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<document id="DCC1391AB8FE020CC55DC4AC5A792AA2" ID-DOI="10.5281/zenodo.5375317" ID-ISSN="1638-9395" ID-Zenodo-Dep="5375317" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630277962741" checkinUser="marcus" docAuthor="Sen, Sevket" docDate="2001" docId="E03A160AFB12FFE5FF79FA413FE3FE8A" docLanguage="en" docName="Geodiversitas.2001.23.2.303-312.pdf" docOrigin="Geodiversitas 23 (2)" docStyle="DocumentStyle:EA833BFD04EE0E3F39962C07E351CEA5.3:Geodiversitas.2000-2008.journal_article.type1" docStyleId="EA833BFD04EE0E3F39962C07E351CEA5" docStyleName="Geodiversitas.2000-2008.journal_article.type1" docStyleVersion="3" docTitle="Hystrix primigenia" docType="treatment" docVersion="4" lastPageNumber="310" masterDocId="1C036E72FB15FFEDFFB6FFE73E03FFAF" masterDocTitle="Early Pliocene porcupine (Mammalia, Rodentia) from Perpignan, France: a new systematic study" masterLastPageNumber="312" masterPageNumber="303" pageNumber="310" updateTime="1699211043790" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="F6B17DFB173535A82EAC5DCA05B65E8F">Early Pliocene porcupine (Mammalia, Rodentia) from Perpignan, France: a new systematic study</mods:title>
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<mods:namePart id="81A9A606347657D33F96651F83097F0A">Sen, Sevket</mods:namePart>
<mods:affiliation id="674C68A32265446514D0E0D6060F3C57">Laboratoire de Paléontologie, Muséum national dHistoire naturelle, UMR 8569 du CNRS, 8 rue Buffon, F- 75231 Paris cedex 05 (France) sen @ mnhn. fr ..</mods:affiliation>
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<mods:title id="8BB702F1477F98BD2DD955FA6ACFFE8E">Geodiversitas</mods:title>
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<mods:date id="7238BA89214ECE3900097D4B6DF0C566">2001</mods:date>
<mods:detail id="F526C051628C9C708FE916FDBD59E380" type="volume">
<mods:number id="6EB53BE2AF303710A180889A19BF6B52">23</mods:number>
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<mods:identifier id="45E70CEB1D38AE061D42A774DBE30FA1" type="DOI">10.5281/zenodo.5375317</mods:identifier>
<mods:identifier id="7E7A0DC07586ACA45A83314F0FF13782" type="ISSN">1638-9395</mods:identifier>
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<treatment id="E03A160AFB12FFE5FF79FA413FE3FE8A" ID-DOI="http://doi.org/10.5281/zenodo.5463927" ID-Zenodo-Dep="5463927" LSID="urn:lsid:plazi:treatment:E03A160AFB12FFE5FF79FA413FE3FE8A" httpUri="http://treatment.plazi.org/id/E03A160AFB12FFE5FF79FA413FE3FE8A" lastPageId="8" lastPageNumber="310" pageId="7" pageNumber="310">
<subSubSection id="2089F497FB12FFEAFF79FA413FC9FA4F" pageId="7" pageNumber="310" type="nomenclature">
<paragraph id="682CA71CFB12FFEAFF79FA413FC9FA4F" blockId="7.[207,627,1446,1504]" pageId="7" pageNumber="310">
<heading id="33641070FB12FFEAFF79FA413FC9FA4F" centered="true" fontSize="11" level="2" pageId="7" pageNumber="310" reason="2">
<taxonomicName id="AF93DC9FFB12FFEAFF79FA413C70FA6D" authority="(Wagner, 1848)" baseAuthorityName="Wagner" baseAuthorityYear="1848" box="[207,627,1446,1474]" class="Mammalia" family="Hystricidae" genus="Hystrix" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="310" phylum="Chordata" rank="species" species="primigenia">
<emphasis id="5AE77B0EFB12FFEAFF79FA413F1CFA6F" bold="true" box="[207,287,1446,1472]" italics="true" pageId="7" pageNumber="310">Hystrix</emphasis>
cf.
