390 lines
48 KiB
XML
390 lines
48 KiB
XML
<document id="DF5DE4E03FAB7817144CC1D6D1890C4F" ID-DOI="10.35463/j.apr.2022.01.04" ID-ISSN="1842-371x" ID-Zenodo-Dep="10520851" IM.bibliography_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1705451523398" checkinUser="felipe" docAuthor="Iamandei, Stanila, Iamandei, Eugenia & Grădinaru, Eugen" docDate="2022" docId="341887851D48FFBF948FE2E7FAE058A9" docLanguage="en" docName="ActaPalaeontolRom.18.1.25-48.pdf" docOrigin="Acta Palaeontologica Romaniae 18 (1)" docSource="http://dx.doi.org/10.35463/j.apr.2022.01.04" docStyle="DocumentStyle:CA474AAD92E97C533D63E0C5C0F159F1.1:ActaPalaeontolRom.2019-.journal_article" docStyleId="CA474AAD92E97C533D63E0C5C0F159F1" docStyleName="ActaPalaeontolRom.2019-.journal_article" docStyleVersion="1" docTitle="Protojuniperoxylon holbavicum Iamandei & Iamandei & Grădinaru 2022, sp. nov." docType="treatment" docVersion="3" lastPageNumber="31" masterDocId="C821FFFD1D4CFFB99479E069FFD45A6E" masterDocTitle="Contributions To The Study Of The Early Jurassic Petrified Forest Of Holbav And Cristian Areas (Brașov Region, South Carpathians, Romania), 2 Part." masterLastPageNumber="48" masterPageNumber="25" pageNumber="29" updateTime="1713182580132" updateUser="juliana" zenodo-license-document="CC-BY-4.0">
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<mods:title id="ABDF95AF9FF341C54712C0EEEA0DD862">Contributions To The Study Of The Early Jurassic Petrified Forest Of Holbav And Cristian Areas (Brașov Region, South Carpathians, Romania), 2 Part.</mods:title>
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<mods:namePart id="F4B21A68F587A9AC0B71D9A095C449B0">Iamandei, Stanila</mods:namePart>
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<mods:affiliation id="30BBD6C2BC27718E7C6584899DA48F33">Geological Institute of Romania, 1 st, Caransebeş Street, 012271 – Bucharest, Romania.</mods:affiliation>
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<mods:namePart id="EE1470790350FEE9FCBB6EC23F37E453">Iamandei, Eugenia</mods:namePart>
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<mods:affiliation id="0FD0D45376B77F9938AF3FB22C466532">Geological Institute of Romania, 1 st, Caransebeş Street, 012271 - Bucharest, Romania. iamandei @ gmail. com</mods:affiliation>
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<mods:nameIdentifier id="7790285EE97BA4B7C31B4FA788968903" type="email">iamandei@gmail.com</mods:nameIdentifier>
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<mods:namePart id="09725ADD6A19F95D35FB616D42AF026D">Grădinaru, Eugen</mods:namePart>
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<mods:affiliation id="F8FB9E2608B721FEEE8C47B4271E364B">University of Bucharest, Department of Geology, 1, Bălcescu Nicolae Blvd., 010041, Bucharest, Romania. egradinaru @ gmail. com</mods:affiliation>
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<mods:nameIdentifier id="5466A50B4BEF98FC2C7F09D75499830A" type="email">egradinaru@gmail.com</mods:nameIdentifier>
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<mods:title id="6203AE00BC0860724D3064FAEE42C71B">Acta Palaeontologica Romaniae</mods:title>
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<mods:date id="F780BBE8097AA043E9AA7C841C04AFA0">2022</mods:date>
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<treatment id="341887851D48FFBF948FE2E7FAE058A9" LSID="urn:lsid:plazi:treatment:341887851D48FFBF948FE2E7FAE058A9" httpUri="http://treatment.plazi.org/id/341887851D48FFBF948FE2E7FAE058A9" lastPageId="6" lastPageNumber="31" pageId="4" pageNumber="29">
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<subSubSection id="F4AB65181D48FFBD948FE2E7FD7158C8" box="[246,677,654,678]" pageId="4" pageNumber="29" type="nomenclature">
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<paragraph id="BC0E36931D48FFBD948FE2E7FD7158C8" blockId="4.[246,677,654,711]" box="[246,677,654,678]" pageId="4" pageNumber="29">
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<heading id="E74681FF1D48FFBD948FE2E7FD7158C8" box="[246,677,654,678]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="29" reason="5">
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<taxonomicName id="7BB14D101D48FFBD948FE2E7FD9D58C8" authority="Iamandei & Iamandei & Grădinaru, 2022" authorityName="Iamandei & Iamandei & Grădinaru" authorityYear="2022" box="[246,585,654,678]" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="29" phylum="Tracheophyta" rank="species" species="holbavicum" status="sp. nov.">
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<emphasis id="8EC5EA811D48FFBD948FE2E7FD9D58C8" box="[246,585,654,678]" italics="true" pageId="4" pageNumber="29">Protojuniperoxylon holbavicum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="95F657FA1D48FFBD9629E2E6FD7158C8" box="[592,677,655,678]" pageId="4" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<subSubSection id="F4AB65181D48FFBD95EFE2C6FDD158A9" box="[406,517,687,711]" pageId="4" pageNumber="29" type="description">
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<paragraph id="BC0E36931D48FFBD95EFE2C6FDD158A9" blockId="4.[246,677,654,711]" box="[406,517,687,711]" pageId="4" pageNumber="29">
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<figureCitation id="248A2A161D48FFBD95EFE2C6FE0258A9" box="[406,470,687,711]" captionStart="Fig" captionStartId="5.[169,204,1925,1946]" captionTargetBox="[169,1412,154,1916]" captionTargetId="figure-19@5.[169,1412,154,1916]" captionTargetPageId="5" captionText="Fig. 2 Protojuniperoxylon holbavicum sp. nov. (inventory no. 27706). a-c: Cross section - tracheids polygonal with rounded corners, and linear rays; d-f: Tangential section - uniseriate rays with biseriate storeys with polygonal ray-cells, with lateral empty spaces; unpitted tracheids; g-j: Radial section –uniseriate radial pitting on tracheids as hexagonal pits; araucarioid cross-fields with vertical pairs of pits or slightly irregular; end-wall with juniperoid thickenings (see arrow)." figureDoi="http://doi.org/10.5281/zenodo.10520855" httpUri="https://zenodo.org/record/10520855/files/figure.png" pageId="4" pageNumber="29">Fig. 2</figureCitation>
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, a-j.
