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<mods:title id="C5FC2E42425D5C79C89C26880F0B32ED">The Pachynidae fam. nov. (Crustacea: Amphipoda: Lysianassoidea) 3246</mods:title>
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<paragraph id="CD7636C5363BFF97B8DF0CC9FDA1488E" blockId="30.[151,544,1507,1534]" box="[151,544,1507,1534]" pageId="30" pageNumber="31">
<heading id="963E81A9363BFF97B8DF0CC9FDA1488E" bold="true" box="[151,544,1507,1534]" fontSize="11" level="1" pageId="30" pageNumber="31" reason="1">
<taxonomicName id="0AC94D46363BFF97B8DF0CC9FDA1488E" ID-CoL="7PP39" authority="Stephensen, 1925" authorityName="Stephensen" authorityYear="1925" box="[151,544,1507,1534]" class="Malacostraca" family="Pakynidae" genus="Pachychelium" kingdom="Animalia" order="Amphipoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="FFBDEAD7363BFF97B8DF0CC9FDA1488E" bold="true" box="[151,544,1507,1534]" pageId="30" pageNumber="31">
<emphasis id="FFBDEAD7363BFF97B8DF0CC9FEC2488D" bold="true" box="[151,323,1508,1534]" italics="true" pageId="30" pageNumber="31">Pachychelium</emphasis>
<bibRefCitation id="A9584B34363BFF97B9030CCEFDA1488E" author="Stephensen, K." box="[331,544,1507,1533]" pageId="30" pageNumber="31" pagination="101 - 178" refId="ref40634" refString="Stephensen, K. (1925) Crustacea Malacostraca, VI: (Amphipoda, II). Danish Ingolf-Expedition, 3 (9), 101 - 178." type="journal article" year="1925">Stephensen, 1925</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="85D3654E363BFF97B8DF0F07FC534B2C" pageId="30" pageNumber="31" type="reference_group">
<paragraph id="CD7636C5363BFF97B8DF0F07FC534B2C" blockId="30.[151,1436,1578,1631]" pageId="30" pageNumber="31">
<treatmentCitationGroup id="EDD911EB363BFF97B8DF0F07FC534B2C" pageId="30" pageNumber="31">
<treatmentCitation id="4C6810D4363BFF97B8DF0F07FD944B33" author="Stephensen, K." box="[151,533,1578,1600]" page="121" pageId="30" pageNumber="31" year="1925">
<taxonomicName id="0AC94D46363BFF97B8DF0F07FD944B33" ID-CoL="7PP39" authority="Stephensen, 1925: 121" authorityName="Stephensen" authorityPageNumber="121" authorityYear="1925" box="[151,533,1578,1600]" class="Malacostraca" family="Pakynidae" genus="Pachychelium" kingdom="Animalia" order="Amphipoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="FFBDEAD7363BFF97B8DF0F07FEA74B33" box="[151,294,1578,1600]" italics="true" pageId="30" pageNumber="31">Pachychelium</emphasis>
<bibRefCitation id="A9584B34363BFF97B9640F07FD944B33" author="Stephensen, K." box="[300,533,1578,1600]" pageId="30" pageNumber="31" pagination="101 - 178" refId="ref40634" refString="Stephensen, K. (1925) Crustacea Malacostraca, VI: (Amphipoda, II). Danish Ingolf-Expedition, 3 (9), 101 - 178." type="journal article" year="1925">Stephensen, 1925: 121</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. ―
<treatmentCitation id="4C6810D4363BFF97BA7F0F07FCB14B33" author="Schellenberg, A." box="[567,816,1578,1600]" page="296" pageId="30" pageNumber="31" year="1926">
<bibRefCitation id="A9584B34363BFF97BA7F0F07FCB14B33" author="Schellenberg, A." box="[567,816,1578,1600]" pageId="30" pageNumber="31" pagination="235 - 414" refId="ref40521" refString="Schellenberg, A. (1926) Die Gammariden der deutschen Sudpolar-Expedition 1901 - 1903. Deutsche Sudpolar-Expedition, Zoology, 10, 18, 235 - 414." type="journal article" year="1926">Schellenberg, 1926: 296</bibRefCitation>
</treatmentCitation>
. ―Schellenberg, 1931: 18. ―K.H.
<treatmentCitation id="4C6810D4363BFF97BCD10F07FAD24B33" author="Barnard, K. H." box="[1177,1363,1578,1600]" page="75" pageId="30" pageNumber="31" year="1932">
<bibRefCitation id="A9584B34363BFF97BCD10F07FAD24B33" author="Barnard, K. H." box="[1177,1363,1578,1600]" pageId="30" pageNumber="31" pagination="1 - 326" refId="ref39306" refString="Barnard, K. H. (1932) Amphipoda. ' Discovery' Reports, 5, 1 - 326, pl. 1." type="journal article" year="1932">Barnard, 1932: 75</bibRefCitation>
</treatmentCitation>
. ―J.L.
<treatmentCitation id="4C6810D4363BFF97B88F0F64FE184B2C" author="Barnard, J. L." box="[199,409,1609,1631]" page="354" pageId="30" pageNumber="31" year="1969">
<bibRefCitation id="A9584B34363BFF97B88F0F64FE184B2C" author="Barnard, J. L." box="[199,409,1609,1631]" pageId="30" pageNumber="31" pagination="1 - 535" refId="ref39150" refString="Barnard, J. L. (1969 a) The families and genera of marine gammaridean Amphipoda. United States National Museum Bulletin, 271, 1 - 535." type="journal article" year="1969">Barnard, 1969a: 354</bibRefCitation>
</treatmentCitation>
. ―
<treatmentCitation id="4C6810D4363BFF97B9F40F67FDE64B2C" author="Lowry, J. K." box="[444,615,1609,1631]" page="99" pageId="30" pageNumber="31" year="1984">
<bibRefCitation id="A9584B34363BFF97B9F40F67FDE64B2C" author="Lowry, J. K." box="[444,615,1609,1631]" pageId="30" pageNumber="31" pagination="51 - 105" refId="ref40103" refString="Lowry, J. K. (1984) Systematics of the pachynid group of lysianassoid Amphipoda (Crustacea). Records of the Australian Museum, 36 (2), 51 - 105." type="journal article" year="1984">Lowry, 1984: 99</bibRefCitation>
</treatmentCitation>
. ―
<treatmentCitation id="4C6810D4363BFF97BAC20F64FC4F4B2C" author="Barnard, J. L. &amp; Karaman, G. S." box="[650,974,1609,1631]" page="509" pageId="30" pageNumber="31" year="1991">
<bibRefCitation id="A9584B34363BFF97BAC20F64FC4F4B2C" author="Barnard, J. L. &amp; Karaman, G. S." box="[650,974,1609,1631]" pageId="30" pageNumber="31" pagination="419 - 866" refId="ref39259" refString="Barnard, J. L. &amp; Karaman, G. S. (1991) The families and genera of marine gammaridean Amphipoda (except marine gammaroids). Records of the Australian Museum, Supplement, 13 (2), 419 - 866." type="journal article" year="1991">Barnard &amp; Karaman, 1991: 509</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="85D3654E363BFF97B8DF0FA1FE9A4AB7" pageId="30" pageNumber="31" type="type_taxon">
<paragraph id="CD7636C5363BFF97B8DF0FA1FC254BD7" blockId="30.[151,1436,1675,1988]" box="[151,932,1675,1700]" pageId="30" pageNumber="31">
<emphasis id="FFBDEAD7363BFF97B8DF0FA1FEB34BD0" box="[151,306,1675,1700]" inLineHeading="true" pageId="30" pageNumber="31" reason="1">
<emphasis id="FFBDEAD7363BFF97B8DF0FA1FEAB4BD7" bold="true" box="[151,298,1675,1700]" pageId="30" pageNumber="31">
<typeStatus id="12728867363BFF97B8DF0FA1FF504BD7" box="[151,209,1676,1700]" pageId="30" pageNumber="31">Type</typeStatus>
species
</emphasis>
.
</emphasis>
<taxonomicName id="0AC94D46363BFF97B9710FA1FD7C4BD7" ID-CoL="4BQL4" authority="Stephensen, 1925" authorityName="Stephensen" authorityYear="1925" box="[313,765,1676,1700]" class="Malacostraca" family="Pakynidae" genus="Pachychelium" kingdom="Animalia" order="Amphipoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="davidis">
<emphasis id="FFBDEAD7363BFF97B9710FA1FDAE4BD7" box="[313,559,1676,1700]" italics="true" pageId="30" pageNumber="31">Pachychelium davidis</emphasis>
<bibRefCitation id="A9584B34363BFF97BA7E0FA1FD7C4BD7" author="Stephensen, K." box="[566,765,1676,1700]" pageId="30" pageNumber="31" pagination="101 - 178" refId="ref40634" refString="Stephensen, K. (1925) Crustacea Malacostraca, VI: (Amphipoda, II). Danish Ingolf-Expedition, 3 (9), 101 - 178." type="journal article" year="1925">Stephensen, 1925</bibRefCitation>
</taxonomicName>
, by monotypy.
</paragraph>
<paragraph id="CD7636C5363BFF97B8810F82FE9A4AB7" blockId="30.[151,1436,1675,1988]" pageId="30" pageNumber="31">
<emphasis id="FFBDEAD7363BFF97B8810F82FE5A4BB4" bold="true" box="[201,475,1711,1736]" inLineHeading="true" pageId="30" pageNumber="31" reason="1">Diagnostic description.</emphasis>
Body shape slender.
<emphasis id="FFBDEAD7363BFF97BA830F9DFAAE4BBB" bold="true" box="[715,1327,1711,1736]" pageId="30" pageNumber="31">Head lateral cephalic lobe large, narrowly rounded</emphasis>
. Antenna 1 peduncular article 1 without posterodistal spine; accessory flagellum article 1 long; [callynophore not known for female]. Mandible with accessory setal row absent; left lamina dentata absent; palp attached distally; palp article 2 broad, article 3 with subparallel margins or distally tapered. Maxilla 1 outer plate setal-teeth smooth, at least one inner row setal-tooth absent; palp absent.
<emphasis id="FFBDEAD7363BFF97BA240E12FC4D4A2B" bold="true" box="[620,972,1855,1880]" pageId="30" pageNumber="31">Maxilliped inner plate absent</emphasis>
; outer plate medium size;
<emphasis id="FFBDEAD7363BFF97BCBF0E6DFE154A0F" bold="true" pageId="30" pageNumber="31">palp 3-articulate (article 4 absent)</emphasis>
.
<emphasis id="FFBDEAD7363BFF97B9EA0E4EFC3A4A0F" bold="true" box="[418,955,1891,1916]" pageId="30" pageNumber="31">Gnathopod 1 ischium smaller than propodus</emphasis>
; corner of palm with simple robust seta or without any robust seta. Gnathopod 2 palm obtuse.
<emphasis id="FFBDEAD7363BFF97BA9C0EAAFAB84AD3" bold="true" box="[724,1337,1927,1952]" pageId="30" pageNumber="31">Epimeron 3 posteroventral corner broadly rounded</emphasis>
. Uropod 3 biramous.
</paragraph>
</subSubSection>
<subSubSection id="85D3654E363AFF96B88F09B5FBDB4CD8" pageId="31" pageNumber="32" type="discussion">
<paragraph id="CD7636C5363AFF96B88F09B5FD0B4C10" blockId="31.[151,1436,151,428]" pageId="31" pageNumber="32">
<emphasis id="FFBDEAD7363AFF96B88F09B5FEBA4DC3" bold="true" box="[199,315,152,176]" inLineHeading="true" pageId="31" pageNumber="32" reason="1">Remarks.</emphasis>
With the new species described from the boreal eastern North Pacific and the tropical western South Pacific and the reassignment of some species to
<taxonomicName id="0AC94D46363AFF96BA800991FCEF4DA7" authorityName="Lowry &amp; Stoddart" authorityYear="2012" box="[712,878,188,212]" class="Malacostraca" family="Pakynidae" genus="Ultimachelium" kingdom="Animalia" order="Amphipoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="FFBDEAD7363AFF96BA800991FCEF4DA7" box="[712,878,188,212]" italics="true" pageId="31" pageNumber="32">Ultimachelium</emphasis>
</taxonomicName>
, the distribution of the previously bipolar genus
<taxonomicName id="0AC94D46363AFF96B8DF09CDFEB74D8B" authorityName="Stephensen" authorityYear="1925" box="[151,310,224,248]" class="Malacostraca" family="Pakynidae" genus="Pachychelium" kingdom="Animalia" order="Amphipoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="FFBDEAD7363AFF96B8DF09CDFEB74D8B" box="[151,310,224,248]" italics="true" pageId="31" pageNumber="32">Pachychelium</emphasis>
</taxonomicName>
is changed significantly. The sister taxon of
<taxonomicName id="0AC94D46363AFF96BB6209CDFC484D8B" authorityName="Stephensen" authorityYear="1925" box="[810,969,224,248]" class="Malacostraca" family="Pakynidae" genus="Pachychelium" kingdom="Animalia" order="Amphipoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="FFBDEAD7363AFF96BB6209CDFC484D8B" box="[810,969,224,248]" italics="true" pageId="31" pageNumber="32">Pachychelium</emphasis>
</taxonomicName>
appears to be
<taxonomicName id="0AC94D46363AFF96BC2209CDFA914D8B" authorityName="Lowry &amp; Stoddart" authorityYear="2012" box="[1130,1296,224,248]" class="Malacostraca" family="Pakynidae" genus="Ultimachelium" kingdom="Animalia" order="Amphipoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="FFBDEAD7363AFF96BC2209CDFA914D8B" box="[1130,1296,224,248]" italics="true" pageId="31" pageNumber="32">Ultimachelium</emphasis>
</taxonomicName>
. The genera differ in the number of articles in the maxillipedal palp (article 4 absent in
<taxonomicName id="0AC94D46363AFF96BB900829FBF74C6F" authorityName="Stephensen" authorityYear="1925" box="[984,1142,260,284]" class="Malacostraca" family="Pakynidae" genus="Pachychelium" kingdom="Animalia" order="Amphipoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="FFBDEAD7363AFF96BB900829FBF74C6F" box="[984,1142,260,284]" italics="true" pageId="31" pageNumber="32">Pachychelium</emphasis>
</taxonomicName>
, large and well developed in
<taxonomicName id="0AC94D46363AFF96B8FC0805FEDB4C33" authorityName="Lowry &amp; Stoddart" authorityYear="2012" box="[180,346,296,320]" class="Malacostraca" family="Pakynidae" genus="Ultimachelium" kingdom="Animalia" order="Amphipoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="FFBDEAD7363AFF96B8FC0805FEDB4C33" box="[180,346,296,320]" italics="true" pageId="31" pageNumber="32">Ultimachelium</emphasis>
</taxonomicName>
) and in epimeron 3 which has a broadly rounded posteroventral corner in
<taxonomicName id="0AC94D46363AFF96BCD00805FAB74C33" authorityName="Stephensen" authorityYear="1925" box="[1176,1334,296,320]" class="Malacostraca" family="Pakynidae" genus="Pachychelium" kingdom="Animalia" order="Amphipoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="FFBDEAD7363AFF96BCD00805FAB74C33" box="[1176,1334,296,320]" italics="true" pageId="31" pageNumber="32">Pachychelium</emphasis>
</taxonomicName>
, projecting and narrowly rounded in
