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<document id="B01504AC652273082756CA888BD20F2B" ID-CLB-Dataset="7385" ID-DOI="10.11646/zootaxa.4951.3.5" ID-GBIF-Dataset="caee27e5-0786-4e95-84fb-ee5394fa79e3" ID-ISSN="1175-5326" ID-Zenodo-Dep="4668365" ID-ZooBank="9BEB98F0-DEA7-4131-8E37-DFC4D5DFCB7B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617780432061" checkinUser="plazi" docAuthor="Bogorodsky, Sergey V., Zajonz, Uwe, Saeed, Fouad N. &amp; Weigmann, Simon" docDate="2021" docId="03B85136664FFFA9FF6C1078E272EC3A" docLanguage="en" docName="zootaxa.4951.3.5.pdf" docOrigin="Zootaxa 4951 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pateobatis jenkinsii" docType="treatment" docVersion="6" lastPageNumber="522" masterDocId="FF81294E6645FFA5FFFB1531E100ED7B" masterDocTitle="Notes on batoid fishes of the Socotra Archipelago (north-western Indian Ocean) with four new records" masterLastPageNumber="528" masterPageNumber="511" pageNumber="521" updateTime="1698936112385" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="1EB7317825CE766532E08D52AF906D43">Notes on batoid fishes of the Socotra Archipelago (north-western Indian Ocean) with four new records</mods:title>
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<mods:namePart id="2403D9D2BDFE82840ED223CF846C4D25">Bogorodsky, Sergey V.</mods:namePart>
<mods:affiliation id="8C225788A7AD3B0D70369C953C8AAB95">Senckenberg oesearch fns ṫẚṫuṫe and Museum of ka ṫure (SMc) I Sec ṫẚon fch ṫhyology &amp; Senckenberg B ẚodẚversẚṫy and Cl ẚmaṫe oesearch Cen ṫre (SB ẚh-c) I B ẚogeography-Senckenberganlage 2 RI SMP 2 R crankfur ṫ am Ma ẚnI GermanyK &amp; S ṫaṫẚon of ka ṫuralẚsṫsI OmskI ouss ẚaK ẚc 18 T 19 S @ yandexKru; h ṫṫps: LLorc ẚdKorgL 0000 - 0002 - 8679 - 9735</mods:affiliation>
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<mods:namePart id="BE82C0CE6CC9699399BB9F130EA638F3">Zajonz, Uwe</mods:namePart>
<mods:affiliation id="DF864494BDE82984D6D89C7CCFCFDFC2">Senckenberg oesearch fns ṫẚṫuṫe and Museum of ka ṫure (SMc) I Sec ṫẚon fch ṫhyology &amp; Senckenberg B ẚodẚversẚṫy and Cl ẚmaṫe oesearch Cen ṫre (SB ẚh-c) I B ẚogeography-Senckenberganlage 2 RI SMP 2 R crankfur ṫ am Ma ẚnI GermanyK</mods:affiliation>
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<mods:namePart id="D3E53E733344AD4B9B4D6C89C0E23F4D">Saeed, Fouad N.</mods:namePart>
<mods:affiliation id="273D521BAE52C8DA8E876D3501DE4A72">Senckenberg oesearch fns ṫẚṫuṫe and Museum of ka ṫure (SMc) I Sec ṫẚon fch ṫhyology &amp; Senckenberg B ẚodẚversẚṫy and Cl ẚmaṫe oesearch Cen ṫre (SB ẚh-c) I B ẚogeography-Senckenberganlage 2 RI SMP 2 R crankfur ṫ am Ma ẚnI GermanyK</mods:affiliation>
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<mods:namePart id="D4FC8461D023A60740AC359A429F1F56">Weigmann, Simon</mods:namePart>
<mods:affiliation id="31F0A8D282656A7C8707D9BF22678BB7">blasmo-iabI blasmobranch oesearch iabora ṫoryI eamburgI GermanyK s ẚmonKweẚgmann @ elasmo-labKde;</mods:affiliation>
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<mods:title id="80B9AC9B227A334D422CE46F7D3473C1">Zootaxa</mods:title>