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</paragraph>
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</subSubSection>
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<subSubSection id="F4AB65181D48FFBD94EEE29BFEE55ED6" pageId="4" pageNumber="29" type="materials_examined">
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<paragraph id="BC0E36931D48FFBD94EEE29BFF2E5964" blockId="4.[151,771,754,1208]" box="[151,250,754,778]" pageId="4" pageNumber="29">
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<emphasis id="8EC5EA811D48FFBD94EEE29BFF2E5964" bold="true" box="[151,250,754,778]" pageId="4" pageNumber="29">Material</emphasis>
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</paragraph>
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<paragraph id="BC0E36931D48FFBD94EEE37AFEE55ED6" blockId="4.[151,771,754,1208]" pageId="4" pageNumber="29">
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<materialsCitation id="0CD93CCE1D48FFBD94EEE37AFEF95ED6" collectionCode="NMG" location="Holbav" municipality="Maiului" pageId="4" pageNumber="29" specimenCode="27704, 27705, 27706, 27707, 27708, 27709, 27710" specimenCount="7">
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The studied material is represented by
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<specimenCount id="AAB7FD1A1D48FFBD964EE37AFD905944" box="[567,580,787,810]" pageId="4" pageNumber="29">7</specimenCount>
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samples of silicified wood, collected from the
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<location id="B96E60481D48FFBD9589E35DFDEB5922" LSID="urn:lsid:plazi:treatment:341887851D48FFBF948FE2E7FAE058A9:B96E60481D48FFBD9589E35DFDEB5922" box="[496,575,820,844]" municipality="Maiului" name="Holbav" pageId="4" pageNumber="29">Holbav</location>
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area, on
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<collectingMunicipality id="5C6AACE91D48FFBD96D6E35DFCD75922" box="[687,771,820,844]" pageId="4" pageNumber="29">Maiului</collectingMunicipality>
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brook. All studied samples are fragments of trunk or thick branches, of decimetric size, dark to black color. Viewing through a hand lens, or even by naked eye, it is possible to see the regular fibrous structure without vessels, which indicate a conifer wood. The specimens attributed to this species have very similar details and have the following field numbers within the "Grădinaru Collection": 1078, 1083, 1085, 1087, 1091, 1095, 1110 – and are stored at the
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<collectionCode id="DAA0AE561D48FFBD94C7E434FDD45E1B" box="[190,512,1117,1141]" pageId="4" pageNumber="29">National Museum of Geology</collectionCode>
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, in Bucharest, under the inventory numbers:
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<specimenCode id="EC179EE81D48FFBD950BE416FE6C5EF9" box="[370,440,1151,1175]" pageId="4" pageNumber="29">27704</specimenCode>
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,
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<specimenCode id="EC179EE81D48FFBD95BAE416FDDC5EF9" box="[451,520,1151,1175]" pageId="4" pageNumber="29">27705</specimenCode>
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,
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<specimenCode id="EC179EE81D48FFBD966CE416FD8F5EF9" box="[533,603,1151,1175]" pageId="4" pageNumber="29">27706</specimenCode>
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,
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<specimenCode id="EC179EE81D48FFBD961EE416FD785EF9" box="[615,684,1151,1175]" pageId="4" pageNumber="29">27707</specimenCode>
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,
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<specimenCode id="EC179EE81D48FFBD96C0E416FD2A5EF9" box="[697,766,1151,1175]" pageId="4" pageNumber="29">27708</specimenCode>
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,
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<specimenCode id="EC179EE81D48FFBD94EEE4C9FF085ED6" box="[151,220,1184,1208]" pageId="4" pageNumber="29">27709</specimenCode>
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,
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<specimenCode id="EC179EE81D48FFBD949EE4C9FEF95ED6" box="[231,301,1184,1208]" pageId="4" pageNumber="29">27710</specimenCode>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="F4AB65181D48FFBD94EEE48BFB655EB7" pageId="4" pageNumber="29" type="description">
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<paragraph id="BC0E36931D48FFBD94EEE48BFE7C5E94" blockId="4.[151,772,1250,2068]" box="[151,424,1250,1274]" pageId="4" pageNumber="29">
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<emphasis id="8EC5EA811D48FFBD94EEE48BFE7C5E94" bold="true" box="[151,424,1250,1274]" pageId="4" pageNumber="29">Microscopic description</emphasis>
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</paragraph>
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<paragraph id="BC0E36931D48FFBD94EEE56AFEDE5FAF" blockId="4.[151,772,1250,2068]" pageId="4" pageNumber="29">
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<emphasis id="8EC5EA811D48FFBD94EEE56AFEFF5F75" box="[151,299,1283,1307]" italics="true" pageId="4" pageNumber="29">Growth rings</emphasis>
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– well developed in the studied samples, representing the secondary xylem which is tracheidoxylic, showing a fairly gradual transition but sometimes abrupt, from early- to late-wood, and has distinct growth ring boundaries. Normal or traumatic axial resin canals are absent.
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</paragraph>
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<paragraph id="BC0E36931D48FFBD94EEE5A3FBE65A96" blockId="4.[151,772,1250,2068]" lastBlockId="4.[816,1437,163,1241]" pageId="4" pageNumber="29">
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<emphasis id="8EC5EA811D48FFBD94EEE5A3FED65F8C" box="[151,258,1482,1506]" italics="true" pageId="4" pageNumber="29">Tracheids</emphasis>
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– in cross section appear quasi-polygonal, usually quadrangular, and with moderately thick walls in the early-wood. They have radial / tangential diameters of 25-60 / 25-55 μm in the early- wood and wall thickness of 3-7 μm double wall. When gradual transition is present, the tracheids gradually diminish in section and usually the tracheidal wall in the late-wood becomes slightly thicker, of 7-9 μm in the final 3-8 rows of smaller sized and radially flattened cells, with r/tg diameters of 10-25 / 20-25 μm, marking the growth- ring boundary. Between two successive rays, 2-12 radial rows of tracheids can be counted. The density ranges 750-1216 tracheids / mm
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<superScript id="4BC49BDB1D48FFBD968DE75DFD285D2C" attach="left" box="[756,764,1844,1858]" fontSize="6" pageId="4" pageNumber="29">2</superScript>
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. Longitudinally, the tracheids have, on the radial walls, uniseriate pits, often with biseriate portions, often contiguous, or biseriate of mixed
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<typeStatus id="630A88311D48FFBD95BAE7F3FE275DDC" box="[451,499,1946,1970]" pageId="4" pageNumber="29">type</typeStatus>
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, sometimes separated by crassulae. The pits are of abietinean
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<typeStatus id="630A88311D48FFBD965AE7D2FD805DBD" box="[547,596,1979,2003]" pageId="4" pageNumber="29">type</typeStatus>
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, have 14-21 μm in diameter and round apertures of 4-5 μm. Details on torus not observed. On the tangential walls, pits are usually absent. Organic deposits - absent. Average tracheids length difficult to measure. Helical thickenings usually absent, or rarely appear.
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</paragraph>
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<paragraph id="BC0E36931D48FFBD9749E168FC075BD1" blockId="4.[816,1437,163,1241]" pageId="4" pageNumber="29">
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<emphasis id="8EC5EA811D48FFBD9749E168FC2D5B77" box="[816,1017,257,281]" italics="true" pageId="4" pageNumber="29">Axial parenchyma</emphasis>
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– in the cross sections is absent, or rarely present as dispersed thin-walled cells with dark content. In longitudinal view, the parenchyma appears as vertical rows of rectangular cells, with the horizontal wall thin and smooth or feebly nodular and with some dark remains inside.
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</paragraph>
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<paragraph id="BC0E36931D48FFBD9749E1A0FAFD5EF9" blockId="4.[816,1437,163,1241]" pageId="4" pageNumber="29">
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<emphasis id="8EC5EA811D48FFBD9749E1A0FCB05B8E" box="[816,868,457,480]" italics="true" pageId="4" pageNumber="29">Rays</emphasis>
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– usually thin in cross section, appear in tangential section as uniseriate, but sometimes with some biseriate storeys, or even as 2-3-seriate rays. The ray height varies from 2–32 to up to 56 cells (average height - 16 cells). The ray cells appear ovoid-rectangular, with 26–39 μm in height and 23–30 μm in width. Lateral triangular spaces are present. Their density is 6-9 rays on tangential milimeter. In radial section, the rays appear heterogeneous, because they have thicker walled cells, as ray tracheids, beside the thinner-walled parenchymal body cells, and are all procumbent. The ray cells have 8-20 μm tall, the marginals are slightly taller, of 25-30 μm. The horizontal walls of the ray cells are smooth and usually pitted, and the tangential end-walls are relatively thick, of 1.5-2 μm the simple wall and are nodular, as typical juniperoid thickenings. The ray tracheids are located in the ray-body or marginally, and have smooth or slightly rough thick walls. The indentures are missing or difficult to observe. Spiral thickenings are usually absent. The cross-fields are typical cupressoid, with 1-2-4(-6) cupressoid oculipores of 8-10 μm in diameter, usually in vertical pairs arranged, and with slit-like inclined apertures, of 2-5 μm.
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</paragraph>
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<paragraph id="BC0E36931D48FFBD9749E4C9FB3C5ED6" blockId="4.[816,1437,163,1241]" box="[816,1256,1184,1208]" pageId="4" pageNumber="29">
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<emphasis id="8EC5EA811D48FFBD9749E4C9FB8E5ED6" box="[816,1114,1184,1208]" italics="true" pageId="4" pageNumber="29">Radial or axial resin canals</emphasis>
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– are absent.
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</paragraph>
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<paragraph id="BC0E36931D48FFBD9749E4A8FB655EB7" blockId="4.[816,1437,163,1241]" box="[816,1201,1217,1241]" pageId="4" pageNumber="29">
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<emphasis id="8EC5EA811D48FFBD9749E4A8FC2D5EB7" box="[816,1017,1217,1241]" italics="true" pageId="4" pageNumber="29">Mineral inclusions</emphasis>
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– are not present
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</paragraph>
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</subSubSection>
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<subSubSection id="F4AB65181D48FFBF9749E56DFA475B35" lastPageId="6" lastPageNumber="31" pageId="4" pageNumber="29" type="discussion">
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<paragraph id="BC0E36931D48FFBD9749E56DFB925F72" blockId="4.[816,1437,1284,2069]" box="[816,1094,1284,1308]" pageId="4" pageNumber="29">
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<emphasis id="8EC5EA811D48FFBD9749E56DFB925F72" bold="true" box="[816,1094,1284,1308]" pageId="4" pageNumber="29">Affinities and discussion</emphasis>
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</paragraph>
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<paragraph id="BC0E36931D48FFBD9749E54DFB465CE9" blockId="4.[816,1437,1284,2069]" pageId="4" pageNumber="29">
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The cupressaceous xylotomy of the
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<specimenCount id="AAB7FD1A1D48FFBD90C6E54CFA915F52" box="[1215,1349,1316,1340]" count="7" pageId="4" pageNumber="29" type="generic">7 specimens</specimenCount>
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studied here, assigned to the genus
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<taxonomicName id="7BB14D101D48FFBD901AE52CFA4C5F33" authority="Eckhold" authorityName="Eckhold" authorityYear="1922" box="[1123,1432,1349,1373]" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="29" phylum="Tracheophyta" rank="genus">
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<emphasis id="8EC5EA811D48FFBD901AE52CFAE05F33" box="[1123,1332,1349,1373]" italics="true" pageId="4" pageNumber="29">Protojuniperoxylon</emphasis>
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Eckhold
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</taxonomicName>
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, are definitely pycnoxylic and homoxylic (or tracheidoxylic – sensu
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<bibRefCitation id="D8204B621D48FFBD97B9E5EEFB865FF1" author="Creber, G. T." box="[960,1106,1415,1439]" pageId="4" pageNumber="29" pagination="655 - 661" refId="ref13670" refString="Creber, G. T., 1972. Gymnospermous wood from the Kimmeridgian of East Sutherland and from the Sandringham Sands of Norfolk. Palaeontology, 15 (4): 655 - 661." type="journal article" year="1972">Creber, 1972</bibRefCitation>
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), having radial pitting on tracheids of mixed
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<typeStatus id="630A88311D48FFBD9797E5C0FBCB5FAF" box="[1006,1055,1449,1473]" pageId="4" pageNumber="29">type</typeStatus>
|
||
, no resin canals, rare parenchyma but with nodular horizontal walls, rays exclusively uniseriate and cross-fields with 1-2(4-6) cupressoid pits with slit-like inclined apertures, arranged in 1-2 vertical pairs, or irregularly to slightly alternately and with nodular tangential end-walls of ray cells as typical juniperoid nodules (i.e. juniperoid thickenings).
|
||
</paragraph>
|
||
<paragraph id="BC0E36931D48FFBD9749E6F8FC0C5DFE" blockId="4.[816,1437,1284,2069]" pageId="4" pageNumber="29">
|
||
We have discussed above the current status of “
|
||
<taxonomicName id="7BB14D101D48FFBD9133E6F9FC545CA7" class="Pinopsida" family="Protopinaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="29" phylum="Tracheophyta" rank="family">Protopinaceae</taxonomicName>
|
||
family” and of “Cheirlepidiaceae family” and, lastly, the proposal to be accepted as valid - the family name “
|
||
<taxonomicName id="7BB14D101D48FFBD9704E69DFBF85D62" authorityName="Harris" authorityYear="1979" box="[893,1068,1780,1804]" family="Hirmeriellaceae" pageId="4" pageNumber="29" rank="family">Hirmeriellaceae</taxonomicName>
|
||
” (
|
||
<bibRefCitation id="D8204B621D48FFBD903FE69DFB075D62" author="Harris, T. M." box="[1094,1235,1780,1804]" pageId="4" pageNumber="29" refId="ref14877" refString="Harris, T. M., 1979. The Yorkshire Jurassic Flora. 5. Coniferales. British Museum Natural History, London, 167 pp." type="book" year="1979">Harris, 1979</bibRefCitation>
|
||
), which is considered correct and valid to be used for the Mesozoic gymnosperms characterized by pollen of
|
||
<taxonomicName id="7BB14D101D48FFBD90A2E75FFA8B5D20" authorityName="H.D.Pflug" authorityYear="1953" box="[1243,1375,1846,1870]" class="Pinopsida" family="Cheirolepidiaceae" genus="Classopollis" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="4" pageNumber="29" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="8EC5EA811D48FFBD90A2E75FFA8B5D20" box="[1243,1375,1846,1870]" italics="true" pageId="4" pageNumber="29">Classopollis</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<typeStatus id="630A88311D48FFBD9111E75EFA4C5D21" box="[1384,1432,1847,1871]" pageId="4" pageNumber="29">type</typeStatus>
|
||
, and was proposed for taxonomic conservation (see
|
||
<bibRefCitation id="D8204B621D48FFBD9749E711FC185DFE" author="Doweld, A. B." box="[816,972,1912,1936]" pageId="4" pageNumber="29" pagination="1092 - 1098" refId="ref14259" refString="Doweld, A. B., 2020. The controversial nomenclature of the fossil plant names Cheirolepis, Cheirolepidium and Hirmeriella (Cheirolepidaceae / Cheirolepidiaceae / Hirmeriellaceae). 69 (5): 1092 - 1098." type="journal article" year="2020">Doweld, 2020</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="BC0E36931D48FFBF9749E7F0FE715B54" blockId="4.[816,1437,1284,2069]" lastBlockId="6.[151,771,158,2068]" lastPageId="6" lastPageNumber="31" pageId="4" pageNumber="29">
|
||
The genus name
|
||
<taxonomicName id="7BB14D101D48FFBD978BE7F0FB165DDF" authorityName="Eckhold" authorityYear="1922" box="[1010,1218,1945,1969]" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="29" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="8EC5EA811D48FFBD978BE7F0FB165DDF" box="[1010,1218,1945,1969]" italics="true" pageId="4" pageNumber="29">Protojuniperoxylon</emphasis>
|
||
</taxonomicName>
|
||
was established by Eckhold in his PhD Thesis (
|
||
<bibRefCitation id="D8204B621D48FFBD9019E7D3FAD45DBC" author="Eckhold, W." box="[1120,1280,1978,2002]" pageId="4" pageNumber="29" refId="ref14298" refString="Eckhold, W., 1921. Die Hoftupfel bei rezenten und fossilen Koniferen. [PhD Thesis] Breslau Schlesische Fridrich-Wilhelms Universitat (Univ. of Wroclaw Poland)" type="book" year="1921">Eckhold, 1921</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="D8204B621D48FFBD9175E7D3FA905DBC" author="Eckhold, W." box="[1292,1348,1978,2002]" pageId="4" pageNumber="29" pagination="472 - 505" refId="ref14328" refString="Eckhold, W., 1922. Die Hoftupfel bei rezenten und fossilen Coniferen. Jahrbuch Konigliche Preussische Geologische Landesansalt, 42: 472 - 505." type="journal article" year="1922">1922</bibRefCitation>
|
||
, p.491), to designate Mesozoic cupressaceous wood of
|
||
<taxonomicName id="7BB14D101D48FFBD9152E7B5FA405D9D" box="[1323,1428,2012,2035]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="29" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="8EC5EA811D48FFBD9152E7B5FA405D9D" box="[1323,1428,2012,2035]" italics="true" pageId="4" pageNumber="29">Juniperus</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<typeStatus id="630A88311D48FFBD9749E794FCB5527B" box="[816,865,2045,2069]" pageId="4" pageNumber="29">type</typeStatus>
|
||
, described like this: “Annual rings more or less marked, tracheid pits in various transitional arrangements, ‘Abietineentüpfelung’ absent, but ‘Juniperustüpfelung’ present, no resin canals, cupressoid pits in cross-field, axial parenchyma occasionally present” (
|
||
<bibRefCitation id="D8204B621D4AFFBF96DBE168FF195B54" author="Eckhold, W." pageId="6" pageNumber="31" refId="ref14298" refString="Eckhold, W., 1921. Die Hoftupfel bei rezenten und fossilen Koniferen. [PhD Thesis] Breslau Schlesische Fridrich-Wilhelms Universitat (Univ. of Wroclaw Poland)" type="book" year="1921">Eckhold, 1921</bibRefCitation>
|
||
fide
|
||
<bibRefCitation id="D8204B621D4AFFBF957DE14BFE4D5B54" author="Bodnar, J." box="[260,409,290,314]" pageId="6" pageNumber="31" pagination="1480 - 1481" refId="ref13105" refString="Bodnar, J., 2017. (2575) Proposal to conserve the name Protocupressinoxylon with a conserved type against Protobrachyoxylon (fossil plants). Taxon (Journal of IAPT), 66 (6): 1480 - 1481. https: // doi. org / 10.12705 / 666.25" type="journal article" year="2017">Bodnar, 2017</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="E8CE661B1D49FFBC94D0E7ECFEB5527F" ID-DOI="http://doi.org/10.5281/zenodo.10520855" ID-Zenodo-Dep="10520855" httpUri="https://zenodo.org/record/10520855/files/figure.png" pageId="5" pageNumber="30" startId="5.[169,204,1925,1946]" targetBox="[169,1412,154,1916]" targetPageId="5" targetType="figure">
|
||
<paragraph id="BC0E36931D49FFBC94D0E7ECFC9E5DF4" blockId="5.[169,1429,1924,2065]" box="[169,842,1924,1946]" pageId="5" pageNumber="30">
|
||
<emphasis id="8EC5EA811D49FFBC94D0E7ECFF365DF4" bold="true" box="[169,226,1925,1946]" pageId="5" pageNumber="30">Fig. 2</emphasis>
|
||
<taxonomicName id="7BB14D101D49FFBC9491E7EDFDCE5DF4" authority="Iamandei & Iamandei & Grădinaru, 2022" authorityName="Iamandei & Iamandei & Grădinaru" authorityYear="2022" box="[232,538,1924,1946]" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="30" phylum="Tracheophyta" rank="species" species="holbavicum" status="sp. nov.">
|
||
<emphasis id="8EC5EA811D49FFBC9491E7EDFDCE5DF4" box="[232,538,1924,1946]" italics="true" pageId="5" pageNumber="30">Protojuniperoxylon holbavicum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="95F657FA1D49FFBC9659E7ECFDB85DF4" box="[544,620,1925,1946]" pageId="5" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(inventory no. 27706).
|
||
</paragraph>
|
||
<paragraph id="BC0E36931D49FFBC94D0E7CAFEB5527F" blockId="5.[169,1429,1924,2065]" pageId="5" pageNumber="30">
|
||
<emphasis id="8EC5EA811D49FFBC94D0E7CAFF1C5DD6" bold="true" box="[169,200,1955,1976]" pageId="5" pageNumber="30">a-c</emphasis>
|
||
: Cross section - tracheids polygonal with rounded corners, and linear rays;
|
||
<emphasis id="8EC5EA811D49FFBC97DDE7CAFC155DD6" bold="true" box="[932,961,1955,1976]" pageId="5" pageNumber="30">d-f</emphasis>
|
||
: Tangential section - uniseriate rays with biseriate storeys with polygonal ray-cells, with lateral empty spaces; unpitted tracheids;
|
||
<emphasis id="8EC5EA811D49FFBC97B6E7A9FC3F5DBB" bold="true" box="[975,1003,1984,2005]" pageId="5" pageNumber="30">g-j</emphasis>
|
||
: Radial section –uniseriate radial pitting on tracheids as hexagonal pits; araucarioid cross-fields with vertical pairs of pits or slightly irregular; end-wall with juniperoid thickenings (see arrow).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="BC0E36931D4AFFBF94EEE12DFE735B12" blockId="6.[151,771,158,2068]" pageId="6" pageNumber="31">In fact, few species were described for this genus, sometimes revised, as follows:</paragraph>
|
||
<paragraph id="BC0E36931D4AFFBF94EEE1ECFDBC580A" blockId="6.[151,771,158,2068]" pageId="6" pageNumber="31">
|
||
<taxonomicName id="7BB14D101D4AFFBF94EEE1ECFCD75BF3" authority="(Stopes) Eckhold, 1921" authorityName="Eckhold" authorityYear="1921" baseAuthorityName="Stopes" baseAuthorityYear="1915" box="[151,771,389,413]" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="species" species="maidstonense">
|
||
<emphasis id="8EC5EA811D4AFFBF94EEE1ECFE2B5BF3" box="[151,511,389,413]" italics="true" pageId="6" pageNumber="31">Protojuniperoxylon maidstonense</emphasis>
|
||
(Stopes)
|
||
<bibRefCitation id="D8204B621D4AFFBF961EE1ECFCD75BF3" author="Eckhold, W." box="[615,771,389,413]" pageId="6" pageNumber="31" refId="ref14298" refString="Eckhold, W., 1921. Die Hoftupfel bei rezenten und fossilen Koniferen. [PhD Thesis] Breslau Schlesische Fridrich-Wilhelms Universitat (Univ. of Wroclaw Poland)" type="book" year="1921">Eckhold, 1921</bibRefCitation>
|
||
</taxonomicName>
|
||
was taken as type-species, even if, initially, this wood
|
||
<typeStatus id="630A88311D4AFFBF94EEE1A1FF115B8E" box="[151,197,456,480]" pageId="6" pageNumber="31">type</typeStatus>
|
||
was described from the Cretaceous of Maidstone (
|
||
<collectingCountry id="C4A676031D4AFFBF94E7E180FEE2586F" box="[158,310,489,513]" name="United Kingdom" pageId="6" pageNumber="31">Great Britain</collectingCountry>
|
||
), as a species of
|
||
<taxonomicName id="7BB14D101D4AFFBF967CE180FD55586F" authorityName="G.Kraus" authorityYear="1870" box="[517,641,489,513]" class="Pinopsida" family="Pinaceae" genus="Cedroxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="8EC5EA811D4AFFBF967CE180FD55586F" box="[517,641,489,513]" italics="true" pageId="6" pageNumber="31">Cedroxylon</emphasis>
|
||
</taxonomicName>
|
||
, by
|
||
<bibRefCitation id="D8204B621D4AFFBF96C5E180FF35584C" author="Stopes, M." pageId="6" pageNumber="31" refId="ref17231" refString="Stopes, M., 1915. Catalogue of the Mesozoic plants in the British Museum - The Cretaceous flora, Part 2, Lower Greensand (Aptian) plants of Britain: London, British Museum (Natural History), 360 p." type="book" year="1915">Stopes (1915)</bibRefCitation>
|
||
. It has uniseriate radial pitting on tracheids, few axial parenchyma with nodular end wall, and cupressoid cross-fields (see Bodnar and Artabe, 2007).
|
||
</paragraph>
|
||
<paragraph id="BC0E36931D4AFFBF94EEE204FDEA5887" blockId="6.[151,771,158,2068]" pageId="6" pageNumber="31">
|
||
<taxonomicName id="7BB14D101D4AFFBF94EEE204FED858C8" authority="(Seward and Bancroft) Eckhold, 1921" authorityName="Eckhold" authorityYear="1921" baseAuthorityName="Seward and Bancroft" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="species" species="hornei">
|
||
<emphasis id="8EC5EA811D4AFFBF94EEE204FE6E58EB" box="[151,442,621,645]" italics="true" pageId="6" pageNumber="31">Protojuniperoxylon hornei</emphasis>
|
||
(Seward and Bancroft)
|
||
<bibRefCitation id="D8204B621D4AFFBF96A8E204FED858C8" author="Eckhold, W." pageId="6" pageNumber="31" refId="ref14298" refString="Eckhold, W., 1921. Die Hoftupfel bei rezenten und fossilen Koniferen. [PhD Thesis] Breslau Schlesische Fridrich-Wilhelms Universitat (Univ. of Wroclaw Poland)" type="book" year="1921">Eckhold, 1921</bibRefCitation>
|
||
</taxonomicName>
|
||
was initially described as Jurassic species of
|
||
<taxonomicName id="7BB14D101D4AFFBF94EEE2C6FEC758A9" authorityName="G.Kraus" authorityYear="1870" box="[151,275,687,711]" class="Pinopsida" family="Pinaceae" genus="Cedroxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="8EC5EA811D4AFFBF94EEE2C6FEC758A9" box="[151,275,687,711]" italics="true" pageId="6" pageNumber="31">Cedroxylon</emphasis>
|
||
</taxonomicName>
|
||
and later, of
|
||
<taxonomicName id="7BB14D101D4AFFBF95DFE2C6FD2B58A9" authority="(Vogellehner, 1968)" baseAuthorityName="Vogellehner" baseAuthorityYear="1968" box="[422,767,687,711]" class="Pinopsida" genus="Scotoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="8EC5EA811D4AFFBF95DFE2C6FDCF58A9" box="[422,539,687,711]" italics="true" pageId="6" pageNumber="31">Scotoxylon</emphasis>
|
||
(
|
||
<bibRefCitation id="D8204B621D4AFFBF9652E2C6FD2258A9" author="Vogellehner, D." box="[555,758,687,711]" pageId="6" pageNumber="31" pagination="125 - 162" refId="ref17771" refString="Vogellehner, D., 1968. Zur Anatomie und Phylogenie de Mesozoischer Gymnospermenholzer, 7: Prodromus zu einer Monographie de Protopinaceae. II. Die Protopinoiden Holzer des Jura. Palaeontographica Abt. B, Stuttgart, 124 (4 - 6): 125 - 162." type="journal article" year="1968">Vogellehner, 1968</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, so, it could be doubtful for comparison.
|
||
</paragraph>
|
||
<paragraph id="BC0E36931D4AFFBF94EEE29BFDAF5903" blockId="6.[151,771,158,2068]" pageId="6" pageNumber="31">
|
||
However, Zhang et al. (2000) described a
|
||
<taxonomicName id="7BB14D101D4AFFBF96F7E29BFE3B5945" authority="Zhang and Zengh" authorityName="Zhang and Zengh" class="Pinopsida" genus="Scotoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="species" species="yanqingense">
|
||
<emphasis id="8EC5EA811D4AFFBF96F7E29BFEC85944" italics="true" pageId="6" pageNumber="31">Scotoxylon yanqingense</emphasis>
|
||
Zhang and Zengh
|
||
</taxonomicName>
|
||
, a Late Jurassic Chinese species, as representing secondary wood of
|
||
<taxonomicName id="7BB14D101D4AFFBF9609E35CFF0A5903" authorityName="Eckhold" authorityYear="1922" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="8EC5EA811D4AFFBF9609E35CFF0A5903" italics="true" pageId="6" pageNumber="31">Protojuniperoxylon</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<typeStatus id="630A88311D4AFFBF949EE33FFEC35900" box="[231,279,854,878]" pageId="6" pageNumber="31">type</typeStatus>
|
||
, but devoid of axial parenchyma.
|
||
</paragraph>
|
||
<paragraph id="BC0E36931D4AFFBF94EEE31FFEB35E3A" blockId="6.[151,771,158,2068]" pageId="6" pageNumber="31">
|
||
<taxonomicName id="7BB14D101D4AFFBF94EEE31FFE7C59C1" authority="(Bonetti 1966) Bodnar and Artabe, 2007" authorityName="Bodnar and Artabe" authorityYear="2007" baseAuthorityName="Bonetti" baseAuthorityYear="1966" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="species" species="ischigualastensis">
|
||
<emphasis id="8EC5EA811D4AFFBF94EEE31FFDEC59E0" box="[151,568,886,910]" italics="true" pageId="6" pageNumber="31">Protojuniperoxylon ischigualastensis</emphasis>
|
||
(
|
||
<bibRefCitation id="D8204B621D4AFFBF9623E31FFD2F59E0" author="Bonetti, M. I. R." box="[602,763,886,910]" pageId="6" pageNumber="31" pagination="211 - 218" refId="ref13201" refString="Bonetti, M. I. R., 1966. Protojuniperoxylon ischigualastensis sp. nov. del Triasico de Ischigualasto (San Juan). Ameghiniana, 4: 211 - 218." type="journal article" year="1966">Bonetti 1966</bibRefCitation>
|
||
) Bodnar and Artabe, 2007
|
||
</taxonomicName>
|
||
is a Triassic wood from
|
||
<collectingCountry id="C4A676031D4AFFBF96CEE3FEFF1659BE" name="Argentina" pageId="6" pageNumber="31">Argentina</collectingCountry>
|
||
, which has mixed radial pitting on tracheids, cupressoid cross fields, juniperoid thickenings on the end walls of ray parenchymal cells and nodules on the horizontal walls axial parenchyma, which are typical for
|
||
<taxonomicName id="7BB14D101D4AFFBF9698E475FF335E3A" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="8EC5EA811D4AFFBF9698E475FF335E3A" italics="true" pageId="6" pageNumber="31">Juniperus</emphasis>
|
||
</taxonomicName>
|
||
wood-type.
|
||
</paragraph>
|
||
<paragraph id="BC0E36931D4AFFBF94EEE434FEEF5F52" blockId="6.[151,771,158,2068]" pageId="6" pageNumber="31">
|
||
<taxonomicName id="7BB14D101D4AFFBF94EEE434FDB95E1B" authority="Selling, 1951" authorityName="Selling" authorityYear="1951" box="[151,621,1117,1141]" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="species" species="arcticum">
|
||
<emphasis id="8EC5EA811D4AFFBF94EEE434FE055E1B" box="[151,465,1117,1141]" italics="true" pageId="6" pageNumber="31">Protojuniperoxylon arcticum</emphasis>
|
||
<bibRefCitation id="D8204B621D4AFFBF95A5E434FDB95E1B" author="Selling, O. H." box="[476,621,1117,1141]" pageId="6" pageNumber="31" pagination="538 - 539" refId="ref17060" refString="Selling, O. H., 1951. On Protojuniperoxylon arcticum. Journal of Paleontology, 25 (4): 538 - 539." type="journal article" year="1951">Selling, 1951</bibRefCitation>
|
||
</taxonomicName>
|
||
described on some Mesozoic root-wood remains from Hope Island (
|
||
<collectingCountry id="C4A676031D4AFFBF94E7E4C9FED55ED6" box="[158,257,1184,1208]" name="Svalbard and Jan Mayen" pageId="6" pageNumber="31">Svalbard</collectingCountry>
|
||
,
|
||
<collectingCountry id="C4A676031D4AFFBF956BE4C9FEBD5ED6" box="[274,361,1184,1208]" name="Norway" pageId="6" pageNumber="31">Norway</collectingCountry>
|
||
) is poorly identified, for it is most probably a bennettitalean taxon (
|
||
<taxonomicName id="7BB14D101D4AFFBF9582E4A8FD2C5EB7" authority="Buckland" authorityName="Buckland" authorityYear="1828" box="[507,760,1217,1241]" class="Cycadopsida" family="Cycadeoideaceae" genus="Cycadeoidea" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="8EC5EA811D4AFFBF9582E4A8FD525EB7" box="[507,646,1217,1241]" italics="true" pageId="6" pageNumber="31">Cycadeoidea</emphasis>
|
||
Buckland
|
||
</taxonomicName>
|
||
), showing interactions with chytrid fungi and bacteria (see
|
||
<bibRefCitation id="D8204B621D4AFFBF94EEE56AFE075F75" author="Strullu-Derrien, C. & McLoughlin, S. & Philippe, M. & Mork, A. & Strullu, D. G." box="[151,467,1283,1307]" pageId="6" pageNumber="31" pagination="45 - 58" refId="ref17334" refString="Strullu-Derrien, C., McLoughlin, S., Philippe, M., Mork, A. and Strullu, D. G., 2012. Arthropod interactions with bennettitalean roots in a Triassic permineralized peat from Hopen, Svalbard Archipelago (Arctic). Palaeogeography Palaeoclimatology Palaeoecology 348 - 349: 45 - 58. https: // doi. org / 10.1016 / j. palaeo. 2012.06.006" type="journal article" year="2012">Strullu-Derrien et al., 2012</bibRefCitation>
|
||
; McLoughlin and Strullu-Derrien. 2016).
|
||
</paragraph>
|
||
<paragraph id="BC0E36931D4AFFBF94EEE52CFD0B5C08" blockId="6.[151,771,158,2068]" pageId="6" pageNumber="31">
|
||
<bibRefCitation id="D8204B621D4AFFBF94EEE52CFE4E5F33" author="Ruiz, D. P. & Bodnar, J." box="[151,410,1349,1373]" pageId="6" pageNumber="31" pagination="481 - 488" refId="ref16844" refString="Ruiz, D. P. and Bodnar, J., 2019. The oldest record of Juniperoxylon, a cupressaceous fossil wood from the Middle Triassic of Argentina. Acta Palaeontologica Polonica, 64 (3): 481 - 488." type="journal article" year="2019">Ruiz and Bodnar (2019)</bibRefCitation>
|
||
revised a Mid Triassic species of
|
||
<taxonomicName id="7BB14D101D4AFFBF94EEE50FFE9E5F10" authorityName="Göppert" authorityYear="1850" box="[151,330,1382,1406]" class="Pinopsida" family="Cupressaceae" genus="Cupressinoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="8EC5EA811D4AFFBF94EEE50FFE9E5F10" box="[151,330,1382,1406]" italics="true" pageId="6" pageNumber="31">Cupressinoxylon</emphasis>
|
||
</taxonomicName>
|
||
of Bodnar et al. (2015) as
|
||
<taxonomicName id="7BB14D101D4AFFBF9612E50FFDA45FF1" authority="(Bodnar et al.) Ruiz and Bodnar" authorityName="Ruiz and Bodnar" baseAuthorityName="Bodnar" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="species" species="zamunerae">
|
||
<emphasis id="8EC5EA811D4AFFBF9612E50FFED85FF1" italics="true" pageId="6" pageNumber="31">Juniperoxylon zamunerae</emphasis>
|
||
(Bodnar et al.) Ruiz and Bodnar
|
||
</taxonomicName>
|
||
, and made an extensive discussion on the wood structure of
|
||
<taxonomicName id="7BB14D101D4AFFBF96E8E5C3FD2E5FAF" box="[657,762,1450,1473]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="8EC5EA811D4AFFBF96E8E5C3FD2E5FAF" box="[657,762,1450,1473]" italics="true" pageId="6" pageNumber="31">Juniperus</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<typeStatus id="630A88311D4AFFBF94EEE5A3FF115F8C" box="[151,197,1482,1506]" pageId="6" pageNumber="31">type</typeStatus>
|
||
including in a table the described species of
|
||
<taxonomicName id="7BB14D101D4AFFBF96D0E5A2FEC25C6D" authorityName="Eckhold" authorityYear="1922" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="8EC5EA811D4AFFBF96D0E5A2FEC25C6D" italics="true" pageId="6" pageNumber="31">Protojuniperoxylon</emphasis>
|
||
</taxonomicName>
|
||
together with Mesozoic and Cenozoic species of
|
||
<taxonomicName id="7BB14D101D4AFFBF9497E665FE525C4A" authorityName="C.Houlbert" authorityYear="1910" box="[238,390,1548,1572]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="8EC5EA811D4AFFBF9497E665FE525C4A" box="[238,390,1548,1572]" italics="true" pageId="6" pageNumber="31">Juniperoxylon</emphasis>
|
||
</taxonomicName>
|
||
, all of them characterized by the presence of nodular ray parenchyma end-walls (i.e. ‘Juniperustüpfelung’), details also observed in our material.
|
||
</paragraph>
|
||
<paragraph id="BC0E36931D4AFFBF94EEE606FD0A5D65" blockId="6.[151,771,158,2068]" pageId="6" pageNumber="31">
|
||
Also, very useful is the discussion of
|
||
<bibRefCitation id="D8204B621D4AFFBF963EE606FD2A5CE9" author="Akkemik, U." box="[583,766,1647,1671]" pageId="6" pageNumber="31" pagination="15 - 26" refId="ref12580" refString="Akkemik, U., 2021. A new species of Juniperoxylon from the Early Miocene of Northwestern Turkey. Acta Palaeontologica Romaniae, 17 (1): 15 - 26. https: // doi. org / 10.35463 / j. apr. 2021.01.02" type="journal article" year="2021">Akkemik (2021)</bibRefCitation>
|
||
, who described and identified an Early Miocene species of
|
||
<taxonomicName id="7BB14D101D4AFFBF94EEE6D8FEE45CA7" authorityName="C.Houlbert" authorityYear="1910" box="[151,304,1713,1737]" class="Pinopsida" family="Cupressaceae" genus="Juniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="8EC5EA811D4AFFBF94EEE6D8FEE45CA7" box="[151,304,1713,1737]" italics="true" pageId="6" pageNumber="31">Juniperoxylon</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="C4A676031D4AFFBF9501E6D8FE135CA7" box="[376,455,1713,1737]" name="Turkey" pageId="6" pageNumber="31">Turkey</collectingCountry>
|
||
, as having typical xylotomy of
|
||
<taxonomicName id="7BB14D101D4AFFBF94C0E6BAFEF65C84" box="[185,290,1747,1770]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="8EC5EA811D4AFFBF94C0E6BAFEF65C84" box="[185,290,1747,1770]" italics="true" pageId="6" pageNumber="31">Juniperus</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="630A88311D4AFFBF9554E6BAFE895C85" box="[301,349,1747,1771]" pageId="6" pageNumber="31">type</typeStatus>
|
||
, proposing even an identification key that includes also some species of
|
||
<taxonomicName id="7BB14D101D4AFFBF967EE69DFD0C5D62" authorityName="Eckhold" authorityYear="1922" box="[519,728,1780,1804]" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="8EC5EA811D4AFFBF967EE69DFD0C5D62" box="[519,728,1780,1804]" italics="true" pageId="6" pageNumber="31">Protojuniperoxylon</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="BC0E36931D4AFFBF94EEE77CFA475B35" blockId="6.[151,771,158,2068]" lastBlockId="6.[816,1436,158,347]" pageId="6" pageNumber="31">
|
||
Our material displays the typical xylotomical features of
|
||
<taxonomicName id="7BB14D101D4AFFBF94EEE75EFED45D20" box="[151,256,1847,1870]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="8EC5EA811D4AFFBF94EEE75EFED45D20" box="[151,256,1847,1870]" italics="true" pageId="6" pageNumber="31">Juniperus</emphasis>
|
||
</taxonomicName>
|
||
type as cupressaceous wood with mixed 1-2 seriate radial pitting on tracheids, rare or absent parenchyma, and usually uniseriate rays with cupressoid cross-fields in vertical pairs and juniperoid nodules on the tangential walls of the ray cells. Evaluating these details comparatively with those included in Table 1 from
|
||
<bibRefCitation id="D8204B621D4AFFBF96A9E7B2FEB9527A" author="Ruiz, D. P. & Bodnar, J." pageId="6" pageNumber="31" pagination="481 - 488" refId="ref16844" refString="Ruiz, D. P. and Bodnar, J., 2019. The oldest record of Juniperoxylon, a cupressaceous fossil wood from the Middle Triassic of Argentina. Acta Palaeontologica Polonica, 64 (3): 481 - 488." type="journal article" year="2019">Ruiz and Bodnar (2019)</bibRefCitation>
|
||
, and using the identification key of
|
||
<bibRefCitation id="D8204B621D4AFFBF9749E0F7FC335AD8" author="Akkemik, U." box="[816,999,158,182]" pageId="6" pageNumber="31" pagination="15 - 26" refId="ref12580" refString="Akkemik, U., 2021. A new species of Juniperoxylon from the Early Miocene of Northwestern Turkey. Acta Palaeontologica Romaniae, 17 (1): 15 - 26. https: // doi. org / 10.35463 / j. apr. 2021.01.02" type="journal article" year="2021">Akkemik (2021)</bibRefCitation>
|
||
, we observe that our specimens are not identical to any described species. Thus, we describe a new species named here
|
||
<taxonomicName id="7BB14D101D4AFFBF903FE089FA485A96" authority="Iamandei & Iamandei & Grădinaru, 2022" authorityName="Iamandei & Iamandei & Grădinaru" authorityYear="2022" box="[1094,1436,224,248]" class="Pinopsida" family="Pinaceae" genus="Protojuniperoxylon" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="31" phylum="Tracheophyta" rank="species" species="holbavicum" status="sp. nov.">
|
||
<emphasis id="8EC5EA811D4AFFBF903FE089FA485A96" box="[1094,1436,224,248]" italics="true" pageId="6" pageNumber="31">Protojuniperoxylon holbavicum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="95F657FA1D4AFFBF9749E16BFC5F5B77" box="[816,907,258,281]" pageId="6" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
|
||
The designated
|
||
<typeStatus id="630A88311D4AFFBF9035E168FB7D5B77" box="[1100,1193,257,281]" pageId="6" pageNumber="31" type="holotype">holotype</typeStatus>
|
||
is the sample 27706 (inventory number), kept in “Grădinaru Collection”, hosted by National Geological Museum, from
|
||
<collectingRegion id="7E75F8711D4AFFBF9158E12AFA5B5B35" box="[1313,1423,323,347]" country="Romania" name="Bucuresti" pageId="6" pageNumber="31">Bucharest</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F4AB65181D4AFFBF9749E1EFFAE058A9" pageId="6" pageNumber="31" type="diagnosis">
|
||
<paragraph id="BC0E36931D4AFFBF9749E1EFFBDB5BD1" blockId="6.[816,1437,390,711]" pageId="6" pageNumber="31">
|
||
<emphasis id="8EC5EA811D4AFFBF9749E1EFFA485BF0" bold="true" box="[816,1436,390,414]" pageId="6" pageNumber="31">
|
||
The diagnosis of the species
|
||
<taxonomicName id="7BB14D101D4AFFBF90BCE1EFFA485BF0" authority="Iamandei & Iamandei & Grădinaru, 2022" authorityName="Iamandei & Iamandei & Grădinaru" authorityYear="2022" box="[1221,1436,390,414]" pageId="6" pageNumber="31" status="sp. nov.">
|
||
<emphasis id="8EC5EA811D4AFFBF90BCE1EFFA485BF0" bold="true" box="[1221,1436,390,414]" italics="true" pageId="6" pageNumber="31">Protojuniperoxylon</emphasis>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
<emphasis id="8EC5EA811D4AFFBF9749E1CEFBDB5BD1" bold="true" box="[816,1039,423,447]" pageId="6" pageNumber="31">
|
||
<emphasis id="8EC5EA811D4AFFBF9749E1CEFC655BD1" bold="true" box="[816,945,423,447]" italics="true" pageId="6" pageNumber="31">holbavicum</emphasis>
|
||
<taxonomicNameLabel id="95F657FA1D4AFFBF97C1E1CEFBDB5BD1" box="[952,1039,423,447]" pageId="6" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="BC0E36931D4AFFBF9749E1A1FAE058A9" blockId="6.[816,1437,390,711]" pageId="6" pageNumber="31">
|
||
Secondary tracheidoxylic wood, with thick-walled tracheids with uniseriate pitting, or even biseriate of mixed
|
||
<typeStatus id="630A88311D4AFFBF9749E262FCB5584D" box="[816,865,523,547]" pageId="6" pageNumber="31">type</typeStatus>
|
||
, parenchyma absent or few and with nodular horizontal walls, heterogeneous uniseriate rays with juniperoid nodules on tangential walls or ray cells, cross-fields with 1-2(4-6) small cupressoid pits per field, as 1-2 vertical pairs pits and when more, slightly irregular or alternate, and having slit-like inclined apertures.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |