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<mods:title id="8B1C1267869BD2308CB512806D34AB2E">A checklist and key to the snakes of the genus Geophis (Squamata: Colubridae: Dipsadinae), with commentary on distribution and conservation</mods:title>
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<heading id="AD676B3AFFFEFF8265E4FA49F866878D" bold="true" box="[151,538,1509,1535]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<emphasis id="C4E40044FFFEFF8265E4FA49F866878D" bold="true" box="[151,538,1509,1535]" pageId="15" pageNumber="16">
<keyStep id="4D64C4F3FFFEFF8265E4FA49F866878D" box="[151,538,1509,1535]" pageId="15" pageNumber="16">
<key id="82018B67FFFEFF8265E4FA49F866878D" box="[151,538,1509,1535]" pageId="15" pageNumber="16">
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37.
<taxonomicName id="3190A7D5FFFEFF8265B0FA49F866878D" authority="Cope" authorityName="Cope" box="[195,538,1509,1535]" class="Reptilia" family="Colubridae" genus="Geophis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="rhodogaster">
<emphasis id="C4E40044FFFEFF8265B0FA49FBC2878D" bold="true" box="[195,446,1509,1535]" italics="true" pageId="15" pageNumber="16">Geophis rhodogaster</emphasis>
(Cope)
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</keyLead>
</key>
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</emphasis>
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<subSubSection id="BE8A8FDDFFFEFF8265E4F99CFBE88510" pageId="15" pageNumber="16" type="reference_group">
<paragraph id="F62FDC56FFFEFF8265E4F99CF83A8435" blockId="15.[151,685,1584,1735]" box="[151,582,1584,1607]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="D680FB78FFFEFF8265E4F99CF83A8435" box="[151,582,1584,1607]" pageId="15" pageNumber="16">
<taxonomicName id="3190A7D5FFFEFF8265E4F99CF83E8435" ID-CoL="X84Z" authority="Cope 1868: 130" authorityName="Cope" authorityPageNumber="130" authorityYear="1868" box="[151,578,1584,1607]" class="Squamata" family="Colubridae" genus="Colophrys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="rhodogaster">
<emphasis id="C4E40044FFFEFF8265E4F99CFBF28435" box="[151,398,1584,1607]" italics="true" pageId="15" pageNumber="16">Colophrys rhodogaster</emphasis>
<treatmentCitation id="7731FA47FFFEFF8264E6F99CF83E8435" author="Cope" box="[405,578,1584,1607]" page="130" pageId="15" pageNumber="16" year="1868">
<bibRefCitation id="9201A1A7FFFEFF8264E6F99CF8718435" author="Cope" box="[405,525,1584,1607]" pageId="15" pageNumber="29" refString="Cope, E. D. (1868) An examination of the reptilia and batrachia obtained by the Orton expedition to Ecuador; and the upper Amazon; with notes on other species. Proceedings of the Academy of Natural Sciences, Philadelphia, 20, 96 - 140." type="journal article" year="1868">Cope 1868</bibRefCitation>
: 130
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</taxonomicName>
.
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</paragraph>
<paragraph id="F62FDC56FFFEFF8265E4F9FCF8298415" blockId="15.[151,685,1584,1735]" box="[151,597,1616,1639]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="D680FB78FFFEFF8265E4F9FCF8298415" box="[151,597,1616,1639]" pageId="15" pageNumber="16">
<taxonomicName id="3190A7D5FFFEFF8265E4F9FCF82D8415" authority="Bocourt 1883: 531" authorityName="Bocourt" authorityPageNumber="531" authorityYear="1883" box="[151,593,1616,1639]" class="Reptilia" family="Colubridae" genus="Geophis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="rhodogaster">
<emphasis id="C4E40044FFFEFF8265E4F9FCFB048415" box="[151,376,1616,1639]" italics="true" pageId="15" pageNumber="16">Geophis rhodogaster</emphasis>
:
<treatmentCitation id="7731FA47FFFEFF8264F5F9FCF82D8415" author="Bocourt" box="[390,593,1616,1639]" page="531" pageId="15" pageNumber="16" year="1883">
<bibRefCitation id="9201A1A7FFFEFF8264F5F9FCF8608415" author="Bocourt" box="[390,540,1616,1639]" pageId="15" pageNumber="28" refString="Bocourt, F. M. (1883) In: Dumeril, A. H. A., Bocourt, M. F. &amp; Mocquard, F. 1870 - 1909. Etudes sur les Reptiles. Mission Scientifique au Mexique et dans l'Amerique Centrale. Recherches Zoologiques pour servir a l'Histoire de la Fauna de l'Amerique Centrale et du Mexique. Troisieme Partie. - 1 re Section. Texte. Imprimerie Nationale, Paris. xiv + 1012 pp." type="book chapter" year="1883">Bocourt 1883</bibRefCitation>
: 531
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="F62FDC56FFFEFF8265E4F9DCF84984F5" blockId="15.[151,685,1584,1735]" box="[151,565,1648,1671]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="D680FB78FFFEFF8265E4F9DCF84984F5" box="[151,565,1648,1671]" pageId="15" pageNumber="16">
<taxonomicName id="3190A7D5FFFEFF8265E4F9DCF84D84F5" authority="Gunther 1893: 87" authorityName="Gunther" authorityPageNumber="87" authorityYear="1893" box="[151,561,1648,1671]" class="Reptilia" family="Colubridae" genus="Geophis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="chalybaea">
<emphasis id="C4E40044FFFEFF8265E4F9DCFB1F84F5" box="[151,355,1648,1671]" italics="true" pageId="15" pageNumber="16">Geophis chalybaea</emphasis>
:
<treatmentCitation id="7731FA47FFFEFF826401F9DCF84D84F5" author="Gunther" box="[370,561,1648,1671]" page="87" pageId="15" pageNumber="16" year="1893">
<bibRefCitation id="9201A1A7FFFEFF826401F9DCF87584F5" author="Gunther" box="[370,521,1648,1671]" pageId="15" pageNumber="29" refString="Gunther, A. C. L. F. (1893) Part 109, pages 89 - 96. In: Salvin, O. &amp; Godman, F. D. (Eds.). Biologia Centrali-Americana; or, Contributions to the Knowledge of the Fauna and Flora of Mexico and Central America. R. H. Porter and Dulau &amp; Co., London. xx + 326 pp. + 76 pls." type="book chapter" year="1893">Günther 1893</bibRefCitation>
: 87
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="F62FDC56FFFEFF8265E4F93CF8D184B5" blockId="15.[151,685,1584,1735]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="D680FB78FFFEFF8265E4F93CF8D184B5" pageId="15" pageNumber="16">
<taxonomicName id="3190A7D5FFFEFF8265E4F93CF81A84D5" authority="Amaral 1929: 192" authorityName="Amaral" authorityPageNumber="192" authorityYear="1929" box="[151,614,1680,1703]" class="Agaricomycetes" family="Agaricaceae" genus="Catastoma" kingdom="Animalia" order="Agaricales" pageId="15" pageNumber="16" phylum="Basidiomycota" rank="species" species="rhodogaster">
<emphasis id="C4E40044FFFEFF8265E4F93CFBEF84D5" box="[151,403,1680,1703]" italics="true" pageId="15" pageNumber="16">Catastoma rhodogaster</emphasis>
:
<treatmentCitation id="7731FA47FFFEFF8264D2F93CF81A84D5" author="Amaral" box="[417,614,1680,1703]" page="192" pageId="15" pageNumber="16" year="1929">
<bibRefCitation id="9201A1A7FFFEFF8264D2F93CF84D84D5" author="Amaral" box="[417,561,1680,1703]" pageId="15" pageNumber="28" refString="Amaral, A. d. (1929) Estudos sobre ophidios neotropicos XVIII. Lista remissive dos ophidios da regiao neotropico. Memorias Do Instituto Butantan, 4, 129 - 271." type="journal article" year="1929">Amaral 1929</bibRefCitation>
: 192
</treatmentCitation>
</taxonomicName>
.
<taxonomicName id="3190A7D5FFFEFF8265E4F91CF84284B5" authority="Kohler 1996: 36" authorityName="Kohler" authorityPageNumber="36" authorityYear="1996" box="[151,574,1712,1735]" class="Reptilia" family="Colubridae" genus="Geophis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="fulvoguttatus">
<emphasis id="C4E40044FFFEFF8265E4F91CFBFD84B5" box="[151,385,1712,1735]" italics="true" pageId="15" pageNumber="16">Geophis fulvoguttatus</emphasis>
:
<treatmentCitation id="7731FA47FFFEFF8264E3F91CF84284B5" author="Kohler" box="[400,574,1712,1735]" page="36" pageId="15" pageNumber="16" year="1996">
<bibRefCitation id="9201A1A7FFFEFF8264E3F91CF86584B5" author="Kohler" box="[400,537,1712,1735]" pageId="15" pageNumber="29" refString="Kohler, G. (1996) Notes on a collection of reptiles from El Salvador collected between 1951 - 1956. Senckenbergiana Biologica, 76, 29 - 38." type="journal article" year="1996">Köhler 1996</bibRefCitation>
: 36
</treatmentCitation>
</taxonomicName>
(in error).
</treatmentCitationGroup>
</paragraph>
<paragraph id="F62FDC56FFFEFF8265E4F954FBE88510" blockId="15.[151,1437,1784,2010]" pageId="15" pageNumber="16">
<typeStatus id="292B62F4FFFEFF8265E4F954FB7A8560" box="[151,262,1784,1810]" pageId="15" pageNumber="16" type="syntype">Syntypes</typeStatus>
. Three: Academy of Natural Sciences of Philadelphia (ANSP) 3316­17 and National Museum of Natural History (USNM) 12425, collected by Van Patten.
<bibRefCitation id="9201A1A7FFFEFF826633F88CFE6A8548" author="Downs" box="[832,1046,1824,1850]" pageId="15" pageNumber="29" refString="Downs, F. L. (1967) Intrageneric relationships among colubrid snakes of the genus Geophis Wagler. Miscellaneous Publications of the Museum of Zoology, University of Michigan, 131, 1 - 193." type="journal article" year="1967">Downs (1967: 92)</bibRefCitation>
designated ANSP 3317, an adult female, the
<typeStatus id="292B62F4FFFEFF826453F8E4FBF38510" box="[288,399,1864,1890]" pageId="15" pageNumber="16" type="lectotype">lectotype</typeStatus>
.
</paragraph>
</subSubSection>
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<paragraph id="F62FDC56FFFEFF8265B4F8DCF9BD85A8" blockId="15.[151,1437,1784,2010]" pageId="15" pageNumber="16">
Type­locality. “…elevated country in the neighborhood of the city of
<collectingCountry id="8E879CC6FFFEFF826686F8DCFE0785F8" box="[1013,1147,1904,1930]" name="Guatemala" pageId="15" pageNumber="16">Guatemala</collectingCountry>
.” Distribution. The mountains of southwestern Chiapas,
<collectingCountry id="8E879CC6FFFEFF82662AF834F9CA85C0" box="[857,950,1944,1970]" name="Mexico" pageId="15" pageNumber="16">Mexico</collectingCountry>
, east through southern
<collectingCountry id="8E879CC6FFFEFF8261BCF834FF3285C0" box="[1231,1358,1944,1970]" name="Guatemala" pageId="15" pageNumber="16">Guatemala</collectingCountry>
to the Cordillera Metapán, Santa Ana,
<collectingCountry id="8E879CC6FFFEFF826761F86CF8E185A8" box="[530,669,1984,2010]" name="El Salvador" pageId="15" pageNumber="16">El Salvador</collectingCountry>
,
<quantity id="316871B3FFFEFF8267DAF86CF93B85A8" box="[681,839,1984,2010]" metricMagnitude="3" metricUnit="m" metricValue="2.122" metricValueMax="2.7439999999999998" metricValueMin="1.5" pageId="15" pageNumber="16" unit="m" value="2122.0" valueMax="2744.0" valueMin="1500.0">1500­2744 m</quantity>
elevation.
</paragraph>
<paragraph id="F62FDC56FFE1FF9D65B4FF34F8FD82C0" blockId="16.[151,1437,152,418]" box="[199,641,152,178]" pageId="16" pageNumber="17">
Species group.
<taxonomicName id="3190A7D5FFE1FF9D6408FF34F80182C0" authority="Downs 1967" authorityName="Downs" authorityYear="1967" box="[379,637,152,178]" class="Reptilia" family="Colubridae" genus="Atractus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="dubius">
<emphasis id="C4E40044FFE1FF9D6408FF34FBB582C3" box="[379,457,152,177]" italics="true" pageId="16" pageNumber="17">dubius</emphasis>
(
<bibRefCitation id="9201A1A7FFE1FF9D64ABFF34F80F82C0" author="Downs" box="[472,627,152,178]" pageId="16" pageNumber="29" refString="Downs, F. L. (1967) Intrageneric relationships among colubrid snakes of the genus Geophis Wagler. Miscellaneous Publications of the Museum of Zoology, University of Michigan, 131, 1 - 193." type="journal article" year="1967">Downs 1967</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="F62FDC56FFE1FF9D65B4FF6CFBE283D0" blockId="16.[151,1437,152,418]" pageId="16" pageNumber="17">
Systematic references.
<bibRefCitation id="9201A1A7FFE1FF9D64A5FF6CF81182A8" author="Cope" box="[470,621,192,218]" pageId="16" pageNumber="29" refString="Cope, E. D. (1868) An examination of the reptilia and batrachia obtained by the Orton expedition to Ecuador; and the upper Amazon; with notes on other species. Proceedings of the Academy of Natural Sciences, Philadelphia, 20, 96 - 140." type="journal article" year="1868">Cope (1868)</bibRefCitation>
,
<bibRefCitation id="9201A1A7FFE1FF9D670AFF6CF95882A8" author="Downs" box="[633,804,192,218]" pageId="16" pageNumber="29" refString="Downs, F. L. (1967) Intrageneric relationships among colubrid snakes of the genus Geophis Wagler. Miscellaneous Publications of the Museum of Zoology, University of Michigan, 131, 1 - 193." type="journal article" year="1967">Downs (1967)</bibRefCitation>
,
<bibRefCitation id="9201A1A7FFE1FF9D6643FF6CF9B382A8" author="Smith" box="[816,975,192,218]" pageId="16" pageNumber="30" refString="Smith, E. N. (1995) Geophis rhodogaster (Colubridae), an addition to the snake fauna of Mexico. Southwestern Naturalist, 40, 123 - 124." type="journal article" year="1995">Smith (1995)</bibRefCitation>
, Townsend (2006). Remarks.
<bibRefCitation id="9201A1A7FFE1FF9D6431FF44FB808370" author="Gunther" box="[322,508,232,258]" pageId="16" pageNumber="29" refString="Gunther, A. C. L. F. (1893) Part 109, pages 89 - 96. In: Salvin, O. &amp; Godman, F. D. (Eds.). Biologia Centrali-Americana; or, Contributions to the Knowledge of the Fauna and Flora of Mexico and Central America. R. H. Porter and Dulau &amp; Co., London. xx + 326 pp. + 76 pls." type="book chapter" year="1893">Günther (1893)</bibRefCitation>
reported
<taxonomicName id="3190A7D5FFE1FF9D6700FF44F95A8373" box="[627,806,232,257]" class="Agaricomycetes" family="Agaricaceae" genus="Catastoma" kingdom="Animalia" order="Agaricales" pageId="16" pageNumber="17" phylum="Basidiomycota" rank="species" species="rhodogaster">
<emphasis id="C4E40044FFE1FF9D6700FF44F95A8373" box="[627,806,232,257]" italics="true" pageId="16" pageNumber="17">G. rhodogaster</emphasis>
</taxonomicName>
from
<collectingCountry id="8E879CC6FFE1FF9D6606FF44F9818370" box="[885,1021,232,258]" name="Costa Rica" pageId="16" pageNumber="17">Costa Rica</collectingCountry>
.
<bibRefCitation id="9201A1A7FFE1FF9D617FFF44FEA38370" author="Boulenger" box="[1036,1247,232,258]" pageId="16" pageNumber="28" refString="Boulenger, G. A. (1894) Catalogue of snakes in the British Museum (Natural History). London, British Museum 2, i - xi, 1 - 382, 20 pls." type="journal article" year="1894">Boulenger (1894)</bibRefCitation>
apparently followed
<bibRefCitation id="9201A1A7FFE1FF9D6595FEBCFBE08358" author="Gunther" box="[230,412,272,298]" pageId="16" pageNumber="29" refString="Gunther, A. C. L. F. (1893) Part 109, pages 89 - 96. In: Salvin, O. &amp; Godman, F. D. (Eds.). Biologia Centrali-Americana; or, Contributions to the Knowledge of the Fauna and Flora of Mexico and Central America. R. H. Porter and Dulau &amp; Co., London. xx + 326 pp. + 76 pls." type="book chapter" year="1893">Günther (1893)</bibRefCitation>
and included
<collectingCountry id="8E879CC6FFE1FF9D6732FEBCF8BE8358" box="[577,706,272,298]" name="Costa Rica" pageId="16" pageNumber="17">Costa Rica</collectingCountry>
in the range of
<taxonomicName id="3190A7D5FFE1FF9D6608FEBCFE54835B" box="[891,1064,272,297]" class="Agaricomycetes" family="Agaricaceae" genus="Catastoma" kingdom="Animalia" order="Agaricales" pageId="16" pageNumber="17" phylum="Basidiomycota" rank="species" species="rhodogaster">
<emphasis id="C4E40044FFE1FF9D6608FEBCFE54835B" box="[891,1064,272,297]" italics="true" pageId="16" pageNumber="17">G. rhodogaster</emphasis>
</taxonomicName>
.
<bibRefCitation id="9201A1A7FFE1FF9D6144FEBCFEA58358" author="Taylor" box="[1079,1241,272,298]" pageId="16" pageNumber="31" refString="Taylor, E. H. (1951) A brief review of the snakes of Costa Rica. University of Kansas Science Bulletin, 34, 1 - 188." type="journal article" year="1951">Taylor (1951)</bibRefCitation>
also included
<taxonomicName id="3190A7D5FFE1FF9D60F0FEBCFB5A8323" class="Agaricomycetes" family="Agaricaceae" genus="Catastoma" kingdom="Animalia" order="Agaricales" pageId="16" pageNumber="17" phylum="Basidiomycota" rank="species" species="rhodogaster">
<emphasis id="C4E40044FFE1FF9D60F0FEBCFB5A8323" italics="true" pageId="16" pageNumber="17">G. rhodogaster</emphasis>
</taxonomicName>
in his review of
<collectingCountry id="8E879CC6FFE1FF9D649BFE94F8148320" box="[488,616,312,338]" name="Costa Rica" pageId="16" pageNumber="17">Costa Rica</collectingCountry>
snakes, but noted “I regard this record as somewhat doubtful.”
<bibRefCitation id="9201A1A7FFE1FF9D6038FE94FA988308" author="Kohler" pageId="16" pageNumber="29" refString="Kohler, G. (1996) Notes on a collection of reptiles from El Salvador collected between 1951 - 1956. Senckenbergiana Biologica, 76, 29 - 38." type="journal article" year="1996">Köhler (1996)</bibRefCitation>
gave measurements and scutellation data for a specimen of
<taxonomicName id="3190A7D5FFE1FF9D66B0FECCFE09830B" box="[963,1141,352,377]" class="Agaricomycetes" family="Agaricaceae" genus="Catastoma" kingdom="Animalia" order="Agaricales" pageId="16" pageNumber="17" phylum="Basidiomycota" rank="species" species="rhodogaster">
<emphasis id="C4E40044FFE1FF9D66B0FECCFE09830B" box="[963,1141,352,377]" italics="true" pageId="16" pageNumber="17">G. rhodogaster</emphasis>
</taxonomicName>
(SMF 77413) under the name
<taxonomicName id="3190A7D5FFE1FF9D65AEFE24FBEB83D3" box="[221,407,392,417]" class="Reptilia" family="Colubridae" genus="Geophis" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="fulvoguttatus">
<emphasis id="C4E40044FFE1FF9D65AEFE24FBEB83D3" box="[221,407,392,417]" italics="true" pageId="16" pageNumber="17">G. fulvoguttatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>