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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.430.8094" ID-PMC="PMC4141162" ID-Pensoft-Pub="1313-2970-430-1" ID-Pensoft-UUID="FFE6FF8CFFD43B66335AD2203774DB49" ID-PubMed="25152663" ID-Zenodo-Dep="578441" ID-ZooBank="86F760563B8B49FB9C86FAD0DB0CBE8C" ModsDocID="1313-2970-430-1" checkinTime="1451245551152" checkinUser="pensoft" docAuthor="Erwin, Terry L. &amp; Zamorano, Laura S." docDate="2014" docId="1A81B99166B5C70D3F1DCF2DDBEDD5F2" docLanguage="en" docName="ZooKeys 430: 1-108" docOrigin="ZooKeys 430" docPubDate="2014-08-01" docSource="http://dx.doi.org/10.3897/zookeys.430.8094" docTitle="Asklepia adisi Erwin &amp; Zamorano 2014, sp. n." docType="treatment" docUuid="01A360AD-0E9E-4626-B2FC-D90DB089F19F" docUuidSource="ZooBank" docVersion="5" id="FFE6FF8CFFD43B66335AD2203774DB49" lastPageNumber="47" masterDocId="FFE6FF8CFFD43B66335AD2203774DB49" masterDocTitle="A synopsis of the tribe Lachnophorini, with a new genus of Neotropical distribution and a revision of the Neotropical genus Asklepia Liebke, 1938 (Insecta, Coleoptera, Carabidae)" masterLastPageNumber="108" masterPageNumber="1" pageNumber="46" updateTime="1668159067277" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A synopsis of the tribe Lachnophorini, with a new genus of Neotropical distribution and a revision of the Neotropical genus Asklepia Liebke, 1938 (Insecta, Coleoptera, Carabidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Erwin, Terry L.</mods:namePart>
<mods:affiliation>Hyper-diversity Group, Department of Entomology, MRC- 187, National Museum of Natural History, Smithsonian Institution, Washington, P. O. Box 37012, DC 20013 - 7012, USA</mods:affiliation>
<mods:nameIdentifier type="email">erwint@si.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zamorano, Laura S.</mods:namePart>
<mods:affiliation>Research Student, MRC- 187, National Museum of Natural History, Smithsonian Institution, Washington, P. O. Box 37012, DC 20013 - 7012, USA &amp; Laboratorio de Zoologia Acuatica LAZOEA, Departamento de Ciencias Biologicas, Universidad de los Andes, Colombia</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="pubDate">
<mods:number>2014-08-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>430</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>108</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.430.8094</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.430.8094</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-430-1</mods:identifier>
<mods:identifier type="ZooBank">86F760563B8B49FB9C86FAD0DB0CBE8C</mods:identifier>
<mods:identifier type="Pensoft-UUID">FFE6FF8CFFD43B66335AD2203774DB49</mods:identifier>
<mods:identifier type="Zenodo-Dep">578441</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152054778" LSID="urn:lsid:zoobank.org:act:01A360AD-0E9E-4626-B2FC-D90DB089F19F" httpUri="http://treatment.plazi.org/id/1A81B99166B5C70D3F1DCF2DDBEDD5F2" lastPageId="47" lastPageNumber="47" pageId="45" pageNumber="46">
<subSubSection pageId="45" pageNumber="46" type="nomenclature">
<paragraph pageId="45" pageNumber="46">
<taxonomicName LSID="http://zoobank.org/01A360AD-0E9E-4626-B2FC-D90DB089F19F" authority="Erwin &amp; Zamorano" authorityName="Erwin &amp; Zamorano" authorityYear="2014" class="Insecta" family="Carabidae" genus="Asklepia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Asklepia adisi" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="adisi" status="sp. n.">Asklepia adisi Erwin &amp; Zamorano</taxonomicName>
