421 lines
52 KiB
XML
421 lines
52 KiB
XML
<document id="37AA3CFC32660F655B85862C5D09E4FE" ID-CLB-Dataset="30789" ID-DOI="10.5852/ejt.2018.402" ID-GBIF-Dataset="8f1510f6-1b79-467f-9e8d-f193ac82cc4b" ID-Zenodo-Dep="1169559" ID-ZooBank="A3CE68AA-C5C6-40B7-B57C-6EF6D949149B" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1518136191129" checkinUser="plazi" docAuthor="Ruiz-C, Raquel I., Román-Valencia, César, Taphorn, Donald C., Buckup, Paulo A. & Ortega, Hernán" docDate="2018" docId="2C0787E7FFC0D435FD5CFDF3FF50FEC4" docLanguage="en" docName="ejt-402_Ruiz-C_Roman-Valencia.pdf.imf" docOrigin="European Journal of Taxonomy 402" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Astyanax embera Ruiz-C & Román-Valencia & Taphorn & Buckup & Ortega 2018, sp. nov." docType="treatment" docVersion="14" lastPageNumber="10" masterDocId="D03EFF9FFFC5D43CFFDCFFD0FE0FFFBE" masterDocTitle="Revision of the Astyanax orthodus species-group (Teleostei: Characidae) with descriptions of three new species" masterLastPageNumber="45" masterPageNumber="1" originalUpdateDomain="Boston" originalUpdateTime="1547298528796" originalUpdateUser="yanell" pageNumber="6" updateTime="1698520653973" updateUser="ExternalLinkService">
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<mods:titleInfo id="76A95D5F767AD4D05FD6C07E8510B18B">
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<mods:title id="956BA448AA292A07E34DE6B09A149411">Revision of the Astyanax orthodus species-group (Teleostei: Characidae) with descriptions of three new species</mods:title>
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<mods:namePart id="56BD234582301967D55D36B3E805A5B4">Ruiz-C, Raquel I.</mods:namePart>
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<mods:namePart id="06CFC18258549806515427E862DD2867">Román-Valencia, César</mods:namePart>
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<mods:namePart id="37441E8561399848E496521926C0B08F">Taphorn, Donald C.</mods:namePart>
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<mods:namePart id="B4F5EAED59B1D0CC8308B3AFA944115D">Buckup, Paulo A.</mods:namePart>
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<mods:namePart id="F9008AE59E2DA53561A3A1B74347EB2E">Ortega, Hernán</mods:namePart>
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<mods:title id="3FA3DED732DE18251B736D5A20FA1A1C">European Journal of Taxonomy</mods:title>
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<mods:part id="84A0DDD98C3ED4A0D600ED9B252A7303">
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<mods:date id="B3B8CB562B624ADB6F54D0A759795936">2018</mods:date>
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<mods:number id="ABBEEA582504B18592CA9E4E7D18592B">2018-02-08</mods:number>
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<treatment id="2C0787E7FFC0D435FD5CFDF3FF50FEC4" ID-DOI="http://doi.org/10.5281/zenodo.5622610" ID-GBIF-Taxon="140562876" ID-Zenodo-Dep="5622610" LSID="urn:lsid:plazi:treatment:2C0787E7FFC0D435FD5CFDF3FF50FEC4" httpUri="http://treatment.plazi.org/id/2C0787E7FFC0D435FD5CFDF3FF50FEC4" lastPageId="9" lastPageNumber="10" pageId="5" pageNumber="6">
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<subSubSection id="ECB4657AFFC0D439FD5CFDF3FAB3FDDE" pageId="5" pageNumber="6" type="nomenclature">
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<paragraph id="A41136F1FFC0D439FD5CFDF3FDBBFD83" blockId="5.[375,1212,546,643]" box="[640,948,546,573]" pageId="5" pageNumber="6">
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<heading id="FF59819DFFC0D439FD5CFDF3FDBBFD83" box="[640,948,546,573]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="6" reason="2">
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<taxonomicName id="63AE4D72FFC0D439FD5CFDF3FD5DFD83" authority="Ruiz-C & Román-Valencia & Taphorn & Buckup & Ortega, 2018" authorityName="Ruiz-C & Román-Valencia & Taphorn & Buckup & Ortega" authorityYear="2018" box="[640,850,546,573]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="embera" status="sp. nov.">
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<emphasis id="96DAEAE3FFC0D439FD5CFDF3FD5DFD83" bold="true" box="[640,850,546,573]" italics="true" pageId="5" pageNumber="6">Astyanax embera</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="8DE95798FFC0D439FC85FDF3FDBBFD83" box="[857,948,547,573]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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||
</paragraph>
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||
<paragraph id="A41136F1FFC0D439FEABFD96FAB3FDDE" blockId="5.[375,1212,546,643]" box="[375,1212,582,608]" pageId="5" pageNumber="6">urn:lsid:zoobank.org:act:9B1792A8-70D6-4167-9136-0DD74844B040</paragraph>
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</subSubSection>
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<subSubSection id="ECB4657AFFC0D439FD40FDB9FD97FD3D" box="[668,920,617,643]" pageId="5" pageNumber="6" type="description">
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<paragraph id="A41136F1FFC0D439FD40FDB9FD97FD3D" blockId="5.[375,1212,546,643]" box="[668,920,617,643]" pageId="5" pageNumber="6">
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<figureCitation id="3C952A74FFC0D439FD40FDB9FCE8FD3D" box="[668,743,617,643]" captionStart="Fig. 2" captionStartId="5.[189,232,1928,1954]" captionTargetBox="[189,1398,1338,1902]" captionTargetId="figure@5.[189,1398,1338,1902]" captionTargetPageId="5" captionText="Fig. 2. Astyanax embera sp. nov., holotype, 83.0 mm SL, Telembí River at mouth of Yamunde River, 1 km below Barbacoas, Municipality of Barbacoas, Nariño, Pacific coast of Colombia (IUQ 3614). Scale bar = 1 cm." httpUri="https://zenodo.org/record/1169563/files/figure.png" pageId="5" pageNumber="6">Figs 2</figureCitation>
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,
