes of cerci, dorsal view; 96. descendant plate of anterior cercus and hypoproct, posterior view; 97. subepandrial sclerite and bacilliform sclerite, dorsal view; 98. ejaculatory apodeme, phallus and hypandrium, lateral view; 99. hypandrium, ventral view." httpUri="https://zenodo.org/record/252531/files/figure.png" pageId="19" pageNumber="521">Fig. 90</figureCitation>
).
<emphasis id="B92BF93BE066FF8EC546FC60AA7713D4" bold="true" box="[534,598,906,931]" pageId="19" pageNumber="521">Head</emphasis>
: Face parallel-sided, dark brown with gray pruinescence visible in ventral view, about
<date id="FFE103E9E066FF8EC670FC45A96313B0" box="[288,322,943,967]" pageId="19" pageNumber="521">4X</date>
longer than lower width. Ocellar tubercle protuberant, dark brown with brown puinescence; ocelli brown. Antenna yellow with black bristles; postpedicel about 1.5X longer than pedicel; stylus about
<date id="FFE103E9E066FF8EC3A5FC39AD39139C" box="[1269,1304,979,1003]" pageId="19" pageNumber="521">2X</date>
longer than postpedicel. Proboscis yellow shorter than head height.
<emphasis id="B92BF93BE066FF8EC449FC1CAB531478" bold="true" box="[793,882,1014,1039]" pageId="19" pageNumber="521">Thorax</emphasis>
yellow, shiny (
<figureCitation id="136439ACE066FF8EC375FC1CAC5D1478" box="[1061,1148,1014,1039]" captionStart="FIGURES 90 99" captionStartId="20.[151,264,1816,1839]" captionTargetBox="[164,1411,193,1785]" captionTargetId="figure@20.[164,1422,193,1796]" captionTargetPageId="20" captionText="FIGURES 90 99. Opeatocerata hadrophallus sp. nov., holotype ♂. 90. habitus, lateral view; 91. wing; 92. abdomen from tergite 6 to terminalia, lateral view; 93. tergites 4 8, dorsal view; 94. terminalia, lateral view; 95. lobes of cerci, dorsal view; 96. descendant plate of anterior cercus and hypoproct, posterior view; 97. subepandrial sclerite and bacilliform sclerite, dorsal view; 98. ejaculatory apodeme, phallus and hypandrium, lateral view; 99. hypandrium, ventral view." httpUri="https://zenodo.org/record/252531/files/figure.png" pageId="19" pageNumber="521">Fig. 90</figureCitation>
). Legs yellow, except by hind femur and tibia with black apical ring and all tarsomeres 45 black; hind tibia and tarsus with anterodorsal and posterodorsal rows of longer bristles.
<emphasis id="B92BF93BE066FF8EC511FBD4AAA31420" bold="true" box="[577,642,1086,1111]" pageId="19" pageNumber="521">Wing</emphasis>
(
<figureCitation id="136439ACE066FF8EC5C2FBD4AAC51420" box="[658,740,1086,1111]" captionStart="FIGURES 90 99" captionStartId="20.[151,264,1816,1839]" captionTargetBox="[164,1411,193,1785]" captionTargetId="figure@20.[164,1422,193,1796]" captionTargetPageId="20" captionText="FIGURES 90 99. Opeatocerata hadrophallus sp. nov., holotype ♂. 90. habitus, lateral view; 91. wing; 92. abdomen from tergite 6 to terminalia, lateral view; 93. tergites 4 8, dorsal view; 94. terminalia, lateral view; 95. lobes of cerci, dorsal view; 96. descendant plate of anterior cercus and hypoproct, posterior view; 97. subepandrial sclerite and bacilliform sclerite, dorsal view; 98. ejaculatory apodeme, phallus and hypandrium, lateral view; 99. hypandrium, ventral view." httpUri="https://zenodo.org/record/252531/files/figure.png" pageId="19" pageNumber="521">Fig. 91</figureCitation>
) hyaline, pterostigma brown, conspicuous, about 2.2X longer than wide. Halter yellow.
<emphasis id="B92BF93BE066FF8EC692FB88AA16140C" bold="true" box="[450,567,1122,1147]" pageId="19" pageNumber="521">Abdomen</emphasis>
yellow, shiny; tergites 36 with paramedian subtriangular black spots (
<figureCitation id="136439ACE066FF8EC23EFB88A89714E8" captionStart="FIGURES 90 99" captionStartId="20.[151,264,1816,1839]" captionTargetBox="[164,1411,193,1785]" captionTargetId="figure@20.[164,1422,193,1796]" captionTargetPageId="20" captionText="FIGURES 90 99. Opeatocerata hadrophallus sp. nov., holotype ♂. 90. habitus, lateral view; 91. wing; 92. abdomen from tergite 6 to terminalia, lateral view; 93. tergites 4 8, dorsal view; 94. terminalia, lateral view; 95. lobes of cerci, dorsal view; 96. descendant plate of anterior cercus and hypoproct, posterior view; 97. subepandrial sclerite and bacilliform sclerite, dorsal view; 98. ejaculatory apodeme, phallus and hypandrium, lateral view; 99. hypandrium, ventral view." httpUri="https://zenodo.org/record/252531/files/figure.png" pageId="19" pageNumber="521">Fig. 93</figureCitation>
); tergite 8 divided in two subrectangular plates; sternite 8 divided in two trapezoidal plates.
<emphasis id="B92BF93BE066FF8EC25FFB6CADBD14E8" bold="true" box="[1295,1436,1158,1183]" pageId="19" pageNumber="521">Terminalia:</emphasis>
Anterior cercus with a proclinate dorsal projection in lateral view (
<figureCitation id="136439ACE066FF8EC4D2FB40AC2214B4" box="[898,1027,1194,1219]" captionStart="FIGURES 90 99" captionStartId="20.[151,264,1816,1839]" captionTargetBox="[164,1411,193,1785]" captionTargetId="figure@20.[164,1422,193,1796]" captionTargetPageId="20" captionText="FIGURES 90 99. Opeatocerata hadrophallus sp. nov., holotype ♂. 90. habitus, lateral view; 91. wing; 92. abdomen from tergite 6 to terminalia, lateral view; 93. tergites 4 8, dorsal view; 94. terminalia, lateral view; 95. lobes of cerci, dorsal view; 96. descendant plate of anterior cercus and hypoproct, posterior view; 97. subepandrial sclerite and bacilliform sclerite, dorsal view; 98. ejaculatory apodeme, phallus and hypandrium, lateral view; 99. hypandrium, ventral view." httpUri="https://zenodo.org/record/252531/files/figure.png" pageId="19" pageNumber="521">Figs 92, 94</figureCitation>
), posterior margin rounded in dorsal view (
<figureCitation id="136439ACE066FF8EC7B1FB24A91B1490" box="[225,314,1230,1255]" captionStart="FIGURES 90 99" captionStartId="20.[151,264,1816,1839]" captionTargetBox="[164,1411,193,1785]" captionTargetId="figure@20.[164,1422,193,1796]" captionTargetPageId="20" captionText="FIGURES 90 99. Opeatocerata hadrophallus sp. nov., holotype ♂. 90. habitus, lateral view; 91. wing; 92. abdomen from tergite 6 to terminalia, lateral view; 93. tergites 4 8, dorsal view; 94. terminalia, lateral view; 95. lobes of cerci, dorsal view; 96. descendant plate of anterior cercus and hypoproct, posterior view; 97. subepandrial sclerite and bacilliform sclerite, dorsal view; 98. ejaculatory apodeme, phallus and hypandrium, lateral view; 99. hypandrium, ventral view." httpUri="https://zenodo.org/record/252531/files/figure.png" pageId="19" pageNumber="521">Fig. 95</figureCitation>
); descendant plate subtrapezoidal with a small median constriction in posterior view (
<figureCitation id="136439ACE066FF8EC267FB24ADB11490" box="[1335,1424,1230,1255]" captionStart="FIGURES 90 99" captionStartId="20.[151,264,1816,1839]" captionTargetBox="[164,1411,193,1785]" captionTargetId="figure@20.[164,1422,193,1796]" captionTargetPageId="20" captionText="FIGURES 90 99. Opeatocerata hadrophallus sp. nov., holotype ♂. 90. habitus, lateral view; 91. wing; 92. abdomen from tergite 6 to terminalia, lateral view; 93. tergites 4 8, dorsal view; 94. terminalia, lateral view; 95. lobes of cerci, dorsal view; 96. descendant plate of anterior cercus and hypoproct, posterior view; 97. subepandrial sclerite and bacilliform sclerite, dorsal view; 98. ejaculatory apodeme, phallus and hypandrium, lateral view; 99. hypandrium, ventral view." httpUri="https://zenodo.org/record/252531/files/figure.png" pageId="19" pageNumber="521">Fig. 96</figureCitation>
). Posterior cercus rounded at apex in lateral view (
<figureCitation id="136439ACE066FF8EC594FB18AB68157C" box="[708,841,1266,1291]" captionStart="FIGURES 90 99" captionStartId="20.[151,264,1816,1839]" captionTargetBox="[164,1411,193,1785]" captionTargetId="figure@20.[164,1422,193,1796]" captionTargetPageId="20" captionText="FIGURES 90 99. Opeatocerata hadrophallus sp. nov., holotype ♂. 90. habitus, lateral view; 91. wing; 92. abdomen from tergite 6 to terminalia, lateral view; 93. tergites 4 8, dorsal view; 94. terminalia, lateral view; 95. lobes of cerci, dorsal view; 96. descendant plate of anterior cercus and hypoproct, posterior view; 97. subepandrial sclerite and bacilliform sclerite, dorsal view; 98. ejaculatory apodeme, phallus and hypandrium, lateral view; 99. hypandrium, ventral view." httpUri="https://zenodo.org/record/252531/files/figure.png" pageId="19" pageNumber="521">Figs 92, 94</figureCitation>
), with dorsal, subbasal, pointed projection (
<figureCitation id="136439ACE066FF8EC26AFB18ADAE157C" box="[1338,1423,1266,1291]" captionStart="FIGURES 90 99" captionStartId="20.[151,264,1816,1839]" captionTargetBox="[164,1411,193,1785]" captionTargetId="figure@20.[164,1422,193,1796]" captionTargetPageId="20" captionText="FIGURES 90 99. Opeatocerata hadrophallus sp. nov., holotype ♂. 90. habitus, lateral view; 91. wing; 92. abdomen from tergite 6 to terminalia, lateral view; 93. tergites 4 8, dorsal view; 94. terminalia, lateral view; 95. lobes of cerci, dorsal view; 96. descendant plate of anterior cercus and hypoproct, posterior view; 97. subepandrial sclerite and bacilliform sclerite, dorsal view; 98. ejaculatory apodeme, phallus and hypandrium, lateral view; 99. hypandrium, ventral view." httpUri="https://zenodo.org/record/252531/files/figure.png" pageId="19" pageNumber="521">Fig. 95</figureCitation>
). Hypoproct subtriangular in lateral view (
<figureCitation id="136439ACE066FF8EC532FAFCAA921558" box="[610,691,1302,1327]" captionStart="FIGURES 90 99" captionStartId="20.[151,264,1816,1839]" captionTargetBox="[164,1411,193,1785]" captionTargetId="figure@20.[164,1422,193,1796]" captionTargetPageId="20" captionText="FIGURES 90 99. Opeatocerata hadrophallus sp. nov., holotype ♂. 90. habitus, lateral view; 91. wing; 92. abdomen from tergite 6 to terminalia, lateral view; 93. tergites 4 8, dorsal view; 94. terminalia, lateral view; 95. lobes of cerci, dorsal view; 96. descendant plate of anterior cercus and hypoproct, posterior view; 97. subepandrial sclerite and bacilliform sclerite, dorsal view; 98. ejaculatory apodeme, phallus and hypandrium, lateral view; 99. hypandrium, ventral view." httpUri="https://zenodo.org/record/252531/files/figure.png" pageId="19" pageNumber="521">Fig. 94</figureCitation>
) and comma-shaped in posterior view (
<figureCitation id="136439ACE066FF8EC322FAFCACE61558" box="[1138,1223,1302,1327]" captionStart="FIGURES 90 99" captionStartId="20.[151,264,1816,1839]" captionTargetBox="[164,1411,193,1785]" captionTargetId="figure@20.[164,1422,193,1796]" captionTargetPageId="20" captionText="FIGURES 90 99. Opeatocerata hadrophallus sp. nov., holotype ♂. 90. habitus, lateral view; 91. wing; 92. abdomen from tergite 6 to terminalia, lateral view; 93. tergites 4 8, dorsal view; 94. terminalia, lateral view; 95. lobes of cerci, dorsal view; 96. descendant plate of anterior cercus and hypoproct, posterior view; 97. subepandrial sclerite and bacilliform sclerite, dorsal view; 98. ejaculatory apodeme, phallus and hypandrium, lateral view; 99. hypandrium, ventral view." httpUri="https://zenodo.org/record/252531/files/figure.png" pageId="19" pageNumber="521">Fig. 96</figureCitation>
). Epandrial ventral lobe rounded at apex (
<figureCitation id="136439ACE066FF8EC6DFFAD0AA331524" box="[399,530,1338,1363]" captionStart="FIGURES 90 99" captionStartId="20.[151,264,1816,1839]" captionTargetBox="[164,1411,193,1785]" captionTargetId="figure@20.[164,1422,193,1796]" captionTargetPageId="20" captionText="FIGURES 90 99. Opeatocerata hadrophallus sp. nov., holotype ♂. 90. habitus, lateral view; 91. wing; 92. abdomen from tergite 6 to terminalia, lateral view; 93. tergites 4 8, dorsal view; 94. terminalia, lateral view; 95. lobes of cerci, dorsal view; 96. descendant plate of anterior cercus and hypoproct, posterior view; 97. subepandrial sclerite and bacilliform sclerite, dorsal view; 98. ejaculatory apodeme, phallus and hypandrium, lateral view; 99. hypandrium, ventral view." httpUri="https://zenodo.org/record/252531/files/figure.png" pageId="19" pageNumber="521">Figs 94, 94</figureCitation>
); spiniform bristles mesially and longer bristles apically. Hypandrium wider than long with rounded apex and small median sinus (
<figureCitation id="136439ACE066FF8EC5ECFAB4AB2E1500" box="[700,783,1374,1399]" captionStart="FIGURES 90 99" captionStartId="20.[151,264,1816,1839]" captionTargetBox="[164,1411,193,1785]" captionTargetId="figure@20.[164,1422,193,1796]" captionTargetPageId="20" captionText="FIGURES 90 99. Opeatocerata hadrophallus sp. nov., holotype ♂. 90. habitus, lateral view; 91. wing; 92. abdomen from tergite 6 to terminalia, lateral view; 93. tergites 4 8, dorsal view; 94. terminalia, lateral view; 95. lobes of cerci, dorsal view; 96. descendant plate of anterior cercus and hypoproct, posterior view; 97. subepandrial sclerite and bacilliform sclerite, dorsal view; 98. ejaculatory apodeme, phallus and hypandrium, lateral view; 99. hypandrium, ventral view." httpUri="https://zenodo.org/record/252531/files/figure.png" pageId="19" pageNumber="521">Fig. 99</figureCitation>