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<mods:date id="067A69E28C7C9F1E3D66DB06E78D0C1F">2021</mods:date>
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<mods:number id="368B51C0B24F0936B3D5B00292669FD7">4951</mods:number>
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<mods:identifier id="BD5B3EDD324C3EEEC50AC5C7E48D94A7" type="DOI">10.11646/zootaxa.4951.3.5</mods:identifier>
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<treatment id="03B85136664FFFA9FF6C1078E272EC3A" ID-DOI="http://doi.org/10.5281/zenodo.4668405" ID-GBIF-Taxon="181003425" ID-Zenodo-Dep="4668405" LSID="urn:lsid:plazi:treatment:03B85136664FFFA9FF6C1078E272EC3A" httpUri="http://treatment.plazi.org/id/03B85136664FFFA9FF6C1078E272EC3A" lastPageId="12" lastPageNumber="522" pageId="10" pageNumber="521">
<subSubSection id="C30BB3AB664FFFAFFF6C1078E055E8F3" pageId="10" pageNumber="521" type="nomenclature">
<paragraph id="8BAEE020664FFFAFFF6C1078E371E818" blockId="10.[151,625,1353,1452]" box="[151,625,1353,1380]" pageId="10" pageNumber="521">
<heading id="D0E6574C664FFFAFFF6C1078E371E818" bold="true" box="[151,625,1353,1380]" fontSize="11" level="1" pageId="10" pageNumber="521" reason="1">
<taxonomicName id="4C119BA3664FFFAFFF6C1078E371E818" ID-CoL="765YF" authority="(Annandale, 1909)" baseAuthorityName="Annandale" baseAuthorityYear="1909" box="[151,625,1353,1380]" class="Elasmobranchii" family="Dasyatidae" genus="Pateobatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="10" pageNumber="521" phylum="Chordata" rank="species" species="jenkinsii">
<emphasis id="B9653C32664FFFAFFF6C1078E371E818" bold="true" box="[151,625,1353,1380]" pageId="10" pageNumber="521">
<emphasis id="B9653C32664FFFAFFF6C1078E084E81F" bold="true" box="[151,388,1353,1380]" italics="true" pageId="10" pageNumber="521">Pateobatis jenkinsii</emphasis>
(
<bibRefCitation id="EF809DD1664FFFAFFE6F1078E369E818" author="Annandale, N." box="[404,617,1353,1379]" pageId="10" pageNumber="521" pagination="1 - 58" refId="ref8226" refString="Annandale, N. (1909) Report on the fishes taken by the Bengal fisheries steamer &quot; Golden Crown. &quot; Part I, Batoidei. Memo ẚrs of the fnd ẚan Museum, 2 (1), 1 - 58. https: // doi. org / 10.5962 / bhl. part. 29058" type="journal article" year="1909">Annandale, 1909</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BAEE020664FFFAFFF6C105FE055E8F3" blockId="10.[151,625,1353,1452]" box="[151,341,1390,1416]" pageId="10" pageNumber="521">Jenkins whipray</paragraph>
</subSubSection>
<subSubSection id="C30BB3AB664FFFAFFF6C10A3E563E895" pageId="10" pageNumber="521" type="description">
<paragraph id="8BAEE020664FFFAFFF6C10A3E1D9E8D7" blockId="10.[151,625,1353,1452]" box="[151,217,1426,1452]" pageId="10" pageNumber="521">
<figureCitation id="132AFCA5664FFFAFFF6C10A3E1D9E8D7" box="[151,217,1426,1452]" captionStart="FIGURE 9" captionStartId="11.[151,250,1567,1591]" captionTargetBox="[151,1435,273,1543]" captionTargetId="figure-65@11.[151,1435,273,1543]" captionTargetPageId="11" captionText="FIGURE 9. maṫeobaṫẚs jenkẚnsẚẚ, underwater photographs. A: Ras Bidou, Socotra Island, 14 m depth; B: Roosh, Socotra Island, 3 m depth. Photos by S.V. Bogorodsky." figureDoi="http://doi.org/10.5281/zenodo.4668395" httpUri="https://zenodo.org/record/4668395/files/figure.png" pageId="10" pageNumber="521">Fig. 9</figureCitation>
</paragraph>
<paragraph id="8BAEE020664FFFAFFF6C10E6E563E895" blockId="10.[151,1123,1494,1519]" box="[151,1123,1494,1519]" pageId="10" pageNumber="521">
<emphasis id="B9653C32664FFFAFFF6C10E6E03BE895" box="[151,315,1494,1518]" italics="true" pageId="10" pageNumber="521">Trygon jenkẚnsẚẚ</emphasis>
<materialsCitation id="3B79EA7D664FFFAFFEBB10E7E55EE895" ID-GBIF-Occurrence="3071284302" box="[320,1118,1494,1519]" country="India" location="Orissa State" pageId="10" pageNumber="521" specimenCount="1" stateProvince="Odisha" typeStatus="holotype">
<bibRefCitation id="EF809DD1664FFFAFFEBB10E7E317E895" author="Annandale, N." box="[320,535,1494,1518]" pageId="10" pageNumber="521" pagination="1 - 58" refId="ref8226" refString="Annandale, N. (1909) Report on the fishes taken by the Bengal fisheries steamer &quot; Golden Crown. &quot; Part I, Batoidei. Memo ẚrs of the fnd ẚan Museum, 2 (1), 1 - 58. https: // doi. org / 10.5962 / bhl. part. 29058" type="journal article" year="1909">Annandale, 1909: 28</bibRefCitation>
;
<typeStatus id="54AA5E82664FFFAFFDDA10E7E37EE895" box="[545,638,1494,1518]" pageId="10" pageNumber="521" type="holotype">holotype</typeStatus>
: ZSI F2473/1; type locality:
<collectingRegion id="49D52EC2664FFFAFFC5A10E7E51DE895" box="[929,1053,1494,1518]" country="India" name="Odisha" pageId="10" pageNumber="521">Orissa State</collectingRegion>
,
<collectingCountry id="F306A0B0664FFFAFFBDC10E7E55EE895" box="[1063,1118,1494,1518]" name="India" pageId="10" pageNumber="521">India</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30BB3AB664FFFAFFF6C132DE0FFEA00" pageId="10" pageNumber="521" type="materials_examined">
<paragraph id="8BAEE020664FFFAFFF6C132DE558EB20" blockId="10.[151,1437,1564,2023]" pageId="10" pageNumber="521">
<emphasis id="B9653C32664FFFAFFF6C132DE084EB4D" bold="true" box="[151,388,1564,1590]" pageId="10" pageNumber="521">Material examined:</emphasis>
<materialsCitation id="3B79EA7D664FFFAFFE77132CE014EB20" ID-GBIF-Occurrence="3071284311" collectingDate="2019-04-17" country="Socotra Island" elevation="14" location="No" municipality="Ras Bidou" pageId="10" pageNumber="521" specimenCount="1">
<location id="8ECEB6FB664FFFAFFE77132CE0AEEB4C" LSID="urn:lsid:plazi:treatment:03B85136664FFFA9FF6C1078E272EC3A:8ECEB6FB664FFFAFFE77132CE0AEEB4C" box="[396,430,1565,1591]" country="Socotra Island" municipality="Ras Bidou" name="No" pageId="10" pageNumber="521">No</location>
specimen collected, underwater photographs,
<collectingCountry id="F306A0B0664FFFAFFC40132CE55FEB4D" box="[955,1119,1564,1591]" name="Socotra Island" pageId="10" pageNumber="521">Socotra Island</collectingCountry>
,
<collectingMunicipality id="6BCA7A5A664FFFAFFB97132CE5E5EB4D" box="[1132,1253,1564,1591]" pageId="10" pageNumber="521">Ras Bidou</collectingMunicipality>
,
<elevation id="003C0713664FFFAFFB0A132CE474EB4D" box="[1265,1396,1564,1591]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="10" pageNumber="521" unit="m" value="14.0">
<quantity id="4CE94DC5664FFFAFFB0A132CE42BEB4C" box="[1265,1323,1565,1591]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="10" pageNumber="521" unit="m" value="14.0">14 m</quantity>
depth
</elevation>
,
<date id="FFAFC6E0664FFFAFFA7B132CE014EB20" pageId="10" pageNumber="521" value="2019-04-17">
<collectingDate id="EFEB3F08664FFFAFFA7B132CE014EB20" pageId="10" pageNumber="521" value="2019-04-17">17 April 2019</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B79EA7D664FFFAFFEDB1371E555EB20" ID-GBIF-Occurrence="3071284303" box="[288,1109,1600,1627]" collectingDate="2019-04-13" country="Socotra Island" elevation="3" location="Roosh" pageId="10" pageNumber="521" specimenCount="1">
underwater photograph,
<collectingCountry id="F306A0B0664FFFAFFDC81370E3D6EB21" box="[563,726,1600,1627]" name="Socotra Island" pageId="10" pageNumber="521">Socotra Island</collectingCountry>
,
<location id="8ECEB6FB664FFFAFFD191371E22CEB21" LSID="urn:lsid:plazi:treatment:03B85136664FFFA9FF6C1078E272EC3A:8ECEB6FB664FFFAFFD191371E22CEB21" box="[738,812,1600,1626]" country="Socotra Island" name="Roosh" pageId="10" pageNumber="521">Roosh</location>
,
<elevation id="003C0713664FFFAFFCCC1370E2AAEB21" box="[823,938,1600,1627]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="521" unit="m" value="3.0">
<quantity id="4CE94DC5664FFFAFFCCC1370E262EB20" box="[823,866,1601,1627]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="521" unit="m" value="3.0">3 m</quantity>
depth
</elevation>
,
<date id="FFAFC6E0664FFFAFFC4E1370E555EB20" box="[949,1109,1600,1627]" pageId="10" pageNumber="521" value="2019-04-13">
<collectingDate id="EFEB3F08664FFFAFFC4E1370E555EB20" box="[949,1109,1600,1627]" pageId="10" pageNumber="521" value="2019-04-13">13 April 2019</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BAEE020664FFFAFFF3C1355E41EEA74" blockId="10.[151,1437,1564,2023]" pageId="10" pageNumber="521">
<emphasis id="B9653C32664FFFAFFF3C1355E0D7EB04" bold="true" box="[199,471,1636,1663]" pageId="10" pageNumber="521">Distinctive characters:</emphasis>
<emphasis id="B9653C32664FFFAFFE1A1357E3C1EBFA" box="[481,705,1638,1665]" italics="true" pageId="10" pageNumber="521">maṫeobaṫẚs jenkẚnsẚẚ</emphasis>
is characterised by having a rhomboidal disc, its width slightly greater than disc length; tip of snout obtuse, slightly projecting; a narrow, asymmetrical well-defined band of denticles on back; a row of enlarged thorns along midline of disc and tail to the base of spine, a character unique within large whiptail stingrays (denticles band and thorns reduced or absent in young); tail relatively long and slender, without cutaneous fold, tail length 1.11.4 times disc width (Last
<emphasis id="B9653C32664FFFAFFC8313C5E2A9EA75" box="[888,937,1780,1807]" italics="true" pageId="10" pageNumber="521">eṫ al</emphasis>
. 2016c; Psomadakis
<emphasis id="B9653C32664FFFAFFB6C13C5E5C8EA75" box="[1175,1224,1780,1807]" italics="true" pageId="10" pageNumber="521">eṫ al</emphasis>
. 2019).
</paragraph>
<paragraph id="8BAEE020664FFFAFFF3C122BE0FFEA00" blockId="10.[151,1437,1564,2023]" pageId="10" pageNumber="521">
<emphasis id="B9653C32664FFFAFFF3C122BE050EA4E" box="[199,336,1818,1845]" italics="true" pageId="10" pageNumber="521">Colouraṫẚon</emphasis>
: Dorsal surface of disc plain uniform greyish, yellowish brown or greenish brown, immature specimens with small dark spots on outer disc margins; tail blackish beyond sting. Ventral surface white (Last
<emphasis id="B9653C32664FFFAFFA9A120DE495EA2D" box="[1377,1429,1852,1879]" italics="true" pageId="10" pageNumber="521">eṫ al</emphasis>
. 2016c; Psomadakis
<emphasis id="B9653C32664FFFAFFE8C1251E0A8EA01" box="[375,424,1888,1915]" italics="true" pageId="10" pageNumber="521">eṫ al</emphasis>
. 2019).
</paragraph>
</subSubSection>
<subSubSection id="C30BB3AB664FFFAEFF3C12B5E093ED82" lastPageId="11" lastPageNumber="522" pageId="10" pageNumber="521" type="distribution">
<paragraph id="8BAEE020664FFFAEFF3C12B5E093ED82" blockId="10.[151,1437,1564,2023]" lastBlockId="11.[151,1436,151,249]" lastPageId="11" lastPageNumber="522" pageId="10" pageNumber="521">
<emphasis id="B9653C32664FFFAFFF3C12B5E063EAE5" bold="true" box="[199,355,1924,1950]" pageId="10" pageNumber="521">Distribution:</emphasis>
Ranging from the inner Gulf of
<collectingRegion id="49D52EC2664FFFAFFD3E12B5E202EAE5" box="[709,770,1924,1950]" country="Yemen" name="'Adan" pageId="10" pageNumber="521">Aden</collectingRegion>
and
<collectingCountry id="F306A0B0664FFFAFFCCE12B4E28BEAE4" box="[821,907,1925,1951]" name="Yemen" pageId="10" pageNumber="521">Socotra</collectingCountry>
Archipelago, possibly to the Gulf, southward to
<collectingCountry id="F306A0B0664FFFAFFF6C1299E01EEAB9" box="[151,286,1960,1986]" name="Madagascar" pageId="10" pageNumber="521">Madagascar</collectingCountry>
and
<collectingCountry id="F306A0B0664FFFAFFEA81299E0E3EAB9" box="[339,483,1960,1986]" name="South Africa" pageId="10" pageNumber="521">South Africa</collectingCountry>
, eastward to the
<collectingCountry id="F306A0B0664FFFAFFD601299E219EAB9" box="[667,793,1960,1986]" name="Philippines" pageId="10" pageNumber="521">Philippines</collectingCountry>
and
<collectingCountry id="F306A0B0664FFFAFFCB61298E527EAB9" box="[845,1063,1960,1987]" name="Papua New Guinea" pageId="10" pageNumber="521">Papua New Guinea</collectingCountry>
. In the Arabian region it is known from the
<collectingRegion id="49D52EC2664FFFAFFF0612FDE030EA9D" box="[253,304,1996,2022]" country="Papua New Guinea" name="Gulf" pageId="10" pageNumber="521">Gulf</collectingRegion>
of Tadjoura (Lips
<emphasis id="B9653C32664FFFAFFDFB12FDE330EA9D" box="[512,560,1996,2023]" italics="true" pageId="10" pageNumber="521">eṫ al</emphasis>
. 2016) and
<collectingCountry id="F306A0B0664FFFAFFD4E12FCE3F9EA9C" box="[693,761,1997,2023]" name="Oman" pageId="10" pageNumber="521">Oman</collectingCountry>
(Randall 1995). Eagderi
<emphasis id="B9653C32664FFFAFFBE812FDE544EA9D" box="[1043,1092,1996,2023]" italics="true" pageId="10" pageNumber="521">eṫ al</emphasis>
. (2019) listed the species from the Gulf, but the record should be confirmed in comparison with the similar
<emphasis id="B9653C32664EFFAEFC0815A6E52CEDC8" box="[1011,1068,151,179]" italics="true" pageId="11" pageNumber="522">mK faẚ</emphasis>
(
<bibRefCitation id="EF809DD1664EFFAEFBC015A6E433EDC9" author="Jordan, D. S. &amp; Seale, A." box="[1083,1331,151,178]" pageId="11" pageNumber="522" pagination="173 - 455" refId="ref10583" refString="Jordan, D. S. &amp; Seale, A. (1906) The fishes of Samoa. Description of the species found in the archipelago, with a provisional check-list of the fishes of Oceania. Bulle tẚn of the Bureau of c ẚsherẚes, 25, 173 - 455. [for 1905] https: // doi. org / 10.5962 / bhl. title. 46247" type="journal article" year="1906">
<collectingCountry id="F306A0B0664EFFAEFBC015A6E588EDCA" box="[1083,1160,151,177]" name="Jordan" pageId="11" pageNumber="522">Jordan</collectingCountry>
&amp; Seale, 1906
</bibRefCitation>
). Almojil
<emphasis id="B9653C32664EFFAEFF6C158BE1C9EDAE" box="[151,201,186,213]" italics="true" pageId="11" pageNumber="522">eṫ al</emphasis>
. (2015) included
<emphasis id="B9653C32664EFFAEFE69158AE0CCEDAC" box="[402,460,187,215]" italics="true" pageId="11" pageNumber="522">mK faẚ</emphasis>
in their book with occurrence in the Gulf confirmed by a specimen collected from the
<collectingCountry id="F306A0B0664EFFAEFF6C15EEE08FED82" box="[151,399,223,249]" name="United Arab Emirates" pageId="11" pageNumber="522">United Arab Emirates</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="DF6EB0A8664EFFAEFF6C132EE326EB20" ID-DOI="http://doi.org/10.5281/zenodo.4668395" ID-Zenodo-Dep="4668395" httpUri="https://zenodo.org/record/4668395/files/figure.png" pageId="11" pageNumber="522" startId="11.[151,250,1567,1591]" targetBox="[151,1435,273,1543]" targetPageId="11">
<paragraph id="8BAEE020664EFFAEFF6C132EE326EB20" blockId="11.[151,1436,1566,1627]" pageId="11" pageNumber="522">
<emphasis id="B9653C32664EFFAEFF6C132EE012EB4C" bold="true" box="[151,274,1567,1591]" pageId="11" pageNumber="522">FIGURE 9.</emphasis>
<emphasis id="B9653C32664EFFAEFEE3132FE0DFEB4D" box="[280,479,1566,1590]" italics="true" pageId="11" pageNumber="522">maṫeobaṫẚs jenkẚnsẚẚ</emphasis>
, underwater photographs.
<emphasis id="B9653C32664EFFAEFD10132EE206EB4C" bold="true" box="[747,774,1567,1591]" pageId="11" pageNumber="522">A:</emphasis>
Ras Bidou, Socotra Island, 14 m depth;
<emphasis id="B9653C32664EFFAEFB61132EE5B4EB4C" bold="true" box="[1178,1204,1567,1591]" pageId="11" pageNumber="522">B:</emphasis>
Roosh, Socotra Island, 3 m depth. Photos by S.V. Bogorodsky.
</paragraph>
</caption>
<subSubSection id="C30BB3AB664EFFA9FF3C13B8E272EC3A" lastPageId="12" lastPageNumber="523" pageId="11" pageNumber="522" type="discussion">
<paragraph id="8BAEE020664EFFA9FF3C13B8E272EC3A" blockId="11.[151,1437,1673,2026]" lastBlockId="12.[151,1436,151,321]" lastPageId="12" lastPageNumber="523" pageId="11" pageNumber="522">
<emphasis id="B9653C32664EFFAEFF3C13B8E03DEBD8" bold="true" box="[199,317,1673,1699]" pageId="11" pageNumber="522">Remarks:</emphasis>
The photographed individuals match well all characters of
<emphasis id="B9653C32664EFFAEFC1413BAE5CFEBDD" box="[1007,1231,1675,1702]" italics="true" pageId="11" pageNumber="522">maṫeobaṫẚs jenkẚnsẚẚ</emphasis>
provided by Last
<emphasis id="B9653C32664EFFAEFF6C139CE1CAEBBC" box="[151,202,1709,1736]" italics="true" pageId="11" pageNumber="522">eṫ al</emphasis>
. (2016c), particularly showing the characteristic row of prominent thorns along mid-line of disc and tail and the small dark spots on rear margins of disc typical for immature specimens. The species inhabits sandy bottoms of shallow lagoons, bays and seaward reefs, often close to coral reefs, at depths of
<quantity id="4CE94DC5664EFFAEFBB413C4E5A6EA6B" box="[1103,1190,1781,1808]" metricMagnitude="1" metricUnit="m" metricValue="4.65" metricValueMax="9.0" metricValueMin="0.3" pageId="11" pageNumber="522" unit="m" value="46.5" valueMax="90.0" valueMin="3.0">390 m</quantity>
following
<bibRefCitation id="EF809DD1664EFFAEFADF13C4E1E2EA4F" author="Weigmann, S." pageId="11" pageNumber="522" pagination="837 - 1037" refId="ref13064" refString="Weigmann, S. (2016) Annotated checklist of the living sharks, batoids and chimaeras (Chondrichthyes) of the world, with a focus on biogeographical diversity. gournal of c ẚsh B ẚology, 88 (3), 837 - 1037. https: // doi. org / 10.1111 / jfb. 12874" type="journal article" year="2016">Weigmann (2016)</bibRefCitation>
whereby the minimum depth is based on a newly photographed individual at Roosh. The present confirmed record from
<collectingCountry id="F306A0B0664EFFAEFEDD120CE07CEA2C" box="[294,380,1853,1879]" name="Yemen" pageId="11" pageNumber="522">Socotra</collectingCountry>
is based on individuals observed on silty sand bottom close to large rocks with overhangs in Ras Bidou, western part of
<collectingCountry id="F306A0B0664EFFAEFE3D1250E369EA00" box="[454,617,1889,1915]" name="Socotra Island" pageId="11" pageNumber="522">Socotra Island</collectingCountry>
, at a depth of
<quantity id="4CE94DC5664EFFAEFCF81250E23DEA07" box="[771,829,1889,1916]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="11" pageNumber="522" unit="m" value="14.0">14 m</quantity>
, and an individual found under a ledge of a large stone at Roosh, north-eastern part of the island, at a depth of
<specimenCount id="9D172BA9664EFFAEFCE312B4E24CEADB" box="[792,844,1925,1952]" pageId="11" pageNumber="522" type="generic">
<quantity id="4CE94DC5664EFFAEFCE312B4E248EADB" box="[792,840,1925,1952]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="522" unit="m" value="3.0">3 m</quantity>
.
</specimenCount>
Subadults are characterised by having small brown spots on edges of posterior half of disc. This growth stage was described as a distinct species,
<emphasis id="B9653C32664EFFAEFB4C129BE41AEAB8" box="[1207,1306,1961,1987]" italics="true" pageId="11" pageNumber="522">eK draco</emphasis>
<bibRefCitation id="EF809DD1664EFFAEFADA1298E06BEA9C" author="Compagno, L. J. V. &amp; Heemstra, P. C." pageId="11" pageNumber="522" pagination="1 - 17" refId="ref8895" refString="Compagno, L. J. V. &amp; Heemstra, P. C. (1984) e ẚmantura draco, a new species of stingray (Myliobatiformes: Dasyatidae) from South Africa, with a key to the Dasyatidae and the first record of aasya tẚs kuhl ẚẚ (Muller &amp; Henle, 1841) from southern Africa. The gKiKB Sm ẚth fns tẚtute of fch thyology Spec ẚal mubl ẚcatẚon, 33, 1 - 17." type="journal article" year="1984">Compagno &amp; Heemstra, 1984</bibRefCitation>
, but subsequently placed in synonymy with
<emphasis id="B9653C32664EFFAEFC9E12FFE2E8EA91" box="[869,1000,1998,2026]" italics="true" pageId="11" pageNumber="522">eK jenkẚnsẚẚ</emphasis>
(
<bibRefCitation id="EF809DD1664EFFAEFC0312FCE410EA9C" author="Last, P. R. &amp; Compagno, L. J. V." box="[1016,1296,1997,2023]" pageId="11" pageNumber="522" pagination="1398 - 2068" refId="ref11281" refString="Last, P. R. &amp; Compagno, L. J. V. (1999) Dasyatidae. fn: Carpenter, K. E. &amp; Niem, V. H. (Eds.), cAO Spec ẚes fden tẚfẚcatẚon Gu ẚde for c ẚshery murposesK The i ẚvẚng Mar ẚne oesources of the tes tern Cen tral mac ẚfẚcK solK PK Ba toẚd f ẚshesI ch ẚmeras and bony f ẚshes par t 1 (blop ẚdae to i ẚnophrynẚdae). FAO, Rome, pp. 1398 - 2068." type="book chapter" year="1999">Last &amp; Compagno 1999</bibRefCitation>
).
<emphasis id="B9653C32664EFFA9FADF12FEE1F6EDC8" italics="true" lastPageId="12" lastPageNumber="523" pageId="11" pageNumber="522">maṫeobaṫẚs jenkẚnsẚẚ</emphasis>
is included as vulnerable in IUCN Red List Assessments and urgently requires conservation management (Manjaji Matsumoto
<emphasis id="B9653C326649FFA9FE7D158BE0B7EDAE" box="[390,439,186,213]" italics="true" pageId="12" pageNumber="523">eṫ al</emphasis>
. 2016b).
<emphasis id="B9653C326649FFA9FDD9158DE3BFEDAC" box="[546,703,188,215]" italics="true" pageId="12" pageNumber="523">maṫeobaṫẚs faẚ</emphasis>
, reported from the Red Sea to the Marquesas Islands, is a similar species, which possibly also occurs off
<collectingCountry id="F306A0B06649FFA9FDA715EEE3B2ED82" box="[604,690,223,249]" name="Yemen" pageId="12" pageNumber="523">Socotra</collectingCountry>
Archipelago. It might be confused with
<emphasis id="B9653C326649FFA9FB7815EEE5FFED80" box="[1155,1279,223,251]" italics="true" pageId="12" pageNumber="523">mK jenkẚnsẚẚ</emphasis>
but differs in having only few slightly enlarged thorny denticles mid-centrally on disc and a mid-row of enlarged rounded tubercles at base of tail (Last
<emphasis id="B9653C326649FFA9FE301417E0FCEC3A" box="[459,508,294,321]" italics="true" pageId="12" pageNumber="523">eṫ al</emphasis>
. 2016c; Psomadakis
<emphasis id="B9653C326649FFA9FD111417E21BEC3A" box="[746,795,294,321]" italics="true" pageId="12" pageNumber="523">eṫ al</emphasis>
. 2019).
</paragraph>
</subSubSection>
</treatment>
</document>