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<document ID-CLB-Dataset="25290" ID-DOI="http://dx.doi.org/10.3897/zookeys.878.37403" ID-GBIF-Dataset="83f095d1-8441-49cd-be4b-71f0ddfa311c" ID-PMC="PMC6791902" ID-Pensoft-Pub="1313-2970-878-73" ID-Pensoft-UUID="C93F9CBB14D25FEAB2ACCBC3E6282D28" ID-PubMed="31632177" ID-ZooBank="192214DE1D38467BA577ECD16EC5EAB5" ModsDocID="1313-2970-878-73" checkinTime="1570497196116" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo &amp; Balke, Michael" docDate="2019" docId="733BAC0C78D75CBFBC8D78BA8FBAC482" docLanguage="en" docName="ZooKeys 878: 73-143" docOrigin="ZooKeys 878" docSource="http://dx.doi.org/10.3897/zookeys.878.37403" docTitle="Exocelina takime Balke 1998" docType="treatment" docVersion="8" id="C93F9CBB14D25FEAB2ACCBC3E6282D28" lastPageNumber="73" masterDocId="C93F9CBB14D25FEAB2ACCBC3E6282D28" masterDocTitle="Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="143" masterPageNumber="73" pageNumber="73" updateTime="1732929340104" updateUser="ExternalLinkService">
<mods:mods id="262312E00549B5A6E278BD499EDCE561" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="6E466A14A604B9E1026FB797F73EA01F">Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
</mods:titleInfo>
<mods:name id="D487E993DA75E4CDF961E699792F3762" type="personal">
<mods:role id="5F2DF3523F3FDE3A1AE49356DF6DA871">
<mods:roleTerm id="4923A88ED81A9631CDB6209447A3749A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="25BF39097BE90103F5688765CD696E89">Shaverdo, Helena</mods:namePart>
</mods:name>
<mods:name id="0A577D07C5D12969FB3AB6CCE180790A" type="personal">
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<mods:namePart id="065199EA1400F8708C39EBF325BCACB1">Surbakti, Suriani</mods:namePart>
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<mods:namePart id="9C73F14BD64B6570691921A9D5A6477C">Warikar, Evie L.</mods:namePart>
</mods:name>
<mods:name id="A6F4E09DD3FAD96ACD20810442A40C2B" type="personal">
<mods:role id="F5F308C105D218B9579A07ED308E5738">
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<mods:namePart id="15D3177A23EE7B7D12C51B3D36134B37">Sagata, Katayo</mods:namePart>
</mods:name>
<mods:name id="9DA9BFD8C6C36211A7325705308EB11E" type="personal">
<mods:role id="6526AC834C01DFD125EB39CFBE4AA390">
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<mods:namePart id="726935CBD8453059EC2DB10A5C2226C6">Balke, Michael</mods:namePart>
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<mods:typeOfResource id="66FD044F83BC9A594028908D44BB8C9C">text</mods:typeOfResource>
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<mods:title id="027B6CD25EFD011A4919297D36E66FDD">ZooKeys</mods:title>
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<mods:part id="87F0BF1D4B0161E94191D37946A51F7F">
<mods:date id="5712D5B6C3786D97135D5D8264634F50">2019</mods:date>
<mods:detail id="B75BE19D5A807FAD9DC120EC7196AF72" type="volume">
<mods:number id="B1AAC3A7DF32A8411FD99A71121BA829">878</mods:number>
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<mods:start id="ED0F3569F8FD010B08C527A2942671A3">73</mods:start>
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<mods:url id="81C0A26EE0AFFAB85A47CCFA7730578B">http://dx.doi.org/10.3897/zookeys.878.37403</mods:url>
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<mods:classification id="D6051D51FE8CC1ABCBB4D38664B7B082">journal article</mods:classification>
<mods:identifier id="C0F4CB30671C07561ABD66D3E8A0D0D1" type="DOI">http://dx.doi.org/10.3897/zookeys.878.37403</mods:identifier>
<mods:identifier id="81DB0849A7427245B5A701B99B762F42" type="Pensoft-Pub">1313-2970-878-73</mods:identifier>
<mods:identifier id="DC7E57E688231F7AE5B4363E2E4D576C" type="ZooBank">192214DE1D38467BA577ECD16EC5EAB5</mods:identifier>
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<treatment id="733BAC0C78D75CBFBC8D78BA8FBAC482" ID-GBIF-Taxon="159654835" LSID="urn:lsid:plazi:treatment:733BAC0C78D75CBFBC8D78BA8FBAC482" httpUri="http://treatment.plazi.org/id/733BAC0C78D75CBFBC8D78BA8FBAC482" lastPageNumber="73" pageId="0" pageNumber="73" scope_family="Dytiscidae" scope_order="Coleoptera" scope_subFamily="Copelatinae">
<subSubSection id="6AB5F8DE58F97245DC8689BACC991B64" pageId="0" pageNumber="73" type="nomenclature">
<paragraph id="6DEE303C99A54ECF2F305260F2B4C882" pageId="0" pageNumber="73">
18.
<taxonomicName id="162B86F02D94F5420FDDA96CCDB880E9" ID-CoL="3DLRK" LSID="733BAC0C-78D7-5CBF-BC8D-78BA8FBAC482" authority="Balke, 1998" authorityName="Balke" authorityYear="1998" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina takime" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="takime">Exocelina takime (Balke, 1998)</taxonomicName>
<figureCitation id="8FE0F2348BE441704A1A10801E7A3389" captionStart="Figures 35–39" captionStartId="F18" captionText="Figures 35 - 39. Habitus and colouration 35 Exocelina larsoni (Balke, 1998) 36 E. nomax (J. Balfour-Browne, 1939) A holotype B specimen from Tapini 37 E. warahulenensis sp. nov. 38 E. mianminensis sp. nov. 39 E. takime (Balke, 1998)." figureDoi="10.3897/zookeys.878.37403.figures35-39" httpUri="https://binary.pensoft.net/fig/344986" pageId="0" pageNumber="73" tableDoi="10.3897/zookeys.878.37403.figures35-39">Figs 39</figureCitation>
,
<figureCitation id="49236C6EB4A89FF52C57665D8AC74731" captionStart="Figures 43, 44" captionStartId="F21" captionText="Figures 43, 44. 43 Exocelina mianminensis sp. nov. 44 E. takime (Balke, 1998) A median lobe in ventral view B median lobe in lateral view C paramere in external view D male protarsomeres 4 - 5 in ventral view." figureDoi="10.3897/zookeys.878.37403.figures43-44" httpUri="https://binary.pensoft.net/fig/344989" pageId="0" pageNumber="73" tableDoi="10.3897/zookeys.878.37403.figures43-44">44</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="0B27185E8F3D0DDE32308DD4FB2305F7" pageId="0" pageNumber="73" type="reference_group">
<paragraph id="865D347EFD0214130B2693DB017F5349" pageId="0" pageNumber="73">
<taxonomicName id="131FF073B92C74FE0415B009941B6684" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus (Papuadytes) takime" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="takime" subGenus="Papuadytes">Copelatus (Papuadytes) takime</taxonomicName>
Balke, 1998: 336;
<bibRefCitation id="E41A4E8766AB3C70B8DF139495465DC9" pageId="0" pageNumber="73" refId="B9">Nilsson 2001</bibRefCitation>
: 77 (catalogue).
</paragraph>
<paragraph id="8DC36EE31B0AAC762ED5250FCC8E9319" pageId="0" pageNumber="73">
<taxonomicName id="9D8526B395AEC52B4C7CFD242CA28AB2" class="Insecta" family="Dytiscidae" genus="Papuadytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Papuadytes takime" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="takime">Papuadytes takime</taxonomicName>
(Balke, 1998):
<bibRefCitation id="83208E10A7385A19529CC4B03BCA5E27" author="Nilsson, AN" journalOrPublisher="Koleopterologische Rundschau" pageId="0" pageNumber="73" pagination="55 - 74" refId="B11" refString="Nilsson, AN, Fery, H, 2006. World Catalogue of Dytiscidae - corrections and additions, 3 (Coleoptera: Dytiscidae). Koleopterologische Rundschau 76: 55 - 74" title="World Catalogue of Dytiscidae - corrections and additions, 3 (Coleoptera: Dytiscidae)." volume="76" year="2006">Nilsson and Fery 2006</bibRefCitation>
: 56 (comb. nov.).
</paragraph>
<paragraph id="A834FEF75F43255B5E1628FFADD48F06" pageId="0" pageNumber="73">
<taxonomicName id="F0842A3AE37D9AEE710165DED304354B" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina takime" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="takime">Exocelina takime</taxonomicName>
(Balke, 1998):
<bibRefCitation id="00D952CBF864E969EBCC22F720CBC64C" author="Nilsson, AN" journalOrPublisher="Latissimus" pageId="0" pageNumber="73" pagination="33 - 34" refId="B10" refString="Nilsson, AN, 2007. Exocelina Broun, 1886, is the valid name of Papuadytes Balke, 1998. Latissimus 23: 33 - 34" title="Exocelina Broun, 1886, is the valid name of Papuadytes Balke, 1998." volume="23" year="2007">Nilsson 2007</bibRefCitation>
: 34 (comb. nov.).
</paragraph>
</subSubSection>
<subSubSection id="7AACD5E81A067307F5BD9CED3E1E7776" pageId="0" pageNumber="73" type="type locality">
<paragraph id="42A1E454B62586B83418125E30EA2580" pageId="0" pageNumber="73">Type locality.</paragraph>
<paragraph id="FBD2D1ED5F42676A39808CF56C0CE34D" pageId="0" pageNumber="73">
Indonesia: Papua Province: Pegunungan Bintang Regency, Bime, ca.
<geoCoordinate id="CBFC4E34AD42237A2FF880048A64F7AA" degrees="04" direction="south" minutes="20" orientation="latitude" precision="925" value="-4.3333335">04°20'S</geoCoordinate>
,
<geoCoordinate id="38C4512DF8E6F2DFF8223BDCCBEF6B6A" degrees="140" direction="east" minutes="12" orientation="longitude" precision="925" value="140.2">140°12'E</geoCoordinate>
, 1400 m a.s.l.
</paragraph>
</subSubSection>
<subSubSection id="02D60C6422C286DB763806111A479DC2" pageId="0" pageNumber="73" type="type material">
<paragraph id="1064FD677B0024D17C3690BA4AB65A15" pageId="0" pageNumber="73">Type material studied.</paragraph>
<paragraph id="9E63210D93CF8637CD71EAA456A49CCF" pageId="0" pageNumber="73">
<emphasis id="1970F498D880A9DE43627A708CA2C873" italics="true" pageId="0" pageNumber="73">Holotype</emphasis>
: male &quot;IRIAN JAYA: 11.9.1993 Bime - Calab Gebiet, Bime, 1400m&quot;, &quot;leg. M. Balke (12) ca.
<geoCoordinate id="67CE0F8939E47E2537AFCF16971EF8A9" degrees="140" direction="east" minutes="12" orientation="longitude" precision="925" value="140.2">140°12'E</geoCoordinate>
<geoCoordinate id="B617B2E0F644619783F8F5018C245E12" degrees="04" direction="south" minutes="20" orientation="latitude" precision="925" value="-4.3333335">04°20'S</geoCoordinate>
&quot;,
<normalizedToken id="17EAF873AD78F98199A2680D7032518F" originalValue="“HOLOTYPUS”">&quot;HOLOTYPUS&quot;</normalizedToken>
[red], &quot;
<taxonomicName id="3C5F7F9780DF242C87A09E320E8A0E89" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
<taxonomicName id="44B13290F50CBE9167A92BA2B18C5DA8" lsidName="takime" pageId="0" pageNumber="73" rank="species" species="takime">takime</taxonomicName>
Balke des. 1997&quot; [red] (NHMW).
<emphasis id="FF4D212BAF976AAED2A21F0FFC898B2E" italics="true" pageId="0" pageNumber="73">Paratypes</emphasis>
: 10 males, 14 females with the same label as the holotype, one female with an additional green label &quot;DNA M.Balke 3291&quot; (NHMW). 11 males, 2 females &quot;IRIAN JAYA: 22.9.1993 Bime - Calab Gebiet, Bime, 1400m&quot;, &quot;ca.
<geoCoordinate id="1A31071341CA2857AF9B9C6B722DCCF4" degrees="140" direction="east" minutes="12" orientation="longitude" precision="925" value="140.2">140°12'E</geoCoordinate>
<geoCoordinate id="9F65E2DF8184453276C27C17272A0D4D" degrees="04" direction="south" minutes="20" orientation="latitude" precision="925" value="-4.3333335">04°20'S</geoCoordinate>
, leg. Balke (16)&quot;, one of the males with an additional green label &quot;DNA M.Balke 3290&quot; (NHMW). 3 males, 1 female &quot;IRIAN JAYA Zentralmassiv
<geoCoordinate id="A656837EC87B7F8BF15FB636C74378D4" degrees="140" direction="east" minutes="25" orientation="longitude" precision="925" value="140.41667">140°25'E</geoCoordinate>
<geoCoordinate id="F881CC7CD18ABBFA2AAA338AFF0C4548" degrees="04" direction="south" minutes="24" orientation="latitude" precision="925" value="-4.4">04°24'S</geoCoordinate>
&quot;, &quot;Kali Takime, 900m 18.8.1992 leg. Balke (17)&quot; (NHMW). 7 males, 6 females &quot;IRIAN JAYA: 29.9.1993 Eme Gebiet Emdoman, 800m&quot;, &quot;ca.
<geoCoordinate id="6207A60C4AAF9382078BFC37EFFB9587" degrees="139" direction="east" minutes="55" orientation="longitude" precision="925" value="139.91667">139°55'E</geoCoordinate>
<geoCoordinate id="B392E5849E76DFA177D5D97DCACE703A" degrees="04" direction="south" minutes="14" orientation="latitude" precision="925" value="-4.233333">04°14'S</geoCoordinate>
, leg. M. Balke (24)&quot; (NHMW). All these specimens are with red paratype labels &quot;PARATYPUS
<taxonomicName id="8185C310D7414EB72B917EFD012BF6F4" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
<taxonomicName id="E2BB6040CFE94D274077A64E46835D48" lsidName="takime" pageId="0" pageNumber="73" rank="species" species="takime">takime</taxonomicName>
Balke des. 1997&quot; or &quot;Paratypus
<taxonomicName id="FF3F2CF22346C1D09928D4C4C77042B1" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
<taxonomicName id="874119960C1552053052A5864F5DD572" lsidName="takime" pageId="0" pageNumber="73" rank="species" species="takime">takime</taxonomicName>
Balke des. 1997&quot; [red]. Note: in the original description (
<bibRefCitation id="35D3DE557E77185419F26B040C13816C" pageId="0" pageNumber="73" refId="B2">Balke 1998</bibRefCitation>
), 8 males and 4 females are reported from the locality 16; there are 11 males, 2 females in the NHMW, therefore, some mistakes were made in presenting the type material in the original description or/and during labeling the material.
</paragraph>
</subSubSection>
<subSubSection id="D1F12FF3FE3967AA46959B0406A26DBA" pageId="0" pageNumber="73" type="materials_examined">
<paragraph id="4754A3B8C946A85CBE3301CD2B5E5BD6" pageId="0" pageNumber="73">Additional material.</paragraph>
<paragraph id="40D3BDBFC6DFCB0C9CD5F2E3F472A917" pageId="0" pageNumber="73">
2 females &quot;IRIAN JAYA: 11.9.1993 Bime - Calab Gebiet, Bime, 1400m&quot;, &quot;leg. M. Balke (12) ca.
<geoCoordinate id="C44C44D0A8383C82C86FF8CE47D70A29" degrees="140" direction="east" minutes="12" orientation="longitude" precision="925" value="140.2">140°12'E</geoCoordinate>
<geoCoordinate id="4A2FEF00C854861BCECC1E057153C977" degrees="04" direction="south" minutes="20" orientation="latitude" precision="925" value="-4.3333335">04°20'S</geoCoordinate>
&quot; (NHMW).
</paragraph>
</subSubSection>
<subSubSection id="F5F861E2FA58FB7026702BDCFEE64083" pageId="0" pageNumber="73" type="diagnosis">
<paragraph id="498876F0BF38AD8F58DC4FD1130331BC" pageId="0" pageNumber="73">Diagnosis.</paragraph>
<paragraph id="1FA892714D03BF94A7F614A60325D7BD" pageId="0" pageNumber="73">
For complete description, see
<bibRefCitation id="D717268CBA08A81391817DF05628DBA6" pageId="0" pageNumber="73" refId="B2">Balke (1998</bibRefCitation>
: 336). Beetle medium-sized: TL-H 4.1-4.5 mm; oblong-oval; dark brown to piceous, with brownish pronotal sides and head anteriorly; shiny, but with distinct dorsal punctation and weakly impressed microreticulation; pronotum with narrow lateral bead (
<figureCitation id="42E79579369E37CECC962A3E4FBEA733" captionStart="Figures 35–39" captionStartId="F18" captionText="Figures 35 - 39. Habitus and colouration 35 Exocelina larsoni (Balke, 1998) 36 E. nomax (J. Balfour-Browne, 1939) A holotype B specimen from Tapini 37 E. warahulenensis sp. nov. 38 E. mianminensis sp. nov. 39 E. takime (Balke, 1998)." figureDoi="10.3897/zookeys.878.37403.figures35-39" httpUri="https://binary.pensoft.net/fig/344986" pageId="0" pageNumber="73" tableDoi="10.3897/zookeys.878.37403.figures35-39">Fig. 39</figureCitation>
); male protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta; male protarsomere 5 with anterior band of ca. 40 and posterior row of eight relatively long setae (
<figureCitation id="F5C5894F514AF2BA7FD7986B34B17C72" captionStart="Figures 43, 44" captionStartId="F21" captionText="Figures 43, 44. 43 Exocelina mianminensis sp. nov. 44 E. takime (Balke, 1998) A median lobe in ventral view B median lobe in lateral view C paramere in external view D male protarsomeres 4 - 5 in ventral view." figureDoi="10.3897/zookeys.878.37403.figures43-44" httpUri="https://binary.pensoft.net/fig/344989" pageId="0" pageNumber="73" tableDoi="10.3897/zookeys.878.37403.figures43-44">Fig. 44D</figureCitation>
); median lobe robust, in lateral view, evenly curved to narrowly rounded, not curved downwards apex; in ventral view, with more strongly thickened lateral sides, distinctly expanded in middle and narrowing to broadly pointed apex; paramere with distinct dorsal notch and subdistal part well developed, with long and dense subdistal and inconspicuous proximal setae (
<figureCitation id="ACD1880F597779B22B7905C10D85233B" captionStart="Figures 43, 44" captionStartId="F21" captionText="Figures 43, 44. 43 Exocelina mianminensis sp. nov. 44 E. takime (Balke, 1998) A median lobe in ventral view B median lobe in lateral view C paramere in external view D male protarsomeres 4 - 5 in ventral view." figureDoi="10.3897/zookeys.878.37403.figures43-44" httpUri="https://binary.pensoft.net/fig/344989" pageId="0" pageNumber="73" tableDoi="10.3897/zookeys.878.37403.figures43-44">
Fig. 44
<normalizedToken id="28F8461A1A316CFC5C1533AC8738C9B4" originalValue="A–C">A-C</normalizedToken>
</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="176DD390DFA61312F4C0E7942871D8BF" pageId="0" pageNumber="73" type="affinities">
<paragraph id="3A7C5A65A1DFCCBC4756AED62EACF451" pageId="0" pageNumber="73">Affinities.</paragraph>
<paragraph id="EBA744CCB395777B388B07A0F45909F3" pageId="0" pageNumber="73">
In the area of its distribution,
<taxonomicName id="956E29ADFE109F57B1EA4010C029379D" lsidName="E. takime" pageId="0" pageNumber="73" rank="species" species="takime">
<emphasis id="D37C5016AF536858B8FCFD2FBB749673" italics="true" pageId="0" pageNumber="73">E. takime</emphasis>
</taxonomicName>
co-occurs with
<taxonomicName id="F5F25F288D22B948803DA71A359BC98F" lsidName="E. aipomek" pageId="0" pageNumber="73" rank="species" species="aipomek">
<emphasis id="A46E123DD53FB7226790CE0F2C73ABEC" italics="true" pageId="0" pageNumber="73">E. aipomek</emphasis>
</taxonomicName>
,
<taxonomicName id="EB8301C1E186BF17580EED98BAF484D2" lsidName="E. ascendens" pageId="0" pageNumber="73" rank="species" species="ascendens">
<emphasis id="E7B90EB656277C1053E2CCD69C7A13F3" italics="true" pageId="0" pageNumber="73">E. ascendens</emphasis>
</taxonomicName>
and species of the
<taxonomicName id="A2955AC327CB1370212CE15CF67AD7D6" lsidName="E. bacchusi" pageId="0" pageNumber="73" rank="species" species="bacchusi">
<emphasis id="2464C6D0C4985949D5DCCBEC55F61517" italics="true" pageId="0" pageNumber="73">E. bacchusi</emphasis>
</taxonomicName>
,
<taxonomicName id="76CFE8235076CD1D3A40FDE05DBF698F" lsidName="E. ekari" pageId="0" pageNumber="73" rank="species" species="ekari">
<emphasis id="E57D2D462E7B1F72D71222BB8CBC011E" italics="true" pageId="0" pageNumber="73">E. ekari</emphasis>
</taxonomicName>
,
<taxonomicName id="714C6A9A6D1483B5593001FE11CEC6B9" lsidName="E. aipo" pageId="0" pageNumber="73" rank="species" species="aipo">
<emphasis id="FA1EB31671E1971AD0E26E22EDF4EB0C" italics="true" pageId="0" pageNumber="73">E. aipo</emphasis>
</taxonomicName>
,
<taxonomicName id="0FDB08BEFE49B9444483EAA8BC627C24" lsidName="E. okbapensis" pageId="0" pageNumber="73" rank="species" species="okbapensis">
<emphasis id="DE6AC96062DB4720839D15F911142A31" italics="true" pageId="0" pageNumber="73">E. okbapensis</emphasis>
</taxonomicName>
,
<taxonomicName id="D540071BB2E760F5E5BFD2A7EE121E02" lsidName="E. casuarina" pageId="0" pageNumber="73" rank="species" species="casuarina">
<emphasis id="CD9758394246B822E6A2178349C92A31" italics="true" pageId="0" pageNumber="73">E. casuarina</emphasis>
</taxonomicName>
, and
<taxonomicName id="227B955017FC02BD34CF37527D8BBC2A" lsidName="E. danae" pageId="0" pageNumber="73" rank="species" species="danae">
<emphasis id="FCE8AA8B05A3B4E4D18194EB5AF0080B" italics="true" pageId="0" pageNumber="73">E. danae</emphasis>
</taxonomicName>
groups. From species of the
<taxonomicName id="D9063024B9E6FC037C62224767BD4367" lsidName="E. ekari" pageId="0" pageNumber="73" rank="species" species="ekari">
<emphasis id="7FB44EEAEFE307BBDE24A9BE8A857C2A" italics="true" pageId="0" pageNumber="73">E. ekari</emphasis>
</taxonomicName>
group, the species differs in larger size, evidently stronger dorsal punctation, and the shape of the median lobe. In the latter character,
<taxonomicName id="461518A536655D4B774DB827B10B4BC0" lsidName="E. takime" pageId="0" pageNumber="73" rank="species" species="takime">
<emphasis id="54666D74FB3F34F703E6046696939168" italics="true" pageId="0" pageNumber="73">E. takime</emphasis>
</taxonomicName>
differs also from the species of the remaining groups. For separating it from some of these species, also presence of the pronotal bead and simple male antennae, and shape and setation of the paramere can be used. For affinities within the group, see the
<normalizedToken id="449479580FACE8B701DFB878AE41A429" originalValue="“Key”">&quot;Key&quot;</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection id="1A7F12A59404B24B6BBEC4A10B2CA91C" pageId="0" pageNumber="73" type="distribution">
<paragraph id="D6DF6041BFADE4E3C6F1AC50C782D3C2" pageId="0" pageNumber="73">Distribution.</paragraph>
<paragraph id="923126954CC92A6E21AF0E20E1DE2761" pageId="0" pageNumber="73">
Indonesia: Papua Province: Pegunungan Bintang Regency. The species is known only from the type material, i.e., Borme - Bime - Emdoman area (
<figureCitation id="7393C2953CBD81F2E634DE63552C096F" captionStart="Figure 45" captionStartId="F22" captionText="Figure 45. Map of the eastern part of New Guinea showing distribution of the species of the E. larsoni and takime groups." figureDoi="10.3897/zookeys.878.37403.figure45" httpUri="https://binary.pensoft.net/fig/344990" pageId="0" pageNumber="73" tableDoi="10.3897/zookeys.878.37403.figure45">Fig. 45</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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