), without long bristles. Phallus with narrow base, wider in mostly, pointed apically; short, not extending beyond apex of the hypandrium (
<figureCitation id="136439ACE066FF8EC355FA68AC7915EC" box="[1029,1112,1410,1435]" captionStart="FIGURES 90 99" captionStartId="20.[151,264,1816,1839]" captionTargetBox="[164,1411,193,1785]" captionTargetId="figure@20.[164,1422,193,1796]" captionTargetPageId="20" captionText="FIGURES 90 99. Opeatocerata hadrophallus sp. nov., holotype ♂. 90. habitus, lateral view; 91. wing; 92. abdomen from tergite 6 to terminalia, lateral view; 93. tergites 4 8, dorsal view; 94. terminalia, lateral view; 95. lobes of cerci, dorsal view; 96. descendant plate of anterior cercus and hypoproct, posterior view; 97. subepandrial sclerite and bacilliform sclerite, dorsal view; 98. ejaculatory apodeme, phallus and hypandrium, lateral view; 99. hypandrium, ventral view." httpUri="https://zenodo.org/record/252531/files/figure.png" pageId="19" pageNumber="521">Fig. 98</figureCitation>
). Subepandrial sclerite wider than long, U-shaped (
<figureCitation id="136439ACE066FF8EC6C0FA4CA9C715C8" box="[400,486,1446,1471]" captionStart="FIGURES 90 99" captionStartId="20.[151,264,1816,1839]" captionTargetBox="[164,1411,193,1785]" captionTargetId="figure@20.[164,1422,193,1796]" captionTargetPageId="20" captionText="FIGURES 90 99. Opeatocerata hadrophallus sp. nov., holotype ♂. 90. habitus, lateral view; 91. wing; 92. abdomen from tergite 6 to terminalia, lateral view; 93. tergites 4 8, dorsal view; 94. terminalia, lateral view; 95. lobes of cerci, dorsal view; 96. descendant plate of anterior cercus and hypoproct, posterior view; 97. subepandrial sclerite and bacilliform sclerite, dorsal view; 98. ejaculatory apodeme, phallus and hypandrium, lateral view; 99. hypandrium, ventral view." httpUri="https://zenodo.org/record/252531/files/figure.png" pageId="19" pageNumber="521">Fig. 97</figureCitation>
). Bacilliform sclerite without folds and longer than subepandrial sclerite (
<figureCitation id="136439ACE066FF8EC269FA4CADB115C8" box="[1337,1424,1446,1471]" captionStart="FIGURES 90 99" captionStartId="20.[151,264,1816,1839]" captionTargetBox="[164,1411,193,1785]" captionTargetId="figure@20.[164,1422,193,1796]" captionTargetPageId="20" captionText="FIGURES 90 99. Opeatocerata hadrophallus sp. nov., holotype ♂. 90. habitus, lateral view; 91. wing; 92. abdomen from tergite 6 to terminalia, lateral view; 93. tergites 4 8, dorsal view; 94. terminalia, lateral view; 95. lobes of cerci, dorsal view; 96. descendant plate of anterior cercus and hypoproct, posterior view; 97. subepandrial sclerite and bacilliform sclerite, dorsal view; 98. ejaculatory apodeme, phallus and hypandrium, lateral view; 99. hypandrium, ventral view." httpUri="https://zenodo.org/record/252531/files/figure.png" pageId="19" pageNumber="521">Fig. 97</figureCitation>
).
<emphasis id="B92BF93BE066FF8EC7C7FA20A97A1594" bold="true" box="[151,347,1482,1507]" pageId="19" pageNumber="521">Specimen length</emphasis>
:
<quantity id="4CA788CCE066FF8EC63AFA20A9E21595" box="[362,451,1482,1507]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="19" pageNumber="521" unit="mm" value="2.6">2.6 mm</quantity>
;
<emphasis id="B92BF93BE066FF8EC69FFA20AA781594" bold="true" box="[463,601,1482,1507]" pageId="19" pageNumber="521">wing length</emphasis>
:
<quantity id="4CA788CCE066FF8EC539FA20AAE31595" box="[617,706,1482,1507]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="19" pageNumber="521" unit="mm" value="2.6">2.6 mm</quantity>
.
<emphasis id="B92BF93BE066FF8EC59CFA20AB021594" bold="true" box="[716,803,1482,1507]" pageId="19" pageNumber="521">Female</emphasis>
. Unknown.
</paragraph>
<paragraph id="8BE02529E066FF8EC797FA04AB7C1670" blockId="19.[151,1437,798,2011]" box="[199,861,1518,1543]" pageId="19" pageNumber="521">
<emphasis id="B92BF93BE066FF8EC797FA04A9DC1670" bold="true" box="[199,509,1518,1543]" pageId="19" pageNumber="521">Geographical distribution</emphasis>
.
<collectingCountry id="F34865B9E066FF8EC55BFA04AA6E1670" box="[523,591,1518,1543]" name="Brazil" pageId="19" pageNumber="521">Brazil</collectingCountry>
(Amazonas) (
<figureCitation id="136439ACE066FF8EC5BFFA04AB711670" box="[751,848,1518,1543]" captionStart="FIGURE 211" captionStartId="42.[151,250,1955,1977]" captionTargetBox="[235,1368,193,1934]" captionTargetId="figure@42.[216,1371,193,1934]" captionTargetPageId="42" captionText="FIGURE 211. Geographic distribution of species of Opeatocerata." httpUri="https://zenodo.org/record/252546/files/figure.png" pageId="19" pageNumber="521">Fig. 211</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34576A2E066FF8EC797F9F8A8DA16A8" pageId="19" pageNumber="521" type="materials_examined">
<paragraph id="8BE02529E066FF8EC797F9F8A8C716E0" blockId="19.[151,1437,798,2011]" pageId="19" pageNumber="521">
<emphasis id="B92BF93BE066FF8EC797F9F8A956165C" bold="true" box="[199,375,1554,1579]" pageId="19" pageNumber="521">
<typeStatus id="54E49B8BE066FF8EC797F9F8A923165C" box="[199,258,1554,1579]" pageId="19" pageNumber="521">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="54E49B8BE066FF8EC6DEF9F8AA0C165C" box="[398,557,1554,1579]" pageId="19" pageNumber="521" type="holotype">
<emphasis id="B92BF93BE066FF8EC6DEF9F8AA0C165C" bold="true" box="[398,557,1554,1579]" pageId="19" pageNumber="521">HOLOTYPE</emphasis>
</typeStatus>
♂ labelled:
<collectingCountry id="F34865B9E066FF8EC583F9F8AB38165C" box="[723,793,1554,1579]" name="Brazil" pageId="19" pageNumber="521">Brasil</collectingCountry>
, AM[azonas], Novo Aripuanã, Reserva / Sokagakai,
<date id="FFE103E9E066FF8EC7C7F9DCA9311638" box="[151,272,1590,1615]" pageId="19" pageNumber="521" value="2000-04-23">23.iv.2000</date>
. J. Vidal, Arm.[adilha] Malaise (
<collectionCode id="ED4EBDECE066FF8EC5F3F9DDAAC81638" box="[675,745,1591,1615]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="19" pageNumber="521">INPA</collectionCode>
).
<typeStatus id="54E49B8BE066FF8EC452F9DCAB5B1638" box="[770,890,1590,1615]" pageId="19" pageNumber="521" type="paratype">
<emphasis id="B92BF93BE066FF8EC452F9DCAB5B1638" bold="true" box="[770,890,1590,1615]" pageId="19" pageNumber="521">Paratypes</emphasis>
</typeStatus>
:
<collectingCountry id="F34865B9E066FF8EC4C0F9DCABF81638" box="[912,985,1590,1615]" name="Brazil" pageId="19" pageNumber="521">
<emphasis id="B92BF93BE066FF8EC4C0F9DCABF81638" bold="true" box="[912,985,1590,1615]" pageId="19" pageNumber="521">Brazil</emphasis>
</collectingCountry>
,
<emphasis id="B92BF93BE066FF8EC4BEF9DCAC4B1638" bold="true" box="[1006,1130,1590,1615]" pageId="19" pageNumber="521">Amazonas</emphasis>
. Parnajaú, Rio Papagaio.
<date id="FFE103E9E066FF8EC7C7F9B0A8CE1604" box="[151,239,1626,1651]" pageId="19" pageNumber="521" value="2001-06">vi.2001</date>
.
<collectingCountry id="F34865B9E066FF8EC7AEF9B1A9191605" box="[254,312,1627,1650]" name="Malta" pageId="19" pageNumber="521">Mata</collectingCountry>
terra-firme.
<geoCoordinate id="EE6B43EEE066FF8EC680F9B1AA691604" box="[464,584,1627,1651]" direction="south" orientation="latitude" pageId="19" pageNumber="521" precision="15" value="-2.0008333">2º0003S</geoCoordinate>
<geoCoordinate id="EE6B43EEE066FF8EC504F9B0AAC41604" box="[596,741,1626,1651]" direction="west" orientation="longitude" pageId="19" pageNumber="521" precision="15" value="-62.731667">62º4354W</geoCoordinate>
/ Arm[adilha] malaise. Henriques, Vidal e Silva leg (
<specimenCount id="9D59EEA0E066FF8EC238F9B1ADB61605" box="[1384,1431,1627,1651]" pageId="19" pageNumber="521" type="generic">1 ♂</specimenCount>
,
<collectionCode id="ED4EBDECE066FF8EC7C7F995A8FA16E0" box="[151,219,1663,1687]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="19" pageNumber="521">INPA</collectionCode>
).
</paragraph>
<paragraph id="8BE02529E066FF8EC797F948A8DA16A8" blockId="19.[151,1437,798,2011]" pageId="19" pageNumber="521">
<emphasis id="B92BF93BE066FF8EC797F948A99016CC" bold="true" box="[199,433,1698,1723]" pageId="19" pageNumber="521">
<typeStatus id="54E49B8BE066FF8EC797F948A91516CC" box="[199,308,1698,1723]" pageId="19" pageNumber="521" type="holotype">Holotype</typeStatus>
condition
</emphasis>
. Wing mounted in microslide; left mid leg, hind legs; abdomen in microtube with glycerin.
</paragraph>
</subSubSection>
<subSubSection id="C34576A2E066FF8EC797F900AD2F1774" box="[199,1294,1770,1795]" pageId="19" pageNumber="521" type="etymology">
<paragraph id="8BE02529E066FF8EC797F900AD2F1774" blockId="19.[151,1437,798,2011]" box="[199,1294,1770,1795]" pageId="19" pageNumber="521">
<emphasis id="B92BF93BE066FF8EC797F900A9671774" bold="true" box="[199,326,1770,1795]" pageId="19" pageNumber="521">Etymology</emphasis>
. From the Greek
<emphasis id="B92BF93BE066FF8EC55CF900AA791774" box="[524,600,1770,1795]" italics="true" pageId="19" pageNumber="521">hadros</emphasis>
(robust) and
<emphasis id="B92BF93BE066FF8EC5BFF900AB631774" box="[751,834,1770,1795]" italics="true" pageId="19" pageNumber="521">phallos</emphasis>
(phallus), referring to the robust phallus.
</paragraph>
</subSubSection>
<subSubSection id="C34576A2E066FF8EC797F8E4AD3617AC" pageId="19" pageNumber="521" type="discussion">
<paragraph id="8BE02529E066FF8EC797F8E4AD3617AC" blockId="19.[151,1437,798,2011]" pageId="19" pageNumber="521">
<emphasis id="B92BF93BE066FF8EC797F8E4A9151750" bold="true" box="[199,308,1806,1831]" pageId="19" pageNumber="521">Remarks</emphasis>
.
<taxonomicName id="4C5F5EAAE066FF8EC615F8E5AA5F1750" box="[325,638,1806,1831]" class="Insecta" family="Empididae" genus="Opeatocerata" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="521" phylum="Arthropoda" rank="species" species="hadrophallus" status="sp. nov.">
<emphasis id="B92BF93BE066FF8EC615F8E5AA5F1750" box="[325,638,1806,1831]" italics="true" pageId="19" pageNumber="521">Opeatocerata hadrophallus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2184440E066FF8EC5DAF8E4AAC31750" box="[650,738,1806,1831]" pageId="19" pageNumber="521" rank="species">
<emphasis id="B92BF93BE066FF8EC5DAF8E4AAC31750" bold="true" box="[650,738,1806,1831]" pageId="19" pageNumber="521">sp. nov</emphasis>
</taxonomicNameLabel>
is similar to
<taxonomicName id="4C5F5EAAE066FF8EC4D4F8E5ABD81750" box="[900,1017,1806,1831]" class="Insecta" family="Empididae" genus="Opeatocerata" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="521" phylum="Arthropoda" rank="species" species="stubbsi">
<emphasis id="B92BF93BE066FF8EC4D4F8E5ABD81750" box="[900,1017,1806,1831]" italics="true" pageId="19" pageNumber="521">O. stubbsi</emphasis>
</taxonomicName>
by the short phallus, not extending beyond the apex of the hypandirum (
<figureCitation id="136439ACE066FF8EC56BF8D8AAB2173C" box="[571,659,1842,1867]" captionStart="FIGURES 90 99" captionStartId="20.[151,264,1816,1839]" captionTargetBox="[164,1411,193,1785]" captionTargetId="figure@20.[164,1422,193,1796]" captionTargetPageId="20" captionText="FIGURES 90 99. Opeatocerata hadrophallus sp. nov., holotype ♂. 90. habitus, lateral view; 91. wing; 92. abdomen from tergite 6 to terminalia, lateral view; 93. tergites 4 8, dorsal view; 94. terminalia, lateral view; 95. lobes of cerci, dorsal view; 96. descendant plate of anterior cercus and hypoproct, posterior view; 97. subepandrial sclerite and bacilliform sclerite, dorsal view; 98. ejaculatory apodeme, phallus and hypandrium, lateral view; 99. hypandrium, ventral view." httpUri="https://zenodo.org/record/252531/files/figure.png" pageId="19" pageNumber="521">Figs 98</figureCitation>
,
<figureCitation id="136439ACE066FF8EC5F0F8D9AAEA173C" box="[672,715,1843,1867]" captionStart="FIGURES 170 178" captionStartId="36.[151,264,1905,1928]" captionTargetBox="[164,1404,337,1860]" captionTargetId="figure@36.[164,1404,337,1869]" captionTargetPageId="36" captionText="FIGURES 170 178. Opeatocerata stubbsi Smith, 1989, holotype ♂. 170. habitus, lateral view; 171. wing; 172. tergite 8 and terminalia, lateral view; 173. tergites 4 - 8, dorsal view; 174. terminalia; 175. lobes of cerci, dorsal view; 176. descendant plate of anterior cercus, posterior view; 177. subepandrial sclerite and bacilliform sclerite, dorsal view; 178. ejaculatory apodeme, phallus and hypandrium, lateral view." httpUri="https://zenodo.org/record/252541/files/figure.png" pageId="19" pageNumber="521">178</figureCitation>
) and posterior cercus with a dorsal pointed projection in dorsal view (
<figureCitation id="136439ACE066FF8EC78DF8BCA9151718" box="[221,308,1878,1903]" captionStart="FIGURES 90 99" captionStartId="20.[151,264,1816,1839]" captionTargetBox="[164,1411,193,1785]" captionTargetId="figure@20.[164,1422,193,1796]" captionTargetPageId="20" captionText="FIGURES 90 99. Opeatocerata hadrophallus sp. nov., holotype ♂. 90. habitus, lateral view; 91. wing; 92. abdomen from tergite 6 to terminalia, lateral view; 93. tergites 4 8, dorsal view; 94. terminalia, lateral view; 95. lobes of cerci, dorsal view; 96. descendant plate of anterior cercus and hypoproct, posterior view; 97. subepandrial sclerite and bacilliform sclerite, dorsal view; 98. ejaculatory apodeme, phallus and hypandrium, lateral view; 99. hypandrium, ventral view." httpUri="https://zenodo.org/record/252531/files/figure.png" pageId="19" pageNumber="521">Figs 95</figureCitation>
,
<figureCitation id="136439ACE066FF8EC611F8BCA94E1718" box="[321,367,1878,1903]" captionStart="FIGURES 170 178" captionStartId="36.[151,264,1905,1928]" captionTargetBox="[164,1404,337,1860]" captionTargetId="figure@36.[164,1404,337,1869]" captionTargetPageId="36" captionText="FIGURES 170 178. Opeatocerata stubbsi Smith, 1989, holotype ♂. 170. habitus, lateral view; 171. wing; 172. tergite 8 and terminalia, lateral view; 173. tergites 4 - 8, dorsal view; 174. terminalia; 175. lobes of cerci, dorsal view; 176. descendant plate of anterior cercus, posterior view; 177. subepandrial sclerite and bacilliform sclerite, dorsal view; 178. ejaculatory apodeme, phallus and hypandrium, lateral view." httpUri="https://zenodo.org/record/252541/files/figure.png" pageId="19" pageNumber="521">175</figureCitation>
). Differs from the latter by tergites with paramedian triangular spots (
<figureCitation id="136439ACE066FF8EC3D0F8BCACF41718" box="[1152,1237,1878,1903]" captionStart="FIGURES 90 99" captionStartId="20.[151,264,1816,1839]" captionTargetBox="[164,1411,193,1785]" captionTargetId="figure@20.[164,1422,193,1796]" captionTargetPageId="20" captionText="FIGURES 90 99. Opeatocerata hadrophallus sp. nov., holotype ♂. 90. habitus, lateral view; 91. wing; 92. abdomen from tergite 6 to terminalia, lateral view; 93. tergites 4 8, dorsal view; 94. terminalia, lateral view; 95. lobes of cerci, dorsal view; 96. descendant plate of anterior cercus and hypoproct, posterior view; 97. subepandrial sclerite and bacilliform sclerite, dorsal view; 98. ejaculatory apodeme, phallus and hypandrium, lateral view; 99. hypandrium, ventral view." httpUri="https://zenodo.org/record/252531/files/figure.png" pageId="19" pageNumber="521">Fig. 93</figureCitation>
), posterior cercus without dorsal subapical projection (
<figureCitation id="136439ACE066FF8EC567F890AAAB17E4" box="[567,650,1914,1939]" captionStart="FIGURES 90 99" captionStartId="20.[151,264,1816,1839]" captionTargetBox="[164,1411,193,1785]" captionTargetId="figure@20.[164,1422,193,1796]" captionTargetPageId="20" captionText="FIGURES 90 99. Opeatocerata hadrophallus sp. nov., holotype ♂. 90. habitus, lateral view; 91. wing; 92. abdomen from tergite 6 to terminalia, lateral view; 93. tergites 4 8, dorsal view; 94. terminalia, lateral view; 95. lobes of cerci, dorsal view; 96. descendant plate of anterior cercus and hypoproct, posterior view; 97. subepandrial sclerite and bacilliform sclerite, dorsal view; 98. ejaculatory apodeme, phallus and hypandrium, lateral view; 99. hypandrium, ventral view." httpUri="https://zenodo.org/record/252531/files/figure.png" pageId="19" pageNumber="521">Fig. 95</figureCitation>
) and hypoproct subtriangular in lateral view (
<figureCitation id="136439ACE066FF8EC3C7F890ACCC17E4" box="[1175,1261,1914,1939]" captionStart="FIGURES 90 99" captionStartId="20.[151,264,1816,1839]" captionTargetBox="[164,1411,193,1785]" captionTargetId="figure@20.[164,1422,193,1796]" captionTargetPageId="20" captionText="FIGURES 90 99. Opeatocerata hadrophallus sp. nov., holotype ♂. 90. habitus, lateral view; 91. wing; 92. abdomen from tergite 6 to terminalia, lateral view; 93. tergites 4 8, dorsal view; 94. terminalia, lateral view; 95. lobes of cerci, dorsal view; 96. descendant plate of anterior cercus and hypoproct, posterior view; 97. subepandrial sclerite and bacilliform sclerite, dorsal view; 98. ejaculatory apodeme, phallus and hypandrium, lateral view; 99. hypandrium, ventral view." httpUri="https://zenodo.org/record/252531/files/figure.png" pageId="19" pageNumber="521">Fig. 94</figureCitation>
).
<taxonomicName id="4C5F5EAAE066FF8EC252F891A8C717C0" class="Insecta" family="Empididae" genus="Opeatocerata" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="521" phylum="Arthropoda" rank="species" species="stubbsi">
<emphasis id="B92BF93BE066FF8EC252F891A8C717C0" italics="true" pageId="19" pageNumber="521">Opeatocerata stubbsi</emphasis>
</taxonomicName>
has tergites with paramedian subrectangular spots (
<figureCitation id="136439ACE066FF8EC408F874ABE117C0" box="[856,960,1950,1975]" captionStart="FIGURES 170 178" captionStartId="36.[151,264,1905,1928]" captionTargetBox="[164,1404,337,1860]" captionTargetId="figure@36.[164,1404,337,1869]" captionTargetPageId="36" captionText="FIGURES 170 178. Opeatocerata stubbsi Smith, 1989, holotype ♂. 170. habitus, lateral view; 171. wing; 172. tergite 8 and terminalia, lateral view; 173. tergites 4 - 8, dorsal view; 174. terminalia; 175. lobes of cerci, dorsal view; 176. descendant plate of anterior cercus, posterior view; 177. subepandrial sclerite and bacilliform sclerite, dorsal view; 178. ejaculatory apodeme, phallus and hypandrium, lateral view." httpUri="https://zenodo.org/record/252541/files/figure.png" pageId="19" pageNumber="521">Fig. 173</figureCitation>
), posterior cercus with dorsal subapical subtriangular projection in dorsal view (
<figureCitation id="136439ACE066FF8EC508F828AA9917AC" box="[600,696,1986,2011]" captionStart="FIGURES 170 178" captionStartId="36.[151,264,1905,1928]" captionTargetBox="[164,1404,337,1860]" captionTargetId="figure@36.[164,1404,337,1869]" captionTargetPageId="36" captionText="FIGURES 170 178. Opeatocerata stubbsi Smith, 1989, holotype ♂. 170. habitus, lateral view; 171. wing; 172. tergite 8 and terminalia, lateral view; 173. tergites 4 - 8, dorsal view; 174. terminalia; 175. lobes of cerci, dorsal view; 176. descendant plate of anterior cercus, posterior view; 177. subepandrial sclerite and bacilliform sclerite, dorsal view; 178. ejaculatory apodeme, phallus and hypandrium, lateral view." httpUri="https://zenodo.org/record/252541/files/figure.png" pageId="19" pageNumber="521">Fig. 175</figureCitation>
) and hypoproct subrounded in lateral view (
<figureCitation id="136439ACE066FF8EC3F8F828AD2B17AC" box="[1192,1290,1986,2011]" captionStart="FIGURES 170 178" captionStartId="36.[151,264,1905,1928]" captionTargetBox="[164,1404,337,1860]" captionTargetId="figure@36.[164,1404,337,1869]" captionTargetPageId="36" captionText="FIGURES 170 178. Opeatocerata stubbsi Smith, 1989, holotype ♂. 170. habitus, lateral view; 171. wing; 172. tergite 8 and terminalia, lateral view; 173. tergites 4 - 8, dorsal view; 174. terminalia; 175. lobes of cerci, dorsal view; 176. descendant plate of anterior cercus, posterior view; 177. subepandrial sclerite and bacilliform sclerite, dorsal view; 178. ejaculatory apodeme, phallus and hypandrium, lateral view." httpUri="https://zenodo.org/record/252541/files/figure.png" pageId="19" pageNumber="521">Fig. 174</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>