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<document id="F5F57BAF41982F9A9577A8D0DB31F8F4" ID="10.11646/zootaxa.4269.1.1SLASH11087" ID-CLB-Dataset="33014" ID-DOI="10.11646/zootaxa.4269.1.1" ID-GBIF-Dataset="0b7fd4db-9f34-47ef-b7e8-33363155d866" ID-ISSN="1175-5326" ID-Zenodo-Dep="581259" ID-ZooBank="2AFC0D53-B439-4BBB-BC22-A0C90C08D65D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1495182755873" checkinUser="plazi" docAuthor="Lee, Chi-Feng" docDate="2017" docId="0A03A20CFF8FFFA8FF24F99C534AE944" docLanguage="en" docName="zootaxa.4269.1.1SLASH11087.pdf" docOrigin="Zootaxa 4269 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Doryscus barclayi Lee, 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="32" masterDocId="F63ADA74FF93FFB7FFB3FFB25110EB24" masterDocTitle="Revision of the genus Doryscus Jacoby (Coleoptera: Chrysomelidae: Galerucinae)" masterLastPageNumber="43" masterPageNumber="1" pageNumber="29" updateTime="1698439733711" updateUser="ExternalLinkService">
<mods:mods id="06306AE18FEFC2F840DFAC9D482EB39F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="84BFC96F4D9DDAD7D4E169EF73C50E11">
<mods:title id="ACE479695302E239234B1ADE97E3AE2A">Revision of the genus Doryscus Jacoby (Coleoptera: Chrysomelidae: Galerucinae)</mods:title>
</mods:titleInfo>
<mods:name id="4AC3F03221FC4BAA6C4B6C2B396C5754" type="personal">
<mods:role id="7EA625F30FFC099AE3CACEC043FD51E3">
<mods:roleTerm id="0FCB072346249978EDD59C6C930FCC16">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="03CBDF42429ED8B2E423AD88C5CB8380">Lee, Chi-Feng</mods:namePart>
</mods:name>
<mods:typeOfResource id="1BB04B02ECBCE3E2A7014867C53F62CF">text</mods:typeOfResource>
<mods:relatedItem id="0C41FF6A3B9631D57446C6C4F6CC851D" type="host">
<mods:titleInfo id="C32A1AC4E535F5387F11E88FAEF93D62">
<mods:title id="CEFBC35D8A8391098B6B8ED11B0155A6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="0C258F5A5A8595316123847981CEB6F7">
<mods:date id="0D72903935A349CA416C0E5C6F2B1065">2017</mods:date>
<mods:detail id="529FE3B9F0014903AB1636B11671E06B" type="volume">
<mods:number id="44195B43702008ACA1722BD1FBE97997">4269</mods:number>
</mods:detail>
<mods:detail id="16FF220D0D5D7DC95A30950ABEAB3853" type="issue">
<mods:number id="30E34995BFFD35C9AED573C3AEBC251B">1</mods:number>
</mods:detail>
<mods:extent id="5EFCCB266AD15A39EE870256B5A9AB94" unit="page">
<mods:start id="38BFDC87C4B51508B5510540F9EBE435">1</mods:start>
<mods:end id="52016D132E63B4BB3E2F2EA08FBF2B91">43</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="75283FC20F6DC1313D6E87247C546762">journal article</mods:classification>
<mods:identifier id="FA145DF62B4CEAC3B1314AD24A7966E8" type="CLB-Dataset">33014</mods:identifier>
<mods:identifier id="72AA8E0FBDD75EE23E3C4B5872154142" type="DOI">10.11646/zootaxa.4269.1.1</mods:identifier>
<mods:identifier id="5247100D149296909F3DB4CDDE2871C7" type="GBIF-Dataset">0b7fd4db-9f34-47ef-b7e8-33363155d866</mods:identifier>
<mods:identifier id="E97521D9C8B1244865470507EF22B797" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="0BA8B3E8B3B46030C82720FB71BE6AD5" type="Zenodo-Dep">581259</mods:identifier>
<mods:identifier id="F8ED465D36EF12B68DD5B6C624BB0F56" type="ZooBank">2AFC0D53-B439-4BBB-BC22-A0C90C08D65D</mods:identifier>
</mods:mods>
<treatment id="0A03A20CFF8FFFA8FF24F99C534AE944" ID-DOI="http://doi.org/10.5281/zenodo.6003755" ID-GBIF-Taxon="130810388" ID-Zenodo-Dep="6003755" LSID="urn:lsid:plazi:treatment:0A03A20CFF8FFFA8FF24F99C534AE944" httpUri="http://treatment.plazi.org/id/0A03A20CFF8FFFA8FF24F99C534AE944" lastPageId="31" lastPageNumber="32" pageId="28" pageNumber="29">
<subSubSection id="CAB04091FF8FFFABFF24F99C502BED4E" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="8215131AFF8FFFABFF24F99C502BED4E" blockId="28.[151,465,1582,1642]" pageId="28" pageNumber="29">
<taxonomicName id="45AA6899FF8FFFABFF24F99C5060ED6C" box="[151,368,1582,1608]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="barclayi" status="sp. nov.">
<emphasis id="B0DECF08FF8FFFABFF24F99C5060ED6C" bold="true" box="[151,368,1582,1608]" italics="true" pageId="28" pageNumber="29">Doryscus barclayi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ABED7273FF8FFFABFEC4F99D50C1ED6C" box="[375,465,1583,1608]" pageId="28" pageNumber="29" rank="species">
<emphasis id="B0DECF08FF8FFFABFEC4F99D50C1ED6C" bold="true" box="[375,465,1583,1608]" pageId="28" pageNumber="29">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="1A910F9FFF8FFFABFF2CF9E351E3ED4E" box="[159,243,1617,1642]" captionStart="FIGURE 16" captionStartId="29.[151,250,1942,1964]" captionTargetBox="[155,1432,409,1921]" captionTargetId="figure@29.[155,1432,409,1921]" captionTargetPageId="29" captionText="FIGURE 16. Habitus of Doryscus species, dorsal view. 16 A. D. barclayi Lee, sp. nov.; 16 B. D. boreri Lee, sp. nov.; 16 C. D. boreri Lee, sp. nov., color variation; 16 D. D. boreri Lee, sp. nov., color variation; 16 E. D. javanensis Lee, sp. nov.; 16 F. D. javanensis Lee, sp. nov., color variation." httpUri="https://zenodo.org/record/581275/files/figure.png" pageId="28" pageNumber="29">Figs 16</figureCitation>
A, 17)
</paragraph>
</subSubSection>
<subSubSection id="CAB04091FF8FFFA8FF24F92B53A2EA89" lastPageId="31" lastPageNumber="32" pageId="28" pageNumber="29" type="description">
<paragraph id="8215131AFF8FFFABFF24F92B5351EC66" blockId="28.[151,1437,1689,2038]" pageId="28" pageNumber="29">
<materialsCitation id="32C21947FF8FFFABFF24F92B5008EDF2" ID-GBIF-Occurrence="1503376971" collectingDate="1985-03-10" collectionCode="BMNH" collectorName="Dumoga-Bone N. P." country="Indonesia" elevation="11230" location="Fog" municipality="Tray" pageId="28" pageNumber="29" specimenCount="1" specimenCount-male="1" stateProvince="Sulawesi Utara" typeStatus="holotype">
<emphasis id="B0DECF08FF8FFFABFF24F92B5021ED96" bold="true" box="[151,305,1689,1714]" pageId="28" pageNumber="29">
<typeStatus id="5D11ADB8FF8FFFABFF24F92B51CCED96" box="[151,220,1689,1714]" pageId="28" pageNumber="29">Types</typeStatus>
(n= 5).
</emphasis>
<typeStatus id="5D11ADB8FF8FFFABFE8BF92B50B0ED96" box="[312,416,1689,1714]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
<specimenCount id="94ACD893FF8FFFABFE14F92850ACED95" box="[423,444,1690,1713]" pageId="28" pageNumber="29" type="male">♂</specimenCount>
(
<collectionCode id="E4BB8BDFFF8FFFABFE7FF9285337ED96" box="[460,551,1690,1714]" pageId="28" pageNumber="29">BMNH</collectionCode>
):
<collectingCountry id="FABD538AFF8FFFABFD8AF92B53C7ED96" box="[569,727,1689,1714]" name="Indonesia" pageId="28" pageNumber="29">
<emphasis id="B0DECF08FF8FFFABFD8AF92B53C7ED96" bold="true" box="[569,727,1689,1714]" pageId="28" pageNumber="29">INDONESIA</emphasis>
</collectingCountry>
.
<collectingRegion id="406EDDF8FF8FFFABFD56F92B5284ED95" box="[741,916,1689,1714]" country="Indonesia" name="Sulawesi Utara" pageId="28" pageNumber="29">Sulawesi Utara</collectingRegion>
:
<collectorName id="2F5F76CCFF8FFFABFC2DF9285593ED95" box="[926,1155,1690,1714]" pageId="28" pageNumber="29">Dumoga-Bone N. P.</collectorName>
,
<collectingMunicipality id="62718960FF8FFFABFB3DF92855D2ED96" box="[1166,1218,1690,1714]" pageId="28" pageNumber="29">Tray</collectingMunicipality>
36,
<location id="877545C1FF8FFFABFB40F928540FED96" LSID="urn:lsid:plazi:treatment:0A03A20CFF8FFFA8FF24F99C534AE944:877545C1FF8FFFABFB40F928540FED96" box="[1267,1311,1690,1714]" country="Indonesia" municipality="Tray" name="Fog" pageId="28" pageNumber="29" stateProvince="Sulawesi Utara">Fog</location>
<elevation id="0987F429FF8FFFABFA96F9285487ED95" box="[1317,1431,1690,1714]" metricMagnitude="4" metricUnit="m" metricValue="1.123" pageId="28" pageNumber="29" unit="m" value="11230.0">
<quantity id="4552BEFFFF8FFFABFA96F9285487ED95" box="[1317,1431,1690,1714]" metricMagnitude="4" metricUnit="m" metricValue="1.123" pageId="28" pageNumber="29" unit="m" value="11230.0">11, 230 m</quantity>
</elevation>
,
<collectingDate id="E650CC32FF8FFFABFF24F90F5008EDF2" box="[151,280,1725,1750]" pageId="28" pageNumber="29" value="1985-03-10">
<date id="F61435DAFF8FFFABFF24F90F5008EDF2" box="[151,280,1725,1750]" pageId="28" pageNumber="29" value="1985-03-10">10.III.1985</date>
</collectingDate>
</materialsCitation>
, BMNH Plot A.
<materialsCitation id="32C21947FF8FFFABFE69F90C5506EDF2" ID-GBIF-Occurrence="1503376952" box="[474,1046,1725,1750]" collectionCode="BMNH" country="Indonesia" location="Tray" pageId="28" pageNumber="29" specimenCount="1" specimenCount-male="1" stateProvince="Sulawesi Utara" typeStatus="paratype">
<typeStatus id="5D11ADB8FF8FFFABFE69F90C535BEDF2" box="[474,587,1726,1750]" pageId="28" pageNumber="29" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="94ACD893FF8FFFABFDE4F90C536AEDF2" box="[599,634,1726,1750]" pageId="28" pageNumber="29" type="male">1♂</specimenCount>
(
<collectionCode id="E4BB8BDFFF8FFFABFD38F90C53F5EDF2" box="[651,741,1726,1750]" pageId="28" pageNumber="29">BMNH</collectionCode>
), same but with “
<location id="877545C1FF8FFFABFC18F90C52F1EDF2" LSID="urn:lsid:plazi:treatment:0A03A20CFF8FFFA8FF24F99C534AE944:877545C1FF8FFFABFC18F90C52F1EDF2" box="[939,993,1726,1750]" country="Indonesia" name="Tray" pageId="28" pageNumber="29" stateProvince="Sulawesi Utara">Tray</location>
28”
</materialsCitation>
;
<materialsCitation id="32C21947FF8FFFABFB90F90C531BEDDE" ID-GBIF-Occurrence="1503376967" collectingDate="1985-07-11" collectionCode="BMNH" country="Indonesia" elevation="13230" location="Fog" municipality="Tray" pageId="28" pageNumber="29" specimenCount="1" specimenCount-female="1" stateProvince="Sulawesi Utara" typeStatus="paratype">
<specimenCount id="94ACD893FF8FFFABFB90F90C5556EDF2" box="[1059,1094,1726,1750]" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
(
<collectionCode id="E4BB8BDFFF8FFFABFBE5F90C55A0EDF2" box="[1110,1200,1726,1750]" pageId="28" pageNumber="29">BMNH</collectionCode>
), same locality,
<collectingMunicipality id="62718960FF8FFFABFADBF90C548BEDF2" box="[1384,1435,1726,1750]" pageId="28" pageNumber="29">Tray</collectingMunicipality>
67,
<location id="877545C1FF8FFFABFF77F95051E0EDDE" LSID="urn:lsid:plazi:treatment:0A03A20CFF8FFFA8FF24F99C534AE944:877545C1FF8FFFABFF77F95051E0EDDE" box="[196,240,1762,1786]" municipality="Tray" name="Fog" pageId="28" pageNumber="29">Fog</location>
<elevation id="0987F429FF8FFFABFF4AF9505061EDDD" box="[249,369,1762,1786]" metricMagnitude="4" metricUnit="m" metricValue="1.323" pageId="28" pageNumber="29" unit="m" value="13230.0">
<quantity id="4552BEFFFF8FFFABFF4AF9505061EDDD" box="[249,369,1762,1786]" metricMagnitude="4" metricUnit="m" metricValue="1.323" pageId="28" pageNumber="29" unit="m" value="13230.0">13, 230 m</quantity>
</elevation>
,
<collectingDate id="E650CC32FF8FFFABFE33F953531BEDDE" box="[384,523,1761,1786]" pageId="28" pageNumber="29" value="1985-07-11">
<date id="F61435DAFF8FFFABFE33F953531BEDDE" box="[384,523,1761,1786]" pageId="28" pageNumber="29" value="1985-07-11">11.VII.1985</date>
</collectingDate>
</materialsCitation>
, BMNH Plot A;
<materialsCitation id="32C21947FF8FFFABFD60F950538FEC3A" ID-GBIF-Occurrence="1503376957" collectingDate="1985-05-30" collectingDateMax="1985-06-06" collectingDateMin="1985-05-30" collectionCode="BMNH" collectorName="Plot A" country="Indonesia" elevation="20" location="Lowland forest" municipality="Tray" pageId="28" pageNumber="29" specimenCount="1" specimenCount-female="1" stateProvince="Sulawesi Utara" typeStatus="paratype">
<specimenCount id="94ACD893FF8FFFABFD60F95053E6EDDE" box="[723,758,1762,1786]" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
(
<collectionCode id="E4BB8BDFFF8FFFABFCBBF9505271EDDE" box="[776,865,1762,1786]" pageId="28" pageNumber="29">BMNH</collectionCode>
), same locality,
<collectorName id="2F5F76CCFF8FFFABFBAEF953557BEDDE" box="[1053,1131,1761,1786]" pageId="28" pageNumber="29">Plot A</collectorName>
, ca
<elevation id="0987F429FF8FFFABFB2FF95055CAEDDD" box="[1180,1242,1762,1786]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="28" pageNumber="29" unit="m" value="20.0">
<quantity id="4552BEFFFF8FFFABFB2FF95055CAEDDD" box="[1180,1242,1762,1786]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="28" pageNumber="29" unit="m" value="20.0">20 m</quantity>
</elevation>
,
<location id="877545C1FF8FFFABFB5BF9535489EDDE" LSID="urn:lsid:plazi:treatment:0A03A20CFF8FFFA8FF24F99C534AE944:877545C1FF8FFFABFB5BF9535489EDDE" box="[1256,1433,1761,1786]" name="Lowland forest" pageId="28" pageNumber="29">Lowland forest</location>
,
<collectingMethod id="5BEB6B0DFF8FFFABFF24F8B75033EC39" box="[151,291,1797,1822]" pageId="28" pageNumber="29">Malaise trap</collectingMethod>
up tree,
<collectingDate id="E650CC32FF8FFFABFE35F8B75353EC3A" box="[390,579,1797,1822]" pageId="28" pageNumber="29" value="1985-05-30" valueMax="1985-06-06" valueMin="1985-05-30">
<date id="F61435DAFF8FFFABFE35F8B75353EC3A" box="[390,579,1797,1822]" pageId="28" pageNumber="29" value="1985-05-30" valueMax="1985-06-06" valueMin="1985-05-30">30.V.–6.VI.1985</date>
</collectingDate>
, 132.25
</materialsCitation>
;
<materialsCitation id="32C21947FF8FFFABFD18F8B45351EC66" ID-GBIF-Occurrence="1503376959" collectingDate="1985-03" collectingDateMax="1985-06-06" collectingDateMin="1985-05-30" collectionCode="BMNH" collectorName="Plot A" country="Indonesia" elevation="1140" location="Lower" municipality="Clarke'" pageId="28" pageNumber="29" specimenCount="1" specimenCount-female="1" stateProvince="Sulawesi Utara" typeStatus="paratype">
<specimenCount id="94ACD893FF8FFFABFD18F8B453DEEC3A" box="[683,718,1798,1822]" pageId="28" pageNumber="29" type="female">1♀</specimenCount>
(
<collectionCode id="E4BB8BDFFF8FFFABFD6EF8B45227EC3A" box="[733,823,1798,1822]" pageId="28" pageNumber="29">BMNH</collectionCode>
), same locality, ‘
<collectingMunicipality id="62718960FF8FFFABFC42F8B7555AEC3A" box="[1009,1098,1797,1822]" pageId="28" pageNumber="29">Clarke’</collectingMunicipality>
camp,
<location id="877545C1FF8FFFABFB2EF8B455F6EC39" LSID="urn:lsid:plazi:treatment:0A03A20CFF8FFFA8FF24F99C534AE944:877545C1FF8FFFABFB2EF8B455F6EC39" box="[1181,1254,1798,1821]" municipality="Clarke'" name="Lower" pageId="28" pageNumber="29">Lower</location>
montane forest,
<elevation id="0987F429FF8FFFABFF24F89851FDEC65" box="[151,237,1834,1858]" metricMagnitude="3" metricUnit="m" metricValue="1.14" pageId="28" pageNumber="29" unit="m" value="1140.0">
<quantity id="4552BEFFFF8FFFABFF24F89851FDEC65" box="[151,237,1834,1858]" metricMagnitude="3" metricUnit="m" metricValue="1.14" pageId="28" pageNumber="29" unit="m" value="1140.0">1140 m</quantity>
</elevation>
,
<collectingDate id="E650CC32FF8FFFABFF4AF89B5046EC66" box="[249,342,1833,1858]" pageId="28" pageNumber="29" value="1985-03">
<date id="F61435DAFF8FFFABFF4AF89B5046EC66" box="[249,342,1833,1858]" pageId="28" pageNumber="29" value="1985-03">III.1985</date>
</collectingDate>
, Wr. 132.25, 132.81.
</materialsCitation>
</paragraph>
<paragraph id="8215131AFF8FFFA8FF74F8FF53A2EA89" blockId="28.[151,1437,1689,2038]" lastBlockId="31.[151,1437,151,608]" lastPageId="31" lastPageNumber="32" pageId="28" pageNumber="29">
<emphasis id="B0DECF08FF8FFFABFF74F8FF5049EC42" bold="true" box="[199,345,1869,1894]" pageId="28" pageNumber="29">Description.</emphasis>
Length
<quantity id="4552BEFFFF8FFFABFE08F8FF5357EC41" box="[443,583,1869,1894]" metricMagnitude="-3" metricUnit="m" metricValue="4.45" metricValueMax="5.2" metricValueMin="3.7" pageId="28" pageNumber="29" unit="mm" value="4.45" valueMax="5.2" valueMin="3.7">3.7–5.2 mm</quantity>
, width
<quantity id="4552BEFFFF8FFFABFD2CF8FF523CEC41" box="[671,812,1869,1894]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.3" metricValueMin="1.6" pageId="28" pageNumber="29" unit="mm" value="1.95" valueMax="2.3" valueMin="1.6">1.6–2.3 mm</quantity>
. General color yellowish brown (
<figureCitation id="1A910F9FFF8FFFABFB1CF8FF5414EC42" box="[1199,1284,1869,1894]" captionStart="FIGURE 16" captionStartId="29.[151,250,1942,1964]" captionTargetBox="[155,1432,409,1921]" captionTargetId="figure@29.[155,1432,409,1921]" captionTargetPageId="29" captionText="FIGURE 16. Habitus of Doryscus species, dorsal view. 16 A. D. barclayi Lee, sp. nov.; 16 B. D. boreri Lee, sp. nov.; 16 C. D. boreri Lee, sp. nov., color variation; 16 D. D. boreri Lee, sp. nov., color variation; 16 E. D. javanensis Lee, sp. nov.; 16 F. D. javanensis Lee, sp. nov., color variation." httpUri="https://zenodo.org/record/581275/files/figure.png" pageId="28" pageNumber="29">Fig. 16</figureCitation>
A). Antennae filiform, antennomere I not modified in males (
<figureCitation id="1A910F9FFF8FFFABFD1EF8C353EDECAE" box="[685,765,1905,1930]" captionStart="FIGURE 17" captionStartId="30.[151,250,1938,1960]" captionTargetBox="[162,1431,207,1900]" captionTargetId="figure@30.[151,1436,193,1916]" captionTargetPageId="30" captionText="FIGURE 17. Diagnostic characters of Doryscus barclayi Lee, sp. nov. 17 A. Antenna, male; 17 B. Antenna, female; 17 C. Penis, dorsal view; 17 D. Penis, lateral view; 17 E. Gonocoxae; 17 F. Abdominal ventrite VIII. 17 G. Apical margin of abdominal ventrite V, male; 17 H. Apical margin of abdominal ventrite V, female; 17 I. Spermatheca." httpUri="https://zenodo.org/record/581276/files/figure.png" pageId="28" pageNumber="29">Fig. 17</figureCitation>
A), length ratios of antennomeres I–XI 1.0: 0.4: 0.7: 0.6: 0.7: 0.7: 0.7: 0.7: 0.7: 0.7: 0.8, length to width ratios of antennomeres I–XI 2.9: 1.9: 2.9: 2.6: 3.0: 3.2: 3.6: 3.7: 3.6: 3.5: 4.5; narrower in females (
<figureCitation id="1A910F9FFF8FFFABFDEEF80B53BEECF6" box="[605,686,1977,2002]" captionStart="FIGURE 17" captionStartId="30.[151,250,1938,1960]" captionTargetBox="[162,1431,207,1900]" captionTargetId="figure@30.[151,1436,193,1916]" captionTargetPageId="30" captionText="FIGURE 17. Diagnostic characters of Doryscus barclayi Lee, sp. nov. 17 A. Antenna, male; 17 B. Antenna, female; 17 C. Penis, dorsal view; 17 D. Penis, lateral view; 17 E. Gonocoxae; 17 F. Abdominal ventrite VIII. 17 G. Apical margin of abdominal ventrite V, male; 17 H. Apical margin of abdominal ventrite V, female; 17 I. Spermatheca." httpUri="https://zenodo.org/record/581276/files/figure.png" pageId="28" pageNumber="29">Fig. 17</figureCitation>
B), length ratios of antennomeres I–XI 1.0: 0.4: 0.8: 0.8: 0.8: 0.8: 0.8: 0.8: 0.8: 0.7: 0.8, length to width ratios of antennomeres I–XI 3.8: 1.9: 3.8: 4.0: 4.0: 4.5: 4.6: 4.5: 4.3: 3.9: 5.0. Elytra 1.7 x longer than wide, with longitudinal ridges between punctures, with erect, yellowish brown, dense, elongate setae and coarse punctures. Apical margin of last abdominal ventrite with two extremely shallow incisions in males (
<figureCitation id="1A910F9FFF8EFFAAFEC9FF6D50DFEBDC" box="[378,463,223,248]" captionStart="FIGURE 17" captionStartId="30.[151,250,1938,1960]" captionTargetBox="[162,1431,207,1900]" captionTargetId="figure@30.[151,1436,193,1916]" captionTargetPageId="30" captionText="FIGURE 17. Diagnostic characters of Doryscus barclayi Lee, sp. nov. 17 A. Antenna, male; 17 B. Antenna, female; 17 C. Penis, dorsal view; 17 D. Penis, lateral view; 17 E. Gonocoxae; 17 F. Abdominal ventrite VIII. 17 G. Apical margin of abdominal ventrite V, male; 17 H. Apical margin of abdominal ventrite V, female; 17 I. Spermatheca." httpUri="https://zenodo.org/record/581276/files/figure.png" pageId="29" pageNumber="30">Fig. 17</figureCitation>
G); but medially depressed in females (
<figureCitation id="1A910F9FFF8EFFAAFC2EFF6D52E2EBDC" box="[925,1010,223,248]" captionStart="FIGURE 17" captionStartId="30.[151,250,1938,1960]" captionTargetBox="[162,1431,207,1900]" captionTargetId="figure@30.[151,1436,193,1916]" captionTargetPageId="30" captionText="FIGURE 17. Diagnostic characters of Doryscus barclayi Lee, sp. nov. 17 A. Antenna, male; 17 B. Antenna, female; 17 C. Penis, dorsal view; 17 D. Penis, lateral view; 17 E. Gonocoxae; 17 F. Abdominal ventrite VIII. 17 G. Apical margin of abdominal ventrite V, male; 17 H. Apical margin of abdominal ventrite V, female; 17 I. Spermatheca." httpUri="https://zenodo.org/record/581276/files/figure.png" pageId="29" pageNumber="30">Fig. 17</figureCitation>
H). Penis (
<figureCitation id="1A910F9FFF8EFFAAFBDCFF6D55D6EBDC" box="[1135,1222,223,248]" captionStart="FIGURE 17" captionStartId="30.[151,250,1938,1960]" captionTargetBox="[162,1431,207,1900]" captionTargetId="figure@30.[151,1436,193,1916]" captionTargetPageId="30" captionText="FIGURE 17. Diagnostic characters of Doryscus barclayi Lee, sp. nov. 17 A. Antenna, male; 17 B. Antenna, female; 17 C. Penis, dorsal view; 17 D. Penis, lateral view; 17 E. Gonocoxae; 17 F. Abdominal ventrite VIII. 17 G. Apical margin of abdominal ventrite V, male; 17 H. Apical margin of abdominal ventrite V, female; 17 I. Spermatheca." httpUri="https://zenodo.org/record/581276/files/figure.png" pageId="29" pageNumber="30">Figs 17</figureCitation>
C–17D) extremely slender, 7.7 x longer than wide, gradually widened towards base, apex of dorsal surface membranous; apex of ventral surface slightly depressed; straight in lateral view. Internal sac with endophallic spiculae short, about 0.6 x as long as penis, apices trilobed, median lobe extremely slender, strongly and dorso-ventrally curved at middle, with cluster of setae at middle, lateral lobes apically tapering, apices horn-like; with one pair of extremely elongate spiculae from base to midpoint, two pair of short, horn-like spiculae between elongate spiculae. Gonocoxae (
<figureCitation id="1A910F9FFF8CFFA8FADCFF0E51A2EBDC" captionStart="FIGURE 17" captionStartId="30.[151,250,1938,1960]" captionTargetBox="[162,1431,207,1900]" captionTargetId="figure@30.[151,1436,193,1916]" captionTargetPageId="30" captionText="FIGURE 17. Diagnostic characters of Doryscus barclayi Lee, sp. nov. 17 A. Antenna, male; 17 B. Antenna, female; 17 C. Penis, dorsal view; 17 D. Penis, lateral view; 17 E. Gonocoxae; 17 F. Abdominal ventrite VIII. 17 G. Apical margin of abdominal ventrite V, male; 17 H. Apical margin of abdominal ventrite V, female; 17 I. Spermatheca." httpUri="https://zenodo.org/record/581276/files/figure.png" pageId="31" pageNumber="32">Fig. 17</figureCitation>
E) slender, 6.2 x longer than wide, sides slightly narrowed at apical 1/10; apices narrowed rounded; inner margins narrowly approximate from apex to midpoint, apex with nine setae; slender from midpoint to base. Ventrite VIII (
<figureCitation id="1A910F9FFF8CFFA8FE88FE95509CEA64" box="[315,396,295,320]" captionStart="FIGURE 17" captionStartId="30.[151,250,1938,1960]" captionTargetBox="[162,1431,207,1900]" captionTargetId="figure@30.[151,1436,193,1916]" captionTargetPageId="30" captionText="FIGURE 17. Diagnostic characters of Doryscus barclayi Lee, sp. nov. 17 A. Antenna, male; 17 B. Antenna, female; 17 C. Penis, dorsal view; 17 D. Penis, lateral view; 17 E. Gonocoxae; 17 F. Abdominal ventrite VIII. 17 G. Apical margin of abdominal ventrite V, male; 17 H. Apical margin of abdominal ventrite V, female; 17 I. Spermatheca." httpUri="https://zenodo.org/record/581276/files/figure.png" pageId="31" pageNumber="32">Fig. 17</figureCitation>
F) weakly sclerotized; spiculum elongate; apex square, apical margin truncate, with sparse short setae and several long setae inside; disc with several long setae at sides. Receptacle of spermatheca (
<figureCitation id="1A910F9FFF8CFFA8FA8AFEFE549AEA40" box="[1337,1418,332,357]" captionStart="FIGURE 17" captionStartId="30.[151,250,1938,1960]" captionTargetBox="[162,1431,207,1900]" captionTargetId="figure@30.[151,1436,193,1916]" captionTargetPageId="30" captionText="FIGURE 17. Diagnostic characters of Doryscus barclayi Lee, sp. nov. 17 A. Antenna, male; 17 B. Antenna, female; 17 C. Penis, dorsal view; 17 D. Penis, lateral view; 17 E. Gonocoxae; 17 F. Abdominal ventrite VIII. 17 G. Apical margin of abdominal ventrite V, male; 17 H. Apical margin of abdominal ventrite V, female; 17 I. Spermatheca." httpUri="https://zenodo.org/record/581276/files/figure.png" pageId="31" pageNumber="32">
Fig.
<date id="F61435DAFF8CFFA8FADDFEFE549AEA40" box="[1390,1418,332,356]" pageId="31" pageNumber="32" value="1917-01">17</date>
</figureCitation>
I) narrower than pump, narrowly separated from pump; pump wide and strongly curved; proximal spermathecal duct slender, shallowly projecting into receptaculum.
</paragraph>
</subSubSection>
<caption id="D6D54392FF8EFFAAFF24F8245324ECCD" httpUri="https://zenodo.org/record/581275/files/figure.png" pageId="29" pageNumber="30" targetBox="[155,1432,409,1921]" targetPageId="29">
<paragraph id="8215131AFF8EFFAAFF24F8245324ECCD" blockId="29.[151,1437,1942,2025]" pageId="29" pageNumber="30">
<emphasis id="B0DECF08FF8EFFAAFF24F8245030EC88" bold="true" box="[151,288,1942,1964]" pageId="29" pageNumber="30">FIGURE 16.</emphasis>
Habitus of
<taxonomicName id="45AA6899FF8EFFAAFE2BF82550E8EC88" box="[408,504,1943,1964]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B0DECF08FF8EFFAAFE2BF82550E8EC88" box="[408,504,1943,1964]" italics="true" pageId="29" pageNumber="30">Doryscus</emphasis>
</taxonomicName>
species, dorsal view. 16A.
<taxonomicName id="45AA6899FF8EFFAAFCA3F82552A2EC88" authority="Lee" authorityName="Lee" box="[784,946,1942,1964]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="barclayi" status="sp. nov.">
<emphasis id="B0DECF08FF8EFFAAFCA3F8255293EC88" box="[784,899,1942,1964]" italics="true" pageId="29" pageNumber="30">D. barclayi</emphasis>
Lee
</taxonomicName>
,
<taxonomicNameLabel id="ABED7273FF8EFFAAFC0EF824551EEC88" box="[957,1038,1942,1964]" pageId="29" pageNumber="30" rank="species">
<emphasis id="B0DECF08FF8EFFAAFC0EF824551EEC88" bold="true" box="[957,1038,1942,1964]" pageId="29" pageNumber="30">sp. nov.</emphasis>
</taxonomicNameLabel>
; 16B.
<taxonomicName id="45AA6899FF8EFFAAFBE7F82555F2EC88" authority="Lee" authorityName="Lee" box="[1108,1250,1942,1964]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="boreri" status="sp. nov.">
<emphasis id="B0DECF08FF8EFFAAFBE7F82555A2EC88" box="[1108,1202,1942,1964]" italics="true" pageId="29" pageNumber="30">D. boreri</emphasis>
Lee
</taxonomicName>
,
<taxonomicNameLabel id="ABED7273FF8EFFAAFB5FF824542DEC88" box="[1260,1341,1942,1964]" pageId="29" pageNumber="30" rank="species">
<emphasis id="B0DECF08FF8EFFAAFB5FF824542DEC88" bold="true" box="[1260,1341,1942,1964]" pageId="29" pageNumber="30">sp. nov.</emphasis>
</taxonomicNameLabel>
; 16C.
<taxonomicName id="45AA6899FF8EFFAAFA30F8255015ECEF" authority="Lee" authorityName="Lee" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="boreri" status="sp. nov.">
<emphasis id="B0DECF08FF8EFFAAFA30F82551C5ECEF" italics="true" pageId="29" pageNumber="30">D. boreri</emphasis>
Lee
</taxonomicName>
,
<taxonomicNameLabel id="ABED7273FF8EFFAAFEA2F8075073ECEF" box="[273,355,1973,1995]" pageId="29" pageNumber="30" rank="species">
<emphasis id="B0DECF08FF8EFFAAFEA2F8075073ECEF" bold="true" box="[273,355,1973,1995]" pageId="29" pageNumber="30">sp. nov.</emphasis>
</taxonomicNameLabel>
, color variation; 16D.
<taxonomicName id="45AA6899FF8EFFAAFDE3F80453F0ECEF" authority="Lee" authorityName="Lee" box="[592,736,1973,1995]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="boreri" status="sp. nov.">
<emphasis id="B0DECF08FF8EFFAAFDE3F80453BFECEF" box="[592,687,1973,1995]" italics="true" pageId="29" pageNumber="30">D. boreri</emphasis>
Lee
</taxonomicName>
,
<taxonomicNameLabel id="ABED7273FF8EFFAAFD58F807522DECEF" box="[747,829,1973,1995]" pageId="29" pageNumber="30" rank="species">
<emphasis id="B0DECF08FF8EFFAAFD58F807522DECEF" bold="true" box="[747,829,1973,1995]" pageId="29" pageNumber="30">sp. nov.</emphasis>
</taxonomicNameLabel>
, color variation; 16E.
<taxonomicName id="45AA6899FF8EFFAAFB94F80455F3ECEF" authority="Lee" authorityName="Lee" box="[1063,1251,1974,1995]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="javanensis" status="sp. nov.">
<emphasis id="B0DECF08FF8EFFAAFB94F80455A3ECEF" box="[1063,1203,1974,1995]" italics="true" pageId="29" pageNumber="30">D. javanensis</emphasis>
Lee
</taxonomicName>
,
<taxonomicNameLabel id="ABED7273FF8EFFAAFB5DF8075450ECEF" box="[1262,1344,1973,1995]" pageId="29" pageNumber="30" rank="species">
<emphasis id="B0DECF08FF8EFFAAFB5DF8075450ECEF" bold="true" box="[1262,1344,1973,1995]" pageId="29" pageNumber="30">sp. nov.</emphasis>
</taxonomicNameLabel>
; 16F.
<taxonomicName id="45AA6899FF8EFFAAFA30F8045020ECCD" authority="Lee" authorityName="Lee" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="javanensis" status="sp. nov.">
<emphasis id="B0DECF08FF8EFFAAFA30F804548DECEF" box="[1411,1437,1974,1995]" italics="true" pageId="29" pageNumber="30">D.</emphasis>
<emphasis id="B0DECF08FF8EFFAAFF24F8665012ECCD" box="[151,258,2004,2025]" italics="true" pageId="29" pageNumber="30">javanensis</emphasis>
Lee
</taxonomicName>
,
<taxonomicNameLabel id="ABED7273FF8EFFAAFE88F861509BECCD" box="[315,395,2003,2025]" pageId="29" pageNumber="30" rank="species">
<emphasis id="B0DECF08FF8EFFAAFE88F861509BECCD" bold="true" box="[315,395,2003,2025]" pageId="29" pageNumber="30">sp. nov.</emphasis>
</taxonomicNameLabel>
, color variation.
</paragraph>
</caption>
<caption id="D6D54392FF8DFFA9FF24F820550BECC1" httpUri="https://zenodo.org/record/581276/files/figure.png" pageId="30" pageNumber="31" targetBox="[162,1431,207,1900]" targetPageId="30">
<paragraph id="8215131AFF8DFFA9FF24F820550BECC1" blockId="30.[151,1436,1938,2021]" pageId="30" pageNumber="31">
<emphasis id="B0DECF08FF8DFFA9FF24F820500FEC8C" bold="true" box="[151,287,1938,1960]" pageId="30" pageNumber="31">FIGURE 17.</emphasis>
Diagnostic characters of
<taxonomicName id="45AA6899FF8DFFA9FDACF8215217EC8C" authority="Lee" authorityName="Lee" box="[543,775,1938,1960]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="barclayi" status="sp. nov.">
<emphasis id="B0DECF08FF8DFFA9FDACF821536FEC8C" box="[543,639,1939,1960]" italics="true" pageId="30" pageNumber="31">Doryscus</emphasis>
<emphasis id="B0DECF08FF8DFFA9FD36F82053C9EC8C" box="[645,729,1938,1960]" italics="true" pageId="30" pageNumber="31">barclayi</emphasis>
Lee
</taxonomicName>
,
<taxonomicNameLabel id="ABED7273FF8DFFA9FCA3F8205270EC8C" box="[784,864,1938,1960]" pageId="30" pageNumber="31" rank="species">
<emphasis id="B0DECF08FF8DFFA9FCA3F8205270EC8C" bold="true" box="[784,864,1938,1960]" pageId="30" pageNumber="31">sp. nov.</emphasis>
</taxonomicNameLabel>
17A. Antenna, male; 17B. Antenna, female; 17C. Penis, dorsal view; 17D. Penis, lateral view; 17E. Gonocoxae; 17F. Abdominal ventrite VIII. 17G. Apical margin of abdominal ventrite V, male; 17H. Apical margin of abdominal ventrite V, female; 17I. Spermatheca.
</paragraph>
</caption>
<subSubSection id="CAB04091FF8CFFA8FF74FE05529AEAD1" pageId="31" pageNumber="32" type="diagnosis">
<paragraph id="8215131AFF8CFFA8FF74FE05529AEAD1" blockId="31.[151,1437,151,608]" pageId="31" pageNumber="32">
<emphasis id="B0DECF08FF8CFFA8FF74FE0550DEEAF4" bold="true" box="[199,462,439,464]" pageId="31" pageNumber="32">Differential diagnosis.</emphasis>
<taxonomicName id="45AA6899FF8CFFA8FE67FE0B53B1EAF4" box="[468,673,439,464]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="barclayi" status="sp. nov.">
<emphasis id="B0DECF08FF8CFFA8FE67FE0B53B1EAF4" box="[468,673,439,464]" italics="true" pageId="31" pageNumber="32">Doryscus barclayi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ABED7273FF8CFFA8FD1AFE055213EAF4" box="[681,771,439,464]" pageId="31" pageNumber="32" rank="species">
<emphasis id="B0DECF08FF8CFFA8FD1AFE055213EAF4" bold="true" box="[681,771,439,464]" pageId="31" pageNumber="32">sp. nov.</emphasis>
</taxonomicNameLabel>
adults are easily recognized by their longitudinal ridges on the elytra and comparatively wider antennae in males than females.
</paragraph>
</subSubSection>
<subSubSection id="CAB04091FF8CFFA8FF74FE4D5071E919" pageId="31" pageNumber="32" type="etymology">
<paragraph id="8215131AFF8CFFA8FF74FE4D5071E919" blockId="31.[151,1437,151,608]" pageId="31" pageNumber="32">
<emphasis id="B0DECF08FF8CFFA8FF74FE4D505EE93C" bold="true" box="[199,334,511,536]" pageId="31" pageNumber="32">Etymology.</emphasis>
This new species is dedicated to Maxwell V. L. Barclay (Curator, BMNH), who encouraged me to study leaf beetles.
</paragraph>
</subSubSection>
<subSubSection id="CAB04091FF8CFFA8FF74FDF5534AE944" box="[199,602,583,608]" pageId="31" pageNumber="32" type="distribution">
<paragraph id="8215131AFF8CFFA8FF74FDF5534AE944" blockId="31.[151,1437,151,608]" box="[199,602,583,608]" pageId="31" pageNumber="32">
<emphasis id="B0DECF08FF8CFFA8FF74FDF55070E944" bold="true" box="[199,352,583,608]" pageId="31" pageNumber="32">Distribution.</emphasis>
<collectingCountry id="FABD538AFF8CFFA8FED4FDF550C4E944" box="[359,468,583,608]" name="Indonesia" pageId="31" pageNumber="32">Indonesia</collectingCountry>
(
<collectingRegion id="406EDDF8FF8CFFA8FE50FDF5535EE944" box="[483,590,583,608]" country="Indonesia" name="Sulawesi Utara" pageId="31" pageNumber="32">Sulawesi</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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