<taxonomicName id="0AC94D46363AFF96B9940861FD034C17" authorityName="Lowry &amp; Stoddart" authorityYear="2012" box="[476,642,332,356]" class="Malacostraca" family="Pakynidae" genus="Ultimachelium" kingdom="Animalia" order="Amphipoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="FFBDEAD7363AFF96B9940861FD034C17" box="[476,642,332,356]" italics="true" pageId="31" pageNumber="32">Ultimachelium</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="CD7636C5363AFF96B88F0842FAEF4CFB" blockId="31.[151,1436,151,428]" box="[199,1390,367,392]" pageId="31" pageNumber="32">
<emphasis id="FFBDEAD7363AFF96B88F0842FE3C4CFB" bold="true" box="[199,445,367,392]" inLineHeading="true" pageId="31" pageNumber="32" reason="1">Species composition.</emphasis>
<taxonomicName id="0AC94D46363AFF96B98C085DFC084CF4" authority="Stephensen, 1925" authorityName="Stephensen" authorityYear="1925" box="[452,905,367,392]" class="Malacostraca" family="Pakynidae" genus="Pachychelium" kingdom="Animalia" order="Amphipoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="davidis">
<emphasis id="FFBDEAD7363AFF96B98C085DFD3B4CFB" box="[452,698,368,392]" italics="true" pageId="31" pageNumber="32">Pachychelium davidis</emphasis>
<bibRefCitation id="A9584B34363AFF96BA890842FC084CF4" author="Stephensen, K." box="[705,905,367,391]" pageId="31" pageNumber="32" pagination="101 - 178" refId="ref40634" refString="Stephensen, K. (1925) Crustacea Malacostraca, VI: (Amphipoda, II). Danish Ingolf-Expedition, 3 (9), 101 - 178." type="journal article" year="1925">Stephensen, 1925</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="0AC94D46363AFF96BBDD085CFB994CFB" authority="Lowry &amp; Stoddart, 2012" authorityName="Lowry &amp; Stoddart" authorityYear="2012" box="[917,1048,368,392]" class="Malacostraca" family="Pakynidae" genus="Pachychelium" kingdom="Animalia" order="Amphipoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="fucaensis" status="sp. nov.">
<emphasis id="FFBDEAD7363AFF96BBDD085CFB994CFB" box="[917,1048,368,392]" italics="true" pageId="31" pageNumber="32">P. fucaensis</emphasis>
</taxonomicName>
<emphasis id="FFBDEAD7363AFF96BC57085DFBF84CFB" bold="true" box="[1055,1145,368,392]" pageId="31" pageNumber="32">
<taxonomicNameLabel id="E48E57AC363AFF96BC57085DFBF84CFB" box="[1055,1145,368,392]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<taxonomicName id="0AC94D46363AFF96BCC0085CFA8C4CFB" authority="Lowry &amp; Stoddart, 2012" authorityName="Lowry &amp; Stoddart" authorityYear="2012" box="[1160,1293,368,392]" class="Malacostraca" family="Pakynidae" genus="Pachychelium" kingdom="Animalia" order="Amphipoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="tropicalis" status="sp. nov.">
<emphasis id="FFBDEAD7363AFF96BCC0085CFA8C4CFB" box="[1160,1293,368,392]" italics="true" pageId="31" pageNumber="32">P. tropicalis</emphasis>
</taxonomicName>
<emphasis id="FFBDEAD7363AFF96BD5C085DFAEF4CFB" bold="true" box="[1300,1390,368,392]" pageId="31" pageNumber="32">
<taxonomicNameLabel id="E48E57AC363AFF96BD5C085DFAEF4CFB" box="[1300,1390,368,392]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="CD7636C5363AFF96B8DF08B9FBDB4CD8" blockId="31.[151,1436,151,428]" box="[151,1114,404,428]" pageId="31" pageNumber="32">
<taxonomicName id="0AC94D46363AFF96B8DF08B9FD254CDF" authority="Schellenberg, 1926" authorityName="Schellenberg" authorityYear="1926" box="[151,676,404,428]" class="Malacostraca" family="Pakynidae" genus="Pachychelium" kingdom="Animalia" order="Amphipoda" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="antarcticum">
<emphasis id="FFBDEAD7363AFF96B8DF08B9FE434CDF" box="[151,450,404,428]" italics="true" pageId="31" pageNumber="32">Pachychelium antarcticum</emphasis>
<bibRefCitation id="A9584B34363AFF96B98208B9FD254CDF" author="Schellenberg, A." box="[458,676,404,428]" pageId="31" pageNumber="32" pagination="235 - 414" refId="ref40521" refString="Schellenberg, A. (1926) Die Gammariden der deutschen Sudpolar-Expedition 1901 - 1903. Deutsche Sudpolar-Expedition, Zoology, 10, 18, 235 - 414." type="journal article" year="1926">Schellenberg, 1926</bibRefCitation>
</taxonomicName>
, is regarded as an unidentifiable taxon.
</paragraph>
</subSubSection>
</treatment>
</document>