188 lines
20 KiB
XML
188 lines
20 KiB
XML
<document id="988713DC730141DBB4201E8E71DE402E" ID-CLB-Dataset="20625" ID-DOI="10.11646/zootaxa.5099.5.5" ID-GBIF-Dataset="97d8226a-7b23-4ce7-9ab3-0566525e8f46" ID-ISSN="1175-5326" ID-Zenodo-Dep="6110289" ID-ZooBank="858A6F93-F7D6-4DCE-95BF-8918BBA4F70C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1644998252578" checkinUser="plazi" docAuthor="Simonsen, Thomas J., Archibald, S. Bruce, Rasmussen, Jan A., Sylvestersen, René L., Olsen, Kent & Ware, Jessica L." docDate="2022" docId="D822CB2BFF96FFA60FF6FE7AFF71F258" docLanguage="en" docName="zootaxa.5099.5.5.pdf" docOrigin="Zootaxa 5099 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Danowhetaksa birgitteae Simonsen, Ware & Archibald 2022, new species" docType="treatment" docVersion="8" lastPageNumber="589" masterDocId="241BB353FF94FFA50F61FFA0FF94F649" masterDocTitle="Danowhetaksa gen. nov. with two species from the early Eocene Ølst Formation from Denmark, the first Palearctic Whetwhetaksidae (Odonata: Cephalozygoptera)" masterLastPageNumber="592" masterPageNumber="586" pageNumber="588" updateTime="1699312110636" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="5E29498DE32866C1E38083F5DC9B22BD">
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<mods:title id="028E9E97D785379F8B91F7FCD4514184">Danowhetaksa gen. nov. with two species from the early Eocene Ølst Formation from Denmark, the first Palearctic Whetwhetaksidae (Odonata: Cephalozygoptera)</mods:title>
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<mods:name id="75C9F64AB93B443F49936D5E3676103A" type="personal">
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<mods:namePart id="9B9AB3AC4FCB403F7326FEF2525A324B">Simonsen, Thomas J.</mods:namePart>
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<mods:affiliation id="6D50E9947BC7B5D06676941D67CCF6E7">Natural History Museum Aarhus, Wilhelm Meyers Allé 10, Aarhus, DK- 8000 Aarhus C, Denmark</mods:affiliation>
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<mods:namePart id="0CEB1A3AEEFE3C88A71827126E4F8C03">Archibald, S. Bruce</mods:namePart>
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<mods:affiliation id="22137815BBA958F4D002D686BB36E1A8">Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada; Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America; Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada.</mods:affiliation>
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<mods:name id="3C77A40FD39AAD61795AB8B0C42B8E72" type="personal">
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<mods:roleTerm id="2DA8C8FB288BE775BC2D37E0694CBEA7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B6966D8418B3CB753A09619A56D8AA50">Rasmussen, Jan A.</mods:namePart>
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<mods:affiliation id="56DA95F5BF5DA5757F011519FD0470E0">Museum Mors, Skarrehagevej 8, DK- 7950 NykØbing Mors, Denmark & Natural History Museum of Denmark, Øster Voldgade 5 - 7, DK- 1350 Copenhagen K, Denmark.</mods:affiliation>
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</mods:name>
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<mods:namePart id="480943EF38DE69E5C4A1F27ACA509E34">Sylvestersen, René L.</mods:namePart>
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<mods:affiliation id="4720D1257B05B64B636D4C2DE3BAED73">Fur Museum, Nederby 28, DK- 7884 Fur, Denmark.</mods:affiliation>
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<mods:namePart id="27E91B50B08146260B1B4A60FB2EF70F">Olsen, Kent</mods:namePart>
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<mods:affiliation id="053D7E393FCF0D16E2B20ED3C6ECE22D">Natural History Museum Aarhus, Wilhelm Meyers Allé 10, Aarhus, DK- 8000 Aarhus C, Denmark.</mods:affiliation>
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<mods:namePart id="976F80F5D75FE3B2E7F12FDB8D2C2C6C">Ware, Jessica L.</mods:namePart>
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<mods:affiliation id="2DE3C63A6C1ABD3FDB3035771BCA234D">Division of Invertebrate Zoology, American Museum of Natural History, NY, NY, 10024, USA.</mods:affiliation>
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<mods:title id="56B30E140CAC7D58C1167C82DE7AD914">Zootaxa</mods:title>
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<mods:date id="A05A1A8586CC9C846EA1BFF1B67EE1E5">2022</mods:date>
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<treatment id="D822CB2BFF96FFA60FF6FE7AFF71F258" ID-DOI="http://doi.org/10.5281/zenodo.6311199" ID-GBIF-Taxon="193380037" ID-Zenodo-Dep="6311199" LSID="urn:lsid:plazi:treatment:D822CB2BFF96FFA60FF6FE7AFF71F258" httpUri="http://treatment.plazi.org/id/D822CB2BFF96FFA60FF6FE7AFF71F258" lastPageId="3" lastPageNumber="589" pageId="2" pageNumber="588">
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<subSubSection id="189129B6FF96FFA70FF6FE7AFC4CF7BD" box="[151,984,474,501]" pageId="2" pageNumber="588" type="nomenclature">
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<paragraph id="50347A3DFF96FFA70FF6FE7AFC4CF7BD" blockId="2.[151,984,474,537]" box="[151,984,474,501]" pageId="2" pageNumber="588">
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<heading id="0B7CCD51FF96FFA70FF6FE7AFC4CF7BD" bold="true" box="[151,984,474,501]" fontSize="11" level="1" pageId="2" pageNumber="588" reason="1">
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<emphasis id="62FFA62FFF96FFA70FF6FE7AFC4CF7BD" bold="true" box="[151,984,474,501]" pageId="2" pageNumber="588">
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<taxonomicName id="978B01BEFF96FFA70FF6FE7AFCAFF7BD" authority="Simonsen, Ware & Archibald" authorityName="Simonsen, Ware & Archibald" authorityYear="2022" box="[151,827,474,501]" class="Insecta" family="Sieblosiidae" genus="Danowhetaksa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="588" phylum="Arthropoda" rank="species" species="birgitteae" status="sp. nov.">
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<emphasis id="62FFA62FFF96FFA70FF6FE7AFE50F7BC" bold="true" box="[151,452,474,501]" italics="true" pageId="2" pageNumber="588">Danowhetaksa birgitteae</emphasis>
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Simonsen, Ware & Archibald
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</taxonomicName>
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,
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<taxonomicNameLabel id="79CC1B54FF96FFA70C28FE7BFC4CF7BD" box="[841,984,474,501]" pageId="2" pageNumber="588" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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</subSubSection>
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<subSubSection id="189129B6FF96FFA70FF6FE5FFF61F450" box="[151,245,511,537]" pageId="2" pageNumber="588" type="description">
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<paragraph id="50347A3DFF96FFA70FF6FE5FFF61F450" blockId="2.[151,984,474,537]" box="[151,245,511,537]" pageId="2" pageNumber="588">
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<figureCitation id="C8B066B8FF96FFA70FF6FE5FFF61F450" box="[151,245,511,537]" captionStart="FIGURE 1" captionStartId="2.[151,250,1725,1749]" captionTargetBox="[151,1436,702,1662]" captionTargetId="figure-156@2.[151,1436,702,1698]" captionTargetPageId="2" captionText="FIGURE 1. Danowhetaksa birgitteae n. sp., holotype wing [FUM-N-17515]. A, photograph of part; B, drawing from part and counterpart. Short arrows indicate crossveins below pterostigma (see text). Scale bar is 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6110293" httpUri="https://zenodo.org/record/6110293/files/figure.png" pageId="2" pageNumber="588">Figure 1</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="189129B6FF96FFA70FF6FDE6FB8EF4E0" pageId="2" pageNumber="588" type="materials_examined">
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<paragraph id="50347A3DFF96FFA70FF6FDE6FB8EF4E0" blockId="2.[151,1436,582,681]" pageId="2" pageNumber="588">
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<emphasis id="62FFA62FFF96FFA70FF6FDE6FE92F429" bold="true" box="[151,262,582,608]" pageId="2" pageNumber="588">Material.</emphasis>
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<materialsCitation id="E0E37060FF96FFA70E6DFDE7FB8EF4E0" ID-GBIF-Occurrence="3467095302" collectingDate="1996" collectorName="Birgitte Munk" pageId="2" pageNumber="588" specimenCode="FUM-M-17515" specimenCount="1" typeStatus="holotype">
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<typeStatus id="8F30C49FFF96FFA70E6DFDE7FEE0F428" box="[268,372,583,609]" pageId="2" pageNumber="588" type="holotype">Holotype</typeStatus>
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[
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<specimenCode id="002DD246FF96FFA70E1EFDE6FDACF428" box="[383,568,582,609]" pageId="2" pageNumber="588">FUM-M-17515</specimenCode>
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]: an isolated wing preserved in a concretion block; the part is missing the basalmost and apical-posterior portions, and the counterpart is missing the apical-most portion including the pterostigma; collected by
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<collectorName id="FD7E1FEBFF96FFA70E47FD2FFE5CF4E0" box="[294,456,655,681]" pageId="2" pageNumber="588">Birgitte Munk</collectorName>
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,
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<collectingDate id="3471A515FF96FFA70EB2FD2FFD9AF4E0" box="[467,526,655,681]" pageId="2" pageNumber="588">1996</collectingDate>
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, Fur Stolleklint; deposited in the Fur Museum.