<emphasis id="5AE77B0EFB12FFEAFEFEFA413FBCFA6F" bold="true" box="[328,447,1446,1472]" italics="true" pageId="7" pageNumber="310">primigenia</emphasis>
(Wagner, 1848)
</taxonomicName>
(
<figureCitation id="F0A8BB99FB12FFEAFE37FA213FC2FA4F" box="[385,449,1478,1504]" captionStart="FIG" captionStartId="8.[714,725,938,955]" captionTargetBox="[741,1205,216,888]" captionTargetId="figure-467@8.[724,1212,206,877]" captionTargetPageId="8" captionText="FIG. 6. — Hystrix cf. primigenia (Wagner, 1848) from?Serrat den Vacquer; A, B, M1 or M2, inverted; A, occlusal view; B, labial view; C, D, m1 or m2, from the left side; C, occlusal view; D, labial view. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5375329" httpUri="https://zenodo.org/record/5375329/files/figure.png" pageId="7" pageNumber="310">Fig. 6</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2089F497FB12FFEAFF21F9E93FA6F92F" pageId="7" pageNumber="310" type="materials_examined">
<paragraph id="682CA71CFB12FFEAFF21F9E93C93F9EF" blockId="7.[151,683,1548,1666]" pageId="7" pageNumber="310">MATERIAL EXAMINED. — M1 or M2 dex (10.3 × 10.0) (PR-153) and m1 or m2 sin (10.7 × 9.1) (PR-156).</paragraph>
<paragraph id="682CA71CFB12FFEAFF21F9B73FA6F92F" blockId="7.[151,683,1548,1666]" pageId="7" pageNumber="310">
<collectionCode id="0E823FD9FB12FFEAFF21F9B73F00F9C9" box="[151,259,1616,1639]" pageId="7" pageNumber="310">LOCALITY</collectionCode>
. — Serrat den Vacquer, Perpignan, Pyrénées orientales,
<collectingCountry id="1084E78CFB12FFEAFEEAF98E3FA2F92F" box="[348,417,1641,1664]" name="France" pageId="7" pageNumber="310">France</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="2089F497FB12FFE5FD7CFD6D3FE3FE8A" lastPageId="8" lastPageNumber="311" pageId="7" pageNumber="310" type="description">
<paragraph id="682CA71CFB12FFEAFD7CFD6D3A21FD0D" blockId="7.[714,1247,650,1669]" box="[714,1058,650,676]" pageId="7" pageNumber="310">DESCRIPTION AND COMPARISON</paragraph>
<paragraph id="682CA71CFB12FFEAFD7CFD4C3AA1FC0A" blockId="7.[714,1247,650,1669]" pageId="7" pageNumber="310">
Perpignan collection includes
<specimenCount id="7E956C95FB12FFEAFBB9FD4D3AAEFD6A" box="[1039,1197,682,709]" pageId="7" pageNumber="310" type="generic">two specimens</specimenCount>
that are different in size and morphology from the other specimens described above as
<taxonomicName id="AF93DC9FFB12FFEAFB3FFD0D3D20FC8A" authority="Sen, 2001" authorityName="Sen" authorityYear="2001" class="Mammalia" family="Hystricidae" genus="Hystrix" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="310" phylum="Chordata" rank="species" species="depereti" status="sp. nov.">
<emphasis id="5AE77B0EFB12FFEAFB3FFD0D3D20FC8A" italics="true" pageId="7" pageNumber="310">Hystrix depereti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="41D4C675FB12FFEAFC98FCED3D73FC8B" box="[814,880,778,804]" pageId="7" pageNumber="310" rank="species">n. sp.</taxonomicNameLabel>
Their colour is also different: brown in these two teeth while all other
<taxonomicName id="AF93DC9FFB12FFEAFB24FCCD3ADDFCEC" authorityName="Linnaeus" authorityYear="1758" box="[1170,1246,810,835]" class="Mammalia" family="Hystricidae" genus="Hystrix" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="310" phylum="Chordata" rank="genus">
<emphasis id="5AE77B0EFB12FFEAFB24FCCD3ADDFCEC" box="[1170,1246,810,835]" italics="true" pageId="7" pageNumber="310">Hystrix</emphasis>
</taxonomicName>
remains from Serrat den Vacquer are whitish or beige in colour. Thus, they may have been erroneously mixed with the Perpignan material.
</paragraph>
<paragraph id="682CA71CFB12FFEAFD7CFC4C3D4BFAAB" blockId="7.[714,1247,650,1669]" pageId="7" pageNumber="310">
These two teeth are first characterized by their smaller size. P4 has a rectangular occlusal outline although it represents a very adult individual (age class II or IV). Its occlusal surface has the usual pattern of
<taxonomicName id="AF93DC9FFB12FFEAFC8AFBCD3D8BFBEC" authorityName="Linnaeus" authorityYear="1758" box="[828,904,1066,1091]" class="Mammalia" family="Hystricidae" genus="Hystrix" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="310" phylum="Chordata" rank="genus">
<emphasis id="5AE77B0EFB12FFEAFC8AFBCD3D8BFBEC" box="[828,904,1066,1091]" italics="true" pageId="7" pageNumber="310">Hystrix</emphasis>
</taxonomicName>
, with a deep lingual hypoflexus and four labial synclines. Its typical feature is the depth of the hypoflexus which starts, in lingual view, at
<quantity id="AF6B0AF9FB12FFEAFC9BFB6C3D8BFB0B" box="[813,904,1162,1189]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="7" pageNumber="310" unit="mm" value="4.6">4.6 mm</quantity>
above the base of the crown. The large lingual root is grooved and subdivided into two cavities. In addition, there are two small labial roots.
</paragraph>
<paragraph id="682CA71CFB12FFEAFD7CFAEC3D8BF98B" blockId="7.[714,1247,650,1669]" pageId="7" pageNumber="310">
The m1 or m2 is also differentiated by its smaller size compare to other
<taxonomicName id="AF93DC9FFB12FFEAFC6AFACD3A32FAEC" authorityName="Linnaeus" authorityYear="1758" box="[988,1073,1322,1347]" class="Mammalia" family="Hystricidae" genus="Hystrix" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="310" phylum="Chordata" rank="genus">
<emphasis id="5AE77B0EFB12FFEAFC6AFACD3A32FAEC" box="[988,1073,1322,1347]" italics="true" pageId="7" pageNumber="310">Hystrix</emphasis>
</taxonomicName>
remains from Perpignan. The pattern of the occlusal surface is as in
<taxonomicName id="AF93DC9FFB12FFEAFCBFFA8D3DA6FA2C" baseAuthorityName="Wagner" baseAuthorityYear="1848" box="[777,933,1386,1411]" class="Mammalia" family="Hystricidae" genus="Hystrix" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="310" phylum="Chordata" rank="species" species="primigenia">
<emphasis id="5AE77B0EFB12FFEAFCBFFA8D3DA6FA2C" box="[777,933,1386,1411]" italics="true" pageId="7" pageNumber="310">H. primigenia</emphasis>
</taxonomicName>
and
<taxonomicName id="AF93DC9FFB12FFEAFC57FA8D3A8DFA2A" authority="Sen, 2001" authorityName="Sen" authorityYear="2001" box="[993,1166,1386,1413]" class="Mammalia" family="Hystricidae" genus="Hystrix" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="310" phylum="Chordata" rank="species" species="depereti" status="sp. nov.">
<emphasis id="5AE77B0EFB12FFEAFC57FA8D3A8DFA2A" box="[993,1166,1386,1413]" italics="true" pageId="7" pageNumber="310">Hystrix depereti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="41D4C675FB12FFEAFB2EFA8D3AD9FA2B" box="[1176,1242,1386,1412]" pageId="7" pageNumber="310" rank="species">n. sp.</taxonomicNameLabel>
: one labial hypoflexid and four labial synclines which are rather oblique. From the labial view, the hypoflexid is deep; it starts at
<quantity id="AF6B0AF9FB12FFEAFBF6FA2C3A9BFA4B" box="[1088,1176,1482,1509]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="7" pageNumber="310" unit="mm" value="7.3">7.3 mm</quantity>
above the base of the crown. There are two labial and two lingual roots.
</paragraph>
<paragraph id="682CA71CFB12FFE5FD7CF9CC3FE3FE8A" blockId="7.[714,1247,650,1669]" lastBlockId="8.[151,684,171,293]" lastPageId="8" lastPageNumber="311" pageId="7" pageNumber="310">
These two teeth cannot be attributed to
<taxonomicName id="AF93DC9FFB12FFEAFB24F9CD3D1EF9CA" authority="Sen, 2001" authorityName="Sen" authorityYear="2001" class="Mammalia" family="Hystricidae" genus="Hystrix" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="310" phylum="Chordata" rank="species" species="depereti" status="sp. nov.">
<emphasis id="5AE77B0EFB12FFEAFB24F9CD3D1EF9CA" italics="true" pageId="7" pageNumber="310">Hystrix depereti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="41D4C675FB12FFEAFC90F9AD3D66F9CB" box="[806,869,1610,1636]" pageId="7" pageNumber="310" rank="species">n. sp.</taxonomicNameLabel>
because of their smaller size, less hypsodont crown, and deeper hypoflexus (id). The characters mentioned above fit better with those of
<taxonomicName id="AF93DC9FFB1DFFE5FF40FF2D3F8FFF4C" baseAuthorityName="Wagner" baseAuthorityYear="1848" box="[246,396,202,227]" class="Mammalia" family="Hystricidae" genus="Hystrix" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="311" phylum="Chordata" rank="species" species="primigenia">
<emphasis id="5AE77B0EFB1DFFE5FF40FF2D3F8FFF4C" box="[246,396,202,227]" italics="true" pageId="8" pageNumber="311">H. primigenia</emphasis>
</taxonomicName>
, when compared with the specimens from Pikermi (
<collectingCountry id="1084E78CFB1DFFE5FE04FF0C3C00FEAA" box="[434,515,235,261]" name="Greece" pageId="8" pageNumber="311">Greece</collectingCountry>
), Kalimanci IV and Hadji Dimovo (
<collectingCountry id="1084E78CFB1DFFE5FEC2FEEC3FD6FE8A" box="[372,469,267,293]" name="Bulgaria" pageId="8" pageNumber="311">Bulgaria</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>