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<document id="5A292E97D74C835E0BC93EF9B76A5D32" ID="10.11646/zootaxa.4269.1.1SLASH11087" ID-CLB-Dataset="33014" ID-DOI="10.11646/zootaxa.4269.1.1" ID-GBIF-Dataset="0b7fd4db-9f34-47ef-b7e8-33363155d866" ID-ISSN="1175-5326" ID-Zenodo-Dep="581259" ID-ZooBank="2AFC0D53-B439-4BBB-BC22-A0C90C08D65D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1495182755873" checkinUser="plazi" docAuthor="Lee, Chi-Feng" docDate="2017" docId="0A03A20CFF82FFA3FF24F92350CDEF34" docLanguage="en" docName="zootaxa.4269.1.1SLASH11087.pdf" docOrigin="Zootaxa 4269 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Doryscus testaceus Jacoby 1887" docType="treatment" docVersion="7" lastPageNumber="21" masterDocId="F63ADA74FF93FFB7FFB3FFB25110EB24" masterDocTitle="Revision of the genus Doryscus Jacoby (Coleoptera: Chrysomelidae: Galerucinae)" masterLastPageNumber="43" masterPageNumber="1" pageNumber="18" updateTime="1698439733711" updateUser="ExternalLinkService">
<mods:mods id="F95AD2B572FA0B6F20D7552A79E29AEC" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="52455516969BB164A32AEA8DC0492AD6">Revision of the genus Doryscus Jacoby (Coleoptera: Chrysomelidae: Galerucinae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="CE722AA7D56B816CC8ABEE99E95008C1">Lee, Chi-Feng</mods:namePart>
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<mods:title id="16A154A4A3B20EF3F48CAA56F31D5A46">Zootaxa</mods:title>
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<mods:date id="9FBF926484EC0BA9E0EEA83F575097C7">2017</mods:date>
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<mods:number id="38A5970584935E4BD24AAA026FADF293">4269</mods:number>
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<mods:identifier id="32CC82C0471A07EEB40E59F212D8173D" type="DOI">10.11646/zootaxa.4269.1.1</mods:identifier>
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<treatment id="0A03A20CFF82FFA3FF24F92350CDEF34" ID-DOI="http://doi.org/10.5281/zenodo.6003739" ID-GBIF-Taxon="130810394" ID-Zenodo-Dep="6003739" LSID="urn:lsid:plazi:treatment:0A03A20CFF82FFA3FF24F92350CDEF34" httpUri="http://treatment.plazi.org/id/0A03A20CFF82FFA3FF24F92350CDEF34" lastPageId="20" lastPageNumber="21" pageId="17" pageNumber="18">
<subSubSection id="CAB04091FF82FFA6FF24F923504CEDE8" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="8215131AFF82FFA6FF24F9235332ED8F" blockId="17.[151,546,1681,1741]" box="[151,546,1681,1707]" pageId="17" pageNumber="18">
<heading id="D95DA476FF82FFA6FF24F9235332ED8F" bold="true" box="[151,546,1681,1707]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
<taxonomicName id="45AA6899FF82FFA6FF24F9235332ED8F" authority="Jacoby, 1887" authorityName="Jacoby" authorityYear="1887" box="[151,546,1681,1707]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="testaceus">
<emphasis id="B0DECF08FF82FFA6FF24F9235332ED8F" bold="true" box="[151,546,1681,1707]" pageId="17" pageNumber="18">
<emphasis id="B0DECF08FF82FFA6FF24F923506BED8F" bold="true" box="[151,379,1681,1707]" italics="true" pageId="17" pageNumber="18">Doryscus testaceus</emphasis>
<bibRefCitation id="E63B6EEBFF82FFA6FE31F9235332ED8F" author="Jacoby" box="[386,546,1681,1707]" pageId="17" pageNumber="18" refString="Jacoby, M. (1887) Descriptions of the phytophagous Coleoptera of Ceylon, obtain by Mr. George Lewis during the years 1881 - 1882. Proceedings of the Scientific Meetings of the Zoological Society of London, 1887, 65 - 118." type="proceedings paper" year="1887">Jacoby, 1887</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8215131AFF82FFA6FF24F906504CEDE8" blockId="17.[151,546,1681,1741]" box="[151,348,1716,1741]" pageId="17" pageNumber="18">
(
<figureCitation id="1A910F9FFF82FFA6FF2CF90651F5EDE8" box="[159,229,1716,1741]" captionStart="FIGURE 9" captionStartId="18.[151,250,1870,1892]" captionTargetBox="[155,1432,337,1849]" captionTargetId="figure@18.[155,1432,337,1849]" captionTargetPageId="18" captionText="FIGURE 9. Habitus of Doryscus species, dorsal view. 9 A. D. testaceus Jacoby; 9 B. D. testaceus Jacoby, color variation; 9 C. D. nepalensis Lee, sp. nov.; 9 D. D. nepalensis Lee, sp. nov., color variation; 9 E. D. nepalensis Lee, sp. nov., color variation; 9 F. D. niger Medvedev &amp; Sprecher-Uebersax." httpUri="https://zenodo.org/record/581268/files/figure.png" pageId="17" pageNumber="18">Figs 9</figureCitation>
A–9B, 10)
</paragraph>
</subSubSection>
<subSubSection id="CAB04091FF82FFA6FF24F94F547DECBB" pageId="17" pageNumber="18" type="reference_group">
<paragraph id="8215131AFF82FFA6FF24F94F53CDEC36" blockId="17.[151,733,1788,1810]" box="[151,733,1788,1810]" pageId="17" pageNumber="18">
<treatmentCitationGroup id="A2BA3434FF82FFA6FF24F94F53CDEC36" box="[151,733,1788,1810]" pageId="17" pageNumber="18">
<taxonomicName id="45AA6899FF82FFA6FF24F94F5309EC36" authority="Jacoby, 1887: 115" authorityName="Jacoby" authorityPageNumber="115" authorityYear="1887" box="[151,537,1788,1810]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="testaceus">
<emphasis id="B0DECF08FF82FFA6FF24F94F504AEC36" box="[151,346,1789,1810]" italics="true" pageId="17" pageNumber="18">Doryscus testaceus</emphasis>
<treatmentCitation id="030B350BFF82FFA6FED3F94E5309EC36" author="Jacoby" box="[352,537,1788,1810]" page="115" pageId="17" pageNumber="18" year="1887">
<bibRefCitation id="E63B6EEBFF82FFA6FED3F94E50F7EC36" author="Jacoby" box="[352,487,1788,1810]" pageId="17" pageNumber="18" refString="Jacoby, M. (1887) Descriptions of the phytophagous Coleoptera of Ceylon, obtain by Mr. George Lewis during the years 1881 - 1882. Proceedings of the Scientific Meetings of the Zoological Society of London, 1887, 65 - 118." type="proceedings paper" year="1887">Jacoby, 1887</bibRefCitation>
: 115
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="030B350BFF82FFA6FD90F94E53C9EC36" author="Kimoto" box="[547,729,1788,1810]" page="31" pageId="17" pageNumber="18" year="2003">
<bibRefCitation id="E63B6EEBFF82FFA6FD90F94E53A3EC36" author="Kimoto" box="[547,691,1788,1810]" pageId="17" pageNumber="18" refString="Kimoto, S. (2003) The Chrysomelidae (Insecta: Coleoptera) collected by Dr. Akio Otake, on the occasion of his entomological survey in Sri Lanka from 1973 to 1975. Bulletin of the Kitakyushu Museum of Natural History and Human History, Series A, 1, 23 - 43." type="journal article" year="2003">Kimoto, 2003</bibRefCitation>
: 31
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8215131AFF82FFA6FF24F88C547DECBB" blockId="17.[151,1437,1854,2023]" pageId="17" pageNumber="18">
<materialsCitation id="32C21947FF82FFA6FF24F88C52E5EC5F" ID-GBIF-Occurrence="1503376913" collectionCode="BMNH" collectorName="G. Lewis &amp; H. Takizawa" country="Sri Lanka" location="Ceylon" pageId="17" pageNumber="18" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
<emphasis id="B0DECF08FF82FFA6FF24F88C5023EC73" bold="true" box="[151,307,1854,1879]" pageId="17" pageNumber="18">
<typeStatus id="5D11ADB8FF82FFA6FF24F88C51CCEC73" box="[151,220,1854,1879]" pageId="17" pageNumber="18">Types</typeStatus>
(n= 2).
</emphasis>
<typeStatus id="5D11ADB8FF82FFA6FE89F88D50BAEC73" box="[314,426,1855,1879]" pageId="17" pageNumber="18" type="lectotype">Lectotype</typeStatus>
<specimenCount id="94ACD893FF82FFA6FE02F88D50D6EC72" box="[433,454,1855,1878]" pageId="17" pageNumber="18" type="female">♀</specimenCount>
(
<collectionCode id="E4BB8BDFFF82FFA6FE66F88D533FEC73" box="[469,559,1855,1879]" pageId="17" pageNumber="18">BMNH</collectionCode>
), here designated, labeled: “
<collectingCountry id="FABD538AFF82FFA6FCD8F88C52D1EC73" box="[875,961,1854,1879]" name="Sri Lanka" pageId="17" pageNumber="18">Ceylon</collectingCountry>
. /
<collectorName id="2F5F76CCFF82FFA6FC6FF88D5552EC73" box="[988,1090,1854,1879]" pageId="17" pageNumber="18">G. Lewis</collectorName>
, 1910-320 [p, w] //
<taxonomicName id="45AA6899FF82FFA6FA90F88D549DEC72" box="[1315,1421,1855,1878]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B0DECF08FF82FFA6FA90F88D549DEC72" box="[1315,1421,1855,1878]" italics="true" pageId="17" pageNumber="18">Doryscus</emphasis>
</taxonomicName>
/
<emphasis id="B0DECF08FF82FFA6FF24F8D6501CEC5F" box="[151,268,1892,1915]" italics="true" pageId="17" pageNumber="18">testaceous</emphasis>
[sic!] / Jac. [h, b] //
<typeStatus id="5D11ADB8FF82FFA6FE47F8D1533CEC5F" box="[500,556,1891,1915]" pageId="17" pageNumber="18">Type</typeStatus>
/
<collectorName id="2F5F76CCFF82FFA6FDF0F8D1536CEC5E" box="[579,636,1891,1914]" name="H. Takizawa" pageId="17" pageNumber="18">H. T.</collectorName>
[p, circle label with red border]”
</materialsCitation>
.
<materialsCitation id="32C21947FF82FFA6FBB3F8D0547DECBB" ID-GBIF-Occurrence="1503376906" collectionCode="MCZC" country="Sri Lanka" location="Ceylon" pageId="17" pageNumber="18" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<typeStatus id="5D11ADB8FF82FFA6FBB3F8D0558DEC5F" box="[1024,1181,1890,1915]" pageId="17" pageNumber="18" type="paralectotype">Paralectotype</typeStatus>
:
<specimenCount id="94ACD893FF82FFA6FB1BF8D155DBEC5F" box="[1192,1227,1891,1915]" pageId="17" pageNumber="18" type="female">1♀</specimenCount>
(
<collectionCode id="E4BB8BDFFF82FFA6FB69F8D1543FEC5F" box="[1242,1327,1891,1915]" pageId="17" pageNumber="18">MCZC</collectionCode>
), labeled: “
<collectingCountry id="FABD538AFF82FFA6FF12F83451E4ECBB" box="[161,244,1926,1951]" name="Sri Lanka" pageId="17" pageNumber="18">Ceylon</collectingCountry>
/ Lewis [h, w] / 1st Jacoby / Coll. [p, w] //
<taxonomicName id="45AA6899FF82FFA6FD6BF8355252ECBA" box="[728,834,1927,1950]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B0DECF08FF82FFA6FD6BF8355252ECBA" box="[728,834,1927,1950]" italics="true" pageId="17" pageNumber="18">Doryscus</emphasis>
</taxonomicName>
/
<taxonomicName id="45AA6899FF82FFA6FCE4F83552AEECBA" box="[855,958,1927,1950]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="testaceus">
<emphasis id="B0DECF08FF82FFA6FCE4F83552AEECBA" box="[855,958,1927,1950]" italics="true" pageId="17" pageNumber="18">testaceus</emphasis>
</taxonomicName>
/ Jac. [h, b] //
<typeStatus id="5D11ADB8FF82FFA6FBECF8355587ECBB" box="[1119,1175,1927,1951]" pageId="17" pageNumber="18">Type</typeStatus>
[p] / 18501 [h, r]”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="CAB04091FF82FFA5FF74F818508AEA38" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="8215131AFF82FFA5FF74F818508AEA38" blockId="17.[151,1437,1854,2023]" lastBlockId="18.[151,1436,151,285]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<emphasis id="B0DECF08FF82FFA6FF74F81853F8ECE7" bold="true" box="[199,744,1962,1987]" pageId="17" pageNumber="18">Additional specimens (n= 13). SRI LANKA</emphasis>
. 1♂ (BMNH), Babapitiya,
<date id="F61435DAFF82FFA6FB81F8195582ECE7" box="[1074,1170,1963,1987]" pageId="17" pageNumber="18" value="1892-06">VI.1892</date>
, leg. H. P. Green; 1♂ (NHMB), Danbulla env.,
<date id="F61435DAFF82FFA6FE0BF87D5361ECC3" box="[440,625,1999,2023]" pageId="17" pageNumber="18" value="1991-04-19" valueMax="1991-05-09" valueMin="1991-04-19">19.IV.–9.V.1991</date>
, leg. J. Kallbáč; 2♂♂ (BMNH), Kandy,
<date id="F61435DAFF82FFA6FB8EF87D558CECC3" box="[1085,1180,1999,2023]" pageId="17" pageNumber="18" value="1908-06">VI.1908</date>
, leg. C. B. Bryant;
<materialsCitation id="32C21947FF82FFA5FACAF87D5249EB94" ID-GBIF-Occurrence="1503376914" collectingDate="1954-02-04" collectionCode="NHMB" collectorName="L. Horton's &amp; F. Keiser" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1">
<specimenCount id="94ACD893FF82FFA6FACAF87D548CECC3" box="[1401,1436,1999,2023]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
(
<collectionCode id="E4BB8BDFFF81FFA5FF2CFF2A51E9EB94" box="[159,249,152,176]" country="Switzerland" httpUri="http://grbio.org/cool/1d5a-4efs" name="Naturhistorisches Museum, Basel" pageId="18" pageNumber="19">NHMB</collectionCode>
), same locality,
<collectingDate id="E650CC32FF81FFA5FE19FF255304EB94" box="[426,532,151,176]" pageId="18" pageNumber="19" value="1954-02-04">
<date id="F61435DAFF81FFA5FE19FF255304EB94" box="[426,532,151,176]" pageId="18" pageNumber="19" value="1954-02-04">4.II.1954</date>
</collectingDate>
, leg.
<collectorName id="2F5F76CCFF81FFA5FDFDFF2A53DEEB94" box="[590,718,152,176]" pageId="18" pageNumber="19">L. Horton’s</collectorName>
&amp;
<collectorName id="2F5F76CCFF81FFA5FD42FF2A5249EB94" box="[753,857,151,176]" pageId="18" pageNumber="19">F. Keiser</collectorName>
</materialsCitation>
;
<materialsCitation id="32C21947FF81FFA5FCD0FF2A51E2EBF1" ID-GBIF-Occurrence="1503376917" collectingDate="1991-04-01" collectingDateMax="1991-04-18" collectingDateMin="1991-04-01" collectionCode="NHMB" collectorName="J. Kallbac" pageId="18" pageNumber="19" specimenCount="4" specimenCount-male="4">
<specimenCount id="94ACD893FF81FFA5FCD0FF2A528BEB94" box="[867,923,152,176]" pageId="18" pageNumber="19" type="male">4♂♂</specimenCount>
(
<collectionCode id="E4BB8BDFFF81FFA5FC19FF2A5513EB94" box="[938,1027,152,176]" country="Switzerland" httpUri="http://grbio.org/cool/1d5a-4efs" name="Naturhistorisches Museum, Basel" pageId="18" pageNumber="19">NHMB</collectionCode>
), same locality,
<collectingDate id="E650CC32FF81FFA5FB06FF2A545FEB94" box="[1205,1359,152,176]" pageId="18" pageNumber="19" value="1991-04-01" valueMax="1991-04-18" valueMin="1991-04-01">
<date id="F61435DAFF81FFA5FB06FF2A545FEB94" box="[1205,1359,152,176]" pageId="18" pageNumber="19" value="1991-04-01" valueMax="1991-04-18" valueMin="1991-04-01">1–18.IV.1991</date>
</collectingDate>
, leg.
<collectorName id="2F5F76CCFF81FFA5FA39FF2A51E2EBF1" pageId="18" pageNumber="19">J. Kallbáč</collectorName>
</materialsCitation>
; 1♀ (EUMJ), Kandy: Hills South of the Lake,
<date id="F61435DAFF81FFA5FC94FF0E528EEBF0" box="[807,926,188,212]" pageId="18" pageNumber="19" value="1974-02-18">18.II.1974</date>
, leg. A. Otake;
<materialsCitation id="32C21947FF81FFA5FBD2FF0E50BEEBDC" ID-GBIF-Occurrence="1503376921" collectingDate="1975-03-04" collectingDateMax="1991-04-18" collectingDateMin="1991-04-01" collectionCode="EUMJ" collectorName="A. Otake" pageId="18" pageNumber="19" specimenCount="1" specimenCount-male="1">
<specimenCount id="94ACD893FF81FFA5FBD2FF0E5594EBF0" box="[1121,1156,188,212]" pageId="18" pageNumber="19" type="male">1♂</specimenCount>
(
<collectionCode id="E4BB8BDFFF81FFA5FB24FF0E55F6EBF0" box="[1175,1254,188,212]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="18" pageNumber="19">EUMJ</collectionCode>
), same locality,
<collectingDate id="E650CC32FF81FFA5FF24FF6D5019EBDC" box="[151,265,223,248]" pageId="18" pageNumber="19" value="1975-03-04">
<date id="F61435DAFF81FFA5FF24FF6D5019EBDC" box="[151,265,223,248]" pageId="18" pageNumber="19" value="1975-03-04">4.III.1975</date>
</collectingDate>
, leg.
<collectorName id="2F5F76CCFF81FFA5FEF5FF5250BEEBDC" box="[326,430,223,248]" pageId="18" pageNumber="19">A. Otake</collectorName>
</materialsCitation>
; 1♂ (EUMJ), Kandy: Uda wattekele Sanctuary,
<date id="F61435DAFF81FFA5FC7DFF52552FEBDC" box="[974,1087,223,248]" pageId="18" pageNumber="19" value="1975-04-01">1.IV.1975</date>
, leg. A. Otake; 1♀ (MNHUB), Nalanda, leg. W. Horn.
</paragraph>
</subSubSection>
<caption id="D6D54392FF81FFA5FF24F8FC537BEC85" httpUri="https://zenodo.org/record/581268/files/figure.png" pageId="18" pageNumber="19" targetBox="[155,1432,337,1849]" targetPageId="18">
<paragraph id="8215131AFF81FFA5FF24F8FC537BEC85" blockId="18.[151,1436,1870,1953]" pageId="18" pageNumber="19">
<emphasis id="B0DECF08FF81FFA5FF24F8FC5003EC40" bold="true" box="[151,275,1870,1892]" pageId="18" pageNumber="19">FIGURE 9.</emphasis>
Habitus of
<taxonomicName id="45AA6899FF81FFA5FE3EF8FD50FDEC40" box="[397,493,1871,1892]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B0DECF08FF81FFA5FE3EF8FD50FDEC40" box="[397,493,1871,1892]" italics="true" pageId="18" pageNumber="19">Doryscus</emphasis>
</taxonomicName>
species, dorsal view. 9A.
<taxonomicName id="45AA6899FF81FFA5FD49F8FD52D8EC40" authority="Jacoby" authorityName="Jacoby" box="[762,968,1870,1892]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="testaceus">
<emphasis id="B0DECF08FF81FFA5FD49F8FD5267EC40" box="[762,887,1871,1892]" italics="true" pageId="18" pageNumber="19">D. testaceus</emphasis>
Jacoby
</taxonomicName>
; 9B.
<taxonomicName id="45AA6899FF81FFA5FC4DF8FD55DAEC40" authority="Jacoby" authorityName="Jacoby" box="[1022,1226,1870,1892]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="testaceus">
<emphasis id="B0DECF08FF81FFA5FC4DF8FD556AEC40" box="[1022,1146,1871,1892]" italics="true" pageId="18" pageNumber="19">D. testaceus</emphasis>
Jacoby
</taxonomicName>
, color variation; 9C.
<taxonomicName id="45AA6899FF81FFA5FF24F8DC5042ECA7" authority="Lee" authorityName="Lee" box="[151,338,1901,1923]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="nepalensis" status="sp. nov.">
<emphasis id="B0DECF08FF81FFA5FF24F8DC5032ECA7" box="[151,290,1901,1923]" italics="true" pageId="18" pageNumber="19">D. nepalensis</emphasis>
Lee
</taxonomicName>
,
<taxonomicNameLabel id="ABED7273FF81FFA5FEEFF8DF50BEECA7" box="[348,430,1901,1923]" pageId="18" pageNumber="19" rank="species">
<emphasis id="B0DECF08FF81FFA5FEEFF8DF50BEECA7" bold="true" box="[348,430,1901,1923]" pageId="18" pageNumber="19">sp. nov.</emphasis>
</taxonomicNameLabel>
; 9D.
<taxonomicName id="45AA6899FF81FFA5FE5AF8DC53B3ECA7" authority="Lee" authorityName="Lee" box="[489,675,1901,1923]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="nepalensis" status="sp. nov.">
<emphasis id="B0DECF08FF81FFA5FE5AF8DC5363ECA7" box="[489,627,1901,1923]" italics="true" pageId="18" pageNumber="19">D. nepalensis</emphasis>
Lee
</taxonomicName>
,
<taxonomicNameLabel id="ABED7273FF81FFA5FD1DF8DF53EFECA7" box="[686,767,1901,1923]" pageId="18" pageNumber="19" rank="species">
<emphasis id="B0DECF08FF81FFA5FD1DF8DF53EFECA7" bold="true" box="[686,767,1901,1923]" pageId="18" pageNumber="19">sp. nov.</emphasis>
</taxonomicNameLabel>
, color variation; 9E.
<taxonomicName id="45AA6899FF81FFA5FC68F8DC5585ECA7" authority="Lee" authorityName="Lee" box="[987,1173,1901,1923]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="nepalensis" status="sp. nov.">
<emphasis id="B0DECF08FF81FFA5FC68F8DC5576ECA7" box="[987,1126,1901,1923]" italics="true" pageId="18" pageNumber="19">D. nepalensis</emphasis>
Lee
</taxonomicName>
,
<taxonomicNameLabel id="ABED7273FF81FFA5FB13F8DF55E1ECA7" box="[1184,1265,1901,1923]" pageId="18" pageNumber="19" rank="species">
<emphasis id="B0DECF08FF81FFA5FB13F8DF55E1ECA7" bold="true" box="[1184,1265,1901,1923]" pageId="18" pageNumber="19">sp. nov.</emphasis>
</taxonomicNameLabel>
, color variation; 9F.
<taxonomicName id="45AA6899FF81FFA5FF0FF83E537BEC85" authority="Medvedev &amp; Sprecher-Uebersax." authorityName="Medvedev &amp; Sprecher-Uebersax." box="[188,619,1931,1953]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="B0DECF08FF81FFA5FF0FF83E5000EC85" box="[188,272,1932,1953]" italics="true" pageId="18" pageNumber="19">D. niger</emphasis>
Medvedev &amp; Sprecher-Uebersax.
</taxonomicName>
</paragraph>
</caption>
<caption id="D6D54392FF80FFA4FF24F826550BECC3" httpUri="https://zenodo.org/record/581269/files/figure.png" pageId="19" pageNumber="20" targetBox="[162,1412,208,1909]" targetPageId="19">
<paragraph id="8215131AFF80FFA4FF24F826550BECC3" blockId="19.[151,1436,1940,2023]" pageId="19" pageNumber="20">
<emphasis id="B0DECF08FF80FFA4FF24F8265032EC8E" bold="true" box="[151,290,1940,1962]" pageId="19" pageNumber="20">FIGURE 10.</emphasis>
Diagnostic characters of
<taxonomicName id="45AA6899FF80FFA4FD9CF8275259EC8E" authority="Jacoby." authorityName="Jacoby." box="[559,841,1940,1962]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="testaceus">
<emphasis id="B0DECF08FF80FFA4FD9CF82753E4EC8E" box="[559,756,1941,1962]" italics="true" pageId="19" pageNumber="20">Doryscus testaceus</emphasis>
Jacoby.
</taxonomicName>
10A. Antenna, male; 10B. Antenna, female; 10C. Penis, dorsal view; 10D. Penis, lateral view; 10E. Gonocoxae; 10F. Abdominal ventrite VIII. 10G. Apical margin of abdominal ventrite V, male; 10H. Apical margin of abdominal ventrite V, female; 10I. Spermatheca.
</paragraph>
</caption>
<subSubSection id="CAB04091FF87FFA3FF74FF255344E881" pageId="20" pageNumber="21" type="description">
<paragraph id="8215131AFF87FFA3FF74FF255344E881" blockId="20.[151,1437,151,1040]" pageId="20" pageNumber="21">
<emphasis id="B0DECF08FF87FFA3FF74FF255064EB94" bold="true" box="[199,372,151,176]" pageId="20" pageNumber="21">Redescription.</emphasis>
Length
<quantity id="4552BEFFFF87FFA3FE63FF255349EB8B" box="[464,601,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="4.8" metricValueMin="3.5" pageId="20" pageNumber="21" unit="mm" value="4.15" valueMax="4.8" valueMin="3.5">3.5–4.8 mm</quantity>
, width 1.5–2.0 mm. General color yellow, but suture and lateral margins of elytra black (
<figureCitation id="1A910F9FFF87FFA3FE9EFF0E5063EBF0" box="[301,371,188,213]" captionStart="FIGURE 9" captionStartId="18.[151,250,1870,1892]" captionTargetBox="[155,1432,337,1849]" captionTargetId="figure@18.[155,1432,337,1849]" captionTargetPageId="18" captionText="FIGURE 9. Habitus of Doryscus species, dorsal view. 9 A. D. testaceus Jacoby; 9 B. D. testaceus Jacoby, color variation; 9 C. D. nepalensis Lee, sp. nov.; 9 D. D. nepalensis Lee, sp. nov., color variation; 9 E. D. nepalensis Lee, sp. nov., color variation; 9 F. D. niger Medvedev &amp; Sprecher-Uebersax." httpUri="https://zenodo.org/record/581268/files/figure.png" pageId="20" pageNumber="21">Fig. 9</figureCitation>
A); some individuals with black stripes reduced (
<figureCitation id="1A910F9FFF87FFA3FC15FF0E52FBEBF0" box="[934,1003,188,213]" captionStart="FIGURE 9" captionStartId="18.[151,250,1870,1892]" captionTargetBox="[155,1432,337,1849]" captionTargetId="figure@18.[155,1432,337,1849]" captionTargetPageId="18" captionText="FIGURE 9. Habitus of Doryscus species, dorsal view. 9 A. D. testaceus Jacoby; 9 B. D. testaceus Jacoby, color variation; 9 C. D. nepalensis Lee, sp. nov.; 9 D. D. nepalensis Lee, sp. nov., color variation; 9 E. D. nepalensis Lee, sp. nov., color variation; 9 F. D. niger Medvedev &amp; Sprecher-Uebersax." httpUri="https://zenodo.org/record/581268/files/figure.png" pageId="20" pageNumber="21">Fig. 9</figureCitation>
B). First antennomere not modified in males (
<figureCitation id="1A910F9FFF87FFA3FF54FF6D5029EBDC" box="[231,313,223,248]" captionStart="FIGURE 10" captionStartId="19.[151,250,1940,1962]" captionTargetBox="[162,1412,208,1909]" captionTargetId="figure@19.[155,1427,193,1923]" captionTargetPageId="19" captionText="FIGURE 10. Diagnostic characters of Doryscus testaceus Jacoby. 10 A. Antenna, male; 10 B. Antenna, female; 10 C. Penis, dorsal view; 10 D. Penis, lateral view; 10 E. Gonocoxae; 10 F. Abdominal ventrite VIII. 10 G. Apical margin of abdominal ventrite V, male; 10 H. Apical margin of abdominal ventrite V, female; 10 I. Spermatheca." httpUri="https://zenodo.org/record/581269/files/figure.png" pageId="20" pageNumber="21">Fig. 10</figureCitation>
A), length ratios of antennomeres I–XI 1.0: 0.4: 0.6: 0.9: 0.8: 0.8: 0.9: 0.8: 0.8: 0.8: 1.0, length to width ratios of antennomeres I–XI 4.0: 1.7: 2.8: 3.7: 3.7: 4.1: 4.2: 4.9: 5.0: 4.6: 5.7; first antennomere in female more slender (
<figureCitation id="1A910F9FFF87FFA3FE39FE9550CCEA64" box="[394,476,295,320]" captionStart="FIGURE 10" captionStartId="19.[151,250,1940,1962]" captionTargetBox="[162,1412,208,1909]" captionTargetId="figure@19.[155,1427,193,1923]" captionTargetPageId="19" captionText="FIGURE 10. Diagnostic characters of Doryscus testaceus Jacoby. 10 A. Antenna, male; 10 B. Antenna, female; 10 C. Penis, dorsal view; 10 D. Penis, lateral view; 10 E. Gonocoxae; 10 F. Abdominal ventrite VIII. 10 G. Apical margin of abdominal ventrite V, male; 10 H. Apical margin of abdominal ventrite V, female; 10 I. Spermatheca." httpUri="https://zenodo.org/record/581269/files/figure.png" pageId="20" pageNumber="21">Fig. 10</figureCitation>
B), length ratios of antennomeres I–X (XI lost) 1.0: 0.4: 0.6: 0.8: 0.8: 0.8: 0.8: 0.8: 0.8: 0.8, length to width ratios of antennomeres I–X 4.5: 1.9: 3.0: 3.9: 3.9: 4.2: 4.3: 4.2: 4.2: 4.2. Elytra 1.9 x longer than wide; surface with erect, pale, dense, elongate setae; and with coarse punctures. Apical margin of last abdominal ventrite with two well developed incisions in male (
<figureCitation id="1A910F9FFF87FFA3FCDAFE2652ADEA88" box="[873,957,404,429]" captionStart="FIGURE 10" captionStartId="19.[151,250,1940,1962]" captionTargetBox="[162,1412,208,1909]" captionTargetId="figure@19.[155,1427,193,1923]" captionTargetPageId="19" captionText="FIGURE 10. Diagnostic characters of Doryscus testaceus Jacoby. 10 A. Antenna, male; 10 B. Antenna, female; 10 C. Penis, dorsal view; 10 D. Penis, lateral view; 10 E. Gonocoxae; 10 F. Abdominal ventrite VIII. 10 G. Apical margin of abdominal ventrite V, male; 10 H. Apical margin of abdominal ventrite V, female; 10 I. Spermatheca." httpUri="https://zenodo.org/record/581269/files/figure.png" pageId="20" pageNumber="21">Fig. 10</figureCitation>
G); but medially and slightly depressed in females (
<figureCitation id="1A910F9FFF87FFA3FF4DFE055040EAF4" box="[254,336,439,464]" captionStart="FIGURE 10" captionStartId="19.[151,250,1940,1962]" captionTargetBox="[162,1412,208,1909]" captionTargetId="figure@19.[155,1427,193,1923]" captionTargetPageId="19" captionText="FIGURE 10. Diagnostic characters of Doryscus testaceus Jacoby. 10 A. Antenna, male; 10 B. Antenna, female; 10 C. Penis, dorsal view; 10 D. Penis, lateral view; 10 E. Gonocoxae; 10 F. Abdominal ventrite VIII. 10 G. Apical margin of abdominal ventrite V, male; 10 H. Apical margin of abdominal ventrite V, female; 10 I. Spermatheca." httpUri="https://zenodo.org/record/581269/files/figure.png" pageId="20" pageNumber="21">Fig. 10</figureCitation>
H). Penis (
<figureCitation id="1A910F9FFF87FFA3FE74FE05530BEAF4" box="[455,539,439,464]" captionStart="FIGURE 10" captionStartId="19.[151,250,1940,1962]" captionTargetBox="[162,1412,208,1909]" captionTargetId="figure@19.[155,1427,193,1923]" captionTargetPageId="19" captionText="FIGURE 10. Diagnostic characters of Doryscus testaceus Jacoby. 10 A. Antenna, male; 10 B. Antenna, female; 10 C. Penis, dorsal view; 10 D. Penis, lateral view; 10 E. Gonocoxae; 10 F. Abdominal ventrite VIII. 10 G. Apical margin of abdominal ventrite V, male; 10 H. Apical margin of abdominal ventrite V, female; 10 I. Spermatheca." httpUri="https://zenodo.org/record/581269/files/figure.png" pageId="20" pageNumber="21">Figs 10</figureCitation>
C–10D) extremely slender, 8.1 x longer than wide, gradually narrowed towards apex; apex of dorsal surface with moderately deep notch; apex of ventral surface with shallow notch; almost straight in lateral view, apex with groove between dorsal and ventral surface; ventral surface smooth. Endophallic spiculae extremely elongate, 0.8 x as long as penis, two pairs of median spiculae, one pair extremely slender, apices filiform and curved upwards, the other pair short and apices with four teeth; one apical endophallic spiculum present between two median endophalic spiculae, apex curved downwards, medially widened; two pairs of dorsal spiculae at middle of median endophallic spiculae, one pair extremely slender, almost straight, the other pair very short; one pair of ventral endophallic spiculae slender. Gonocoxae (
<figureCitation id="1A910F9FFF87FFA3FC38FD0652CCE9E8" box="[907,988,692,717]" captionStart="FIGURE 10" captionStartId="19.[151,250,1940,1962]" captionTargetBox="[162,1412,208,1909]" captionTargetId="figure@19.[155,1427,193,1923]" captionTargetPageId="19" captionText="FIGURE 10. Diagnostic characters of Doryscus testaceus Jacoby. 10 A. Antenna, male; 10 B. Antenna, female; 10 C. Penis, dorsal view; 10 D. Penis, lateral view; 10 E. Gonocoxae; 10 F. Abdominal ventrite VIII. 10 G. Apical margin of abdominal ventrite V, male; 10 H. Apical margin of abdominal ventrite V, female; 10 I. Spermatheca." httpUri="https://zenodo.org/record/581269/files/figure.png" pageId="20" pageNumber="21">Fig. 10</figureCitation>
E) slender, 6.1 x longer than wide, sides slightly narrowed at apical 1/10; apices narrowed rounded; inner margins closely approximate from apex to apical 1/3, apex with nine setae; extremely slender from midpoint to base but abruptly widened at base. Ventrite VIII (
<figureCitation id="1A910F9FFF87FFA3FADCFD4E51A3E81C" captionStart="FIGURE 10" captionStartId="19.[151,250,1940,1962]" captionTargetBox="[162,1412,208,1909]" captionTargetId="figure@19.[155,1427,193,1923]" captionTargetPageId="19" captionText="FIGURE 10. Diagnostic characters of Doryscus testaceus Jacoby. 10 A. Antenna, male; 10 B. Antenna, female; 10 C. Penis, dorsal view; 10 D. Penis, lateral view; 10 E. Gonocoxae; 10 F. Abdominal ventrite VIII. 10 G. Apical margin of abdominal ventrite V, male; 10 H. Apical margin of abdominal ventrite V, female; 10 I. Spermatheca." httpUri="https://zenodo.org/record/581269/files/figure.png" pageId="20" pageNumber="21">Fig. 10</figureCitation>
F) weakly sclerotized; spiculum elongate; apex square, apical margin truncate, with dense, short setae; disc with several long setae at sides. Receptacle of spermatheca (
<figureCitation id="1A910F9FFF87FFA3FCBAFCF6524BE878" box="[777,859,836,861]" captionStart="FIGURE 10" captionStartId="19.[151,250,1940,1962]" captionTargetBox="[162,1412,208,1909]" captionTargetId="figure@19.[155,1427,193,1923]" captionTargetPageId="19" captionText="FIGURE 10. Diagnostic characters of Doryscus testaceus Jacoby. 10 A. Antenna, male; 10 B. Antenna, female; 10 C. Penis, dorsal view; 10 D. Penis, lateral view; 10 E. Gonocoxae; 10 F. Abdominal ventrite VIII. 10 G. Apical margin of abdominal ventrite V, male; 10 H. Apical margin of abdominal ventrite V, female; 10 I. Spermatheca." httpUri="https://zenodo.org/record/581269/files/figure.png" pageId="20" pageNumber="21">
Fig.
<date id="F61435DAFF87FFA3FC8CFCF6524BE878" box="[831,859,836,860]" pageId="20" pageNumber="21" value="2010-01">10</date>
</figureCitation>
I) slightly swollen, narrowly separated from pump; pump much narrower and strongly curved; proximal spermathecal duct slender and much narrower than receptacle; shallowly projecting into receptaculum.
</paragraph>
</subSubSection>
<subSubSection id="CAB04091FF87FFA3FF74FC1D52B8E8C9" pageId="20" pageNumber="21" type="diagnosis">
<paragraph id="8215131AFF87FFA3FF74FC1D52B8E8C9" blockId="20.[151,1437,151,1040]" pageId="20" pageNumber="21">
<emphasis id="B0DECF08FF87FFA3FF74FC1D50C3E8EC" bold="true" box="[199,467,943,968]" pageId="20" pageNumber="21">Differential diagnosis.</emphasis>
<taxonomicName id="45AA6899FF87FFA3FE53FC0353ADE8EC" box="[480,701,945,968]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="testaceus">
<emphasis id="B0DECF08FF87FFA3FE53FC0353ADE8EC" box="[480,701,945,968]" italics="true" pageId="20" pageNumber="21">Doryscus testaceus</emphasis>
</taxonomicName>
adults are similar to
<taxonomicName id="45AA6899FF87FFA3FC75FC035552E8EC" box="[966,1090,945,968]" class="Insecta" family="Chrysomelidae" genus="Doryscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="varians">
<emphasis id="B0DECF08FF87FFA3FC75FC035552E8EC" box="[966,1090,945,968]" italics="true" pageId="20" pageNumber="21">D. varians</emphasis>
</taxonomicName>
in color patterns but can be distinguished by presence of dense, elongate, erect setae on the elytra.
</paragraph>
</subSubSection>
<subSubSection id="CAB04091FF87FFA3FF74FC4550CDEF34" box="[199,477,1015,1040]" pageId="20" pageNumber="21" type="distribution">
<paragraph id="8215131AFF87FFA3FF74FC4550CDEF34" blockId="20.[151,1437,151,1040]" box="[199,477,1015,1040]" pageId="20" pageNumber="21">
<emphasis id="B0DECF08FF87FFA3FF74FC455070EF34" bold="true" box="[199,352,1015,1040]" pageId="20" pageNumber="21">Distribution.</emphasis>
<collectingCountry id="FABD538AFF87FFA3FED4FC4550C9EF34" box="[359,473,1015,1040]" name="Sri Lanka" pageId="20" pageNumber="21">
<collectingRegion id="406EDDF8FF87FFA3FED4FC455098EF34" box="[359,392,1015,1040]" country="United Kingdom" name="Surrey" pageId="20" pageNumber="21">Sri</collectingRegion>
Lanka
</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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