<taxonomicNameLabel pageId="45" pageNumber="46">sp. n.</taxonomicNameLabel>
<normalizedToken originalValue="Adis">Adis'</normalizedToken>
pattern-wing beetle
<figureCitation captionStart="Figure 3740" captionStartId="F11" captionText="Figure 37 - 40. Digital Photo-illustrations, habitus, dorsal aspect of holotypes. 37 Asklepia matomena Zamorano &amp; Erwin, sp. n. ADP 132519, 20 km SW Manaus, Brazil 38 Asklepia adisi Erwin &amp; Zamorano, sp. n. Adis 000691, Ilha de Marchantaria, Lago Camaleao, Brazil 39 Asklepia asuncionensis Erwin &amp; Zamorano, sp. n. ADP 130036, Asuncion, Rio Paraguay, Paraguay 40 Asklepia biolat Erwin &amp; Zamorano, sp. n. ADP 132509, Pakitza, Peru. Scale line = 1 mm." httpUri="https://binary.pensoft.net/fig/30641" pageId="45" pageNumber="46">Figs 38</figureCitation>
<figureCitation captionStart="Figure 6368" captionStartId="F18" captionText="Figure 63 - 68. Illustrations, male aedeagus, dorsal, ventral, left lateral aspects. 63 Asklepia matomena Zamorano &amp; Erwin, sp. n. ADP 132527, 20 km SW Manaus, Brazil 64 Asklepia adisi Erwin &amp; Zamorano, sp. n. Adis 001335, Ilha de Marchantaria, Lago Camaleao, Brazil 65 Asklepia biolat Erwin &amp; Zamorano, sp. n. ADP 132480, Pakitza, Peru 66 Asklepia bracheia Zamorano &amp; Erwin, sp. n. ADP 067304, Boca del Rio Samiria Peru 67 Asklepia ecuadoriana Erwin &amp; Zamorano, sp. n. ADP 132468, Limoncocha, Ecuador. Endophallus not everted 68 Asklepia ecuadoriana Erwin &amp; Zamorano, sp. n. ADP 132468, Limoncocha, Ecuador. Endophallus everted. Scale line = 0.25 mm." httpUri="https://binary.pensoft.net/fig/30648" pageId="45" pageNumber="46">, 64</figureCitation>
<figureCitation captionStart="Figure 78" captionStartId="F22" captionText="Figure 78. Distribution map for known localities of Asklepia species of the pulchripennis group." httpUri="https://binary.pensoft.net/fig/30652" pageId="45" pageNumber="46">, 78</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="45" pageNumber="46" type="holotype">
<paragraph pageId="45" pageNumber="46">Holotype.</paragraph>
<paragraph pageId="45" pageNumber="46">
<emphasis bold="true" pageId="45" pageNumber="46">Brazil</emphasis>
, Amazonas, Rio
<normalizedToken originalValue="Solimões">Solimoes</normalizedToken>
, Ilha de Marchantaria, Lago
<normalizedToken originalValue="Camaleão">Camaleao</normalizedToken>
,
<geoCoordinate degrees="3.2488" direction="south" orientation="latitude" precision="5" value="-3.2488">3.2488°S</geoCoordinate>
,
<geoCoordinate degrees="59.9556" direction="west" orientation="longitude" precision="5" value="-59.9556">59.9556°W</geoCoordinate>
, 7m, 11 April 1981 (NMNH: ADIS # 000691, female).
</paragraph>
</subSubSection>
<subSubSection pageId="45" pageNumber="46" type="derivation of specific epithet">
<paragraph pageId="45" pageNumber="46">Derivation of specific epithet.</paragraph>
<paragraph pageId="45" pageNumber="46">
The specific epithet,
<emphasis italics="true" pageId="45" pageNumber="46">adisi</emphasis>
, is an eponym, masculine, genitive case, based on the family name of Joachim Adis
<emphasis bold="true" pageId="45" pageNumber="46"></emphasis>
who collected the type series.
</paragraph>
</subSubSection>
<subSubSection pageId="45" pageNumber="46" type="proposed english vernacular name">
<paragraph pageId="45" pageNumber="46">Proposed english vernacular name.</paragraph>
<paragraph pageId="45" pageNumber="46">
<normalizedToken originalValue="Adis">Adis'</normalizedToken>
pattern-wing beetle.
</paragraph>
</subSubSection>
<subSubSection pageId="45" pageNumber="46" type="diagnosis">
<paragraph pageId="45" pageNumber="46">Diagnosis.</paragraph>
<paragraph pageId="45" pageNumber="46">
With the attributes of the genus
<taxonomicName authorityName="Liebke" authorityYear="1938" class="Insecta" family="Carabidae" genus="Asklepia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="45" pageNumber="46">Asklepia</emphasis>
</taxonomicName>
as described by
<bibRefCitation author="Liebke, M" journalOrPublisher="Embrik Strand" pageId="81" pageNumber="82" pagination="37 - 141" refId="B54" refString="Liebke, M, 1938. Denkschrift ueber die Carabiden-Tribus Colliurini. Festschrift zum 60. Geburtstage von Profesor Dr. Embrik Strand 4: 37 - 141" title="Denkschrift ueber die Carabiden-Tribus Colliurini. Festschrift zum 60. Geburtstage von Profesor Dr." volume="4" year="1938">Liebke (1938)</bibRefCitation>
and as noted above under the generic diagnosis, and medium-sized for the genus (SBL = 2.542-2.688 mm). Adults with head brunneus, prothorax aurantiacus, elytral maculae aurantiacus; elytron fuscous with a broad triangular aurantiacus macula entirely covering proximal basal quadrant and humeral area of lateral basal quadrant, medial quadrants largely aurantiacus, sutural area fuscous, lateral margin, apical margin and sutural area aurantiacus; metasternum fulvous, abdominal sterna with III-VI, and epipleuron fulvous, abdominal sternum VII fuscous; legs flavotestaceous; antennal scape, pedicel, antennomere 3 and basal half of 4 testaceous, apical half of antennomere 4, 5-6 deeply infuscated, 7-11 white. Dorsal surface devoid of microsculpture, surface luster very shiny. Pronotum markedly convex with lateral margin effaced except just anterior to hind angle and there a simple bead; hind angle moderately prominent; median line feebly defined. Elytral interneurs evident as short discontinuous rows of widely spaced coarse punctures.
</paragraph>
</subSubSection>
<subSubSection lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="description">
<paragraph pageId="45" pageNumber="46">Description.</paragraph>
<paragraph lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
(
<figureCitation captionStart="Figure 3740" captionStartId="F11" captionText="Figure 37 - 40. Digital Photo-illustrations, habitus, dorsal aspect of holotypes. 37 Asklepia matomena Zamorano &amp; Erwin, sp. n. ADP 132519, 20 km SW Manaus, Brazil 38 Asklepia adisi Erwin &amp; Zamorano, sp. n. Adis 000691, Ilha de Marchantaria, Lago Camaleao, Brazil 39 Asklepia asuncionensis Erwin &amp; Zamorano, sp. n. ADP 130036, Asuncion, Rio Paraguay, Paraguay 40 Asklepia biolat Erwin &amp; Zamorano, sp. n. ADP 132509, Pakitza, Peru. Scale line = 1 mm." httpUri="https://binary.pensoft.net/fig/30641" pageId="45" pageNumber="46">Fig. 38</figureCitation>
,
<figureCitation captionStart="Figure 6368" captionStartId="F18" captionText="Figure 63 - 68. Illustrations, male aedeagus, dorsal, ventral, left lateral aspects. 63 Asklepia matomena Zamorano &amp; Erwin, sp. n. ADP 132527, 20 km SW Manaus, Brazil 64 Asklepia adisi Erwin &amp; Zamorano, sp. n. Adis 001335, Ilha de Marchantaria, Lago Camaleao, Brazil 65 Asklepia biolat Erwin &amp; Zamorano, sp. n. ADP 132480, Pakitza, Peru 66 Asklepia bracheia Zamorano &amp; Erwin, sp. n. ADP 067304, Boca del Rio Samiria Peru 67 Asklepia ecuadoriana Erwin &amp; Zamorano, sp. n. ADP 132468, Limoncocha, Ecuador. Endophallus not everted 68 Asklepia ecuadoriana Erwin &amp; Zamorano, sp. n. ADP 132468, Limoncocha, Ecuador. Endophallus everted. Scale line = 0.25 mm." httpUri="https://binary.pensoft.net/fig/30648" pageId="45" pageNumber="46">64</figureCitation>
).
<emphasis bold="true" pageId="46" pageNumber="47">
<emphasis italics="true" pageId="46" pageNumber="47">
<pageBreakToken pageId="46" pageNumber="47" start="start">Habitus</pageBreakToken>
</emphasis>
:
</emphasis>
(
<figureCitation captionStart="Figure 3740" captionStartId="F11" captionText="Figure 37 - 40. Digital Photo-illustrations, habitus, dorsal aspect of holotypes. 37 Asklepia matomena Zamorano &amp; Erwin, sp. n. ADP 132519, 20 km SW Manaus, Brazil 38 Asklepia adisi Erwin &amp; Zamorano, sp. n. Adis 000691, Ilha de Marchantaria, Lago Camaleao, Brazil 39 Asklepia asuncionensis Erwin &amp; Zamorano, sp. n. ADP 130036, Asuncion, Rio Paraguay, Paraguay 40 Asklepia biolat Erwin &amp; Zamorano, sp. n. ADP 132509, Pakitza, Peru. Scale line = 1 mm." httpUri="https://binary.pensoft.net/fig/30641" pageId="46" pageNumber="47">Fig. 38</figureCitation>
)
<emphasis bold="true" pageId="46" pageNumber="47">
<emphasis italics="true" pageId="46" pageNumber="47">Size</emphasis>
:
</emphasis>
[See also
<tableCitation captionStart="Table 10" captionStartId="T10" captionText="Table 10. Asklepia adisi Erwin &amp; Zamorano, sp. n." httpUri="http://table.plazi.org/id/4BD3E244343B91BBC7865309AABABC2F" pageId="46" pageNumber="47" tableUuid="4BD3E244343B91BBC7865309AABABC2F">Table 10</tableCitation>
] Medium-size for the genus; ABL = 2.582-3.199 mm, SBL = 2.542-2.688 mm, TW (total width) 1.341-1.473 mm, LP = 0.513-0.591 mm, WP = 0.660-0.717 mm, LE = 1.578-1.712 mm.
<emphasis bold="true" pageId="46" pageNumber="47">
<emphasis italics="true" pageId="46" pageNumber="47">Color</emphasis>
:
</emphasis>
See diagnosis above.
<emphasis bold="true" pageId="46" pageNumber="47">
<emphasis italics="true" pageId="46" pageNumber="47">Luster</emphasis>
:
</emphasis>
See diagnosis above.
<emphasis bold="true" italics="true" pageId="46" pageNumber="47">Head</emphasis>
(
<figureCitation captionStart="Figure 3740" captionStartId="F11" captionText="Figure 37 - 40. Digital Photo-illustrations, habitus, dorsal aspect of holotypes. 37 Asklepia matomena Zamorano &amp; Erwin, sp. n. ADP 132519, 20 km SW Manaus, Brazil 38 Asklepia adisi Erwin &amp; Zamorano, sp. n. Adis 000691, Ilha de Marchantaria, Lago Camaleao, Brazil 39 Asklepia asuncionensis Erwin &amp; Zamorano, sp. n. ADP 130036, Asuncion, Rio Paraguay, Paraguay 40 Asklepia biolat Erwin &amp; Zamorano, sp. n. ADP 132509, Pakitza, Peru. Scale line = 1 mm." httpUri="https://binary.pensoft.net/fig/30641" pageId="46" pageNumber="47">Fig. 38</figureCitation>
): as in description for genus above.
<emphasis bold="true" pageId="46" pageNumber="47">
<emphasis italics="true" pageId="46" pageNumber="47">Prothorax</emphasis>
.
</emphasis>
Pronotum (
<figureCitation captionStart="Figure 3740" captionStartId="F11" captionText="Figure 37 - 40. Digital Photo-illustrations, habitus, dorsal aspect of holotypes. 37 Asklepia matomena Zamorano &amp; Erwin, sp. n. ADP 132519, 20 km SW Manaus, Brazil 38 Asklepia adisi Erwin &amp; Zamorano, sp. n. Adis 000691, Ilha de Marchantaria, Lago Camaleao, Brazil 39 Asklepia asuncionensis Erwin &amp; Zamorano, sp. n. ADP 130036, Asuncion, Rio Paraguay, Paraguay 40 Asklepia biolat Erwin &amp; Zamorano, sp. n. ADP 132509, Pakitza, Peru. Scale line = 1 mm." httpUri="https://binary.pensoft.net/fig/30641" pageId="46" pageNumber="47">Fig. 38</figureCitation>
) moderately broad, slightly narrower than head across eyes (WH/WP, mean both sexes: 1.103), longer than head (LP/LH, mean both sexes: 1.327), wider than longer (WP/LP, mean both sexes: 1.226); markedly cordiform and convex, lateral margin effaced with seta at anterior third on slightly raised area; apex markedly constricted; anterior angle feebly produced, hind angle slightly produced and setose; median line feebly define as an infuscate line, transverse impressions punctate, punctures infuscate; surface smooth throughout.
<emphasis bold="true" pageId="46" pageNumber="47">
<emphasis italics="true" pageId="46" pageNumber="47">Pterothorax</emphasis>
.
</emphasis>
Normal for genus, see description for genus above. Elytra markedly convex; at apical third twice as wide as head across eyes (WH/TW, mean both sexes: 0.535) and pronotum (WP/TW, mean both sexes: 0.520), longer than wide. Elytral interneurs evident as short discontinuous rows of widely spaced coarse punctures. Hind wings fully developed.
<emphasis bold="true" pageId="46" pageNumber="47">
<emphasis italics="true" pageId="46" pageNumber="47">Legs</emphasis>
.
</emphasis>
Overall, normal for genus, see description for genus above.
<emphasis bold="true" pageId="46" pageNumber="47">
<emphasis italics="true" pageId="46" pageNumber="47">Abdominal sterna</emphasis>
.
</emphasis>
Overall, normal for genus, see description for genus above.
<emphasis bold="true" italics="true" pageId="46" pageNumber="47">Male genitalia</emphasis>
(
<figureCitation captionStart="Figure 6368" captionStartId="F18" captionText="Figure 63 - 68. Illustrations, male aedeagus, dorsal, ventral, left lateral aspects. 63 Asklepia matomena Zamorano &amp; Erwin, sp. n. ADP 132527, 20 km SW Manaus, Brazil 64 Asklepia adisi Erwin &amp; Zamorano, sp. n. Adis 001335, Ilha de Marchantaria, Lago Camaleao, Brazil 65 Asklepia biolat Erwin &amp; Zamorano, sp. n. ADP 132480, Pakitza, Peru 66 Asklepia bracheia Zamorano &amp; Erwin, sp. n. ADP 067304, Boca del Rio Samiria Peru 67 Asklepia ecuadoriana Erwin &amp; Zamorano, sp. n. ADP 132468, Limoncocha, Ecuador. Endophallus not everted 68 Asklepia ecuadoriana Erwin &amp; Zamorano, sp. n. ADP 132468, Limoncocha, Ecuador. Endophallus everted. Scale line = 0.25 mm." httpUri="https://binary.pensoft.net/fig/30648" pageId="46" pageNumber="47">Fig. 64</figureCitation>
, see
<figureCitation captionStart="Figure 5762" captionStartId="F17" captionText="Figure 57 - 62. Illustrations, male aedeagus, dorsal, ventral, left lateral aspects. 57 Asklepia geminata (Bates, 1871) ADP 109186, Rio Samiria, Boca Cano Ingles Camp, Peru 58 Asklepia campbellorum Zamorano &amp; Erwin, sp. n. ADP 109196, 20 km SW Manaus, Brazil 59 Asklepia demiti Erwin &amp; Zamorano, sp. n. ADP 132585, Rio Demiti, Brazil 60 Asklepia grammechrysea Zamorano &amp; Erwin, sp. n. ADP 052565, Rio Sucusari, Peru 61 Asklepia laetitia Zamorano &amp; Erwin, sp. n. ADP 109190, Leticia, Colombia. Legend: a apical area; bl basal lobe; bo basal orifice; lp left paramere; ml median lobe; om ostial membrane; oo ostial opening; rp right paramere; sh shaft; pb phallobase; ps phallobase shaft; ms medial spine; ds distal spine 62 Asklepia lebioides (Bates, 1871), comb. n., ADP 109208, Rio Demiti, Brazil. Scale line = 0.25 mm." httpUri="https://binary.pensoft.net/fig/30647" pageId="46" pageNumber="47">Fig. 61</figureCitation>
for attribute labels). Median lobe with phallobase moderate in length, about a fourth the length of shaft, basal opening large, oriented parallel to the central part of the shaft. Shaft broad, slightly twisted ventrally, dorsally sclerotized except for short ostium; in ventral aspect tapered toward rather broad apex, in lateral aspect, a thick rounded apex. Left paramere very large and broad, right small and triangular; apex of left paramere lobate much longer than right paramere, about two-thirds the length of shaft (measured in left lateral aspect). Endophallus with one median spine, and one large distal spine.
<emphasis bold="true" pageId="46" pageNumber="47">
<emphasis italics="true" pageId="46" pageNumber="47">Female genitalia</emphasis>
.
</emphasis>
Not investigated, presumably similar to that of
<taxonomicName authorityName="Erwin &amp; Zamorano" authorityYear="2014" class="Insecta" family="Carabidae" genus="Asklepia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="demiti">
<emphasis italics="true" pageId="46" pageNumber="47">Asklepia demiti</emphasis>
</taxonomicName>
sp. n.
</paragraph>
</subSubSection>
<subSubSection pageId="46" pageNumber="47" type="dispersal potential">
<paragraph pageId="46" pageNumber="47">Dispersal potential.</paragraph>
<paragraph pageId="46" pageNumber="47">These beetles are macropterous and capable of flight as they have been captured at lights. They are moderately swift and agile runners.</paragraph>
</subSubSection>
<subSubSection pageId="46" pageNumber="47" type="distribution">
<paragraph pageId="46" pageNumber="47">Distribution.</paragraph>
<paragraph pageId="46" pageNumber="47">
(
<figureCitation captionStart="Figure 78" captionStartId="F22" captionText="Figure 78. Distribution map for known localities of Asklepia species of the pulchripennis group." httpUri="https://binary.pensoft.net/fig/30652" pageId="46" pageNumber="47">Fig. 78</figureCitation>
). This species has been found at locations on both the clear-water and black-water systems of the upper and middle Amazon River drainage system. But that does not at all indicate its real distribution: as has been pointed out above, very small beetles are inadequately sampled, especially in the Neotropics.
</paragraph>
</subSubSection>
<subSubSection pageId="46" pageNumber="47" type="way of life">
<paragraph pageId="46" pageNumber="47">Way of life.</paragraph>
<paragraph pageId="46" pageNumber="47">
See
<bibRefCitation author="Erwin, TL" journalOrPublisher="Revista Peruana de Entomologia" pageId="79" pageNumber="80" pagination="1 - 85" refId="B22" refString="Erwin, TL, 1991. Natural history of the carabid beetles at the BIOLAT Rio Manu Biological Station, Pakitza, Peru. Revista Peruana de Entomologia 33: 1 - 85" title="Natural history of the carabid beetles at the BIOLAT Rio Manu Biological Station, Pakitza, Peru." volume="33" year="1991">Erwin (1991)</bibRefCitation>
for a general description. Adults of this species are active in the rainy season in the Varzea rainforest along the main course of the Rio
<normalizedToken originalValue="Solimões">Solimoes</normalizedToken>
. They occur at the Varzea forest edge on the floating macrophyte,
<taxonomicName authorityName="Solms-Laubach" authorityYear="1883" baseAuthorityName="Mart." class="Liliopsida" family="Pontederiaceae" genus="Eichornia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Commelinales" pageId="46" pageNumber="47" phylum="Tracheophyta" rank="species" species="crassipes">
<emphasis italics="true" pageId="46" pageNumber="47">Eichornia crassipes</emphasis>
</taxonomicName>
(Mart.) Solms and on culms of the creeping river grass,
<taxonomicName class="Monocotyledoneae" family="Poaceae" genus="Echinochloa" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Graminales" pageId="46" pageNumber="47" phylum="Angiospermae" rank="species" species="polystachya">
<emphasis italics="true" pageId="46" pageNumber="47">Echinochloa polystachya</emphasis>
</taxonomicName>
(Kunth) Hitchc. In addition, they are climbers and have been found in Varzea forest, some 3.6 m above ground, on
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Macrolobium" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Fabales" pageId="46" pageNumber="47" phylum="Magnoliophyta" rank="species" species="acaciifolium">
<emphasis italics="true" pageId="46" pageNumber="47">Macrolobium acaciifolium</emphasis>
</taxonomicName>
(Benth.) Benth., a small tree in the
<taxonomicName genus="Fabaceae" lsidName="" pageId="46" pageNumber="47" rank="genus">Fabaceae</taxonomicName>
. In
<normalizedToken originalValue="Perú">Peru</normalizedToken>
, one individual was found on a black water river (
<normalizedToken originalValue="Igapó">Igapo</normalizedToken>
) and was attracted to lights on our boat.
</paragraph>
</subSubSection>
<subSubSection lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="other specimens examined">
<paragraph pageId="46" pageNumber="47">Other specimens examined.</paragraph>
<paragraph lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
<emphasis bold="true" pageId="46" pageNumber="47">Brazil</emphasis>
, Amazonas, Rio
<normalizedToken originalValue="Solimões">Solimoes</normalizedToken>
, Ilha de Marchantaria, Lago
<normalizedToken originalValue="Camaleão">Camaleao</normalizedToken>
,
<geoCoordinate degrees="3.2488" direction="south" orientation="latitude" precision="5" value="-3.2488">3.2488°S</geoCoordinate>
,
<geoCoordinate degrees="59.9556" direction="west" orientation="longitude" precision="5" value="-59.9556">59.9556°W</geoCoordinate>
, 7m, 11 April 1981 (NMNH: ADIS # 001149, female paratype), 28 July 1981 (NMNH: ADIS # 001650, female); 14 September 1981 (NMNH: ADIS # 000356, female paratype); 1 October 1981 (J. Adis)(NMNH: ADIS # 000662, ADIS # 000621, ADIS # 000667, ADIS # 000664, ADIS # 000666, ADIS # 000233, ADIS # 000239, ADIS # 000236, ADIS # 000241, ADIS # 000231, female paratypes, ADIS # 000670, ADIS # 000622, ADIS # 000659, ADIS # 000226, ADIS # 000232, ADIS # 000238, ADIS # 000240, ADIS # 000228, ADIS # 000265, ADIS # 000653, male paratypes), 1 October 1981 (NMNH: Adis # 000656, male, not paratype, damaged, Adis # 000185, female, not paratype, damaged), 20 October 1981 (NMNH: ADIS # 000750, ADIS # 000727, female paratypes, ADIS # 001335, ADIS # 001106, ADIS # 001623, ADIS # 001344, ADIS # 001418, male paratypes) 4 November 1981 (NMNH: ADIS # 001106, females); 13-17 September 1991 (C. Martius, A. Rebello)(NMNH: ADIS # 000849, male paratype);
<normalizedToken originalValue="Pará">Para</normalizedToken>
,
<normalizedToken originalValue="Santarém">Santarem</normalizedToken>
,
<normalizedToken originalValue="Río">Rio</normalizedToken>
Tapajos,
<geoCoordinate degrees="2.4079" direction="south" orientation="latitude" precision="5" value="-2.4079">2.4079°S</geoCoordinate>
,
<geoCoordinate degrees="54.7969" direction="west" orientation="longitude" precision="5" value="-54.7969">54.7969°W</geoCoordinate>
, 30m, 27-28 December 1967 (H. Reichardt)(MZUSP: ADP132811, ADP132817, ADP132809, male paratypes, ADP132807, ADP132805, ADP133655, ADP132819, ADP132813, fe
<pageBreakToken pageId="47" pageNumber="48" start="start">male</pageBreakToken>
paratypes, Pacoval, Rio
<normalizedToken originalValue="Curuá">Curua</normalizedToken>
,
<geoCoordinate degrees="1.7733" direction="south" orientation="latitude" precision="5" value="-1.7733">1.7733°S</geoCoordinate>
,
<geoCoordinate degrees="54.9971" direction="west" orientation="longitude" precision="5" value="-54.9971">54.9971°W</geoCoordinate>
, 12m, 16 February 1968 (H. Reichardt)(MZUSP: ADP132815, female paratype).
<emphasis bold="true" pageId="47" pageNumber="48">
<normalizedToken originalValue="Perú">Peru</normalizedToken>
</emphasis>
, Loreto, Boca del
<normalizedToken originalValue="Río">Rio</normalizedToken>
Samiria, 1 km SW Vigilante post No. 1,
<geoCoordinate degrees="4.5005" direction="south" orientation="latitude" precision="5" value="-4.5005">4.5005°S</geoCoordinate>
,
<geoCoordinate degrees="74.0659" direction="west" orientation="longitude" precision="5" value="-74.0659">74.0659°W</geoCoordinate>
, 99m, 6 May 1990 (T.L. Erwin)(MUSM: ADP093365, female).
</paragraph>
</subSubSection>
</treatment>
</document>