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<figureCitation id="3C952A74FFC0D439FD2FFDB9FD17FD3D" box="[755,792,617,643]" captionStart="Fig. 3" captionStartId="6.[189,232,1964,1990]" captionTargetBox="[462,1020,848,1930]" captionTargetId="figure@6.[446,1142,837,1938]" captionTargetPageId="6" captionText="Fig. 3. Shape of caudal-peduncle spot in species of the Astyanax orthodus species-group. A. Astyanax orthodus Eigenmann, 1907. B. Astyanax embera sp. nov. Scale bar = 1 cm." httpUri="https://zenodo.org/record/1169565/files/figure.png" pageId="5" pageNumber="6">3B</figureCitation>
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,
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<figureCitation id="3C952A74FFC0D439FCF8FDB9FD3AFD3D" box="[804,821,617,643]" captionStart="Fig. 4" captionStartId="8.[189,232,2000,2026]" captionTargetBox="[189,1392,764,1921]" captionTargetId="figure@8.[151,1417,755,1932]" captionTargetPageId="8" captionText="Fig. 4.Distribution of speciesof the Astyanaxorthodus species-group(symbol includestypelocalities)." httpUri="https://zenodo.org/record/1169567/files/figure.png" pageId="5" pageNumber="6">4</figureCitation>
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,
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<tableCitation id="E92C034AFFC0D439FC9DFDB9FD97FD3D" box="[833,920,617,643]" captionStart="Table 1" captionStartId="11.[145,214,599,625]" captionTargetPageId="11" captionText="Table 1. Morphometric data for some members of the Astyanax orthodus species-group." httpUri="http://table.plazi.org/id/F0D16679FFCED437FF4DFD87FA92FDCF" pageId="5" pageNumber="6" tableUuid="F0D16679FFCED437FF4DFD87FA92FDCF">Table 1</tableCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="ECB4657AFFC0D439FF61FD7AFB2AFBEA" pageId="5" pageNumber="6" type="diagnosis">
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||
<paragraph id="A41136F1FFC0D439FF61FD7AFF39FD7A" blockId="5.[189,310,682,708]" box="[189,310,682,708]" pageId="5" pageNumber="6">
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||
<heading id="FF59819DFFC0D439FF61FD7AFF39FD7A" bold="true" box="[189,310,682,708]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
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<emphasis id="96DAEAE3FFC0D439FF61FD7AFF39FD7A" bold="true" box="[189,310,682,708]" pageId="5" pageNumber="6">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="A41136F1FFC0D439FF61FD09FB2AFBEA" blockId="5.[189,1399,729,1108]" pageId="5" pageNumber="6">
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<taxonomicName id="63AE4D72FFC0D439FF61FD09FF87FD4D" authority="Ruiz-C & Román-Valencia & Taphorn & Buckup & Ortega, 2018" authorityName="Ruiz-C & Román-Valencia & Taphorn & Buckup & Ortega" authorityYear="2018" box="[189,392,729,755]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="embera" status="sp. nov.">
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<emphasis id="96DAEAE3FFC0D439FF61FD09FF87FD4D" box="[189,392,729,755]" italics="true" pageId="5" pageNumber="6">Astyanax embera</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="8DE95798FFC0D439FE4CFD09FFE4FD4D" box="[400,491,729,755]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
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is a member of the
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<taxonomicName id="63AE4D72FFC0D439FD0AFD09FD31FD4D" authorityName="Eigenmann in Eigenmann & Ogle" authorityYear="1907" box="[726,830,729,755]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="orthodus">
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<emphasis id="96DAEAE3FFC0D439FD0AFD09FD31FD4D" box="[726,830,729,755]" italics="true" pageId="5" pageNumber="6">orthodus</emphasis>
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</taxonomicName>
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species-group of
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<taxonomicName id="63AE4D72FFC0D439FBCDFD09FA72FD4D" authorityName="Baird & Girard" authorityYear="1854" box="[1041,1149,729,755]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
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<emphasis id="96DAEAE3FFC0D439FBCDFD09FA72FD4D" box="[1041,1149,729,755]" italics="true" pageId="5" pageNumber="6">Astyanax</emphasis>
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</taxonomicName>
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, differing from other members of the group as follows: from
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<emphasis id="96DAEAE3FFC0D439FD7CFD2DFD22FCA8" box="[672,813,764,791]" italics="true" pageId="5" pageNumber="6">A. villwocki</emphasis>
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,
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<taxonomicName id="63AE4D72FFC0D439FCE2FD2DFDC4FCA8" authorityName="Myers" authorityYear="1942" box="[830,971,764,791]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="superbus">
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<emphasis id="96DAEAE3FFC0D439FCE2FD2DFDC4FCA8" box="[830,971,764,791]" italics="true" pageId="5" pageNumber="6">A. superbus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="63AE4D72FFC0D439FC00FD2DFA62FCA8" authority="Ruiz-C & Román-Valencia & Taphorn & Buckup & Ortega, 2018" authorityName="Ruiz-C & Román-Valencia & Taphorn & Buckup & Ortega" authorityYear="2018" box="[988,1133,764,791]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="gandhiae" status="sp. nov.">
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<emphasis id="96DAEAE3FFC0D439FC00FD2DFA62FCA8" box="[988,1133,764,791]" italics="true" pageId="5" pageNumber="6">A. gandhiae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="8DE95798FFC0D439FBABFD2DFAD9FCA9" box="[1143,1238,765,791]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
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,
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<taxonomicName id="63AE4D72FFC0D439FB38FD2DFB78FCA8" authorityName="Ruiz-C & Román-Valencia & Taphorn & Buckup & Ortega" authorityYear="2018" box="[1252,1399,764,791]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="bopiensis" status="nom. nov.">
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<emphasis id="96DAEAE3FFC0D439FB38FD2DFB78FCA8" box="[1252,1399,764,791]" italics="true" pageId="5" pageNumber="6">A. bopiensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="8DE95798FFC0D439FF61FCF0FF3CFC84" box="[189,307,800,826]" pageId="5" pageNumber="6">nom. nov.</taxonomicNameLabel>
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,
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<taxonomicName id="63AE4D72FFC0D439FEE2FCF0FFD5FC87" authority="Ruiz-C & Román-Valencia & Taphorn & Buckup & Ortega, 2018" authorityName="Ruiz-C & Román-Valencia & Taphorn & Buckup & Ortega" authorityYear="2018" box="[318,474,799,826]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="boliviensis" status="sp. nov.">
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<emphasis id="96DAEAE3FFC0D439FEE2FCF0FFD5FC87" box="[318,474,799,826]" italics="true" pageId="5" pageNumber="6">A. boliviensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="8DE95798FFC0D439FE3DFCF0FC35FC84" box="[481,570,800,826]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
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and
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<taxonomicName id="63AE4D72FFC0D439FDADFCF0FCF2FC87" authorityName="Ruiz-C & Román-Valencia & Taphorn & Buckup & Ortega" authorityYear="2018" baseAuthorityName="Torres-Mejia, Hernandez & Senechal" baseAuthorityYear="2012" box="[625,765,800,826]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="yariguies" status="comb. nov.">
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<emphasis id="96DAEAE3FFC0D439FDADFCF0FCF2FC87" box="[625,765,800,826]" italics="true" pageId="5" pageNumber="6">A. yariguies</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="8DE95798FFC0D439FCD8FCCFFD8BFC84" box="[772,900,799,826]" pageId="5" pageNumber="6">comb. nov.</taxonomicNameLabel>
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by the absence of a dark stripe on the sides of the body (vs dark lateral stripe present or anterior prolongation of the caudal peduncle spot extending anteriorly over the silvery lateral stripe). It differs from
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<taxonomicName id="63AE4D72FFC0D439FC92FCB6FDD8FC3E" authorityName="Eigenmann in Eigenmann & Ogle" authorityYear="1907" box="[846,983,870,896]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="orthodus">
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<emphasis id="96DAEAE3FFC0D439FC92FCB6FDD8FC3E" box="[846,983,870,896]" italics="true" pageId="5" pageNumber="6">A. orthodus</emphasis>
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</taxonomicName>
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in having the caudal peduncle spot extending onto the ventral and dorsal margins of the caudal peduncle, covering 4–6 horizontal scale rows (vs covering 2–4 horizontal scale rows on the caudal peduncle,
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<figureCitation id="3C952A74FFC0D439FC76FC7CFDFFFC78" box="[938,1008,940,966]" captionStart="Fig. 3" captionStartId="6.[189,232,1964,1990]" captionTargetBox="[462,1020,848,1930]" captionTargetId="figure@6.[446,1142,837,1938]" captionTargetPageId="6" captionText="Fig. 3. Shape of caudal-peduncle spot in species of the Astyanax orthodus species-group. A. Astyanax orthodus Eigenmann, 1907. B. Astyanax embera sp. nov. Scale bar = 1 cm." httpUri="https://zenodo.org/record/1169565/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
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). It also differs from
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<taxonomicName id="63AE4D72FFC0D439FB31FC7DFB79FC78" authorityName="Eigenmann in Eigenmann & Ogle" authorityYear="1907" box="[1261,1398,940,967]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="orthodus">
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<emphasis id="96DAEAE3FFC0D439FB31FC7DFB79FC78" box="[1261,1398,940,967]" italics="true" pageId="5" pageNumber="6">A. orthodus</emphasis>
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</taxonomicName>
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in having an unpigmented band across the anal-fin rays in both sexes (vs pigmentation uniform on analfin rays). It differs from
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<taxonomicName id="63AE4D72FFC0D439FE03FC23FC41FBB2" authorityName="Ruiz-C & Román-Valencia & Taphorn & Buckup & Ortega" authorityYear="2018" baseAuthorityName="Boulenger" baseAuthorityYear="1892" box="[479,590,1011,1037]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="moorii" status="comb. nov.">
|
||
<emphasis id="96DAEAE3FFC0D439FE03FC23FC41FBB2" box="[479,590,1011,1037]" italics="true" pageId="5" pageNumber="6">A. moorii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8DE95798FFC0D439FD8BFC22FCD5FBB3" box="[599,730,1010,1037]" pageId="5" pageNumber="6">comb. nov.</taxonomicNameLabel>
|
||
in the body depth> 39% SL (vs <39% SL), the dorsalpectoral fin distance> 40.5% SL (vs <40.5% SL), maxilla length <33.5% HL (vs> 33.5% SH), upper jaw length <32% HL (vs> 47% SH) and by the number of lateral-line scales 36–41 (vs 44–50).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="ECB4657AFFC0D439FF61FBABFD5BFB59" pageId="5" pageNumber="6" type="etymology">
|
||
<paragraph id="A41136F1FFC0D439FF61FBABFF4BFB2B" blockId="5.[189,324,1147,1173]" box="[189,324,1147,1173]" pageId="5" pageNumber="6">
|
||
<heading id="FF59819DFFC0D439FF61FBABFF4BFB2B" bold="true" box="[189,324,1147,1173]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
|
||
<emphasis id="96DAEAE3FFC0D439FF61FBABFF4BFB2B" bold="true" box="[189,324,1147,1173]" pageId="5" pageNumber="6">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="A41136F1FFC0D439FF61FB79FD5BFB59" blockId="5.[189,1399,1193,1255]" pageId="5" pageNumber="6">
|
||
The specific epithet
|
||
<taxonomicName id="63AE4D72FFC0D439FE6DFB79FC06FB7D" authorityName="Ruiz-C & Román-Valencia & Taphorn & Buckup & Ortega" authorityYear="2018" box="[433,521,1193,1219]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="embera">
|
||
<emphasis id="96DAEAE3FFC0D439FE6DFB79FC06FB7D" box="[433,521,1193,1219]" italics="true" pageId="5" pageNumber="6">embera</emphasis>
|
||
</taxonomicName>
|
||
is a noun in apposition and refers to the native Americans known as the Embera, who occupy the
|
||
<typeStatus id="7B158853FFC0D439FE34FB1DFC14FB59" box="[488,539,1229,1255]" pageId="5" pageNumber="6">type</typeStatus>
|
||
locality of this new taxon.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="F0D16679FFC0D439FF61F858FF85F857" httpUri="https://zenodo.org/record/1169563/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1928,1954]" targetBox="[189,1398,1338,1902]" targetPageId="5">
|
||
<paragraph id="A41136F1FFC0D439FF61F858FF85F857" blockId="5.[189,1399,1928,2025]" pageId="5" pageNumber="6">
|
||
<emphasis id="96DAEAE3FFC0D439FF61F858FF03F81C" bold="true" box="[189,268,1928,1954]" pageId="5" pageNumber="6">Fig. 2.</emphasis>
|
||
<taxonomicName id="63AE4D72FFC0D439FEC9F859FFEEF81D" authority="Ruiz-C & Román-Valencia & Taphorn & Buckup & Ortega, 2018" authorityName="Ruiz-C & Román-Valencia & Taphorn & Buckup & Ortega" authorityYear="2018" box="[277,481,1929,1955]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="embera" status="sp. nov.">
|
||
<emphasis id="96DAEAE3FFC0D439FEC9F859FFEEF81D" box="[277,481,1929,1955]" italics="true" pageId="5" pageNumber="6">Astyanax embera</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8DE95798FFC0D439FE36F859FC46F81D" box="[490,585,1929,1955]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, 83.0 mm SL, Telembí River at mouth of Yamunde River, 1 km below Barbacoas, Municipality of Barbacoas, Nariño, Pacific coast of Colombia (IUQ 3614). Scale bar = 1 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="ECB4657AFFC3D43AFF61FEDBFF49FCA7" pageId="6" pageNumber="7" type="materials_examined">
|
||
<paragraph id="A41136F1FFC3D43AFF61FEDBFFA5FE9B" blockId="6.[189,426,267,293]" box="[189,426,267,293]" pageId="6" pageNumber="7">
|
||
<heading id="FF59819DFFC3D43AFF61FEDBFFA5FE9B" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
|
||
<emphasis id="96DAEAE3FFC3D43AFF61FEDBFFA5FE9B" bold="true" box="[189,426,267,293]" pageId="6" pageNumber="7">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="A41136F1FFC3D43AFF36FEEAFF53FEEA" blockId="6.[189,1399,314,446]" box="[234,348,314,340]" pageId="6" pageNumber="7">
|
||
<emphasis id="96DAEAE3FFC3D43AFF36FEEAFF53FEEA" bold="true" box="[234,348,314,340]" pageId="6" pageNumber="7">Holotype</emphasis>
|
||
</paragraph>
|
||
<paragraph id="A41136F1FFC3D43AFF61FE8DFD1AFE00" blockId="6.[189,1399,314,446]" pageId="6" pageNumber="7">
|
||
<materialsCitation id="14C63CACFFC3D43AFF61FE8DFD1AFE00" ID-GBIF-Occurrence="1806442493" collectionCode="IUQ" country="Colombia" county="Barbacoas Municipality" elevation="61" latitude="1.6655555" location="Patía River Basin (Pacific Coast), Telembí River drainage, in Telembí River at mouth of Yamunde River, 1 km below Barbacoas" longLatPrecision="21" longitude="-78.153336" municipality="Patia River Basin" pageId="6" pageNumber="7" specimenCode="IUQ 3614" specimenCount="1" stateProvince="Nariño Department">
|
||
<collectingCountry id="DCB97661FFC3D43AFF61FE8DFF51FEC9" box="[189,350,349,375]" name="Colombia" pageId="6" pageNumber="7">COLOMBIA</collectingCountry>
|
||
: 83.0 mm SL,
|
||
<collectingRegion id="666AF813FFC3D43AFDC4FE8CFD0DFEC9" box="[536,770,348,375]" country="Colombia" name="Narino" pageId="6" pageNumber="7">Nariño Department</collectingRegion>
|
||
,
|
||
<collectingCounty id="4D704E7DFFC3D43AFCCDFE8CFA20FEC8" box="[785,1071,348,374]" pageId="6" pageNumber="7">Barbacoas Municipality</collectingCounty>
|
||
,
|
||
<location id="A171602AFFC3D43AFBE2FE8DFB7DFE24" LSID="urn:lsid:plazi:treatment:2C0787E7FFC0D435FD5CFDF3FF50FEC4:A171602AFFC3D43AFBE2FE8DFB7DFE24" country="Colombia" county="Barbacoas Municipality" latitude="1.6655555" longLatPrecision="21" longitude="-78.153336" municipality="Patia River Basin" name="Patía River Basin (Pacific Coast), Telembí River drainage, in Telembí River at mouth of Yamunde River, 1 km below Barbacoas" pageId="6" pageNumber="7" stateProvince="Nariño Department">
|
||
Patía River Basin (Pacific Coast), Telembí River drainage, in Telembí River at mouth of Yamunde River,
|
||
<quantity id="63569B14FFC3D43AFBBDFE50FA92FE24" box="[1121,1181,384,410]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="7" unit="km" value="1.0">1 km</quantity>
|
||
below Barbacoas
|
||
</location>
|
||
,
|
||
<geoCoordinate id="C19A5036FFC3D43AFF61FE71FF5CFE00" box="[189,339,417,446]" direction="north" orientation="latitude" pageId="6" pageNumber="7" precision="15" value="1.6655555">01°39′56″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="C19A5036FFC3D43AFE83FE71FFF7FE00" box="[351,504,417,446]" direction="west" orientation="longitude" pageId="6" pageNumber="7" precision="15" value="-78.153336">78°09′12″ W</geoCoordinate>
|
||
,
|
||
<elevation id="2F83D1C2FFC3D43AFDDFFE73FC72FE03" box="[515,637,419,446]" metricMagnitude="1" metricUnit="m" metricValue="6.1" pageId="6" pageNumber="7" unit="m" value="61.0">
|
||
<quantity id="63569B14FFC3D43AFDDFFE73FC30FE00" box="[515,575,419,446]" metricMagnitude="1" metricUnit="m" metricValue="6.1" pageId="6" pageNumber="7" unit="m" value="61.0">61 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
(
|
||
<specimenCode id="F4089E8AFFC3D43AFD50FE73FD06FE00" box="[652,777,419,446]" collectionCode="IUQ" country="Colombia" httpUri="http://grbio.org/cool/8xqv-ibyz" name="Laboratorio de Ictiologia" pageId="6" pageNumber="7">IUQ 3614</specimenCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="A41136F1FFC3D43AFF36FE35FF68FE41" blockId="6.[189,1399,485,793]" box="[234,359,485,511]" pageId="6" pageNumber="7">
|
||
<emphasis id="96DAEAE3FFC3D43AFF36FE35FF68FE41" bold="true" box="[234,359,485,511]" pageId="6" pageNumber="7">Paratypes</emphasis>
|
||
</paragraph>
|
||
<paragraph id="A41136F1FFC3D43AFF61FDD8FF49FCA7" blockId="6.[189,1399,485,793]" pageId="6" pageNumber="7">
|
||
<materialsCitation id="14C63CACFFC3D43AFF61FDD8FFC8FDD7" ID-GBIF-Occurrence="1806442499" collectionCode="IUQ" collectorName="La Tundera Brook & Barbacoas" country="Colombia" elevation="41" latitude="-1.6508334" location="Barbacoas Municipality, Patía River Basin, Telembi River Drainage" longLatPrecision="21" longitude="-78.17583" pageId="6" pageNumber="7" specimenCode="IUQ 2264" specimenCount="9" specimenCount-male="9" stateProvince="Nariño Department">
