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<document id="F058AB78D14AC2C076BF98B4719C8EC8" ID-DOI="10.1649/1144.1" ID-ISSN="1938-4394" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630077134113" checkinUser="valdenar" docAuthor="Bitar, Alberto &amp; Delgado, Leonardo" docDate="2009" docId="03EF87D1393FFFDF4175AAD3FC63FB1D" docLanguage="en" docName="1144.1.pdf.imf" docOrigin="The Coleopterists Bulletin 63 (2)" docSource="http://dx.doi.org/10.1649/1144.1" docStyle="DocumentStyle:07711681861DB98305E87147CAB6DAD5.4:TheColeopBull.2003-2009.journal_article.0cover" docStyleId="07711681861DB98305E87147CAB6DAD5" docStyleName="TheColeopBull.2003-2009.journal_article.0cover" docStyleVersion="4" docTitle="Xyloryctes guatemalensis Bitar and Delgado, new species" docType="treatment" docVersion="3" lastPageNumber="215" masterDocId="FFD6FFA9393FFFDD4058AE51FFE6FFC5" masterDocTitle="A New Species of Xyloryctes Hope (Scarabaeidae: Dynastinae) from Guatemala, with a Key to the Species" masterLastPageNumber="217" masterPageNumber="213" pageNumber="213" updateTime="1699141271781" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="5EEFD3D98A6F2BD54F5CBAD5EC2FCAA5">A New Species of Xyloryctes Hope (Scarabaeidae: Dynastinae) from Guatemala, with a Key to the Species</mods:title>
</mods:titleInfo>
<mods:name id="E0412A63A06556018C0690127C71E06C" type="personal">
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<mods:namePart id="388893CD33DD96ECBB11DB854F8214E1">Bitar, Alberto</mods:namePart>
</mods:name>
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<mods:namePart id="E6880CFEF680E9F3276176378B626489">Delgado, Leonardo</mods:namePart>
</mods:name>
<mods:typeOfResource id="FAFBD6F0A24FA7D7524E76F4FBDA5449">text</mods:typeOfResource>
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<mods:title id="618D1DC8DCDAD9B45ACF4B9DFB6DE679">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="0F23B0F3EFB90A692E66288BF3E84CBF">
<mods:date id="32F4F96155FA88EEBB4449B3F354B5D3">2009</mods:date>
<mods:detail id="4DDF50A37A0B9ED910EE5578BD4A495F" type="pubDate">
<mods:number id="55EDEBD95230287F0F1B5A238960B9DD">2009-06-30</mods:number>
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<mods:number id="D29FB3C877AC59558703440BEEDCA855">63</mods:number>
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<mods:identifier id="90FFF3AF843578C1E69D22D3CAE7EAF9" type="DOI">10.1649/1144.1</mods:identifier>
<mods:identifier id="3DDC99B1DF69DB48C9B6000B9E99DB29" type="ISSN">1938-4394</mods:identifier>
</mods:mods>
<treatment id="03EF87D1393FFFDF4175AAD3FC63FB1D" ID-DOI="http://doi.org/10.5281/zenodo.5301779" ID-Zenodo-Dep="5301779" LSID="urn:lsid:plazi:treatment:03EF87D1393FFFDF4175AAD3FC63FB1D" httpUri="http://treatment.plazi.org/id/03EF87D1393FFFDF4175AAD3FC63FB1D" lastPageId="2" lastPageNumber="215" pageId="0" pageNumber="213">
<subSubSection id="C35C654C393FFFDD4175AAD3FD74FB76" pageId="0" pageNumber="213" type="nomenclature">
<paragraph id="8BF936C7393FFFDD4175AAD3FC88FB5D" blockId="0.[301,878,1154,1203]" box="[301,878,1154,1176]" pageId="0" pageNumber="213">
<heading id="D0B181AB393FFFDD4175AAD3FC88FB5D" box="[301,878,1154,1176]" centered="true" fontSize="9" level="2" pageId="0" pageNumber="213" reason="2">
<emphasis id="B932EAD5393FFFDD4175AAD3FC88FB5D" bold="true" box="[301,878,1154,1176]" pageId="0" pageNumber="213">
<taxonomicName id="4C464D44393FFFDD4175AAD3FD09FB5D" authority="Bitar and Delgado" authorityName="Bitar and Delgado" box="[301,751,1154,1176]" class="Insecta" family="Dynastidae" genus="Xyloryctes" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="213" phylum="Arthropoda" rank="species" species="guatemalensis" status="sp. nov.">
<emphasis id="B932EAD5393FFFDD4175AAD3FDCCFB5D" bold="true" box="[301,554,1154,1176]" italics="true" pageId="0" pageNumber="213">Xyloryctes guatemalensis</emphasis>
Bitar and Delgado
</taxonomicName>
,
<taxonomicNameLabel id="A20157AE393FFFDD42A4AAD3FC88FB5D" box="[764,878,1154,1176]" pageId="0" pageNumber="213" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF936C7393FFFDD4252AACCFD74FB76" blockId="0.[301,878,1154,1203]" box="[522,658,1181,1203]" pageId="0" pageNumber="213">(Figs. 1, 34)</paragraph>
</subSubSection>
<subSubSection id="C35C654C393FFFDD409FAA98FDB8F994" pageId="0" pageNumber="213" type="materials_examined">
<paragraph id="8BF936C7393FFFDD409FAA98FDB7FA5E" blockId="0.[175,1005,1225,1570]" pageId="0" pageNumber="213">
<emphasis id="B932EAD5393FFFDD409FAA98FEB8FB1A" bold="true" box="[199,350,1225,1247]" pageId="0" pageNumber="213">Type Material.</emphasis>
<materialsCitation id="3B2E3C9A393FFFDD413DAA98FDC6FB3F" collectorName="F. Jimenez" country="Guatemala" location="San Jose Pinula" pageId="0" pageNumber="213" specimenCount="1" typeStatus="holotype">
<typeStatus id="54FD8865393FFFDD413DAA98FE23FB1A" box="[357,453,1225,1247]" pageId="0" pageNumber="213" type="holotype">Holotype</typeStatus>
male labeled:
<collectingCountry id="F3517657393FFFDD4238AA98FCEAFB1A" box="[608,780,1225,1247]" name="Guatemala" pageId="0" pageNumber="213">GUATEMALA</collectingCountry>
, Guatemala,
<location id="8E99601C393FFFDD43CCAA98FF15FB3F" LSID="urn:lsid:plazi:treatment:03EF87D1393FFFDF4175AAD3FC63FB1D:8E99601C393FFFDD43CCAA98FF15FB3F" country="Guatemala" name="San Jose Pinula" pageId="0" pageNumber="213">San José Pinula</location>
, 27-IX-93.
<collectorName id="26B35311393FFFDD4134AAB5FE39FB3F" box="[364,479,1252,1274]" pageId="0" pageNumber="213">F. Jiménez</collectorName>
, col.
</materialsCitation>
.
<materialsCitation id="3B2E3C9A393FFFDD4275AAB5FE50FAEA" collectingDate="1989-05-13" collectorName="E. Cano" country="Guatemala" county="Alameda" location="Ciudad Capital" municipality="Zona" pageId="0" pageNumber="213" specimenCount="30" typeStatus="allotype">
<typeStatus id="54FD8865393FFFDD4275AAB5FD63FB3F" box="[557,645,1252,1274]" pageId="0" pageNumber="213" type="allotype">Allotype</typeStatus>
female labeled:
<collectingCountry id="F3517657393FFFDD4364AAB5FC0EFB3F" box="[828,1000,1252,1274]" name="Guatemala" pageId="0" pageNumber="213">GUATEMALA</collectingCountry>
, Guatemala,
<location id="8E99601C393FFFDD4175AAAEFE2BFAD0" LSID="urn:lsid:plazi:treatment:03EF87D1393FFFDF4175AAD3FC63FB1D:8E99601C393FFFDD4175AAAEFE2BFAD0" box="[301,461,1279,1301]" country="Guatemala" county="Alameda" municipality="Zona" name="Ciudad Capital" pageId="0" pageNumber="213">Ciudad Capital</location>
,
<collectingMunicipality id="6B9DACBD393FFFDD418FAAAEFDEBFAD0" box="[471,525,1279,1301]" pageId="0" pageNumber="213">Zona</collectingMunicipality>
18,
<collectingCounty id="62984E4B393FFFDD4261AAAEFD73FAD0" box="[569,661,1279,1301]" pageId="0" pageNumber="213">Alameda</collectingCounty>
1,
<date id="FFF81007393FFFDD42ECAAAEFCF9FAD0" box="[692,799,1279,1301]" pageId="0" pageNumber="213" value="1989-05-13">
<collectingDate id="EFBCE9EF393FFFDD42ECAAAEFCF9FAD0" box="[692,799,1279,1301]" pageId="0" pageNumber="213" value="1989-05-13">13-V-1989</collectingDate>
</date>
, alumbrado público 9:
<specimenCount id="9D40FD4E393FFFDD40E7AB48FF1CFAEA" box="[191,250,1305,1327]" pageId="0" pageNumber="213" type="generic">30 p.</specimenCount>
m.
<collectorName id="26B35311393FFFDD417BAB48FE98FAEA" box="[291,382,1305,1327]" pageId="0" pageNumber="213">E. Cano</collectorName>
col
</materialsCitation>
.
<materialsCitation id="3B2E3C9A393FFFDD4193AB48FEF6FAA0" collectingDate="1985-09-17" collectorName="M. Zepeda" country="Guatemala" county="Alameda" location="OiOi" municipality="Same" pageId="0" pageNumber="213" specimenCount="1" specimenCount-female="1" stateProvince="Guatemala" typeStatus="paratype">
<typeStatus id="54FD8865393FFFDD4193AB48FDC9FAEA" box="[459,559,1305,1327]" pageId="0" pageNumber="213" type="paratype">Paratypes</typeStatus>
labeled:
<collectingMunicipality id="6B9DACBD393FFFDD42C1AB48FD36FAEA" box="[665,720,1305,1327]" pageId="0" pageNumber="213">Same</collectingMunicipality>
data as allotype (2
<location id="8E99601C393FFFDD43E4AB48FC38FAEA" LSID="urn:lsid:plazi:treatment:03EF87D1393FFFDF4175AAD3FC63FB1D:8E99601C393FFFDD43E4AB48FC38FAEA" box="[956,990,1305,1327]" country="Guatemala" municipality="Same" name="OiOi" pageId="0" pageNumber="213" stateProvince="Guatemala">OiOi</location>
);
<collectingCountry id="F3517657393FFFDD40E1AB65FE80FA8F" box="[185,358,1332,1354]" name="Guatemala" pageId="0" pageNumber="213">GUATEMALA</collectingCountry>
: Guatemala,
<collectingRegion id="4982F825393FFFDD41AFAB65FD03FA8F" box="[503,741,1332,1354]" country="Guatemala" name="Guatemala" pageId="0" pageNumber="213">Ciudad de Guatemala</collectingRegion>
.
<date id="FFF81007393FFFDD42ABAB65FC8FFA8F" box="[755,873,1332,1354]" pageId="0" pageNumber="213" value="1985-09-17">
<collectingDate id="EFBCE9EF393FFFDD42ABAB65FC8FFA8F" box="[755,873,1332,1354]" pageId="0" pageNumber="213" value="1985-09-17">17-IX-1985</collectingDate>
</date>
,
<collectorName id="26B35311393FFFDD4320AB65FC0AFA8F" box="[888,1004,1332,1354]" pageId="0" pageNumber="213">M. Zepeda</collectorName>
col. (
<specimenCount id="9D40FD4E393FFFDD40A9AB1EFEE1FAA0" box="[241,263,1359,1381]" pageId="0" pageNumber="213" type="female">1♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B2E3C9A393FFFDD4144AB1EFCCAFAA0" box="[284,812,1359,1381]" collectingDate="1986-05" collectorName="L. Rodriguez &amp; Oi" country="Guatemala" county="Alameda" location="OiOi" municipality="Same" pageId="0" pageNumber="213" specimenCount="1" stateProvince="Guatemala" typeStatus="paratype">
same data except:
<date id="FFF81007393FFFDD41B8AB1EFDC9FAA0" box="[480,559,1359,1381]" pageId="0" pageNumber="213" value="1986-05">
<collectingDate id="EFBCE9EF393FFFDD41B8AB1EFDC9FAA0" box="[480,559,1359,1381]" pageId="0" pageNumber="213" value="1986-05">V-1986</collectingDate>
</date>
,
<collectorName id="26B35311393FFFDD4261AB1EFD27FAA0" box="[569,705,1359,1381]" pageId="0" pageNumber="213">L. Rodriguez</collectorName>
col. (1
<collectorName id="26B35311393FFFDD434BAB1EFCC2FAA0" box="[787,804,1359,1381]" pageId="0" pageNumber="213">Oi</collectorName>
)
</materialsCitation>
;
<materialsCitation id="3B2E3C9A393FFFDD4360AB1EFDE9FA45" collectingDate="1983-06-15" collectionCode="R" collectorName="Perez" country="Guatemala" county="Alameda" location="OiOi" municipality="Same" pageId="0" pageNumber="213" specimenCount="1" specimenCount-female="1" stateProvince="Guatemala" typeStatus="paratype">
same data except:
<date id="FFF81007393FFFDD40E1AB3BFEDDFA45" box="[185,315,1386,1408]" pageId="0" pageNumber="213" value="1983-06-15">
<collectingDate id="EFBCE9EF393FFFDD40E1AB3BFEDDFA45" box="[185,315,1386,1408]" pageId="0" pageNumber="213" value="1983-06-15">15-Jun-1983</collectingDate>
</date>
,
<collectionCode id="ED57AE02393FFFDD4110AB3BFEBBFA45" box="[328,349,1386,1408]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="0" pageNumber="213">R</collectionCode>
.
<collectorName id="26B35311393FFFDD4133AB3BFE44FA45" box="[363,418,1386,1408]" pageId="0" pageNumber="213">Pérez</collectorName>
col. (
<specimenCount id="9D40FD4E393FFFDD41B7AB3BFDE0FABA" box="[495,518,1385,1408]" pageId="0" pageNumber="213" type="female">1♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B2E3C9A393FFFDD4246AB3BFDABFA5E" collectingDate="1985-09-12" collectorName="C. Granados &amp; Oi" country="Guatemala" location="San Lucas" pageId="0" pageNumber="213" specimenCount="1" specimenCount-female="1" stateProvince="Sacatepequez" typeStatus="paratype">
<collectingCountry id="F3517657393FFFDD427FAB3BFD35FA45" box="[551,723,1386,1408]" name="Guatemala" pageId="0" pageNumber="213">GUATEMALA</collectingCountry>
,
<collectingRegion id="4982F825393FFFDD42B9AB3BFC8BFA45" box="[737,877,1386,1408]" country="Guatemala" name="Sacatepequez" pageId="0" pageNumber="213">Sacatepéquez</collectingRegion>
,
<location id="8E99601C393FFFDD4321AB3BFC0EFA45" LSID="urn:lsid:plazi:treatment:03EF87D1393FFFDF4175AAD3FC63FB1D:8E99601C393FFFDD4321AB3BFC0EFA45" box="[889,1000,1386,1408]" country="Guatemala" name="San Lucas" pageId="0" pageNumber="213" stateProvince="Sacatepequez">San Lucas</location>
,
<date id="FFF81007393FFFDD40F7ABD4FEC0FA5E" box="[175,294,1413,1435]" pageId="0" pageNumber="213" value="1985-09-12">
<collectingDate id="EFBCE9EF393FFFDD40F7ABD4FEC0FA5E" box="[175,294,1413,1435]" pageId="0" pageNumber="213" value="1985-09-12">12-IX-1985</collectingDate>
</date>
,
<collectorName id="26B35311393FFFDD4169ABD4FE51FA5E" box="[305,439,1413,1435]" pageId="0" pageNumber="213">C. Granados</collectorName>
col. (1
<collectorName id="26B35311393FFFDD4254ABD5FDFBFA5F" box="[524,541,1412,1434]" pageId="0" pageNumber="213">Oi</collectorName>
,
<specimenCount id="9D40FD4E393FFFDD4273ABD4FDA5FA5F" box="[555,579,1412,1435]" pageId="0" pageNumber="213" type="female">1♀</specimenCount>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF936C7393FFFDD409FABF0FE64F9E7" blockId="0.[175,1005,1225,1570]" pageId="0" pageNumber="213">
<materialsCitation id="3B2E3C9A393FFFDD409FABF0FDBEFA28" country="Guatemala" location="Universidad del Valle" municipality="University" pageId="0" pageNumber="213" specimenCount="3" stateProvince="Nebraska" typeStatus="holotype">
The
<typeStatus id="54FD8865393FFFDD40AEABF0FEB6FA72" box="[246,336,1441,1463]" pageId="0" pageNumber="213" type="holotype">holotype</typeStatus>
and
<typeStatus id="54FD8865393FFFDD41DCABF0FE30FA72" box="[388,470,1441,1463]" pageId="0" pageNumber="213" type="allotype">allotype</typeStatus>
are deposited in the entomological collection of the
<location id="8E99601C393FFFDD40F7ABEDFE75FA17" LSID="urn:lsid:plazi:treatment:03EF87D1393FFFDF4175AAD3FC63FB1D:8E99601C393FFFDD40F7ABEDFE75FA17" box="[175,403,1468,1490]" country="Guatemala" municipality="University" name="Universidad del Valle" pageId="0" pageNumber="213" stateProvince="Nebraska">Universidad del Valle</location>
de
<collectingCountry id="F3517657393FFFDD41E7ABEDFDD2FA17" box="[447,564,1468,1490]" name="Guatemala" pageId="0" pageNumber="213">Guatemala</collectingCountry>
.
<typeStatus id="54FD8865393FFFDD4219ABEDFD43FA17" box="[577,677,1468,1490]" pageId="0" pageNumber="213" type="paratype">Paratypes</typeStatus>
are deposited in the following collections:
<collectingMunicipality id="6B9DACBD393FFFDD4171AB86FE75FA28" box="[297,403,1495,1517]" pageId="0" pageNumber="213">University</collectingMunicipality>
of
<collectingRegion id="4982F825393FFFDD41E1AB86FDBEFA28" box="[441,600,1495,1517]" country="United States of America" name="Nebraska" pageId="0" pageNumber="213">Nebraska State</collectingRegion>
</materialsCitation>
<materialsCitation id="3B2E3C9A393FFFDD4207AB86FE9BF9E7" collectorName="M. A. Moron &amp; L. Delgado &amp; A. Bitar" country="Mexico" location="Veracruz" pageId="0" pageNumber="213" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="holotype">
Museum (Lincoln, Nebraska, U.S.A.),
<collectorName id="26B35311393FFFDD40F7ABA0FEA4F9C2" box="[175,322,1521,1543]" pageId="0" pageNumber="213">M. A. Morón</collectorName>
(Xalapa,
<collectingRegion id="4982F825393FFFDD41F7ABA0FDE8F9C2" box="[431,526,1521,1543]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="0" pageNumber="213">Veracruz</collectingRegion>
,
<collectingCountry id="F3517657393FFFDD4243ABA0FD8DF9C2" box="[539,619,1521,1543]" name="Mexico" pageId="0" pageNumber="213">Mexico</collectingCountry>
),
<collectorName id="26B35311393FFFDD42D8ABA0FD11F9C2" box="[640,759,1521,1543]" pageId="0" pageNumber="213">L. Delgado</collectorName>
(
<collectingRegion id="4982F825393FFFDD4350ABA0FC68F9C2" box="[776,910,1521,1543]" country="Mexico" name="Distrito Federal" pageId="0" pageNumber="213">Mexico City</collectingRegion>
), and
<collectorName id="26B35311393FFFDD438BABA0FF05F9E7" pageId="0" pageNumber="213">A. Bitar</collectorName>
(
<collectingRegion id="4982F825393FFFDD40A9A85DFE90F9E7" box="[241,374,1548,1570]" country="Mexico" name="Distrito Federal" pageId="0" pageNumber="213">Mexico City</collectingRegion>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF936C7393FFFDD4266A86FFDB8F994" blockId="0.[574,606,1598,1617]" box="[574,606,1598,1617]" pageId="0" pageNumber="213">213</paragraph>
</subSubSection>
<caption id="DF39664F393EFFDC40FFAA57FC73FB8C" pageId="1" pageNumber="214" startId="1.[167,206,1030,1049]" targetBox="[181,940,194,1008]" targetPageId="1">
<paragraph id="8BF936C7393EFFDC40FFAA57FC73FB8C" blockId="1.[146,976,1030,1097]" pageId="1" pageNumber="214">
<emphasis id="B932EAD5393EFFDC40FFAA57FF1BFBDC" bold="true" box="[167,253,1030,1049]" pageId="1" pageNumber="214">Figs. 16</emphasis>
<taxonomicName id="4C464D44393EFFDC414AAA56FE4DFBDC" box="[274,427,1030,1050]" class="Insecta" family="Dynastidae" genus="Xyloryctes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="214" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="B932EAD5393EFFDC414AAA56FE97FBDF" box="[274,369,1031,1050]" italics="true" pageId="1" pageNumber="214">Xyloryctes</emphasis>
spp. 1
</taxonomicName>
) Lateral view of pronotum of
<taxonomicName id="4C464D44393EFFDC429CAA56FCBCFBDF" box="[708,858,1031,1050]" class="Insecta" family="Dynastidae" genus="Xyloryctes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="214" phylum="Arthropoda" rank="species" species="guatemalensis">
<emphasis id="B932EAD5393EFFDC429CAA56FCBCFBDF" box="[708,858,1031,1050]" italics="true" pageId="1" pageNumber="214">X. guatemalensis</emphasis>
</taxonomicName>
; 2) Same,
<taxonomicName id="4C464D44393EFFDC43E2AA56FEE7FBF4" class="Insecta" family="Dynastidae" genus="Xyloryctes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="214" phylum="Arthropoda" rank="species" species="howdenorum">
<emphasis id="B932EAD5393EFFDC43E2AA56FEE7FBF4" italics="true" pageId="1" pageNumber="214">X. howdenorum</emphasis>
</taxonomicName>
; 3) Frontal view of parameres of
<taxonomicName id="4C464D44393EFFDC421DAA4FFD3BFBF4" box="[581,733,1054,1073]" class="Insecta" family="Dynastidae" genus="Xyloryctes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="214" phylum="Arthropoda" rank="species" species="guatemalensis">
<emphasis id="B932EAD5393EFFDC421DAA4FFD3BFBF4" box="[581,733,1054,1073]" italics="true" pageId="1" pageNumber="214">X. guatemalensis</emphasis>
</taxonomicName>
; 4) Same, lateral view; 5) Frontal view of parameres of
<taxonomicName id="4C464D44393EFFDC41FEAA67FDD4FB8C" box="[422,562,1078,1097]" class="Insecta" family="Dynastidae" genus="Xyloryctes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="214" phylum="Arthropoda" rank="species" species="howdenorum">
<emphasis id="B932EAD5393EFFDC41FEAA67FDD4FB8C" box="[422,562,1078,1097]" italics="true" pageId="1" pageNumber="214">X. howdenorum</emphasis>
</taxonomicName>
; 6) Same, lateral view. Scale
<emphasis id="B932EAD5393EFFDC4319AA67FCB5FB8C" bold="true" box="[833,851,1078,1097]" italics="true" pageId="1" pageNumber="214">5</emphasis>
1 mm.
</paragraph>
</caption>
<subSubSection id="C35C654C393EFFDF40F2AA20FC69FE21" lastPageId="2" lastPageNumber="215" pageId="1" pageNumber="214" type="description">
<paragraph id="8BF936C7393EFFDC40F2AA20FE6FF9DB" blockId="1.[146,976,1137,1566]" pageId="1" pageNumber="214">
<emphasis id="B932EAD5393EFFDC40F2AA20FE2BFB42" bold="true" box="[170,461,1137,1159]" pageId="1" pageNumber="214">
Description.
<typeStatus id="54FD8865393EFFDC4177AA20FE6DFB42" box="[303,395,1137,1159]" pageId="1" pageNumber="214" type="holotype">Holotype</typeStatus>
male.
</emphasis>
Total body length:
<quantity id="4CBE9B22393EFFDC42F7AA20FCEAFB42" box="[687,780,1137,1159]" metricMagnitude="-2" metricUnit="m" metricValue="2.58" pageId="1" pageNumber="214" unit="mm" value="25.8">25.8 mm</quantity>
; maximum elytral width
<quantity id="4CBE9B22393EFFDC408CAADDFED7FB67" box="[212,305,1164,1186]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" pageId="1" pageNumber="214" unit="mm" value="13.8">13.8 mm</quantity>
. Head and pronotum shiny black; elytra, legs and ventral region reddish black. Head: Apex of clypeus bilobed and reflexed, lateral margins slightly concave, frons with a short, vertical, upright horn, horn flattened anteroposteriorly. Pronotum: Pronotal declivity vertical, not excavated, occupying apical third; pronotal process absent (Fig. 1); pronotum completely margined, anterior corners at right angles, lateral margins almost equally curved; pronotal cavity densely punctate, punctures becoming sparser towards posterior half, disc of pronotum with micropunctures. Elytra: Sutural stria impressed for entire length; surface smooth, with only some small, scarcely punctuate furrows. Venter: Prosternal process moderately short and with dense setae, projected backward. Pygidium: Surface regularly convex, basal region and sides finely rugose, remaining surface with small, sparse punctures. Legs: Apex of metatibia with three rounded teeth, without incisions between them. Genitalia: Parameres moderately large, apex slightly widened, with large and dense setae along sides and at apex (Figs. 34).
</paragraph>
<paragraph id="8BF936C7393DFFDF409FAE93FCCEFE9B" blockId="2.[175,1005,194,1240]" pageId="2" pageNumber="215">
<emphasis id="B932EAD5393DFFDF409FAE93FE8AFF1D" bold="true" box="[199,364,194,216]" pageId="2" pageNumber="215">
<typeStatus id="54FD8865393DFFDF409FAE93FEFDFF1D" box="[199,283,194,216]" pageId="2" pageNumber="215" type="allotype">Allotype</typeStatus>
female.
</emphasis>
Total body length
<quantity id="4CBE9B22393DFFDF426BAE93FD69FF1D" box="[563,655,194,216]" metricMagnitude="-2" metricUnit="m" metricValue="2.46" pageId="2" pageNumber="215" unit="mm" value="24.6">24.6 mm</quantity>
; maximum elytral width
<quantity id="4CBE9B22393DFFDF43D5AE93FC0FFF1D" box="[909,1001,194,216]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" pageId="2" pageNumber="215" unit="mm" value="13.4">13.4 mm</quantity>
. Similar to
<typeStatus id="54FD8865393DFFDF4142AE8CFE94FF36" box="[282,370,221,243]" pageId="2" pageNumber="215" type="holotype">holotype</typeStatus>
except for the following characters: Head with small tubercule between eyes. Pronotum regularly convex, disc with micropunctures, anterior angles rugopunctate. Elytra with furrows more pronounced. Pygidium wider, convex, with small punctures; last visible sternite not emarginate at apex. Protibial teeth acute. Metatibia with teeth smaller, shorter spur preapically widened.
</paragraph>
<paragraph id="8BF936C7393DFFDF409FAF32FC69FE21" blockId="2.[175,1005,194,1240]" pageId="2" pageNumber="215">
<emphasis id="B932EAD5393DFFDF409FAF32FDE6FEBC" bold="true" box="[199,512,355,377]" pageId="2" pageNumber="215">
Variation (
<specimenCount id="9D40FD4E393DFFDF416EAF32FE63FEBC" box="[310,389,355,377]" pageId="2" pageNumber="215" type="male">4 males</specimenCount>
,
<specimenCount id="9D40FD4E393DFFDF41CBAF32FE10FEBC" box="[403,502,355,377]" pageId="2" pageNumber="215" type="female">3 females</specimenCount>
).
</emphasis>
<materialsCitation id="3B2E3C9A393DFFDF4251AF32FC6DFE21" location="The" municipality="Males" pageId="2" pageNumber="215" specimenCount="1" stateProvince="Guatemala" typeStatus="paratype">
Male
<typeStatus id="54FD8865393DFFDF421EAF32FD4FFEBC" box="[582,681,355,377]" pageId="2" pageNumber="215" type="paratype">paratypes</typeStatus>
with lengths of 23.0
<quantity id="4CBE9B22393DFFDF43D5AF32FC0EFEBC" box="[909,1000,355,377]" metricMagnitude="-2" metricUnit="m" metricValue="2.43" pageId="2" pageNumber="215" unit="mm" value="24.3">24.3 mm</quantity>
; females with lengths of
<quantity id="4CBE9B22393DFFDF41F7AF2FFDA4FE51" box="[431,578,382,404]" metricMagnitude="-2" metricUnit="m" metricValue="2.26" metricValueMax="2.31" metricValueMin="2.21" pageId="2" pageNumber="215" unit="mm" value="22.6" valueMax="23.1" valueMin="22.1">22.123.1 mm</quantity>
.
<collectingMunicipality id="6B9DACBD393DFFDF4217AF2FFD6BFE51" box="[591,653,382,404]" pageId="2" pageNumber="215">Males</collectingMunicipality>
with maximum elytral widths of
<quantity id="4CBE9B22393DFFDF40F7AFC8FEA4FE6A" box="[175,322,409,431]" metricMagnitude="-2" metricUnit="m" metricValue="1.3649999999999998" metricValueMax="1.3699999999999999" metricValueMin="1.3599999999999999" pageId="2" pageNumber="215" unit="mm" value="13.649999999999999" valueMax="13.7" valueMin="13.6">13.613.7 mm</quantity>
; maximum female elytral widths
<quantity id="4CBE9B22393DFFDF42FAAFC8FCD0FE6A" box="[674,822,409,431]" metricMagnitude="-2" metricUnit="m" metricValue="1.305" metricValueMax="1.39" metricValueMin="1.22" pageId="2" pageNumber="215" unit="mm" value="13.05" valueMax="13.9" valueMin="12.2">12.213.9 mm</quantity>
.
<location id="8E99601C393DFFDF431AAFC8FC8CFE6A" LSID="urn:lsid:plazi:treatment:03EF87D1393FFFDF4175AAD3FC63FB1D:8E99601C393DFFDF431AAFC8FC8CFE6A" box="[834,874,409,431]" municipality="Males" name="The" pageId="2" pageNumber="215" stateProvince="Guatemala">The</location>
variation in male punctation is minimal, and in the smallest female specimen (from the region of
<collectingRegion id="4982F825393DFFDF4094AF9FFE95FE21" box="[204,371,462,484]" country="Guatemala" name="Guatemala" pageId="2" pageNumber="215">Guatemala City</collectingRegion>
) the elytral striae and punctation are more marked
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35C654C393DFFDF409FAFBBFCEBFDC5" box="[199,781,490,512]" pageId="2" pageNumber="215" type="materials_examined">
<paragraph id="8BF936C7393DFFDF409FAFBBFCEBFDC5" blockId="2.[175,1005,194,1240]" box="[199,781,490,512]" pageId="2" pageNumber="215">
<emphasis id="B932EAD5393DFFDF409FAFBBFEBDFDC5" bold="true" box="[199,347,490,512]" pageId="2" pageNumber="215">Type Locality.</emphasis>
San José Pinula,
<collectingCountry id="F3517657393DFFDF424DAFBBFD6FFDC5" box="[533,649,490,512]" name="Guatemala" pageId="2" pageNumber="215">Guatemala</collectingCountry>
,
<collectingCountry id="F3517657393DFFDF42CDAFBBFCECFDC5" box="[661,778,490,512]" name="Guatemala" pageId="2" pageNumber="215">Guatemala</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C35C654C393DFFDF409FAC57FE1AFBF3" pageId="2" pageNumber="215" type="etymology">
<paragraph id="8BF936C7393DFFDF409FAC57FC44FDF2" blockId="2.[175,1005,194,1240]" pageId="2" pageNumber="215">
<emphasis id="B932EAD5393DFFDF409FAC57FEDCFDD9" bold="true" box="[199,314,518,540]" pageId="2" pageNumber="215">Etymology.</emphasis>
Named for
<collectingCountry id="F3517657393DFFDF418DAC57FDAFFDD9" box="[469,585,518,540]" name="Guatemala" pageId="2" pageNumber="215">Guatemala</collectingCountry>
, the country where this species was discovered, combined with the suffix -ensis, indicating belonging to.
</paragraph>
<paragraph id="8BF936C7393DFFDF409FAC6DFDA1FCCB" blockId="2.[175,1005,194,1240]" pageId="2" pageNumber="215">
<emphasis id="B932EAD5393DFFDF409FAC6DFE78FD97" bold="true" box="[199,414,572,594]" pageId="2" pageNumber="215">Taxonomic Remarks.</emphasis>
<taxonomicName id="4C464D44393DFFDF41FDAC6DFD47FD94" box="[421,673,572,593]" class="Insecta" family="Dynastidae" genus="Xyloryctes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="215" phylum="Arthropoda" rank="species" species="guatemalensis">
<emphasis id="B932EAD5393DFFDF41FDAC6DFD47FD94" box="[421,673,572,593]" italics="true" pageId="2" pageNumber="215">Xyloryctes guatemalensis</emphasis>
</taxonomicName>
can be distinguished from other species of the genus by the following combination of characters: elytra with distinct sutural striae; apex of metatibiae with three rounded teeth, without incisions between them; males with cephalic horn flattened anteroposteriorly; pronotal declivity vertical, not excavated, occupying apical third of pronotum; pronotal process absent (Fig. 1); and shape of the parameres distinctive (Figs. 3 4). Females have the pygidium almost glabrous and convex, and the shorter spur of the metatibia is preapically widened.
</paragraph>
<paragraph id="8BF936C7393DFFDF409FAD42FCBEFC6A" blockId="2.[175,1005,194,1240]" pageId="2" pageNumber="215">
Males of
<taxonomicName id="4C464D44393DFFDF416CAD42FE05FCED" box="[308,483,787,808]" class="Insecta" family="Dynastidae" genus="Xyloryctes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="215" phylum="Arthropoda" rank="species" species="guatemalensis">
<emphasis id="B932EAD5393DFFDF416CAD42FE05FCED" box="[308,483,787,808]" italics="true" pageId="2" pageNumber="215">X. guatemalensis</emphasis>
</taxonomicName>
are similar to those of
<taxonomicName id="4C464D44393DFFDF42AFAD42FE63FC81" authority="Delgado and Najera" authorityName="Delgado and Najera" class="Insecta" family="Dynastidae" genus="Xyloryctes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="215" phylum="Arthropoda" rank="species" species="howdenorum">
<emphasis id="B932EAD5393DFFDF42AFAD42FC0AFCED" box="[759,1004,787,808]" italics="true" pageId="2" pageNumber="215">Xyloryctes howdenorum</emphasis>
Delgado and Nájera
</taxonomicName>
, but they can be distinguished by the following differences: cephalic horn flattened anteroposteriorly in
<taxonomicName id="4C464D44393DFFDF42DCAD18FCD6FC9B" box="[644,816,841,862]" class="Insecta" family="Dynastidae" genus="Xyloryctes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="215" phylum="Arthropoda" rank="species" species="guatemalensis">
<emphasis id="B932EAD5393DFFDF42DCAD18FCD6FC9B" box="[644,816,841,862]" italics="true" pageId="2" pageNumber="215">X. guatemalensis</emphasis>
</taxonomicName>
(cylindrical in
<taxonomicName id="4C464D44393DFFDF438DAD18FECBFCBC" class="Insecta" family="Dynastidae" genus="Xyloryctes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="215" phylum="Arthropoda" rank="species" species="howdenorum">
<emphasis id="B932EAD5393DFFDF438DAD18FECBFCBC" italics="true" pageId="2" pageNumber="215">X. howdenorum</emphasis>
</taxonomicName>
); pronotal declivity vertical in
<taxonomicName id="4C464D44393DFFDF42C9AD35FCA7FCBC" box="[657,833,868,889]" class="Insecta" family="Dynastidae" genus="Xyloryctes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="215" phylum="Arthropoda" rank="species" species="guatemalensis">
<emphasis id="B932EAD5393DFFDF42C9AD35FCA7FCBC" box="[657,833,868,889]" italics="true" pageId="2" pageNumber="215">X. guatemalensis</emphasis>
</taxonomicName>
(oblique in
<taxonomicName id="4C464D44393DFFDF438DAD35FECBFC51" class="Insecta" family="Dynastidae" genus="Xyloryctes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="215" phylum="Arthropoda" rank="species" species="howdenorum">
<emphasis id="B932EAD5393DFFDF438DAD35FECBFC51" italics="true" pageId="2" pageNumber="215">X. howdenorum</emphasis>
</taxonomicName>
) (Figs. 12); and parameres without a lateral tooth in
<taxonomicName id="4C464D44393DFFDF438DAD2EFEDFFC6B" class="Insecta" family="Dynastidae" genus="Xyloryctes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="215" phylum="Arthropoda" rank="species" species="guatemalensis">
<emphasis id="B932EAD5393DFFDF438DAD2EFEDFFC6B" italics="true" pageId="2" pageNumber="215">X. guatemalensis</emphasis>
</taxonomicName>
(with a lateral tooth in
<taxonomicName id="4C464D44393DFFDF426EADC8FD32FC6B" box="[566,724,921,942]" class="Insecta" family="Dynastidae" genus="Xyloryctes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="215" phylum="Arthropoda" rank="species" species="howdenorum">
<emphasis id="B932EAD5393DFFDF426EADC8FD32FC6B" box="[566,724,921,942]" italics="true" pageId="2" pageNumber="215">X. howdenorum</emphasis>
</taxonomicName>
) (Figs. 36).
</paragraph>
<paragraph id="8BF936C7393DFFDF409FADE4FE1AFBF3" blockId="2.[175,1005,194,1240]" pageId="2" pageNumber="215">
Females of
<taxonomicName id="4C464D44393DFFDF4119ADE4FE0DFC0F" box="[321,491,949,970]" class="Insecta" family="Dynastidae" genus="Xyloryctes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="215" phylum="Arthropoda" rank="species" species="guatemalensis">
<emphasis id="B932EAD5393DFFDF4119ADE4FE0DFC0F" box="[321,491,949,970]" italics="true" pageId="2" pageNumber="215">X. guatemalensis</emphasis>
</taxonomicName>
are similar to those of
<taxonomicName id="4C464D44393DFFDF42BAADE4FC0EFC0E" authority="Bates" authorityName="Bates" authorityYear="1888" box="[738,1000,949,971]" class="Insecta" family="Dynastidae" genus="Xyloryctes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="215" phylum="Arthropoda" rank="species" species="teuthras">
<emphasis id="B932EAD5393DFFDF42BAADE4FC40FC0F" box="[738,934,949,970]" italics="true" pageId="2" pageNumber="215">Xyloryctes teuthras</emphasis>
Bates
</taxonomicName>
, except for the following characters: body length is generally less than 24.0 mm in
<taxonomicName id="4C464D44393DFFDF40F7ADBBFEBDFC3A" box="[175,347,1002,1023]" class="Insecta" family="Dynastidae" genus="Xyloryctes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="215" phylum="Arthropoda" rank="species" species="guatemalensis">
<emphasis id="B932EAD5393DFFDF40F7ADBBFEBDFC3A" box="[175,347,1002,1023]" italics="true" pageId="2" pageNumber="215">X. guatemalensis</emphasis>
</taxonomicName>
(longer than 24.0 mm in
<taxonomicName id="4C464D44393DFFDF422AADBBFD03FC3A" authorityName="Bates" authorityYear="1888" box="[626,741,1002,1023]" class="Insecta" family="Dynastidae" genus="Xyloryctes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="215" phylum="Arthropoda" rank="species" species="teuthras">
<emphasis id="B932EAD5393DFFDF422AADBBFD03FC3A" box="[626,741,1002,1023]" italics="true" pageId="2" pageNumber="215">X. teuthras</emphasis>
</taxonomicName>
); pygidium convex, base almost glabrous in
<taxonomicName id="4C464D44393DFFDF412AAA54FDFCFBDF" box="[370,538,1029,1050]" class="Insecta" family="Dynastidae" genus="Xyloryctes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="215" phylum="Arthropoda" rank="species" species="guatemalensis">
<emphasis id="B932EAD5393DFFDF412AAA54FDFCFBDF" box="[370,538,1029,1050]" italics="true" pageId="2" pageNumber="215">X. guatemalensis</emphasis>
</taxonomicName>
(slightly convex to flattened and with distinct setae at the base in
<taxonomicName id="4C464D44393DFFDF4125AA71FE08FBF0" authorityName="Bates" authorityYear="1888" box="[381,494,1056,1077]" class="Insecta" family="Dynastidae" genus="Xyloryctes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="215" phylum="Arthropoda" rank="species" species="teuthras">
<emphasis id="B932EAD5393DFFDF4125AA71FE08FBF0" box="[381,494,1056,1077]" italics="true" pageId="2" pageNumber="215">X. teuthras</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C35C654C393DFFDF409FAA6AFC63FB1D" pageId="2" pageNumber="215" type="distribution">
<paragraph id="8BF936C7393DFFDF409FAA6AFC03FB67" blockId="2.[175,1005,194,1240]" pageId="2" pageNumber="215">
<emphasis id="B932EAD5393DFFDF409FAA6AFEA0FB94" bold="true" box="[199,326,1083,1105]" pageId="2" pageNumber="215">Distribution.</emphasis>
<taxonomicName id="4C464D44393DFFDF4114AA6DFDA1FB94" box="[332,583,1084,1105]" class="Insecta" family="Dynastidae" genus="Xyloryctes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="215" phylum="Arthropoda" rank="species" species="guatemalensis">
<emphasis id="B932EAD5393DFFDF4114AA6DFDA1FB94" box="[332,583,1084,1105]" italics="true" pageId="2" pageNumber="215">Xyloryctes guatemalensis</emphasis>
</taxonomicName>
is known from three localities situated in the southern region of
<collectingCountry id="F3517657393DFFDF41FAAA07FDF1FBA9" box="[418,535,1110,1132]" name="Guatemala" pageId="2" pageNumber="215">Guatemala</collectingCountry>
. The
<collectingRegion id="4982F825393DFFDF420CAA07FD1CFBA9" box="[596,762,1110,1132]" country="Guatemala" name="Guatemala" pageId="2" pageNumber="215">Guatemala City</collectingRegion>
(
<quantity id="4CBE9B22393DFFDF4352AA07FCB8FBA9" box="[778,862,1110,1132]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="2" pageNumber="215" unit="m" value="1520.0">1,520 m</quantity>
) and the San José Pinula (
<quantity id="4CBE9B22393DFFDF4168AA20FE63FB42" box="[304,389,1137,1159]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="2" pageNumber="215" unit="m" value="1750.0">1,750 m</quantity>
) localities are located in the Department of
<collectingCountry id="F3517657393DFFDF431FAA20FC5DFB42" box="[839,955,1137,1159]" name="Guatemala" pageId="2" pageNumber="215">Guatemala</collectingCountry>
, and the San Lucas locality (
<quantity id="4CBE9B22393DFFDF41FBAADDFE1EFB67" box="[419,504,1164,1186]" metricMagnitude="3" metricUnit="m" metricValue="2.09" pageId="2" pageNumber="215" unit="m" value="2090.0">2,090 m</quantity>
) is located in the Department of
<collectingRegion id="4982F825393DFFDF430CAADDFC06FB67" box="[852,992,1164,1186]" country="Guatemala" name="Sacatepequez" pageId="2" pageNumber="215">Sacatepéquez</collectingRegion>
.
</paragraph>
<paragraph id="8BF936C7393DFFDF409FAAF6FC63FB1D" blockId="2.[175,1005,194,1240]" pageId="2" pageNumber="215">
This new species inhabits regions with premontane and montane wet forests (such as oak and pine-oak forests) (
<emphasis id="B932EAD5393DFFDF427CAA93FDBFFB12" box="[548,601,1218,1239]" italics="true" pageId="2" pageNumber="215">sensu</emphasis>
Campbell and Lamar 1989).
</paragraph>
</subSubSection>
</treatment>
</document>