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<caption id="04F42AB5FF96FFA70FF6F91DFBDBF0B0" ID-DOI="http://doi.org/10.5281/zenodo.6110293" ID-Zenodo-Dep="6110293" httpUri="https://zenodo.org/record/6110293/files/figure.png" pageId="2" pageNumber="588" startId="2.[151,250,1725,1749]" targetBox="[151,1436,702,1662]" targetPageId="2">
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<paragraph id="50347A3DFF96FFA70FF6F91DFBDBF0B0" blockId="2.[151,1437,1725,1785]" pageId="2" pageNumber="588">
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<emphasis id="62FFA62FFF96FFA70FF6F91DFE87F09C" bold="true" box="[151,275,1725,1749]" pageId="2" pageNumber="588">FIGURE 1.</emphasis>
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<taxonomicName id="978B01BEFF96FFA70E7BF91DFD8CF09C" authority="Simonsen & Archibald & Rasmussen & Sylvestersen & Olsen & Ware, 2022" authorityName="Simonsen & Archibald & Rasmussen & Sylvestersen & Olsen & Ware" authorityYear="2022" box="[282,536,1725,1749]" class="Insecta" family="Sieblosiidae" genus="Danowhetaksa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="588" phylum="Arthropoda" rank="species" species="birgitteae" status="sp. nov.">
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<emphasis id="62FFA62FFF96FFA70E7BF91DFD8CF09C" box="[282,536,1725,1749]" italics="true" pageId="2" pageNumber="588">Danowhetaksa birgitteae</emphasis>
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</taxonomicName>
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<emphasis id="62FFA62FFF96FFA70D7FF91DFDCCF09C" bold="true" box="[542,600,1725,1749]" pageId="2" pageNumber="588">
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<taxonomicNameLabel id="79CC1B54FF96FFA70D7FF91DFDCCF09C" box="[542,600,1725,1749]" pageId="2" pageNumber="588" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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, holotype wing [FUM-N-17515]. A, photograph of part; B, drawing from part and counterpart. Short arrows indicate crossveins below pterostigma (see text). Scale bar is 5 mm.
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</paragraph>
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</caption>
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||
<subSubSection id="189129B6FF96FFA60FA6F887FB38F7BC" lastPageId="3" lastPageNumber="589" pageId="2" pageNumber="588" type="description">
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<paragraph id="50347A3DFF96FFA60FA6F887FB38F7BC" blockId="2.[151,1437,1831,2038]" lastBlockId="3.[151,1437,151,1041]" lastPageId="3" lastPageNumber="589" pageId="2" pageNumber="588">
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<emphasis id="62FFA62FFF96FFA70FA6F887FECDF108" bold="true" box="[199,345,1831,1857]" pageId="2" pageNumber="588">Description.</emphasis>
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<typeStatus id="8F30C49FFF96FFA70E03F887FE5EF108" box="[354,458,1831,1857]" pageId="2" pageNumber="588" type="holotype">Holotype</typeStatus>
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wing. Length, arculus to distal end of pterostigma: 28.0 mm; nodus to distal end of pterostigma:
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<quantity id="9773D7D8FF96FFA70E4BF8ECFE06F12F" box="[298,402,1868,1894]" metricMagnitude="-2" metricUnit="m" metricValue="2.03" pageId="2" pageNumber="588" unit="mm" value="20.3">20.3 mm</quantity>
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; arculus to base of pterostigma: max
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<quantity id="9773D7D8FF96FFA70C55F8ECFC08F12F" box="[820,924,1868,1894]" metricMagnitude="-2" metricUnit="m" metricValue="2.34" pageId="2" pageNumber="588" unit="mm" value="23.4">23.4 mm</quantity>
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; nodus to base of pterostigma: max
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<quantity id="9773D7D8FF96FFA70A50F8EBFA0CF12F" box="[1329,1432,1867,1894]" metricMagnitude="-2" metricUnit="m" metricValue="1.6199999999999999" pageId="2" pageNumber="588" unit="mm" value="16.2">16.2 mm</quantity>
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; width:
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<quantity id="9773D7D8FF96FFA70F84F8D0FEA9F1C3" box="[229,317,1904,1930]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="2" pageNumber="588" unit="mm" value="9.3">9.3 mm</quantity>
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. Pterostigma dark (damaged basally), preserved part approximately
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<quantity id="9773D7D8FF96FFA70B52F8CFFBE7F1C3" box="[1075,1139,1903,1930]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="588" unit="mm" value="5.0">5 mm</quantity>
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in length, highly elongate, at least seven times longer than wide, subtends numerous cells, five crossveins detected by preservation, surely many more. Membrane with a transverse, sub-central dark fascia, slightly narrower than length of pterostigma, otherwise hyaline. Twenty-three crossveins preserved in postnodal space,
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<quantity id="9773D7D8FF96FFA70CD1F87CFC7DF1BC" box="[944,1001,2011,2038]" metricMagnitude="-1" metricUnit="m" metricValue="5.588" pageId="2" pageNumber="588" unit="in" value="22.0">22 in</quantity>
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postsubnodal space, only pair aligned. IR1 very poorly preserved, origin probably slightly zigzagged, seven cells distal to origin of RP2. RP2 originates seven cells distal to subnodus. IR2 originates approximately 8.5/10 distance arculus to nodus, preserved part of IR2 close to linear. RP3-4 originates approximately 4/10 distance arculus to nodus, preserved part of RP3-4 close to linear. MA linear in basal 1/3, starts zigzagging slightly about basal margin of darkened wing band, poorly preserved beyond band. MP linear from origin to terminus at wing margin, slightly curved. CuA linear from origin to terminus on wing margin, basal 2/3 subparallel to MA, then curving sharper to wing margin, terminates basal to mid-wing. MA-CuA space two cells wide where CuA starts curving away, widening to at least six cells at wing margin. CuA-A space well preserved, very broad, at least five cells wide. Quadrangle sub-trapezoid, broadest distally, approximately 1.3 times longer than wide. Anterior wing margin not preserved basal to arculus, so Ax1 not preserved. Ax2 approximately 2/10 distance arculus to nodus, just distal to anterodistal corner of quadrangle.
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</paragraph>
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</subSubSection>
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<subSubSection id="189129B6FF97FFA60FA6FE5EFB64F55C" pageId="3" pageNumber="589" type="diagnosis">
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<paragraph id="50347A3DFF97FFA60FA6FE5EFA1FF4B8" blockId="3.[151,1437,151,1041]" pageId="3" pageNumber="589">
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<emphasis id="62FFA62FFF97FFA60FA6FE5EFED6F451" bold="true" box="[199,322,510,536]" pageId="3" pageNumber="589">Diagnosis.</emphasis>
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Distinguished from
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<taxonomicName id="978B01BEFF97FFA60D49FE5FFDE9F451" authorityName="Simonsen & Archibald & Rasmussen & Sylvestersen & Olsen & Ware" authorityYear="2022" box="[552,637,511,537]" class="Insecta" family="Sieblosiidae" genus="Danowhetaksa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="589" phylum="Arthropoda" rank="species" species="rusti">
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<emphasis id="62FFA62FFF97FFA60D49FE5FFDA8F451" box="[552,572,511,536]" italics="true" pageId="3" pageNumber="589">D</emphasis>
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.
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<emphasis id="62FFA62FFF97FFA60D2BFE5FFDE9F451" box="[586,637,511,536]" italics="true" pageId="3" pageNumber="589">rusti</emphasis>
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</taxonomicName>
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by any of: 1, RP1-2–IR2 space one cell wide, RP2–IR2 space to origin of IR1 (not preserved beyond this) [RP1-2–IR2 space becomes two cells wide
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<emphasis id="62FFA62FFF97FFA60B76FD83FBA5F475" box="[1047,1073,547,572]" italics="true" pageId="3" pageNumber="589">ca</emphasis>
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. two cells basal origin of RP2]; 2, IR2–RP3-4 space becomes two cells wide
|
||
<emphasis id="62FFA62FFF97FFA60DEEFDE7FD25F429" box="[655,689,583,608]" italics="true" pageId="3" pageNumber="589">ca.</emphasis>
|
||
mid-way between origins of RP2, IR1 [ca. origin of IR2]; 3, seven crossveins in R1+2–IR2 space between nodus and origin of RP2 [
|
||
<emphasis id="62FFA62FFF97FFA60C1CFDCBFC03F4CD" box="[893,919,619,644]" italics="true" pageId="3" pageNumber="589">ca</emphasis>
|
||
. 9]. Further separated by colouration (sex unknown): narrow, dark fascia mid-wing [almost twice as wide], basally starting
|
||
<emphasis id="62FFA62FFF97FFA60C9FFD2FFB8CF4E1" box="[1022,1048,655,680]" italics="true" pageId="3" pageNumber="589">ca</emphasis>
|
||
. five cells distal to nodus, includes origin of RP2, ends slightly distal to termination of CuA [anterior to R1+2 starts at nodus, posterior R1+2 just basal origin of IR2, ends on posterior margin just basal to termination of CuA, on anterior margin slightly more distal].
|
||
</paragraph>
|
||
<paragraph id="50347A3DFF97FFA60FA6FD5AFB64F55C" blockId="3.[151,1437,151,1041]" box="[199,1264,762,791]" pageId="3" pageNumber="589">
|
||
<emphasis id="62FFA62FFF97FFA60FA6FD5AFE18F55C" bold="true" box="[199,396,762,789]" pageId="3" pageNumber="589">Deposit and age.</emphasis>
|
||
Stolleklint clay, Ølst Formation, Stolleklint, Fur,
|
||
<collectingCountry id="289C3AADFF97FFA60CD9FD5BFBB0F55C" box="[952,1060,763,789]" name="Denmark" pageId="3" pageNumber="589">Denmark</collectingCountry>
|
||
; earliest Ypresian.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="189129B6FF97FFA60FA6FCBFFE48F514" pageId="3" pageNumber="589" type="etymology">
|
||
<paragraph id="50347A3DFF97FFA60FA6FCBFFE48F514" blockId="3.[151,1437,151,1041]" pageId="3" pageNumber="589">
|
||
<emphasis id="62FFA62FFF97FFA60FA6FCBFFEDAF570" bold="true" box="[199,334,799,825]" pageId="3" pageNumber="589">Etymology.</emphasis>
|
||
An eponym formed from the given name of Birgitte Munk, who found and donated the
|
||
<typeStatus id="8F30C49FFF97FFA60A52FCBFFA0CF570" box="[1331,1432,799,825]" pageId="3" pageNumber="589" type="holotype">holotype</typeStatus>
|
||
, recognising her contribution.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="189129B6FF97FFA60FA6FCC7FF71F258" pageId="3" pageNumber="589" type="discussion">
|
||
<paragraph id="50347A3DFF97FFA60FA6FCC7FEBDF580" blockId="3.[151,1437,151,1041]" pageId="3" pageNumber="589">
|
||
<emphasis id="62FFA62FFF97FFA60FA6FCC7FEAFF5C8" bold="true" box="[199,315,871,897]" pageId="3" pageNumber="589">Remarks.</emphasis>
|
||
Colouration is provided as supplementary in the diagnosis, as it is unknown if there are differences in colouration due to sexual dimorphism, variation between forewings and hind wings, or polymorphism in
|
||
<taxonomicName id="978B01BEFF97FFA60A37FC2BFEB1F580" authorityName="Archibald & Cannings" authorityYear="2021" class="Insecta" family="Whetwhetaksidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="589" phylum="Arthropoda" rank="family">Whetwhetaksidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="50347A3DFF97FFA60FA6FC73FF71F258" blockId="3.[151,1437,151,1041]" pageId="3" pageNumber="589">Although the wing base is not preserved, we estimate CuA to terminate on the margin just over half wing length.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |