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<document id="DF463507C092C1BE3C397FD9AB9BC1C0" ID-CLB-Dataset="28984" ID-DOI="10.11646/zootaxa.4461.2.11" ID-GBIF-Dataset="f86c0afe-5fbf-4b97-b6b6-16dece45e39f" ID-ISSN="1175-5326" ID-Zenodo-Dep="1460010" ID-ZooBank="93CE22AF-840A-44D4-BC3B-B411F4569B47" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539336697524" checkinUser="plazi" docAuthor="Janák, Jiří" docDate="2018" docId="597EC221FFAFFF81FF410ED4099C11E0" docLanguage="en" docName="zootaxa.4461.2.11.pdf" docOrigin="Zootaxa 4461 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tympanophorus Nordmann 1837" docType="key" docVersion="6" lastPageNumber="295" masterDocId="A547BA59FFADFF83FFD60E4C0C001215" masterDocTitle="A new genus of the Tympanophorus lineage of the subtribe Anisolinina from Madagascar (Coleoptera: Staphylinidae: Staphylininae: Staphylinini)" masterLastPageNumber="299" masterPageNumber="293" pageNumber="295" updateTime="1698579679398" updateUser="ExternalLinkService">
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<mods:title id="A3E7E47E02125B54120F9D22A63FCF1F">A new genus of the Tympanophorus lineage of the subtribe Anisolinina from Madagascar (Coleoptera: Staphylinidae: Staphylininae: Staphylinini)</mods:title>
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<mods:date id="E64C9DC566FD54DB7897080EE722A4AA">2018</mods:date>
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<treatment id="597EC221FFAFFF81FF410ED4099C11E0" ID-DOI="http://doi.org/10.5281/zenodo.5952400" ID-GBIF-Taxon="148701886" ID-Zenodo-Dep="5952400" LSID="urn:lsid:plazi:treatment:597EC221FFAFFF81FF410ED4099C11E0" httpUri="http://treatment.plazi.org/id/597EC221FFAFFF81FF410ED4099C11E0" lastPageNumber="295" pageId="2" pageNumber="295">
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Key to genera of the
<taxonomicName id="16D708B4FFAFFF81FE4C0ED40E5F12A7" ID-CoL="8VS6C" authorityName="Nordmann" authorityYear="1837" box="[410,607,152,178]" class="Insecta" family="Staphylinidae" genus="Tympanophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="E3A3AF25FFAFFF81FE4C0ED40E5F12A7" bold="true" box="[410,607,152,178]" italics="true" pageId="2" pageNumber="295">Tympanophorus</emphasis>
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lineage
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<paragraph id="D1687337FFAFFF81FF410EF60E5612C6" blockId="2.[151,703,152,211]" box="[151,598,186,211]" pageId="2" pageNumber="295">
(based on
<bibRefCitation id="B5460EC6FFAFFF81FEDC0EF60DDD12C6" box="[266,477,186,211]" pageId="2" pageNumber="295" refId="ref3599" refString="Brunke, A. J., Schillhammer, H. &amp; Chatzimanolis, S. (2017) The first fossil rove beetle from the middle Eocene Kishenehn Formation (North America) provides evidence for ancient Eocene relicts within the hyperdiverse Staphylinini (Coleoptera: Staphylinidae: Staphylininae). Journal of Systematic Palaeontology, 15 (12), 1015 - 1025. https: // doi. org / 10.1080 / 14772019.2016.1266402" type="journal article">
Brunke
<emphasis id="E3A3AF25FFAFFF81FEB50EF00D9412C6" box="[355,404,186,211]" italics="true" pageId="2" pageNumber="295">et al</emphasis>
. 2017
</bibRefCitation>
, modified)
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1 Apical labial palpomere slender, fusiform...............................................................................................
<taxonomicName id="16D708B4FFAFFF81FDEE0F530F4A1320" ID-CoL="8W48C" authority="Bernhauer, 1923" authorityName="Bernhauer" authorityYear="1923" box="[568,842,287,309]" class="Insecta" family="Staphylinidae" genus="Tolmerinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="E3A3AF25FFAFFF81FDEE0F530EA31321" box="[568,675,287,308]" italics="true" pageId="2" pageNumber="295">Tolmerinus</emphasis>
<bibRefCitation id="B5460EC6FFAFFF81FD7F0F530F4A1320" box="[681,842,287,309]" pageId="2" pageNumber="295" refId="ref3820" refString="Bernhauer, M. (1923) Synonymische Bemerkungen bezuglich mehrfach beschriebener Staphylinidengattungen and Arten. Wiener Entomologische Zeitung, 40, 63. https: // doi. org / 10.5962 / bhl. part. 2586" type="journal article">Bernhauer, 1923</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="16D708B4FFAFFF81FC820F5308641320" ID-CoL="8X79Q" authority="Bernhauer, 1911" authorityName="Bernhauer" authorityYear="1911" box="[852,1124,287,309]" class="Insecta" family="Staphylinidae" genus="Diatrechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="E3A3AF25FFAFFF81FC820F530FBE1321" box="[852,958,287,308]" italics="true" pageId="2" pageNumber="295">Diatrechus</emphasis>
<bibRefCitation id="B5460EC6FFAFFF81FC120F5308641320" box="[964,1124,287,309]" pageId="2" pageNumber="295" refId="ref3763" refString="Bernhauer, M. (1911) Zur Staphylinidenfauna Ostindiens und der Sundainseln. Entomologische Blatter, 7, 55 - 62, 86 - 93." type="journal article">Bernhauer, 1911</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="16D708B4FFAFFF81FBB80F53099B1320" ID-CoL="8XNL8" authority="Cameron, 1944" authorityName="Cameron" authorityYear="1944" box="[1134,1435,287,309]" class="Insecta" family="Staphylinidae" genus="Paradiatrechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="E3A3AF25FFAFFF81FBB80F5309021321" box="[1134,1282,287,308]" italics="true" pageId="2" pageNumber="295">Paradiatrechus</emphasis>
<bibRefCitation id="B5460EC6FFAFFF81FADE0F53099B1320" box="[1288,1435,287,309]" pageId="2" pageNumber="295" refId="ref3858" refString="Cameron, M. (1944) New oriental Staphylinidae (Col.). The Annals and Magazine of Natural History, Series 11, 11 (77), 312 - 322. https: // doi. org / 10.1080 / 00222934408527435" type="journal article">Cameron, 1944</bibRefCitation>
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<paragraph id="D1687337FFAFFF81FF410F71099B137A" blockId="2.[151,1437,257,1013]" pageId="2" pageNumber="295">
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- Apical labial palpomere distinctly dilated apically and almost always securiform (
<figureCitation id="49EC6FB2FFAFFF81FBD60F70085C1347" box="[1024,1116,316,338]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1969,1991]" captionTargetBox="[151,1430,348,1884]" captionTargetId="figure@3.[151,1435,339,1944]" captionTargetPageId="3" captionText="FIGURES 1–6. Janalia jarmilae gen. nov., sp. nov. 1, habitus; 2, forebody; 3, head ventral; 4, head ventro-lateral; 5, mouthparts dorsal; 6, mouthparts ventral." httpUri="https://zenodo.org/record/1460012/files/figure.png" pageId="2" pageNumber="295">Figs. 4–6</figureCitation>
and
<figureCitation id="49EC6FB2FFAFFF81FB580F7008FA1347" box="[1166,1274,316,338]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1969,1991]" captionTargetBox="[151,1430,348,1884]" captionTargetId="figure@3.[151,1435,339,1944]" captionTargetPageId="3" captionText="FIGURES 1–6. Janalia jarmilae gen. nov., sp. nov. 1, habitus; 2, forebody; 3, head ventral; 4, head ventro-lateral; 5, mouthparts dorsal; 6, mouthparts ventral." httpUri="https://zenodo.org/record/1460012/files/figure.png" pageId="2" pageNumber="295">Figs. 4E, F</figureCitation>
in
<bibRefCitation id="B5460EC6FFAFFF81FACA0F700D13137A" pageId="2" pageNumber="295" refId="ref3599" refString="Brunke, A. J., Schillhammer, H. &amp; Chatzimanolis, S. (2017) The first fossil rove beetle from the middle Eocene Kishenehn Formation (North America) provides evidence for ancient Eocene relicts within the hyperdiverse Staphylinini (Coleoptera: Staphylinidae: Staphylininae). Journal of Systematic Palaeontology, 15 (12), 1015 - 1025. https: // doi. org / 10.1080 / 14772019.2016.1266402" type="journal article">
Brunke
<emphasis id="E3A3AF25FFAFFF81FABC0F71099D1347" box="[1386,1437,317,338]" italics="true" pageId="2" pageNumber="295">et al.</emphasis>
2017
</bibRefCitation>
)............................................................................................... 2
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<keyStep id="6A236B92FFAFFF81FF410F3B099B13BF" pageId="2" pageNumber="295">
<paragraph id="D1687337FFAFFF81FF410F3B099B1398" blockId="2.[151,1437,257,1013]" box="[151,1435,375,397]" pageId="2" pageNumber="295">
<keyLead id="6A26D002FFAFFF81FF410F3B099B1398" box="[151,1435,375,397]" pageId="2" pageNumber="295">
2 Apical labial palpomere laterally flattened with narrow apical face (
<figureCitation id="49EC6FB2FFAFFF81FCBC0F3B0FB11398" box="[874,945,375,397]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1969,1991]" captionTargetBox="[151,1430,348,1884]" captionTargetId="figure@3.[151,1435,339,1944]" captionTargetPageId="3" captionText="FIGURES 1–6. Janalia jarmilae gen. nov., sp. nov. 1, habitus; 2, forebody; 3, head ventral; 4, head ventro-lateral; 5, mouthparts dorsal; 6, mouthparts ventral." httpUri="https://zenodo.org/record/1460012/files/figure.png" pageId="2" pageNumber="295">Fig. 4E</figureCitation>
in
<bibRefCitation id="B5460EC6FFAFFF81FC060F3B08841398" box="[976,1156,375,397]" pageId="2" pageNumber="295" refId="ref3599" refString="Brunke, A. J., Schillhammer, H. &amp; Chatzimanolis, S. (2017) The first fossil rove beetle from the middle Eocene Kishenehn Formation (North America) provides evidence for ancient Eocene relicts within the hyperdiverse Staphylinini (Coleoptera: Staphylinidae: Staphylininae). Journal of Systematic Palaeontology, 15 (12), 1015 - 1025. https: // doi. org / 10.1080 / 14772019.2016.1266402" type="journal article">
Brunke
<emphasis id="E3A3AF25FFAFFF81FBCB0F3B08471399" box="[1053,1095,375,396]" italics="true" pageId="2" pageNumber="295">et al</emphasis>
. 2017
</bibRefCitation>
)..................... 3
</keyLead>
</paragraph>
<paragraph id="D1687337FFAFFF81FF410FD9099B13BF" blockId="2.[151,1437,257,1013]" box="[151,1435,404,426]" pageId="2" pageNumber="295">
<keyLead id="6A26D002FFAFFF81FF410FD9099B13BF" box="[151,1435,404,426]" pageId="2" pageNumber="295">
- Apical labial palpomere not laterally flattened, with broad apical face (
<figureCitation id="49EC6FB2FFAFFF81FC510FD80FE313BF" box="[903,995,404,426]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1969,1991]" captionTargetBox="[151,1430,348,1884]" captionTargetId="figure@3.[151,1435,339,1944]" captionTargetPageId="3" captionText="FIGURES 1–6. Janalia jarmilae gen. nov., sp. nov. 1, habitus; 2, forebody; 3, head ventral; 4, head ventro-lateral; 5, mouthparts dorsal; 6, mouthparts ventral." httpUri="https://zenodo.org/record/1460012/files/figure.png" pageId="2" pageNumber="295">Figs. 4–6</figureCitation>
)................................... 4
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<keyStep id="6A236B92FFAFFF81FF410FFD099B104F" pageId="2" pageNumber="295">
<paragraph id="D1687337FFAFFF81FF410FFD099B1017" blockId="2.[151,1437,257,1013]" pageId="2" pageNumber="295">
<keyLead id="6A26D002FFAFFF81FF410FFD099B1017" pageId="2" pageNumber="295">
3 Labrum expanded, lobes less than twice as wide as long (
<figureCitation id="49EC6FB2FFAFFF81FCD30FFD0F4D13D2" box="[773,845,433,455]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1969,1991]" captionTargetBox="[151,1430,348,1884]" captionTargetId="figure@3.[151,1435,339,1944]" captionTargetPageId="3" captionText="FIGURES 1–6. Janalia jarmilae gen. nov., sp. nov. 1, habitus; 2, forebody; 3, head ventral; 4, head ventro-lateral; 5, mouthparts dorsal; 6, mouthparts ventral." httpUri="https://zenodo.org/record/1460012/files/figure.png" pageId="2" pageNumber="295">Fig. 4E</figureCitation>
in
<bibRefCitation id="B5460EC6FFAFFF81FCBD0FFD082213D2" box="[875,1058,433,455]" pageId="2" pageNumber="295" refId="ref3599" refString="Brunke, A. J., Schillhammer, H. &amp; Chatzimanolis, S. (2017) The first fossil rove beetle from the middle Eocene Kishenehn Formation (North America) provides evidence for ancient Eocene relicts within the hyperdiverse Staphylinini (Coleoptera: Staphylinidae: Staphylininae). Journal of Systematic Palaeontology, 15 (12), 1015 - 1025. https: // doi. org / 10.1080 / 14772019.2016.1266402" type="journal article">
Brunke
<emphasis id="E3A3AF25FFAFFF81FC6E0FFE0FE213D2" box="[952,994,434,455]" italics="true" pageId="2" pageNumber="295">et al</emphasis>
. 2017
</bibRefCitation>
); apical maxillary palpomere weakly to moderately expanded subapically or apically, apex never pointed (
<figureCitation id="49EC6FB2FFAFFF81FC8A0F830FBF13F1" box="[860,959,463,485]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1969,1991]" captionTargetBox="[151,1430,348,1884]" captionTargetId="figure@3.[151,1435,339,1944]" captionTargetPageId="3" captionText="FIGURES 1–6. Janalia jarmilae gen. nov., sp. nov. 1, habitus; 2, forebody; 3, head ventral; 4, head ventro-lateral; 5, mouthparts dorsal; 6, mouthparts ventral." httpUri="https://zenodo.org/record/1460012/files/figure.png" pageId="2" pageNumber="295">Fig. 4B, E</figureCitation>
in
<bibRefCitation id="B5460EC6FFAFFF81FC080F83089513F0" box="[990,1173,463,485]" pageId="2" pageNumber="295" refId="ref3599" refString="Brunke, A. J., Schillhammer, H. &amp; Chatzimanolis, S. (2017) The first fossil rove beetle from the middle Eocene Kishenehn Formation (North America) provides evidence for ancient Eocene relicts within the hyperdiverse Staphylinini (Coleoptera: Staphylinidae: Staphylininae). Journal of Systematic Palaeontology, 15 (12), 1015 - 1025. https: // doi. org / 10.1080 / 14772019.2016.1266402" type="journal article">
Brunke
<emphasis id="E3A3AF25FFAFFF81FBFD0F83085513F1" box="[1067,1109,463,484]" italics="true" pageId="2" pageNumber="295">et al</emphasis>
. 2017
</bibRefCitation>
); pronotum punctate across entire disc....................................................................
<taxonomicName id="16D708B4FFAFFF81FBAA0FA1099B1017" ID-CoL="8XJWX" authority="Bernhauer, 1902" authorityName="Bernhauer" authorityYear="1902" box="[1148,1435,492,514]" class="Insecta" family="Staphylinidae" genus="Barygnathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="E3A3AF25FFAFFF81FBAA0FA108F61017" box="[1148,1270,493,514]" italics="true" pageId="2" pageNumber="295">Barygnathus</emphasis>
<bibRefCitation id="B5460EC6FFAFFF81FB2A0FA0099B1017" box="[1276,1435,492,514]" pageId="2" pageNumber="295" refId="ref3741" refString="Bernhauer, M. (1902) Zur Staphyliniden-Fauna von Ceylon. Deutsche Entomologische Zeitschrift, 1902, 17 - 45." type="journal article">Bernhauer, 1902</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="D1687337FFAFFF81FF410C46099B104F" blockId="2.[151,1437,257,1013]" pageId="2" pageNumber="295">
<keyLead id="6A26D002FFAFFF81FF410C46099B104F" pageId="2" pageNumber="295">
<emphasis id="E3A3AF25FFAFFF81FF410C460C9F100A" box="[151,159,522,543]" italics="true" pageId="2" pageNumber="295">-</emphasis>
Labrum not expanded, lobes distinctly more than twice as wide as long (as in
<figureCitation id="49EC6FB2FFAFFF81FC1B0C450845100A" box="[973,1093,521,543]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1969,1991]" captionTargetBox="[151,1430,348,1884]" captionTargetId="figure@3.[151,1435,339,1944]" captionTargetPageId="3" captionText="FIGURES 1–6. Janalia jarmilae gen. nov., sp. nov. 1, habitus; 2, forebody; 3, head ventral; 4, head ventro-lateral; 5, mouthparts dorsal; 6, mouthparts ventral." httpUri="https://zenodo.org/record/1460012/files/figure.png" pageId="2" pageNumber="295">Figs. 2C, 4F</figureCitation>
in
<bibRefCitation id="B5460EC6FFAFFF81FBB50C45091A100A" box="[1123,1306,521,543]" pageId="2" pageNumber="295" refId="ref3599" refString="Brunke, A. J., Schillhammer, H. &amp; Chatzimanolis, S. (2017) The first fossil rove beetle from the middle Eocene Kishenehn Formation (North America) provides evidence for ancient Eocene relicts within the hyperdiverse Staphylinini (Coleoptera: Staphylinidae: Staphylininae). Journal of Systematic Palaeontology, 15 (12), 1015 - 1025. https: // doi. org / 10.1080 / 14772019.2016.1266402" type="journal article">
Brunke
<emphasis id="E3A3AF25FFAFFF81FB660C4608DA100A" box="[1200,1242,522,543]" italics="true" pageId="2" pageNumber="295">et al</emphasis>
. 2017
</bibRefCitation>
); apical maxillary palpomere evenly converging posteriad to pointed apex (
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in
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Brunke
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); pronotum with median impunctate line.........................................................................
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4 Apical maxillary palpomere fusiform, not conspicuously constricted apically, at least twice as long as previous segment (
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).......................................................................... 5
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- Apical maxillary palpomere conspicuously constricted apically, at most slightly longer than previous segment (
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).................................................................................... 7
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5 Head with temples nearly straight and strongly converging to neck; body with areas of long golden pubescence (Fig.
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); male unknown..................................................
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- Head with temples rounded or only slightly converging to neck (
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Brunke
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); body without conspicuous areas of golden pubescence....................................................................... 6
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6 Paramere with long apex and distinct expansion at midlength (
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Figs.
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–19 in
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)...
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- Paramere short and broad (
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)......................................................
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7 Ventral surface of head with gular setae approximate (
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Brunke
<emphasis id="E3A3AF25FFAFFF81FC4A0DCB0FC61189" box="[924,966,903,924]" italics="true" pageId="2" pageNumber="295">et al</emphasis>
. 2017
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); prosternum not projected ventrad..................................................................................
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- Ventral surface of head with gular setae distant (
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in
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Brunke
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. 2017
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); pronotum projected ventrad.........................................................................................
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Nordmann, 1837
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