|
||
<collectingCountry id="DCB97661FFC3D43AFF61FDD8FF51FD9C" box="[189,350,520,546]" name="Colombia" pageId="6" pageNumber="7">COLOMBIA</collectingCountry>
|
||
: all from
|
||
<collectingRegion id="666AF813FFC3D43AFE0EFDD8FCB6FD9C" box="[466,697,520,546]" country="Colombia" name="Narino" pageId="6" pageNumber="7">Nariño Department</collectingRegion>
|
||
,
|
||
<location id="A171602AFFC3D43AFD1AFDD8FF22FDFB" LSID="urn:lsid:plazi:treatment:2C0787E7FFC0D435FD5CFDF3FF50FEC4:A171602AFFC3D43AFD1AFDD8FF22FDFB" country="Colombia" latitude="-1.6508334" longLatPrecision="21" longitude="-78.17583" name="Barbacoas Municipality, Patía River Basin, Telembi River Drainage" pageId="6" pageNumber="7" stateProvince="Nariño Department">Barbacoas Municipality, Patía River Basin, Telembi River Drainage</location>
|
||
:
|
||
<specimenCount id="B2A8FD78FFC3D43AFEE6FDFBFF8FFDFB" box="[314,384,555,581]" pageId="6" pageNumber="7" type="male">9 ♂♂</specimenCount>
|
||
,
|
||
<quantity id="63569B14FFC3D43AFE53FDFBFC34FDFB" box="[399,571,555,582]" metricMagnitude="-2" metricUnit="m" metricValue="4.63" metricValueMax="5.2299999999999995" metricValueMin="4.029999999999999" pageId="6" pageNumber="7" unit="mm" value="46.3" valueMax="52.3" valueMin="40.3">40.3–52.3 mm</quantity>
|
||
SL,
|
||
<location id="A171602AFFC3D43AFDABFDFBFA4DFDFB" LSID="urn:lsid:plazi:treatment:2C0787E7FFC0D435FD5CFDF3FF50FEC4:A171602AFFC3D43AFDABFDFBFA4DFDFB" box="[631,1090,555,581]" country="Colombia" latitude="-1.6508334" longLatPrecision="21" longitude="-78.17583" name="La Tundera Brook, Barbacoas, Nariño" pageId="6" pageNumber="7" stateProvince="Nariño Department">La Tundera Brook, Barbacoas, Nariño</location>
|
||
,
|
||
<geoCoordinate id="C19A5036FFC3D43AFB8CFDF9FAD9FDFB" box="[1104,1238,553,581]" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="15" value="-1.6508334">01°39′03″S</geoCoordinate>
|
||
<geoCoordinate id="C19A5036FFC3D43AFB3CFDF9FB7DFDFB" box="[1248,1394,553,581]" direction="west" orientation="longitude" pageId="6" pageNumber="7" precision="15" value="-78.17583">78°10′33″W</geoCoordinate>
|
||
,
|
||
<elevation id="2F83D1C2FFC3D43AFF61FD9EFF3AFDD6" box="[189,309,590,617]" metricMagnitude="1" metricUnit="m" metricValue="4.1" pageId="6" pageNumber="7" unit="m" value="41.0">
|
||
<quantity id="63569B14FFC3D43AFF61FD9EFEF7FDD7" box="[189,248,590,617]" metricMagnitude="1" metricUnit="m" metricValue="4.1" pageId="6" pageNumber="7" unit="m" value="41.0">41 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
(
|
||
<specimenCode id="F4089E8AFFC3D43AFE9FFD9EFFB0FDD7" box="[323,447,590,617]" collectionCode="IUQ" country="Colombia" httpUri="http://grbio.org/cool/8xqv-ibyz" name="Laboratorio de Ictiologia" pageId="6" pageNumber="7">IUQ 2264</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="14C63CACFFC3D43AFE0EFD9EFC6DFD35" ID-GBIF-Occurrence="1806442517" collectionCode="IUQ" country="Colombia" elevation="41" latitude="-1.6508334" location="Narino" longLatPrecision="21" longitude="-78.17583" pageId="6" pageNumber="7" specimenCode="IUQ 701" specimenCount="9" specimenCount-female="5" specimenCount-male="4" stateProvince="Narino" typeStatus="holotype">
|
||
<specimenCount id="B2A8FD78FFC3D43AFE0EFD9EFC1AFDD7" box="[466,533,590,617]" pageId="6" pageNumber="7" type="male">4 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="B2A8FD78FFC3D43AFDFDFD9EFC6DFDD7" box="[545,610,590,617]" pageId="6" pageNumber="7" type="female">5 ♀♀</specimenCount>
|
||
,
|
||
<quantity id="63569B14FFC3D43AFDB3FD9EFD16FDD7" box="[623,793,590,617]" metricMagnitude="-2" metricUnit="m" metricValue="6.77" metricValueMax="8.98" metricValueMin="4.56" pageId="6" pageNumber="7" unit="mm" value="67.7" valueMax="89.8" valueMin="45.6">45.6–89.8 mm</quantity>
|
||
SL, 4C&S (sex unknown), 52.8–63.0 mm SL, same locality as for
|
||
<typeStatus id="7B158853FFC3D43AFEB3FDA1FFD9FD35" box="[367,470,625,651]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
|
||
(
|
||
<specimenCode id="F4089E8AFFC3D43AFE34FDA1FC57FD35" box="[488,600,625,651]" collectionCode="IUQ" country="Colombia" httpUri="http://grbio.org/cool/8xqv-ibyz" name="Laboratorio de Ictiologia" pageId="6" pageNumber="7">IUQ 701</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="14C63CACFFC3D43AFDADFDA1FD48FD11" ID-GBIF-Occurrence="1806442540" collectionCode="IUQ" location="brook on La Hacienda Ranch" pageId="6" pageNumber="7" specimenCode="IUQ 2252" specimenCount="47" specimenCount-female="11" specimenCount-male="36">
|
||
<specimenCount id="B2A8FD78FFC3D43AFDADFDA1FCC8FD32" box="[625,711,625,652]" pageId="6" pageNumber="7" type="male">36 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="B2A8FD78FFC3D43AFD0AFDA1FD10FD35" box="[726,799,625,651]" pageId="6" pageNumber="7" type="female">11♀♀</specimenCount>
|
||
,
|
||
<quantity id="63569B14FFC3D43AFCEDFDA1FDD1FD32" box="[817,990,625,652]" metricMagnitude="-2" metricUnit="m" metricValue="5.295" metricValueMax="6.27" metricValueMin="4.32" pageId="6" pageNumber="7" unit="mm" value="52.95" valueMax="62.7" valueMin="43.2">43.2–62.7 mm</quantity>
|
||
SL, 4C&S (sex unknown), 39.2– 68.0 mm SL,
|
||
<location id="A171602AFFC3D43AFE87FD44FCA0FD10" LSID="urn:lsid:plazi:treatment:2C0787E7FFC0D435FD5CFDF3FF50FEC4:A171602AFFC3D43AFE87FD44FCA0FD10" box="[347,687,660,687]" name="brook on La Hacienda Ranch" pageId="6" pageNumber="7">brook on La Hacienda Ranch</location>
|
||
(
|
||
<specimenCode id="F4089E8AFFC3D43AFD61FD45FD35FD11" box="[701,826,660,687]" pageId="6" pageNumber="7">IUQ 2252</specimenCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="14C63CACFFC3D43AFC91FD45FDDFFD6C" ID-GBIF-Occurrence="1806442454" collectionCode="IUQ" collectorName="Barbacoas" country="Colombia" elevation="43" latitude="1.6608334" location="Yamunde River" longLatPrecision="21" longitude="-78.151115" pageId="6" pageNumber="7" specimenCode="IUQ 2259" specimenCount="19" specimenCount-female="8" specimenCount-male="11" stateProvince="Narino" typeStatus="holotype">
|
||
<specimenCount id="B2A8FD78FFC3D43AFC91FD45FD92FD11" box="[845,925,661,687]" pageId="6" pageNumber="7" type="male">11 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="B2A8FD78FFC3D43AFC75FD45FDE5FD11" box="[937,1002,661,687]" pageId="6" pageNumber="7" type="female">8 ♀♀</specimenCount>
|
||
,
|
||
<quantity id="63569B14FFC3D43AFC24FD44FAB0FD11" box="[1016,1215,660,687]" metricMagnitude="-2" metricUnit="m" metricValue="4.462000000000001" metricValueMax="5.063000000000001" metricValueMin="3.8609999999999998" pageId="6" pageNumber="7" unit="mm" value="44.62" valueMax="50.63" valueMin="38.61">38.61–50.63 mm</quantity>
|
||
SL,
|
||
<collectorName id="095B5327FFC3D43AFB29FD44FB7DFD10" box="[1269,1394,660,686]" pageId="6" pageNumber="7">Barbacoas</collectorName>
|
||
,
|
||
<location id="A171602AFFC3D43AFF61FD68FF79FD6C" LSID="urn:lsid:plazi:treatment:2C0787E7FFC0D435FD5CFDF3FF50FEC4:A171602AFFC3D43AFF61FD68FF79FD6C" box="[189,374,696,722]" country="Colombia" latitude="1.6608334" longLatPrecision="21" longitude="-78.151115" name="Yamunde River" pageId="6" pageNumber="7" stateProvince="Narino">Yamunde River</location>
|
||
,
|
||
<geoCoordinate id="C19A5036FFC3D43AFE5DFD66FC19FD6C" box="[385,534,694,722]" direction="north" orientation="latitude" pageId="6" pageNumber="7" precision="15" value="1.6608334">01°39′39″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="C19A5036FFC3D43AFDFDFD66FCB5FD6C" box="[545,698,694,722]" direction="west" orientation="longitude" pageId="6" pageNumber="7" precision="15" value="-78.151115">78°09′04″ W</geoCoordinate>
|
||
,
|
||
<elevation id="2F83D1C2FFC3D43AFD19FD68FD31FD6C" box="[709,830,696,722]" metricMagnitude="1" metricUnit="m" metricValue="4.3" pageId="6" pageNumber="7" unit="m" value="43.0">
|
||
<quantity id="63569B14FFC3D43AFD19FD68FD0FFD6C" box="[709,768,696,722]" metricMagnitude="1" metricUnit="m" metricValue="4.3" pageId="6" pageNumber="7" unit="m" value="43.0">43 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
(
|
||
<specimenCode id="F4089E8AFFC3D43AFC90FD68FDC6FD6C" box="[844,969,695,722]" collectionCode="IUQ" country="Colombia" httpUri="http://grbio.org/cool/8xqv-ibyz" name="Laboratorio de Ictiologia" pageId="6" pageNumber="7">IUQ 2259</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="14C63CACFFC3D43AFC00FD68FFFBFD4B" ID-GBIF-Occurrence="1806442452" collectionCode="IUQ" collectorName="Barbacoas" country="Colombia" elevation="43" latitude="1.6608334" location="Yamunde River" longLatPrecision="21" longitude="-78.151115" pageId="6" pageNumber="7" specimenCode="IUQ 989" specimenCount="3" specimenCount-male="3" stateProvince="Narino" typeStatus="holotype">
|
||
<specimenCount id="B2A8FD78FFC3D43AFC00FD68FA12FD6C" box="[988,1053,696,722]" pageId="6" pageNumber="7" type="male">3 ♂♂</specimenCount>
|
||
,
|
||
<quantity id="63569B14FFC3D43AFBF0FD67FAD9FD6C" box="[1068,1238,695,722]" metricMagnitude="-2" metricUnit="m" metricValue="6.904999999999999" metricValueMax="8.55" metricValueMin="5.26" pageId="6" pageNumber="7" unit="mm" value="69.05" valueMax="85.5" valueMin="52.6">52.6–85.5 mm</quantity>
|
||
SL, collected with holotype (
|
||
<specimenCode id="F4089E8AFFC3D43AFEA4FD0BFFE3FD4B" box="[376,492,731,757]" collectionCode="IUQ" country="Colombia" httpUri="http://grbio.org/cool/8xqv-ibyz" name="Laboratorio de Ictiologia" pageId="6" pageNumber="7">IUQ 989</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="14C63CACFFC3D43AFDD9FD0BFF4DFCA7" ID-GBIF-Occurrence="1806442453" collectionCode="IUQ" collectorName="Barbacoas" country="Colombia" elevation="43" latitude="1.6608334" location="Yamunde River" longLatPrecision="21" longitude="-78.151115" pageId="6" pageNumber="7" specimenCode="IUQ1165" specimenCount="1" stateProvince="Narino" typeStatus="holotype">
|
||
<specimenCount id="B2A8FD78FFC3D43AFDD9FD0BFC80FD4B" box="[517,655,731,757]" pageId="6" pageNumber="7" type="generic">1 specimen</specimenCount>
|
||
(sex unknown) C&S,
|
||
<quantity id="63569B14FFC3D43AFC76FD0BFA18FD4B" box="[938,1047,731,758]" metricMagnitude="-2" metricUnit="m" metricValue="5.38" pageId="6" pageNumber="7" unit="mm" value="53.8">53.8 mm</quantity>
|
||
SL, collected with holotype (
|
||
<specimenCode id="F4089E8AFFC3D43AFF19FD2EFF35FCA7" box="[197,314,766,793]" collectionCode="IUQ" country="Colombia" httpUri="http://grbio.org/cool/8xqv-ibyz" name="Laboratorio de Ictiologia" pageId="6" pageNumber="7">IUQ1165</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="F0D16679FFC3D43AFF61F87CFA22F854" httpUri="https://zenodo.org/record/1169565/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1964,1990]" targetBox="[462,1020,848,1930]" targetPageId="6">
|
||
<paragraph id="A41136F1FFC3D43AFF61F87CFA22F854" blockId="6.[189,1399,1964,2026]" pageId="6" pageNumber="7">
|
||
<emphasis id="96DAEAE3FFC3D43AFF61F87CFF04F878" bold="true" box="[189,267,1964,1990]" pageId="6" pageNumber="7">Fig. 3.</emphasis>
|
||
Shape of caudal-peduncle spot in species of the
|
||
<taxonomicName id="63AE4D72FFC3D43AFC93F87DFA24F878" authorityName="Eigenmann in Eigenmann & Ogle" authorityYear="1907" box="[847,1067,1964,1991]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="orthodus">
|
||
<emphasis id="96DAEAE3FFC3D43AFC93F87DFA24F878" box="[847,1067,1964,1991]" italics="true" pageId="6" pageNumber="7">Astyanax orthodus</emphasis>
|
||
</taxonomicName>
|
||
species-group.
|
||
<emphasis id="96DAEAE3FFC3D43AFB3AF87CFAF4F878" bold="true" box="[1254,1275,1964,1990]" pageId="6" pageNumber="7">A</emphasis>
|
||
.
|
||
<taxonomicName id="63AE4D72FFC3D43AFAD7F87DFFF3F854" authority="Eigenmann, 1907" authorityName="Eigenmann in Eigenmann & Ogle" authorityYear="1907" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="orthodus">
|
||
<emphasis id="96DAEAE3FFC3D43AFAD7F87DFF2AF854" italics="true" pageId="6" pageNumber="7">Astyanax orthodus</emphasis>
|
||
Eigenmann, 1907
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="96DAEAE3FFC3D43AFDD6F800FC11F854" bold="true" box="[522,542,2000,2026]" pageId="6" pageNumber="7">B</emphasis>
|
||
.
|
||
<taxonomicName id="63AE4D72FFC3D43AFDF0F800FCF8F854" authority="Ruiz-C & Román-Valencia & Taphorn & Buckup & Ortega, 2018" authorityName="Ruiz-C & Román-Valencia & Taphorn & Buckup & Ortega" authorityYear="2018" box="[556,759,2000,2026]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="embera" status="sp. nov.">
|
||
<emphasis id="96DAEAE3FFC3D43AFDF0F800FCF8F854" box="[556,759,2000,2026]" italics="true" pageId="6" pageNumber="7">Astyanax embera</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8DE95798FFC3D43AFD23F800FD56F854" box="[767,857,2000,2026]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Scale bar = 1 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="ECB4657AFFC2D434FF61FEDBFB45FE41" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
|
||
<paragraph id="A41136F1FFC2D43BFF61FEDBFF41FE9B" blockId="7.[189,334,267,293]" box="[189,334,267,293]" pageId="7" pageNumber="8">
|
||
<heading id="FF59819DFFC2D43BFF61FEDBFF41FE9B" bold="true" box="[189,334,267,293]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
|
||
<emphasis id="96DAEAE3FFC2D43BFF61FEDBFF41FE9B" bold="true" box="[189,334,267,293]" pageId="7" pageNumber="8">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="A41136F1FFC2D43BFF61FEEAFC10FDB4" blockId="7.[189,1399,314,522]" pageId="7" pageNumber="8">Body compressed, greatest body depth at or anterior to dorsal-fin origin. Mouth terminal. Dorsal profile of head straight between snout tip to posterior margin of supraoccipital spine, convex between head and dorsal fin, convex between last dorsal-fin ray and adipose-fin origin. Caudal peduncle arched, with dorsal profile concave and ventral profile convex. Ventral profile convex between snout tip and posterior end of anal-fin base in males; in females convex to pelvic-fin insertions, concave beyond the insertion of pelvic fin to anal-fin origin.</paragraph>
|
||
<paragraph id="A41136F1FFC2D43BFF61FDE2FD9FFD63" blockId="7.[189,1399,562,734]" pageId="7" pageNumber="8">Premaxillary teeth in two series; outer series with four tricuspid teeth covering three most medial teeth of internal series; inner row with five pentacuspid teeth. Maxilla long, of same width along entire length, with two tricuspid teeth set in anterior-most part of ventral margin. Dentary with anterior four teeth pentacuspid, following ten teeth smaller, progressively inclined posteromedially, varying from tri- to unicuspid; proportion of tri- vs unicuspid teeth quite variable.</paragraph>
|
||
<paragraph id="A41136F1FFC2D43BFF61FCD6FCEAFB0E" blockId="7.[189,1399,774,1200]" pageId="7" pageNumber="8">
|
||
Pored lateral-line scales 36(1), 37(11), 38(11), 39(13), 40(1), 41(1) (n = 38); transverse scales from lateral line to dorsal-fin origin 7(13), 8(25) (n = 38); scales from lateral line to anal-fin origin 7(1), 8(29), 9(8) (n = 38); scales from lateral line to pelvic-fin insertion 6(22), 7(15), 8(1) (n = 38). Predorsal midline covered with bilobed medial scales for more than three quarters of its length, naked anteriorly. Dorsalfin rays
|
||
<date id="D0101031FFC2D43BFEC0FC47FF44FC0F" box="[284,331,919,945]" pageId="7" pageNumber="8">iii 9</date>
|
||
(n = 38); first simple ray reduced in size, easily visible only in C&S specimens, detectable with dissecting needle in non-C&S specimens; second simple ray about half length of third simple ray. Distal margin of dorsal fin slightly convex. Adipose-fin origin anterior to vertical through insertion of last anal-fin ray. Pectoral-fin rays
|
||
<date id="D0101031FFC2D43BFD9FFBD4FC61FBA0" box="[579,622,1028,1054]" pageId="7" pageNumber="8" value="2010-01">i 10</date>
|
||
i (2),
|
||
<date id="D0101031FFC2D43BFD6FFBD4FCD2FBA0" box="[691,733,1028,1054]" pageId="7" pageNumber="8" value="2011-01">i 11</date>
|
||
i (23),
|
||
<date id="D0101031FFC2D43BFCECFBD4FD53FBA0" box="[816,860,1028,1054]" pageId="7" pageNumber="8" value="2012-01">i 12</date>
|
||
i (12),
|
||
<date id="D0101031FFC2D43BFC73FBD4FDD5FBA0" box="[943,986,1028,1054]" pageId="7" pageNumber="8" value="2013-01">i 13</date>
|
||
(1) (n = 38). Anal-fin rays
|
||
<date id="D0101031FFC2D43BFACDFBD4FB42FBA1" box="[1297,1357,1028,1055]" pageId="7" pageNumber="8" value="1925-03">iii 25</date>
|
||
(7),
|
||
<date id="D0101031FFC2D43BFF61FBF8FEF5FBFC" box="[189,250,1064,1090]" pageId="7" pageNumber="8" value="1926-03">iii 26</date>
|
||
(9),
|
||
<date id="D0101031FFC2D43BFEF0FBF8FF66FBFC" box="[300,361,1064,1090]" pageId="7" pageNumber="8" value="1927-03">iii 27</date>
|
||
(12),
|
||
<date id="D0101031FFC2D43BFE76FBF8FFE8FBFC" box="[426,487,1064,1090]" pageId="7" pageNumber="8" value="1928-03">iii 28</date>
|
||
(9),
|
||
<date id="D0101031FFC2D43BFDC4FBF8FC59FBFC" box="[536,598,1064,1090]" pageId="7" pageNumber="8" value="1929-03">iii 29</date>
|
||
(1) (n = 38); first simple rays only visible in C&S material. Anal-fin origin posterior to vertical through insertion of last dorsal-fin ray (
|
||
<tableCitation id="E92C034AFFC2D43BFC03FB9CFA37FBD9" box="[991,1080,1100,1127]" captionStart="Table 1" captionStartId="11.[145,214,599,625]" captionTargetPageId="11" captionText="Table 1. Morphometric data for some members of the Astyanax orthodus species-group." httpUri="http://table.plazi.org/id/F0D16679FFCED437FF4DFD87FA92FDCF" pageId="7" pageNumber="8" tableUuid="F0D16679FFCED437FF4DFD87FA92FDCF">Table 1</tableCitation>
|
||
). Principal caudal-fin rays 10(8) associated with four dorsal hypurals, 9(8) associated with three ventral hypurals; dorsal procurrent rays 10–11(8); ventral procurrent rays 9–10(8).
|
||
</paragraph>
|
||
<paragraph id="A41136F1FFC2D43BFF61FB07FDDDFA3D" blockId="7.[189,1399,1239,1411]" pageId="7" pageNumber="8">Total vertebrae 34(8), including those of the Weberian apparatus: precaudal centra 16(2)–17(6), last two without true ribs; caudal centra 17(2)–18(6). Epipleurals 21(2)–22(6). Epineurals 30(2)–31(6); posterior-most epineural may reach anterior surface of urostyle. Hypurals 7(8); first dorsal hypural with anterior margin dotted, without projections without extensions that articulate the urostyle; second and third hypural with anterior margin swollen that contacting urostyle.</paragraph>
|
||
<paragraph id="A41136F1FFC2D43BFF61FA7BFFE9FA7B" blockId="7.[189,486,1451,1477]" box="[189,486,1451,1477]" pageId="7" pageNumber="8">
|
||
<heading id="FF59819DFFC2D43BFF61FA7BFFE9FA7B" bold="true" box="[189,486,1451,1477]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
|
||
<emphasis id="96DAEAE3FFC2D43BFF61FA7BFFE9FA7B" bold="true" box="[189,486,1451,1477]" pageId="7" pageNumber="8">Pigmentation in alcohol</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="A41136F1FFC2D43BFF61FA0BFA90F938" blockId="7.[189,1399,1499,1670]" pageId="7" pageNumber="8">Sides of body yellowish, with reticulated pattern predominant over dorsal region of coelomic cavity. Silvery stripe present from humeral region to caudal-peduncle base, overlain by series of chevronshaped marks formed by dark lines along myosepta between myotomes extending from dorsal region of coelomic cavity to caudal peduncle; pigmented muscle septae forming chevrons not coinciding with scale rows. Chevrons without distal extensions both in juvenile and adult specimens.</paragraph>
|
||
<paragraph id="A41136F1FFC2D434FF61F97EFA2BFEF6" blockId="7.[189,1399,1710,2027]" lastBlockId="8.[189,1399,267,329]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
|
||
Dorsal region of head and body chestnut brown. Sides of cranium and ventral surface of body light brown, not silvery. The pigments of the humeral region form an anterior and posterior humeral spot. The anterior spot is formed by two layers of pigment: brown melanophores distributed in thin superficial layer of the epithelium (Layer 1,
|
||
<figureCitation id="3C952A74FFC2D43BFD9BF8CBFC87F888" box="[583,648,1819,1846]" captionStart="Fig.1" captionStartId="4.[189,232,1859,1885]" captionTargetBox="[189,1398,540,1837]" captionTargetId="figure@4.[189,1398,540,1837]" captionTargetPageId="4" captionText="Fig.1.A.DistributionofpigmentinthehumeralregionofAstyanaxspecies.B.A.bimaculatus(Linnaeus, 1758).C.A.caucanus(Steindachner,1879).D.A.mexicanus(DeFilippi,1853).E.A.courensisBertaco, Carvalho & Jerep, 2010 (modified from Bertaco et al. 2010). The anterior humeral spot consists of two overlapping layers of pigment identified by numbers 1 and 2 (see Román-Valencia et al. 2015: fig. 1): 1. Diffuse vertical bar. 2. Horizontal spot with concentrated pigments." httpUri="https://zenodo.org/record/1169561/files/figure.png" pageId="7" pageNumber="8">Fig.1</figureCitation>
|
||
); deeper layer with dark melanophores (Layer 2,
|
||
<figureCitation id="3C952A74FFC2D43BFB0FF8CBFB17F88B" box="[1235,1304,1819,1845]" captionStart="Fig.1" captionStartId="4.[189,232,1859,1885]" captionTargetBox="[189,1398,540,1837]" captionTargetId="figure@4.[189,1398,540,1837]" captionTargetPageId="4" captionText="Fig.1.A.DistributionofpigmentinthehumeralregionofAstyanaxspecies.B.A.bimaculatus(Linnaeus, 1758).C.A.caucanus(Steindachner,1879).D.A.mexicanus(DeFilippi,1853).E.A.courensisBertaco, Carvalho & Jerep, 2010 (modified from Bertaco et al. 2010). The anterior humeral spot consists of two overlapping layers of pigment identified by numbers 1 and 2 (see Román-Valencia et al. 2015: fig. 1): 1. Diffuse vertical bar. 2. Horizontal spot with concentrated pigments." httpUri="https://zenodo.org/record/1169561/files/figure.png" pageId="7" pageNumber="8">Fig. 1</figureCitation>
|
||
). Layer 2 forming polygon-shaped spot, with two parts, forming vertices on margins, usually with four sides, located from third to sixth or seventh scale of lateral series. Posterior humeral spot situated two or three scales posterior to anterior humeral spot, arc- or sigmoid-shaped, inconspicuous, covering two to three scales above lateral-line. Scales on sides of body with spots or dots. Caudal peduncle spot pentagonal, its posterior tip or margin centered on middle caudal-fin rays, usually not further prolonged on middle caudal-fin rays. Pectoral and pelvic fins mostly hyaline; pelvic fins with melanophores at tips of rays. Dorsal, caudal and anal fins with melanophores on interradial membranes.
|
||
</paragraph>
|
||
<paragraph id="A41136F1FFCDD434FF61FEA0FFBFFE34" blockId="8.[189,432,368,394]" box="[189,432,368,394]" pageId="8" pageNumber="9">
|
||
<heading id="FF59819DFFCDD434FF61FEA0FFBFFE34" bold="true" box="[189,432,368,394]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
|
||
<emphasis id="96DAEAE3FFCDD434FF61FEA0FFBFFE34" bold="true" box="[189,432,368,394]" pageId="8" pageNumber="9">Sexual dimorphism</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="A41136F1FFCDD434FF61FE4EFB45FE41" blockId="8.[189,1399,414,511]" pageId="8" pageNumber="9">Sexual dimorphism was observed in the ventral profile, which is convex from snout tip to posterior end of anal-fin base in males, but in females is convex from snout tip to the pelvic-fin insertions, then concave to the origin of the anal fin. No hooks were detected on fins of examined adult individuals.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="ECB4657AFFCDD434FF61FDF6FC5DFD2D" pageId="8" pageNumber="9" type="distribution">
|
||
<paragraph id="A41136F1FFCDD434FF61FDF6FF59FDFE" blockId="8.[189,342,550,576]" box="[189,342,550,576]" pageId="8" pageNumber="9">
|
||
<heading id="FF59819DFFCDD434FF61FDF6FF59FDFE" bold="true" box="[189,342,550,576]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
|
||
<emphasis id="96DAEAE3FFCDD434FF61FDF6FF59FDFE" bold="true" box="[189,342,550,576]" pageId="8" pageNumber="9">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="A41136F1FFCDD434FF61FD85FC5DFD2D" blockId="8.[189,1398,597,659]" pageId="8" pageNumber="9">
|
||
Middle portion of the Telembí River drainage, which is a tributary of the Patía River, Nariño Department, Pacific coast of Colombia (
|
||
<figureCitation id="3C952A74FFCDD434FE20FDA8FC4EFD2C" box="[508,577,632,658]" captionStart="Fig. 4" captionStartId="8.[189,232,2000,2026]" captionTargetBox="[189,1392,764,1921]" captionTargetId="figure@8.[151,1417,755,1932]" captionTargetPageId="8" captionText="Fig. 4.Distribution of speciesof the Astyanaxorthodus species-group(symbol includestypelocalities)." httpUri="https://zenodo.org/record/1169567/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="F0D16679FFCDD434FF61F800FB78F855" box="[189,1399,2000,2027]" httpUri="https://zenodo.org/record/1169567/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,232,2000,2026]" targetBox="[189,1392,764,1921]" targetPageId="8">
|
||
<paragraph id="A41136F1FFCDD434FF61F800FB78F855" blockId="8.[189,1399,2000,2027]" box="[189,1399,2000,2027]" pageId="8" pageNumber="9">
|
||
<emphasis id="96DAEAE3FFCDD434FF61F800FF05F854" bold="true" box="[189,266,2000,2026]" pageId="8" pageNumber="9">Fig. 4.</emphasis>
|
||
Distribution of species of the
|
||
<taxonomicName id="63AE4D72FFCDD434FDB5F801FD4CF854" authorityName="Eigenmann in Eigenmann & Ogle" authorityYear="1907" box="[617,835,2000,2027]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="orthodus">
|
||
<emphasis id="96DAEAE3FFCDD434FDB5F801FD4CF854" box="[617,835,2000,2027]" italics="true" pageId="8" pageNumber="9">Astyanax orthodus</emphasis>
|
||
</taxonomicName>
|
||
species-group (symbol includes type localities).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="ECB4657AFFCCD435FF61FEDBFF50FEC4" pageId="9" pageNumber="10" type="discussion">
|
||
<paragraph id="A41136F1FFCCD435FF61FEDBFF4BFE9B" blockId="9.[189,324,267,293]" box="[189,324,267,293]" pageId="9" pageNumber="10">
|
||
<heading id="FF59819DFFCCD435FF61FEDBFF4BFE9B" bold="true" box="[189,324,267,293]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
|
||
<emphasis id="96DAEAE3FFCCD435FF61FEDBFF4BFE9B" bold="true" box="[189,324,267,293]" pageId="9" pageNumber="10">Comments</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="A41136F1FFCCD435FF61FEEBFF50FEC4" blockId="9.[189,1398,315,378]" pageId="9" pageNumber="10">
|
||
The stream where the
|
||
<typeStatus id="7B158853FFCCD435FE1DFEECFFFBFEE8" box="[449,500,316,342]" pageId="9" pageNumber="10">type</typeStatus>
|
||
locality of this species is located is subject to mining activity for gold, silver and platinum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |