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<document id="2C366218E199A72AC285D69B3923A039" ID-DOI="10.11646/zootaxa.4092.1.4" ID-GBIF-Dataset="f448a9b2-e7ab-4dbf-87a1-64d0338b8b95" ID-ISSN="1175-5326" ID-Zenodo-Dep="266115" ID-ZooBank="51F44763-E817-4E58-A4AC-525E63B6D27B" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1458036828487" checkinUser="plazi" docAuthor="Fernandez, Julio C. C., Cárdenas, César A., Bravo, Alejandro, Lôbo-Hajdu, Gisele, Willenz, Philippe &amp; Hajdu, Eduardo" docDate="2016" docId="03EF87BEFFC0FFB2419BFDD1FA2574CA" docLanguage="en" docName="15617-5152-1-PB.pdf.imf" docOrigin="Zootaxa 4092 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lissodendoryx (Ectyodoryx) diegoramirezensis Fernandez, Cárdenas, Bravo, Lôbo-Hajdu, Willenz &amp; Hajdu, 2016, sp. nov." docType="treatment" docVersion="13" lastPageNumber="84" masterDocId="FFD6FFC6FFCDFFBD410CFFBDFF5F7320" masterDocTitle="Lissodendoryx (Ectyodoryx) Lundbeck, 1909 (Coelosphaeridae, Poecilosclerida, Demospongiae) from Southern Chile: new species and a discussion of morphologic characters in the subgenus" pageNumber="82" updateTime="1698184538152" updateUser="plazi">
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<mods:title id="8E1BF4E3A93E4B4DCD05776A09423C7C">Lissodendoryx (Ectyodoryx) Lundbeck, 1909 (Coelosphaeridae, Poecilosclerida, Demospongiae) from Southern Chile: new species and a discussion of morphologic characters in the subgenus</mods:title>
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<mods:namePart id="1937394F8CA29FF7E8099DB08D8BFF5E">Fernandez, Julio C. C.</mods:namePart>
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<mods:namePart id="1CA68039A011FFD497B4B37A96D0B395">Cárdenas, César A.</mods:namePart>
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<mods:namePart id="7C29C04A4B29D47B5A5E26065BCFF122">Bravo, Alejandro</mods:namePart>
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<mods:namePart id="FACF257308463309D74388F116D2C8BC">Willenz, Philippe</mods:namePart>
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<treatment id="03EF87BEFFC0FFB2419BFDD1FA2574CA" ID-DOI="http://doi.org/10.5281/zenodo.5615611" ID-GBIF-Taxon="117911994" ID-Zenodo-Dep="5615611" LSID="urn:lsid:plazi:treatment:03EF87BEFFC0FFB2419BFDD1FA2574CA" httpUri="http://treatment.plazi.org/id/03EF87BEFFC0FFB2419BFDD1FA2574CA" lastPageId="15" lastPageNumber="84" pageId="13" pageNumber="82">
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<paragraph id="8BF936A8FFC0FFB0419BFDD1FC6E71A6" blockId="13.[151,817,620,679]" box="[151,817,620,646]" pageId="13" pageNumber="82">
<heading id="D0B181C4FFC0FFB0419BFDD1FC6E71A6" bold="true" box="[151,817,620,646]" fontSize="11" level="1" pageId="13" pageNumber="82" reason="1">
<emphasis id="B932EABAFFC0FFB0419BFDD1FC6E71A6" bold="true" box="[151,817,620,646]" pageId="13" pageNumber="82">
<taxonomicName id="4C464D2BFFC0FFB0419BFDD1FD9471A6" ID-CoL="3VF86" box="[151,715,620,646]" class="Demospongiae" family="Coelosphaeridae" genus="Lissodendoryx" kingdom="Animalia" order="Poecilosclerida" pageId="13" pageNumber="82" phylum="Porifera" rank="species" species="diegoramirezensis" status="sp. nov." subGenus="Ectyodoryx">
<emphasis id="B932EABAFFC0FFB0419BFDD1FE1871A6" bold="true" box="[151,327,620,646]" italics="true" pageId="13" pageNumber="82">Lissodendoryx</emphasis>
(
<emphasis id="B932EABAFFC0FFB04054FDD1FEBE71A6" bold="true" box="[344,481,620,646]" italics="true" pageId="13" pageNumber="82">Ectyodoryx</emphasis>
)
<emphasis id="B932EABAFFC0FFB040FDFDD1FD9471A6" bold="true" box="[497,715,620,646]" italics="true" pageId="13" pageNumber="82">diegoramirezensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20157C1FFC0FFB043DFFDD1FC6E71A6" box="[723,817,620,646]" pageId="13" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF936A8FFC0FFB0419BFD33FE247187" blockId="13.[151,817,620,679]" box="[151,379,654,679]" pageId="13" pageNumber="82">
(
<tableCitation id="C6C40313FFC0FFB04193FD33FFB77187" box="[159,232,654,679]" captionStart="TABLE 2" captionStartId="5.[151,242,152,175]" captionTargetBox="[160,1427,266,1950]" captionTargetPageId="5" captionText="TABLE 2. Comparative table for the spicule micrometries (µm), distributions and depth of occurrence of Lissodendoryx (Ectyodoryx) Lundbeck, 1909 known from the Antarctic, the Subantarctic region, the SW Atlantic and SE Pacific; * (absent),? (no information)." httpUri="http://table.plazi.org/id/DF396620FFC8FFB8419BFF25FE9A73CF" pageId="13" pageNumber="82" tableUuid="DF396620FFC8FFB8419BFF25FE9A73CF">Tab. 2</tableCitation>
;
<figureCitation id="137D2A2DFFC0FFB041F8FD33FE2D7187" box="[244,370,654,679]" captionStart-0="FIGURE 10. A C" captionStart-1="FIGURE 11" captionStart-2="FIGURE 12" captionStart-3="FIGURE 13" captionStartId-0="14.[151,250,1118,1140]" captionStartId-1="14.[151,250,1877,1899]" captionStartId-2="15.[151,250,1123,1145]" captionStartId-3="16.[151,250,1427,1449]" captionTargetBox-0="[188,1398,337,1096]" captionTargetBox-1="[188,1398,1251,1856]" captionTargetBox-2="[188,1398,193,1100]" captionTargetBox-3="[188,1398,195,1405]" captionTargetId-0="figure@14.[188,1398,337,1097]" captionTargetId-1="figure@14.[188,1398,1251,1856]" captionTargetId-2="figure@15.[188,1398,193,1101]" captionTargetId-3="figure@16.[188,1398,195,1405]" captionTargetPageId-0="14" captionTargetPageId-1="14" captionTargetPageId-2="15" captionTargetPageId-3="16" captionText-0="FIGURE 10. A C, Lissodendoryx (Ectyodoryx) diegoramirezensis sp. nov., habit. A C, holotype (IZUA POR 169). A, whole specimen (dry); B, markedly hispid surface in detail; C, surface under transmitted light showing reticulated pattern. Scales: A = 0.5 cm; B C = 0.1 cm." captionText-1="FIGURE 11. Lissodendoryx (Ectyodoryx) diegoramirezensis sp. nov., skeleton. A D, holotype (IZUA POR 169): A, overview of skeletal architecture; B, choanosomal reticulation in detail; C, section of a main tract echinated by smaller acanthostyles, i = larger acanthostyle, ii = intermediate acanthostyle, iii = smaller acanthostyle; D, detail of surface with tylotes scattered in disorder. Scales: A = 500 µm; B D = 200 µm." captionText-2="FIGURE 12. Lissodendoryx (Ectyodoryx) diegoramirezensis sp. nov., electron micrographs. A K, holotype (IZUA POR 169): A, aniso-subtylotes; B, extremities of the aniso-subtylotes; C, acanthostyles I; D, bases of acanthostyles I; E, acanthostyles II; F, bases of acanthostyles II; G, acanthostyles III; H, bases of acanthostyles III, and sigma II; I, arcuate isochelae; J, sigmas I; K, sigmas II. Scales: A, C, E, G = 100 µm; B = 5 µm; D, F, H I, K = 10 µm; J = 20 µm." captionText-3="FIGURE 13. Lissodendoryx (Ectyodoryx) anacantha (Hentschel, 1914), electron micrographs. A I, holotype (ZMH S 2327): A, aniso-subtylotes; B, extremities of aniso-subtylotes; C, acanthostyles I, and sigma I; D, bases of acanthostyles I; E, acanthostyles II, and sigma II; F, bases of acanthostyles II; G, arcuate isochelae; H, sigmas I; I, sigmas II. Scales: A, C, E = 100 µm; B, D, H = 10 µm; F G, I = 5 µm." httpUri-0="https://zenodo.org/record/266125/files/figure.png" httpUri-1="https://zenodo.org/record/266126/files/figure.png" httpUri-2="https://zenodo.org/record/266127/files/figure.png" httpUri-3="https://zenodo.org/record/266128/files/figure.png" pageId="13" pageNumber="82">Figs 1013</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35C6523FFC0FFB0419BFD6AFE2E705F" pageId="13" pageNumber="82" type="materials_examined">
<paragraph id="8BF936A8FFC0FFB0419BFD6AFFBB7017" blockId="13.[151,1437,726,2011]" pageId="13" pageNumber="82">
<emphasis id="B932EABAFFC0FFB0419BFD6AFE5471CE" bold="true" box="[151,267,727,750]" pageId="13" pageNumber="82">
<typeStatus id="54FD880AFFC0FFB0419BFD6AFE5971CE" box="[151,262,727,750]" pageId="13" pageNumber="82" type="holotype">Holotype</typeStatus>
.
</emphasis>
IZUAPOR 169,
<collectingCountry id="F3517638FFC0FFB040E0FD6BFD4B71CF" box="[492,532,726,751]" name="Iceland" pageId="13" pageNumber="82">Isla</collectingCountry>
Diego Ramírez, southern
<collectingCountry id="F3517638FFC0FFB0425BFD6BFCCB71CF" box="[855,916,726,751]" name="Chile" pageId="13" pageNumber="82">Chile</collectingCountry>
(56ºS, off Cape Horn), ca.
<quantity id="4CBE9B4DFFC0FFB045E2FD6AFA1671CE" box="[1262,1353,727,751]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="82" unit="m" value="2000.0">2000 m</quantity>
depth,
<date id="FFF81068FFC0FFB0419BFD47FE0F7033" box="[151,336,762,787]" pageId="13" pageNumber="82" value="2002-09">September 2002</date>
, bycatch from longline fisheries, Leg. A. Bravo. Fragment from
<typeStatus id="54FD880AFFC0FFB04521FD47FBD07033" box="[1069,1167,762,787]" pageId="13" pageNumber="82" type="holotype">holotype</typeStatus>
deposited under MNRJ 10884.
</paragraph>
<paragraph id="8BF936A8FFC0FFB041CBFCFFFE2E705F" blockId="13.[151,1437,726,2011]" pageId="13" pageNumber="82">
<emphasis id="B932EABAFFC0FFB041CBFCFFFE85707B" bold="true" box="[199,474,834,859]" pageId="13" pageNumber="82">Comparative material.</emphasis>
<taxonomicName id="4C464D2BFFC0FFB040E8FCFFFB26707B" authority="Hentschel, 1914" authorityName="Hentschel" authorityYear="1914" box="[484,1145,834,859]" class="Demospongiae" family="Coelosphaeridae" genus="Lissodendoryx" kingdom="Animalia" order="Poecilosclerida" pageId="13" pageNumber="82" phylum="Porifera" rank="species" species="anacantha" subGenus="Ectyodoryx">
<emphasis id="B932EABAFFC0FFB040E8FCFFFDD8707B" box="[484,647,834,859]" italics="true" pageId="13" pageNumber="82">Lissodendoryx</emphasis>
(
<emphasis id="B932EABAFFC0FFB04396FCFFFC46707B" box="[666,793,834,859]" italics="true" pageId="13" pageNumber="82">Ectyodoryx</emphasis>
)
<emphasis id="B932EABAFFC0FFB04221FCFFFCFC707B" box="[813,931,834,859]" italics="true" pageId="13" pageNumber="82">anacantha</emphasis>
(Hentschel, 1914)
</taxonomicName>
(ZMH S 2327, fragment from the
<typeStatus id="54FD880AFFC0FFB041F1FCDBFE3B705F" box="[253,356,870,895]" pageId="13" pageNumber="82" type="holotype">holotype</typeStatus>
).
</paragraph>
</subSubSection>
<subSubSection id="C35C6523FFC0FFB041CBFC37FB6C70CB" pageId="13" pageNumber="82" type="diagnosis">
<paragraph id="8BF936A8FFC0FFB041CBFC37FB6C70CB" blockId="13.[151,1437,726,2011]" pageId="13" pageNumber="82">
<emphasis id="B932EABAFFC0FFB041CBFC37FE1E7083" bold="true" box="[199,321,906,931]" pageId="13" pageNumber="82">Diagnosis.</emphasis>
Seemingly erect
<taxonomicName id="4C464D2BFFC0FFB04308FC37FC1E7083" authority="(Ectyodoryx)" baseAuthorityName="Ectyodoryx" box="[516,833,906,931]" class="Demospongiae" family="Coelosphaeridae" genus="Lissodendoryx" kingdom="Animalia" order="Poecilosclerida" pageId="13" pageNumber="82" phylum="Porifera" rank="genus">
<emphasis id="B932EABAFFC0FFB04308FC37FDF87083" box="[516,679,906,931]" italics="true" pageId="13" pageNumber="82">Lissodendoryx</emphasis>
(
<emphasis id="B932EABAFFC0FFB043BBFC37FC697083" box="[695,822,906,931]" italics="true" pageId="13" pageNumber="82">Ectyodoryx</emphasis>
)
</taxonomicName>
with irregular surface, terminally microspined tylotes (223252/7.29.6), acanthostyles (I. 435602/2431, II. 320415/1921, III. 220242/1417), arcuate isochelae with lateral alae bent towards the shaft (2224), and sigmas (
<date id="FFF81068FFC0FFB0424EFC6FFC3270CB" box="[834,877,978,1003]" pageId="13" pageNumber="82" value="1945-01">I.45</date>
60, II. 18.625).
</paragraph>
</subSubSection>
<subSubSection id="C35C6523FFC0FFB041CBFC4BFD9C744F" pageId="13" pageNumber="82" type="description">
<paragraph id="8BF936A8FFC0FFB041CBFC4BFE6477E3" blockId="13.[151,1437,726,2011]" pageId="13" pageNumber="82">
<emphasis id="B932EABAFFC0FFB041CBFC4BFE06772F" bold="true" box="[199,345,1014,1039]" pageId="13" pageNumber="82">Description.</emphasis>
The
<typeStatus id="54FD880AFFC0FFB04091FC4BFEA0772F" box="[413,511,1014,1039]" pageId="13" pageNumber="82" type="holotype">holotype</typeStatus>
is incomplete and therefore imprecise to describe its gross morphology. It is seemingly erect, possibly stalked (
<figureCitation id="137D2A2DFFC0FFB04324FBA7FDCC7713" box="[552,659,1050,1075]" captionStart="FIGURE 10. A C" captionStartId="14.[151,250,1118,1140]" captionTargetBox="[188,1398,337,1096]" captionTargetId="figure@14.[188,1398,337,1097]" captionTargetPageId="14" captionText="FIGURE 10. A C, Lissodendoryx (Ectyodoryx) diegoramirezensis sp. nov., habit. A C, holotype (IZUA POR 169). A, whole specimen (dry); B, markedly hispid surface in detail; C, surface under transmitted light showing reticulated pattern. Scales: A = 0.5 cm; B C = 0.1 cm." httpUri="https://zenodo.org/record/266125/files/figure.png" pageId="13" pageNumber="82">Fig. 10A</figureCitation>
), ca.
<quantity id="4CBE9B4DFFC0FFB043DBFBA6FC4D7712" box="[727,786,1051,1075]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="82" unit="cm" value="2.0">2 cm</quantity>
high and
<quantity id="4CBE9B4DFFC0FFB04281FBA6FC827712" box="[909,989,1051,1075]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="13" pageNumber="82" unit="cm" value="0.4">0.4 cm</quantity>
thick. Surface rather hispid, irregular, bearing short projections (
<figureCitation id="137D2A2DFFC0FFB040CFFB83FD787777" box="[451,551,1086,1111]" captionStart="FIGURE 10. A C" captionStartId="14.[151,250,1118,1140]" captionTargetBox="[188,1398,337,1096]" captionTargetId="figure@14.[188,1398,337,1097]" captionTargetPageId="14" captionText="FIGURE 10. A C, Lissodendoryx (Ectyodoryx) diegoramirezensis sp. nov., habit. A C, holotype (IZUA POR 169). A, whole specimen (dry); B, markedly hispid surface in detail; C, surface under transmitted light showing reticulated pattern. Scales: A = 0.5 cm; B C = 0.1 cm." httpUri="https://zenodo.org/record/266125/files/figure.png" pageId="13" pageNumber="82">Fig. 10B</figureCitation>
) and roundish openings (ca.
<quantity id="4CBE9B4DFFC0FFB0427AFB82FC9B7776" box="[886,964,1087,1111]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="82" unit="cm" value="0.1">0.1 cm</quantity>
across), which are of doubtful oscular in function. A gross reticulation is apparent in the entire sponge under transmitted light in a stereo microscope (
<figureCitation id="137D2A2DFFC0FFB04463FBDFFF9677BF" captionStart="FIGURE 10. A C" captionStartId="14.[151,250,1118,1140]" captionTargetBox="[188,1398,337,1096]" captionTargetId="figure@14.[188,1398,337,1097]" captionTargetPageId="14" captionText="FIGURE 10. A C, Lissodendoryx (Ectyodoryx) diegoramirezensis sp. nov., habit. A C, holotype (IZUA POR 169). A, whole specimen (dry); B, markedly hispid surface in detail; C, surface under transmitted light showing reticulated pattern. Scales: A = 0.5 cm; B C = 0.1 cm." httpUri="https://zenodo.org/record/266125/files/figure.png" pageId="13" pageNumber="82">Fig. 10C</figureCitation>
). Colour in life not reported, dried
<typeStatus id="54FD880AFFC0FFB04352FB3BFD9F77BF" box="[606,704,1158,1183]" pageId="13" pageNumber="82" type="holotype">holotype</typeStatus>
beige. Consistency relatively compressible and delicate, texture slightly rough.
</paragraph>
<paragraph id="8BF936A8FFC0FFB041CBFB73FBED76BB" blockId="13.[151,1437,726,2011]" pageId="13" pageNumber="82">
<emphasis id="B932EABAFFC0FFB041CBFB73FE6A77C7" bold="true" box="[199,309,1230,1255]" pageId="13" pageNumber="82">Skeleton.</emphasis>
Plumoreticulate with a slight degree of axial compression (
<figureCitation id="137D2A2DFFC0FFB042D5FB73FB7477C7" box="[985,1067,1230,1255]" captionStart="FIGURE 11" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[188,1398,1251,1856]" captionTargetId="figure@14.[188,1398,1251,1856]" captionTargetPageId="14" captionText="FIGURE 11. Lissodendoryx (Ectyodoryx) diegoramirezensis sp. nov., skeleton. A D, holotype (IZUA POR 169): A, overview of skeletal architecture; B, choanosomal reticulation in detail; C, section of a main tract echinated by smaller acanthostyles, i = larger acanthostyle, ii = intermediate acanthostyle, iii = smaller acanthostyle; D, detail of surface with tylotes scattered in disorder. Scales: A = 500 µm; B D = 200 µm." httpUri="https://zenodo.org/record/266126/files/figure.png" pageId="13" pageNumber="82">Figs 11</figureCitation>
AB). Larger acanthostyles core main paucispicular tracts (approx. five spicules across) that are entirely echinated by another two distinct
<typeStatus id="54FD880AFFC0FFB0444EFB4EFA22762A" box="[1346,1405,1267,1290]" pageId="13" pageNumber="82">types</typeStatus>
of acanthostyles (
<figureCitation id="137D2A2DFFC0FFB0404DFAABFECB760F" box="[321,404,1302,1327]" captionStart="FIGURE 11" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[188,1398,1251,1856]" captionTargetId="figure@14.[188,1398,1251,1856]" captionTargetPageId="14" captionText="FIGURE 11. Lissodendoryx (Ectyodoryx) diegoramirezensis sp. nov., skeleton. A D, holotype (IZUA POR 169): A, overview of skeletal architecture; B, choanosomal reticulation in detail; C, section of a main tract echinated by smaller acanthostyles, i = larger acanthostyle, ii = intermediate acanthostyle, iii = smaller acanthostyle; D, detail of surface with tylotes scattered in disorder. Scales: A = 500 µm; B D = 200 µm." httpUri="https://zenodo.org/record/266126/files/figure.png" pageId="13" pageNumber="82">Fig. 11</figureCitation>
C). Some of the larger acanthostyles fan out from the main tracts, interconnecting them. Aniso-subtylotes are scattered at the surface, disposed at various angles to it (
<figureCitation id="137D2A2DFFC0FFB0450AFA87FB097673" box="[1030,1110,1338,1363]" captionStart="FIGURE 11" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[188,1398,1251,1856]" captionTargetId="figure@14.[188,1398,1251,1856]" captionTargetPageId="14" captionText="FIGURE 11. Lissodendoryx (Ectyodoryx) diegoramirezensis sp. nov., skeleton. A D, holotype (IZUA POR 169): A, overview of skeletal architecture; B, choanosomal reticulation in detail; C, section of a main tract echinated by smaller acanthostyles, i = larger acanthostyle, ii = intermediate acanthostyle, iii = smaller acanthostyle; D, detail of surface with tylotes scattered in disorder. Scales: A = 500 µm; B D = 200 µm." httpUri="https://zenodo.org/record/266126/files/figure.png" pageId="13" pageNumber="82">Fig. 11</figureCitation>
D). Isochelae and sigmas are scattered in the ectosome and choanosome. There is no spongin enveloping the fibres and subectosomal lacunae are absent. Choanosomal cavities (ca. 50 µm in diameter) are uncommon, scattered and roundish.
</paragraph>
<paragraph id="8BF936A8FFC0FFB041CBFA1BFC95746B" blockId="13.[151,1437,726,2011]" pageId="13" pageNumber="82">
<emphasis id="B932EABAFFC0FFB041CBFA1BFE6F769F" bold="true" box="[199,304,1446,1471]" pageId="13" pageNumber="82">Spicules.</emphasis>
Megascleres (
<tableCitation id="C6C40313FFC0FFB040DAFA1BFD7C769F" box="[470,547,1446,1471]" captionStart="TABLE 2" captionStartId="5.[151,242,152,175]" captionTargetBox="[160,1427,266,1950]" captionTargetPageId="5" captionText="TABLE 2. Comparative table for the spicule micrometries (µm), distributions and depth of occurrence of Lissodendoryx (Ectyodoryx) Lundbeck, 1909 known from the Antarctic, the Subantarctic region, the SW Atlantic and SE Pacific; * (absent),? (no information)." httpUri="http://table.plazi.org/id/DF396620FFC8FFB8419BFF25FE9A73CF" pageId="13" pageNumber="82" tableUuid="DF396620FFC8FFB8419BFF25FE9A73CF">Tabs 2</tableCitation>
4): Aniso-subtylotes to aniso-strongyles (
<figureCitation id="137D2A2DFFC0FFB04504FA1BFB01769F" box="[1032,1118,1446,1471]" captionStart="FIGURE 12" captionStartId="15.[151,250,1123,1145]" captionTargetBox="[188,1398,193,1100]" captionTargetId="figure@15.[188,1398,193,1101]" captionTargetPageId="15" captionText="FIGURE 12. Lissodendoryx (Ectyodoryx) diegoramirezensis sp. nov., electron micrographs. A K, holotype (IZUA POR 169): A, aniso-subtylotes; B, extremities of the aniso-subtylotes; C, acanthostyles I; D, bases of acanthostyles I; E, acanthostyles II; F, bases of acanthostyles II; G, acanthostyles III; H, bases of acanthostyles III, and sigma II; I, arcuate isochelae; J, sigmas I; K, sigmas II. Scales: A, C, E, G = 100 µm; B = 5 µm; D, F, H I, K = 10 µm; J = 20 µm." httpUri="https://zenodo.org/record/266127/files/figure.png" pageId="13" pageNumber="82">Figs 12</figureCitation>
AB), straight, microspined
<taxonomicName id="4C464D2BFFC0FFB0419BFA77FF9276C3" box="[151,205,1482,1507]" class="Demospongiae" family="Coelosphaeridae" genus="Ectyodoryx" higherTaxonomySource="CoL" kingdom="Animalia" order="Poecilosclerida" pageId="13" pageNumber="82" phylum="Porifera" rank="variety" species="nobilis" variety="ends">ends</taxonomicName>
, 223
<emphasis id="B932EABAFFC0FFB0401FFA76FE0D76C3" box="[275,338,1483,1507]" italics="true" pageId="13" pageNumber="82">233.5</emphasis>
(10.2) 252/7.2
<emphasis id="B932EABAFFC0FFB04315FA76FD6376C3" box="[537,572,1483,1507]" italics="true" pageId="13" pageNumber="82">8.4</emphasis>
(1.3) 9.6. Acanthostyles I (
<figureCitation id="137D2A2DFFC0FFB04285FA77FC8076C3" box="[905,991,1482,1507]" captionStart="FIGURE 12" captionStartId="15.[151,250,1123,1145]" captionTargetBox="[188,1398,193,1100]" captionTargetId="figure@15.[188,1398,193,1101]" captionTargetPageId="15" captionText="FIGURE 12. Lissodendoryx (Ectyodoryx) diegoramirezensis sp. nov., electron micrographs. A K, holotype (IZUA POR 169): A, aniso-subtylotes; B, extremities of the aniso-subtylotes; C, acanthostyles I; D, bases of acanthostyles I; E, acanthostyles II; F, bases of acanthostyles II; G, acanthostyles III; H, bases of acanthostyles III, and sigma II; I, arcuate isochelae; J, sigmas I; K, sigmas II. Scales: A, C, E, G = 100 µm; B = 5 µm; D, F, H I, K = 10 µm; J = 20 µm." httpUri="https://zenodo.org/record/266127/files/figure.png" pageId="13" pageNumber="82">Figs 12</figureCitation>
CD), straight to slightly curved, stout and somewhat fusiform; roundish base, frequently narrower than the central portion of the shaft; gradually thinning point; surface with few straight spines (up to 2.5 µm), mainly over the basal third of the spicule, 435
<emphasis id="B932EABAFFC0FFB0441DF9AEFA64750B" box="[1297,1339,1555,1579]" italics="true" pageId="13" pageNumber="82">536</emphasis>
(52.8) 602/24
<emphasis id="B932EABAFFC0FFB041FFF98AFE7B756F" box="[243,292,1591,1615]" italics="true" pageId="13" pageNumber="82">26.6</emphasis>
(2.4) 31. Acanthostyles II (
<figureCitation id="137D2A2DFFC0FFB04379F98BFD94756F" box="[629,715,1590,1615]" captionStart="FIGURE 12" captionStartId="15.[151,250,1123,1145]" captionTargetBox="[188,1398,193,1100]" captionTargetId="figure@15.[188,1398,193,1101]" captionTargetPageId="15" captionText="FIGURE 12. Lissodendoryx (Ectyodoryx) diegoramirezensis sp. nov., electron micrographs. A K, holotype (IZUA POR 169): A, aniso-subtylotes; B, extremities of the aniso-subtylotes; C, acanthostyles I; D, bases of acanthostyles I; E, acanthostyles II; F, bases of acanthostyles II; G, acanthostyles III; H, bases of acanthostyles III, and sigma II; I, arcuate isochelae; J, sigmas I; K, sigmas II. Scales: A, C, E, G = 100 µm; B = 5 µm; D, F, H I, K = 10 µm; J = 20 µm." httpUri="https://zenodo.org/record/266127/files/figure.png" pageId="13" pageNumber="82">Figs 12</figureCitation>
EF), same overall morphology as the preceding category, but smaller, and bearing more spines, although the apical portion is also totally smooth, 320
<emphasis id="B932EABAFFC0FFB04579F9E6FBC07553" box="[1141,1183,1627,1651]" italics="true" pageId="13" pageNumber="82">391</emphasis>
(29.2) 415/19
<emphasis id="B932EABAFFC0FFB04455F9E6FA2A7553" box="[1369,1397,1627,1651]" italics="true" pageId="13" pageNumber="82">20</emphasis>
(1) 21. Acanthostyles III (
<figureCitation id="137D2A2DFFC0FFB040ADF9C3FEAE75B7" box="[417,497,1662,1687]" captionStart="FIGURE 12" captionStartId="15.[151,250,1123,1145]" captionTargetBox="[188,1398,193,1100]" captionTargetId="figure@15.[188,1398,193,1101]" captionTargetPageId="15" captionText="FIGURE 12. Lissodendoryx (Ectyodoryx) diegoramirezensis sp. nov., electron micrographs. A K, holotype (IZUA POR 169): A, aniso-subtylotes; B, extremities of the aniso-subtylotes; C, acanthostyles I; D, bases of acanthostyles I; E, acanthostyles II; F, bases of acanthostyles II; G, acanthostyles III; H, bases of acanthostyles III, and sigma II; I, arcuate isochelae; J, sigmas I; K, sigmas II. Scales: A, C, E, G = 100 µm; B = 5 µm; D, F, H I, K = 10 µm; J = 20 µm." httpUri="https://zenodo.org/record/266127/files/figure.png" pageId="13" pageNumber="82">Fig. 12</figureCitation>
H) straight to slightly curved; narrow roundish base; gradually thinning point; shaft completely covered with spines (moderately to abundantly), spines frequently straight and usually larger than those of the preceding categories, 220
<emphasis id="B932EABAFFC0FFB04315F97AFD0775FF" box="[537,600,1735,1759]" italics="true" pageId="13" pageNumber="82">232.5</emphasis>
(10) 242/14
<emphasis id="B932EABAFFC0FFB04209F97AFC7E75FF" box="[773,801,1735,1759]" italics="true" pageId="13" pageNumber="82">15</emphasis>
(1.1) 17. Microscleres (
<tableCitation id="C6C40313FFC0FFB04546F97BFBC675FF" box="[1098,1177,1734,1759]" captionStart="TABLE 2" captionStartId="5.[151,242,152,175]" captionTargetBox="[160,1427,266,1950]" captionTargetPageId="5" captionText="TABLE 2. Comparative table for the spicule micrometries (µm), distributions and depth of occurrence of Lissodendoryx (Ectyodoryx) Lundbeck, 1909 known from the Antarctic, the Subantarctic region, the SW Atlantic and SE Pacific; * (absent),? (no information)." httpUri="http://table.plazi.org/id/DF396620FFC8FFB8419BFF25FE9A73CF" pageId="13" pageNumber="82" tableUuid="DF396620FFC8FFB8419BFF25FE9A73CF">Tabs 2</tableCitation>
4): Arcuate isochelae (
<figureCitation id="137D2A2DFFC0FFB04193F957FFAD7423" box="[159,242,1770,1795]" captionStart="FIGURE 12" captionStartId="15.[151,250,1123,1145]" captionTargetBox="[188,1398,193,1100]" captionTargetId="figure@15.[188,1398,193,1101]" captionTargetPageId="15" captionText="FIGURE 12. Lissodendoryx (Ectyodoryx) diegoramirezensis sp. nov., electron micrographs. A K, holotype (IZUA POR 169): A, aniso-subtylotes; B, extremities of the aniso-subtylotes; C, acanthostyles I; D, bases of acanthostyles I; E, acanthostyles II; F, bases of acanthostyles II; G, acanthostyles III; H, bases of acanthostyles III, and sigma II; I, arcuate isochelae; J, sigmas I; K, sigmas II. Scales: A, C, E, G = 100 µm; B = 5 µm; D, F, H I, K = 10 µm; J = 20 µm." httpUri="https://zenodo.org/record/266127/files/figure.png" pageId="13" pageNumber="82">
Fig.
<date id="FFF81068FFC0FFB041D9F957FFAD7423" box="[213,242,1770,1795]" pageId="13" pageNumber="82" value="2012-01">12</date>
</figureCitation>
I), shaft gently curved, lateral alae elongated and curved towards the shaft as a claw, frontal alae simple, 22
<emphasis id="B932EABAFFC0FFB041CDF8B2FFAD7407" box="[193,242,1807,1831]" italics="true" pageId="13" pageNumber="82">23.5</emphasis>
(1) 24. Sigmas I (
<figureCitation id="137D2A2DFFC0FFB040C1F8B3FD407407" box="[461,543,1806,1831]" captionStart="FIGURE 12" captionStartId="15.[151,250,1123,1145]" captionTargetBox="[188,1398,193,1100]" captionTargetId="figure@15.[188,1398,193,1101]" captionTargetPageId="15" captionText="FIGURE 12. Lissodendoryx (Ectyodoryx) diegoramirezensis sp. nov., electron micrographs. A K, holotype (IZUA POR 169): A, aniso-subtylotes; B, extremities of the aniso-subtylotes; C, acanthostyles I; D, bases of acanthostyles I; E, acanthostyles II; F, bases of acanthostyles II; G, acanthostyles III; H, bases of acanthostyles III, and sigma II; I, arcuate isochelae; J, sigmas I; K, sigmas II. Scales: A, C, E, G = 100 µm; B = 5 µm; D, F, H I, K = 10 µm; J = 20 µm." httpUri="https://zenodo.org/record/266127/files/figure.png" pageId="13" pageNumber="82">Fig. 12</figureCitation>
J), contorted, smooth, with sharp
<taxonomicName id="4C464D2BFFC0FFB0429BF8B3FC927407" box="[919,973,1806,1831]" class="Demospongiae" family="Coelosphaeridae" genus="Ectyodoryx" higherTaxonomySource="CoL" kingdom="Animalia" order="Poecilosclerida" pageId="13" pageNumber="82" phylum="Porifera" rank="variety" species="nobilis" variety="ends">ends</taxonomicName>
, 45
<emphasis id="B932EABAFFC0FFB0450FF8B2FB6B7406" box="[1027,1076,1807,1830]" italics="true" pageId="13" pageNumber="82">55.5</emphasis>
(4.9) 60. Sigmas II (
<figureCitation id="137D2A2DFFC0FFB04422F8B3FA227407" box="[1326,1405,1806,1831]" captionStart="FIGURE 12" captionStartId="15.[151,250,1123,1145]" captionTargetBox="[188,1398,193,1100]" captionTargetId="figure@15.[188,1398,193,1101]" captionTargetPageId="15" captionText="FIGURE 12. Lissodendoryx (Ectyodoryx) diegoramirezensis sp. nov., electron micrographs. A K, holotype (IZUA POR 169): A, aniso-subtylotes; B, extremities of the aniso-subtylotes; C, acanthostyles I; D, bases of acanthostyles I; E, acanthostyles II; F, bases of acanthostyles II; G, acanthostyles III; H, bases of acanthostyles III, and sigma II; I, arcuate isochelae; J, sigmas I; K, sigmas II. Scales: A, C, E, G = 100 µm; B = 5 µm; D, F, H I, K = 10 µm; J = 20 µm." httpUri="https://zenodo.org/record/266127/files/figure.png" pageId="13" pageNumber="82">Fig. 12</figureCitation>
K) same morphology as the preceding one, but smaller, 16.8
<emphasis id="B932EABAFFC0FFB04228F88EFC0A746B" box="[804,853,1843,1867]" italics="true" pageId="13" pageNumber="82">20.8</emphasis>
(2.9) 25.
</paragraph>
<paragraph id="8BF936A8FFC0FFB041CBF8EAFD9C744F" blockId="13.[151,1437,726,2011]" box="[199,707,1878,1903]" pageId="13" pageNumber="82">
<emphasis id="B932EABAFFC0FFB041CBF8EAFE73744E" bold="true" box="[199,300,1879,1902]" pageId="13" pageNumber="82">Ecology.</emphasis>
Deep-water habitat (
<quantity id="4CBE9B4DFFC0FFB0431BF8EBFD32744E" box="[535,621,1878,1903]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="82" unit="m" value="2000.0">2000 m</quantity>
depth).
</paragraph>
</subSubSection>
<subSubSection id="C35C6523FFC0FFB041CBF8C7FC0274B3" box="[199,861,1914,1939]" pageId="13" pageNumber="82" type="distribution">
<paragraph id="8BF936A8FFC0FFB041CBF8C7FC0274B3" blockId="13.[151,1437,726,2011]" box="[199,861,1914,1939]" pageId="13" pageNumber="82">
<emphasis id="B932EABAFFC0FFB041CBF8C7FE3F74B3" bold="true" box="[199,352,1914,1939]" pageId="13" pageNumber="82">Distribution.</emphasis>
Currently only known from the
<typeStatus id="54FD880AFFC0FFB043C0F8C6FDA374B2" box="[716,764,1915,1938]" pageId="13" pageNumber="82">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C35C6523FFC0FFB041CBF822FC687497" box="[199,823,1950,1975]" pageId="13" pageNumber="82" type="etymology">
<paragraph id="8BF936A8FFC0FFB041CBF822FC687497" blockId="13.[151,1437,726,2011]" box="[199,823,1950,1975]" pageId="13" pageNumber="82">
<emphasis id="B932EABAFFC0FFB041CBF822FE117496" bold="true" box="[199,334,1951,1974]" pageId="13" pageNumber="82">Etymology.</emphasis>
The species is named after its
<typeStatus id="54FD880AFFC0FFB043AAF822FD897496" box="[678,726,1951,1974]" pageId="13" pageNumber="82">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C35C6523FFC0FFB241CBF87EFA2574CA" lastPageId="15" lastPageNumber="84" pageId="13" pageNumber="82" type="discussion">
<paragraph id="8BF936A8FFC0FFB041CBF87EFAC374FB" blockId="13.[151,1437,726,2011]" box="[199,1436,1986,2011]" pageId="13" pageNumber="82">
<emphasis id="B932EABAFFC0FFB041CBF87EFE6474FA" bold="true" box="[199,315,1987,2010]" pageId="13" pageNumber="82">Remarks.</emphasis>
<taxonomicName id="4C464D2BFFC0FFB04048F87FFC0674FB" box="[324,857,1986,2011]" class="Demospongiae" family="Coelosphaeridae" genus="Lissodendoryx" kingdom="Animalia" order="Poecilosclerida" pageId="13" pageNumber="82" phylum="Porifera" rank="species" species="diegoramirezensis" status="sp. nov." subGenus="Ectyodoryx">
<emphasis id="B932EABAFFC0FFB04048F87FFEB774FB" box="[324,488,1986,2011]" italics="true" pageId="13" pageNumber="82">Lissodendoryx</emphasis>
(
<emphasis id="B932EABAFFC0FFB040F7F87FFD2574FB" box="[507,634,1986,2011]" italics="true" pageId="13" pageNumber="82">Ectyodoryx</emphasis>
)
<emphasis id="B932EABAFFC0FFB04381F87FFC0674FB" box="[653,857,1986,2011]" italics="true" pageId="13" pageNumber="82">diegoramirezensis</emphasis>
</taxonomicName>
<emphasis id="B932EABAFFC0FFB0426FF87EFCE074FA" bold="true" box="[867,959,1987,2010]" pageId="13" pageNumber="82">
<taxonomicNameLabel id="A20157C1FFC0FFB0426FF87EFCE074FA" box="[867,959,1987,2010]" pageId="13" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is distinguished from
<taxonomicName id="4C464D2BFFC0FFB045C8F87FFA3874FB" box="[1220,1383,1986,2011]" class="Demospongiae" family="Coelosphaeridae" genus="Lissodendoryx" kingdom="Animalia" order="Poecilosclerida" pageId="13" pageNumber="82" phylum="Porifera" rank="genus">
<emphasis id="B932EABAFFC0FFB045C8F87FFA3874FB" box="[1220,1383,1986,2011]" italics="true" pageId="13" pageNumber="82">Lissodendoryx</emphasis>
</taxonomicName>
(
<emphasis id="B932EABAFFC0FFB04476F87EFACC74FA" box="[1402,1427,1987,2010]" italics="true" pageId="13" pageNumber="82">E.</emphasis>
)
</paragraph>
<paragraph id="8BF936A8FFC3FFB3419BFF25FFA7723D" blockId="14.[151,1436,151,285]" pageId="14" pageNumber="83">
spp. occurring in the SE Pacific, additional allied biogeographic provinces, as well as
<emphasis id="B932EABAFFC3FFB34560FF25FBDC738F" box="[1132,1155,152,175]" italics="true" pageId="14" pageNumber="83">L.</emphasis>
(
<emphasis id="B932EABAFFC3FFB34598FF25FBFA738F" box="[1172,1189,152,175]" italics="true" pageId="14" pageNumber="83">E</emphasis>
.)
<taxonomicName id="4C464D2BFFC3FFB345B2FF2AFA4D7390" box="[1214,1298,151,176]" class="Demospongiae" family="Coelosphaeridae" genus="Lissodendoryx" kingdom="Animalia" order="Poecilosclerida" pageId="14" pageNumber="83" phylum="Porifera" rank="species" species="ballena" status="sp. nov.">
<emphasis id="B932EABAFFC3FFB345B2FF2AFA4D7390" box="[1214,1298,151,176]" italics="true" pageId="14" pageNumber="83">ballena</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20157C1FFC3FFB34417FF25FA29738F" box="[1307,1398,152,175]" pageId="14" pageNumber="83" rank="species">
<emphasis id="B932EABAFFC3FFB34417FF25FA29738F" bold="true" box="[1307,1398,152,175]" pageId="14" pageNumber="83">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<emphasis id="B932EABAFFC3FFB34489FF25FAC3738F" box="[1413,1436,152,175]" italics="true" pageId="14" pageNumber="83">L.</emphasis>
(
<emphasis id="B932EABAFFC3FFB341ACFF00FFEE73F4" box="[160,177,189,212]" italics="true" pageId="14" pageNumber="83">E</emphasis>
.)
<taxonomicName id="4C464D2BFFC3FFB341DCFF00FE1F73F4" box="[208,320,189,212]" class="Demospongiae" family="Coelosphaeridae" genus="Lissodendoryx" kingdom="Animalia" order="Poecilosclerida" pageId="14" pageNumber="83" phylum="Porifera" rank="species" species="corrugata" status="sp. nov.">
<emphasis id="B932EABAFFC3FFB341DCFF00FE1F73F4" box="[208,320,189,212]" italics="true" pageId="14" pageNumber="83">corrugata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20157C1FFC3FFB34042FF00FEEF73F4" box="[334,432,189,212]" pageId="14" pageNumber="83" rank="species">
<emphasis id="B932EABAFFC3FFB34042FF00FEEF73F4" bold="true" box="[334,432,189,212]" pageId="14" pageNumber="83">sp. nov.</emphasis>
</taxonomicNameLabel>
, and
<emphasis id="B932EABAFFC3FFB340F0FF00FD4C73F4" box="[508,531,189,212]" italics="true" pageId="14" pageNumber="83">L.</emphasis>
(
<emphasis id="B932EABAFFC3FFB34326FF00FD6473F4" box="[554,571,189,212]" italics="true" pageId="14" pageNumber="83">E</emphasis>
.)
<taxonomicName id="4C464D2BFFC3FFB34356FF01FD8173F5" box="[602,734,188,213]" class="Demospongiae" family="Coelosphaeridae" genus="Lissodendoryx" kingdom="Animalia" order="Poecilosclerida" pageId="14" pageNumber="83" phylum="Porifera" rank="species" species="coloanensis" status="sp. nov.">
<emphasis id="B932EABAFFC3FFB34356FF01FD8173F5" box="[602,734,188,213]" italics="true" pageId="14" pageNumber="83">coloanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20157C1FFC3FFB343E0FF00FC1273F4" box="[748,845,189,212]" pageId="14" pageNumber="83" rank="species">
<emphasis id="B932EABAFFC3FFB343E0FF00FC1273F4" bold="true" box="[748,845,189,212]" pageId="14" pageNumber="83">sp. nov.</emphasis>
</taxonomicNameLabel>
, mainly by its possession of three categories of acanthostyles, separable by their dimensions, morphology and placement in the skeletal architecture (
<tableCitation id="C6C40313FFC3FFB34418FF62FA0173D8" box="[1300,1374,223,248]" captionStart="TABLE 2" captionStartId="5.[151,242,152,175]" captionTargetBox="[160,1427,266,1950]" captionTargetPageId="5" captionText="TABLE 2. Comparative table for the spicule micrometries (µm), distributions and depth of occurrence of Lissodendoryx (Ectyodoryx) Lundbeck, 1909 known from the Antarctic, the Subantarctic region, the SW Atlantic and SE Pacific; * (absent),? (no information)." httpUri="http://table.plazi.org/id/DF396620FFC8FFB8419BFF25FE9A73CF" pageId="14" pageNumber="83" tableUuid="DF396620FFC8FFB8419BFF25FE9A73CF">Tab. 2</tableCitation>
;
<figureCitation id="137D2A2DFFC3FFB34460FF62FFED723D" captionStart="FIGURE 12" captionStartId="15.[151,250,1123,1145]" captionTargetBox="[188,1398,193,1100]" captionTargetId="figure@15.[188,1398,193,1101]" captionTargetPageId="15" captionText="FIGURE 12. Lissodendoryx (Ectyodoryx) diegoramirezensis sp. nov., electron micrographs. A K, holotype (IZUA POR 169): A, aniso-subtylotes; B, extremities of the aniso-subtylotes; C, acanthostyles I; D, bases of acanthostyles I; E, acanthostyles II; F, bases of acanthostyles II; G, acanthostyles III; H, bases of acanthostyles III, and sigma II; I, arcuate isochelae; J, sigmas I; K, sigmas II. Scales: A, C, E, G = 100 µm; B = 5 µm; D, F, H I, K = 10 µm; J = 20 µm." httpUri="https://zenodo.org/record/266127/files/figure.png" pageId="14" pageNumber="83">Figs 12</figureCitation>
CH).
</paragraph>
<caption id="DF396620FFC3FFB3419BFBE3FE207791" httpUri="https://zenodo.org/record/266125/files/figure.png" pageId="14" pageNumber="83" targetBox="[188,1398,337,1096]" targetPageId="14">
<paragraph id="8BF936A8FFC3FFB3419BFBE3FE207791" blockId="14.[151,1436,1118,1201]" pageId="14" pageNumber="83">
<emphasis id="B932EABAFFC3FFB3419BFBE3FE407754" bold="true" box="[151,287,1118,1140]" pageId="14" pageNumber="83">FIGURE 10.</emphasis>
AC,
<taxonomicName id="4C464D2BFFC3FFB3406DFBE3FC617754" box="[353,830,1118,1140]" class="Demospongiae" family="Coelosphaeridae" genus="Lissodendoryx" kingdom="Animalia" order="Poecilosclerida" pageId="14" pageNumber="83" phylum="Porifera" rank="species" species="diegoramirezensis" status="sp. nov." subGenus="Ectyodoryx">
<emphasis id="B932EABAFFC3FFB3406DFBE3FEAA7754" box="[353,501,1118,1140]" italics="true" pageId="14" pageNumber="83">Lissodendoryx</emphasis>
(
<emphasis id="B932EABAFFC3FFB34308FBE3FD287754" box="[516,631,1118,1140]" italics="true" pageId="14" pageNumber="83">Ectyodoryx</emphasis>
)
<emphasis id="B932EABAFFC3FFB34389FBE3FC617754" box="[645,830,1118,1140]" italics="true" pageId="14" pageNumber="83">diegoramirezensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20157C1FFC3FFB34248FBE2FCCA7754" box="[836,917,1119,1140]" pageId="14" pageNumber="83" rank="species">
<emphasis id="B932EABAFFC3FFB34248FBE2FCCA7754" bold="true" box="[836,917,1119,1140]" pageId="14" pageNumber="83">sp. nov.</emphasis>
</taxonomicNameLabel>
, habit. AC, holotype (IZUAPOR 169). A, whole specimen (dry); B, markedly hispid surface in detail; C, surface under transmitted light showing reticulated pattern. Scales: A = 0.5 cm; BC = 0.1 cm.
</paragraph>
</caption>
<caption id="DF396620FFC3FFB3419BF8E8FDB874E8" httpUri="https://zenodo.org/record/266126/files/figure.png" pageId="14" pageNumber="83" targetBox="[188,1398,1251,1856]" targetPageId="14">
<paragraph id="8BF936A8FFC3FFB3419BF8E8FDB874E8" blockId="14.[151,1436,1877,1992]" pageId="14" pageNumber="83">
<emphasis id="B932EABAFFC3FFB3419BF8E8FE79744B" bold="true" box="[151,294,1877,1899]" pageId="14" pageNumber="83">FIGURE 11.</emphasis>
<taxonomicName id="4C464D2BFFC3FFB3403FF8E8FC7F744B" box="[307,800,1877,1899]" class="Demospongiae" family="Coelosphaeridae" genus="Lissodendoryx" kingdom="Animalia" order="Poecilosclerida" pageId="14" pageNumber="83" phylum="Porifera" rank="species" species="diegoramirezensis" status="sp. nov." subGenus="Ectyodoryx">
<emphasis id="B932EABAFFC3FFB3403FF8E8FE98744B" box="[307,455,1877,1899]" italics="true" pageId="14" pageNumber="83">Lissodendoryx</emphasis>
(
<emphasis id="B932EABAFFC3FFB340D1F8E8FD0F744B" box="[477,592,1877,1899]" italics="true" pageId="14" pageNumber="83">Ectyodoryx</emphasis>
)
<emphasis id="B932EABAFFC3FFB3436BF8E8FC7F744B" box="[615,800,1877,1899]" italics="true" pageId="14" pageNumber="83">diegoramirezensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20157C1FFC3FFB34222F8EBFCD9744B" box="[814,902,1878,1899]" pageId="14" pageNumber="83" rank="species">
<emphasis id="B932EABAFFC3FFB34222F8EBFCD9744B" bold="true" box="[814,902,1878,1899]" pageId="14" pageNumber="83">sp. nov.</emphasis>
</taxonomicNameLabel>
, skeleton. AD, holotype (IZUAPOR 169): A, overview of skeletal architecture; B, choanosomal reticulation in detail; C, section of a main tract echinated by smaller acanthostyles, i = larger acanthostyle, ii = intermediate acanthostyle, iii = smaller acanthostyle; D, detail of surface with tylotes scattered in disorder. Scales: A = 500 µm; BD = 200 µm.
</paragraph>
</caption>
<caption id="DF396620FFC2FFB2419BFBDEFBAE77F4" httpUri="https://zenodo.org/record/266127/files/figure.png" pageId="15" pageNumber="84" targetBox="[188,1398,193,1100]" targetPageId="15">
<paragraph id="8BF936A8FFC2FFB2419BFBDEFBAE77F4" blockId="15.[151,1436,1123,1236]" pageId="15" pageNumber="84">
<emphasis id="B932EABAFFC2FFB2419BFBDEFE7B7759" bold="true" box="[151,292,1123,1145]" pageId="15" pageNumber="84">FIGURE 12.</emphasis>
<taxonomicName id="4C464D2BFFC2FFB24023FBDEFC4A7759" box="[303,789,1123,1145]" class="Demospongiae" family="Coelosphaeridae" genus="Lissodendoryx" kingdom="Animalia" order="Poecilosclerida" pageId="15" pageNumber="84" phylum="Porifera" rank="species" species="diegoramirezensis" status="sp. nov." subGenus="Ectyodoryx">
<emphasis id="B932EABAFFC2FFB24023FBDEFE9C7759" box="[303,451,1123,1145]" italics="true" pageId="15" pageNumber="84">Lissodendoryx</emphasis>
(
<emphasis id="B932EABAFFC2FFB240DAFBDEFD167759" box="[470,585,1123,1145]" italics="true" pageId="15" pageNumber="84">Ectyodoryx</emphasis>
)
<emphasis id="B932EABAFFC2FFB24350FBDEFC4A7759" box="[604,789,1123,1145]" italics="true" pageId="15" pageNumber="84">diegoramirezensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20157C1FFC2FFB2422CFBDEFC2A7758" box="[800,885,1123,1144]" pageId="15" pageNumber="84" rank="species">
<emphasis id="B932EABAFFC2FFB2422CFBDEFC2A7758" bold="true" box="[800,885,1123,1144]" pageId="15" pageNumber="84">sp. nov.</emphasis>
</taxonomicNameLabel>
, electron micrographs. AK, holotype (IZUAPOR 169): A, aniso-subtylotes; B, extremities of the aniso-subtylotes; C, acanthostyles I; D, bases of acanthostyles I; E, acanthostyles II; F, bases of acanthostyles II; G, acanthostyles III; H, bases of acanthostyles III, and sigma II; I, arcuate isochelae; J, sigmas I; K, sigmas II. Scales: A, C, E, G =100 µm; B = 5 µm; D, F, HI, K = 10 µm; J = 20 µm.
</paragraph>
</caption>
<paragraph id="8BF936A8FFC2FFB241CBFABCFFB6745E" blockId="15.[151,1437,1281,2026]" pageId="15" pageNumber="84">
<taxonomicName id="4C464D2BFFC2FFB241CBFABCFD88763A" box="[199,727,1281,1306]" class="Demospongiae" family="Coelosphaeridae" genus="Lissodendoryx" kingdom="Animalia" order="Poecilosclerida" pageId="15" pageNumber="84" phylum="Porifera" rank="species" species="diegoramirezensis" status="sp. nov." subGenus="Ectyodoryx">
<emphasis id="B932EABAFFC2FFB241CBFABCFE34763A" box="[199,363,1281,1306]" italics="true" pageId="15" pageNumber="84">Lissodendoryx</emphasis>
(
<emphasis id="B932EABAFFC2FFB24077FABCFEA5763A" box="[379,506,1281,1306]" italics="true" pageId="15" pageNumber="84">Ectyodoryx</emphasis>
)
<emphasis id="B932EABAFFC2FFB24306FABCFD88763A" box="[522,727,1281,1306]" italics="true" pageId="15" pageNumber="84">diegoramirezensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20157C1FFC2FFB243D1FABFFC687639" box="[733,823,1282,1305]" pageId="15" pageNumber="84" rank="species">
<emphasis id="B932EABAFFC2FFB243D1FABFFC687639" bold="true" box="[733,823,1282,1305]" pageId="15" pageNumber="84">sp. nov.</emphasis>
</taxonomicNameLabel>
has arcuate isochelae of uncommon morphology, with lateral alae of both extremities curved towards the shaft (
<figureCitation id="137D2A2DFFC2FFB2423EFA98FCD8761E" box="[818,903,1317,1342]" captionStart="FIGURE 12" captionStartId="15.[151,250,1123,1145]" captionTargetBox="[188,1398,193,1100]" captionTargetId="figure@15.[188,1398,193,1101]" captionTargetPageId="15" captionText="FIGURE 12. Lissodendoryx (Ectyodoryx) diegoramirezensis sp. nov., electron micrographs. A K, holotype (IZUA POR 169): A, aniso-subtylotes; B, extremities of the aniso-subtylotes; C, acanthostyles I; D, bases of acanthostyles I; E, acanthostyles II; F, bases of acanthostyles II; G, acanthostyles III; H, bases of acanthostyles III, and sigma II; I, arcuate isochelae; J, sigmas I; K, sigmas II. Scales: A, C, E, G = 100 µm; B = 5 µm; D, F, H I, K = 10 µm; J = 20 µm." httpUri="https://zenodo.org/record/266127/files/figure.png" pageId="15" pageNumber="84">
Fig.
<date id="FFF81068FFC2FFB24265FA98FCD8761E" box="[873,903,1317,1342]" pageId="15" pageNumber="84" value="2012-01">12</date>
</figureCitation>
I). Precisely this isochelae morphotype can be found in
<emphasis id="B932EABAFFC2FFB241F7FAF7FE4D7641" box="[251,274,1354,1377]" italics="true" pageId="15" pageNumber="84">L.</emphasis>
(
<emphasis id="B932EABAFFC2FFB2402DFAF7FE657641" box="[289,314,1354,1377]" italics="true" pageId="15" pageNumber="84">E.</emphasis>
)
<taxonomicName id="4C464D2BFFC2FFB24045FAF4FEE07642" box="[329,447,1353,1378]" class="Demospongiae" family="Coelosphaeridae" genus="Lissodendoryx" kingdom="Animalia" order="Poecilosclerida" pageId="15" pageNumber="84" phylum="Porifera" rank="species" species="anacantha">
<emphasis id="B932EABAFFC2FFB24045FAF4FEE07642" box="[329,447,1353,1378]" italics="true" pageId="15" pageNumber="84">anacantha</emphasis>
</taxonomicName>
(Hentschel, 1914: p.107, Taf. VII,
<figureCitation id="137D2A2DFFC2FFB2424AFAF4FCC97642" box="[838,918,1353,1378]" captionStart="FIGURE 12" captionStartId="15.[151,250,1123,1145]" captionTargetBox="[188,1398,193,1100]" captionTargetId="figure@15.[188,1398,193,1101]" captionTargetPageId="15" captionText="FIGURE 12. Lissodendoryx (Ectyodoryx) diegoramirezensis sp. nov., electron micrographs. A K, holotype (IZUA POR 169): A, aniso-subtylotes; B, extremities of the aniso-subtylotes; C, acanthostyles I; D, bases of acanthostyles I; E, acanthostyles II; F, bases of acanthostyles II; G, acanthostyles III; H, bases of acanthostyles III, and sigma II; I, arcuate isochelae; J, sigmas I; K, sigmas II. Scales: A, C, E, G = 100 µm; B = 5 µm; D, F, H I, K = 10 µm; J = 20 µm." httpUri="https://zenodo.org/record/266127/files/figure.png" pageId="15" pageNumber="84">Fig. 12</figureCitation>
), a species amply distributed around
<collectingCountry id="F3517638FFC2FFB24438FAF4FAC37642" box="[1332,1436,1353,1378]" name="Antarctica" pageId="15" pageNumber="84">Antarctic</collectingCountry>
and the Subantarctic Region, and in another
<collectingCountry id="F3517638FFC2FFB243AFFAD0FC5476A6" box="[675,779,1389,1414]" name="Antarctica" pageId="15" pageNumber="84">Antarctic</collectingCountry>
species,
<taxonomicName id="4C464D2BFFC2FFB24275FAD3FAC376A6" authority="Hentschel, 1914" authorityName="Hentschel" authorityYear="1914" box="[889,1436,1389,1414]" class="Demospongiae" family="Coelosphaeridae" genus="Lissodendoryx" kingdom="Animalia" order="Poecilosclerida" pageId="15" pageNumber="84" phylum="Porifera" rank="species" species="styloderma" subGenus="Lissodendoryx">
<emphasis id="B932EABAFFC2FFB24275FAD3FCCF76A5" box="[889,912,1390,1413]" italics="true" pageId="15" pageNumber="84">L.</emphasis>
(
<emphasis id="B932EABAFFC2FFB242A8FAD0FB1876A6" box="[932,1095,1389,1414]" italics="true" pageId="15" pageNumber="84">Lissodendoryx</emphasis>
)
<emphasis id="B932EABAFFC2FFB24557FAD0FB8776A6" box="[1115,1240,1389,1414]" italics="true" pageId="15" pageNumber="84">styloderma</emphasis>
Hentschel, 1914
</taxonomicName>
(Hentschel, 1914: p.101, Taf. VII,
<figureCitation id="137D2A2DFFC2FFB24310FA2CFD3F768A" box="[540,608,1425,1450]" captionStart="FIGURE 7" captionStartId="10.[151,250,1571,1593]" captionTargetBox="[188,1398,339,1549]" captionTargetId="figure@10.[188,1398,339,1549]" captionTargetPageId="10" captionText="FIGURE 7. Lissodendoryx (Ectyodoryx) corrugata sp. nov., electron micrographs of spicules. A H, holotype (IZUA POR 167): A, ectosomal tylotes; B, detail of extremities of tylotes; C, acanthostyles I; D, detail of base of acanthostyles I; E, acanthostyles II; F, detail of base of acanthostyles II; G, arcuate isochelae I; H, arcuate isochelae II. I P, paratype (MNRJ 17398): I, ectosomal tylotes; J, detail of extremities of the tylotes; K, acanthostyles I; L, detail of base of acanthostyles I; M, acanthostyles II; N, detail of base of acanthostyles II; O, arcuate isochelae I; P, arcuate isochelae II. Scales: A, C, E, I, K, M = 50 µm; B, J = 5 µm; D, F G, H, L, N P = 10 µm." httpUri="https://zenodo.org/record/266122/files/figure.png" pageId="15" pageNumber="84">Fig. 7</figureCitation>
). This latter species can be readily distinguished from the new species by its lack of sigmas, possession of styles in the ectosome, and acanthostyles of a single category only. However, the similarity of the isochelae of the new species and those seen in
<emphasis id="B932EABAFFC2FFB24262FA67FCDA76D1" box="[878,901,1498,1521]" italics="true" pageId="15" pageNumber="84">L.</emphasis>
(
<emphasis id="B932EABAFFC2FFB2429AFA67FCF076D1" box="[918,943,1498,1521]" italics="true" pageId="15" pageNumber="84">E.</emphasis>
)
<taxonomicName id="4C464D2BFFC2FFB242CCFA64FB6976D2" box="[960,1078,1497,1522]" class="Demospongiae" family="Coelosphaeridae" genus="Lissodendoryx" kingdom="Animalia" order="Poecilosclerida" pageId="15" pageNumber="84" phylum="Porifera" rank="species" species="anacantha">
<emphasis id="B932EABAFFC2FFB242CCFA64FB6976D2" box="[960,1078,1497,1522]" italics="true" pageId="15" pageNumber="84">anacantha</emphasis>
</taxonomicName>
might indicate a closer affinity between both species, as they further share similar habit and additional spicule characteristics. Nevertheless, both species are considered distinct on the basis of the presence of a third category of acanthostyles in the new species, while its acanthostyles, as a rule, are also more densely spined or with spines more widespread over the shaft. Furthermore,
<emphasis id="B932EABAFFC2FFB2403DF9D7FE1775A1" box="[305,328,1642,1665]" italics="true" pageId="15" pageNumber="84">L.</emphasis>
(
<emphasis id="B932EABAFFC2FFB2405BF9D7FE2F75A1" box="[343,368,1642,1665]" italics="true" pageId="15" pageNumber="84">E.</emphasis>
)
<taxonomicName id="4C464D2BFFC2FFB24073F9D4FEAA75A2" box="[383,501,1641,1666]" class="Demospongiae" family="Coelosphaeridae" genus="Lissodendoryx" kingdom="Animalia" order="Poecilosclerida" pageId="15" pageNumber="84" phylum="Porifera" rank="species" species="anacantha">
<emphasis id="B932EABAFFC2FFB24073F9D4FEAA75A2" box="[383,501,1641,1666]" italics="true" pageId="15" pageNumber="84">anacantha</emphasis>
</taxonomicName>
has ectosomal megascleres that can be perfect styles (
<figureCitation id="137D2A2DFFC2FFB24559F9D4FBFC75A2" box="[1109,1187,1641,1666]" captionStart="FIGURE 13" captionStartId="16.[151,250,1427,1449]" captionTargetBox="[188,1398,195,1405]" captionTargetId="figure@16.[188,1398,195,1405]" captionTargetPageId="16" captionText="FIGURE 13. Lissodendoryx (Ectyodoryx) anacantha (Hentschel, 1914), electron micrographs. A I, holotype (ZMH S 2327): A, aniso-subtylotes; B, extremities of aniso-subtylotes; C, acanthostyles I, and sigma I; D, bases of acanthostyles I; E, acanthostyles II, and sigma II; F, bases of acanthostyles II; G, arcuate isochelae; H, sigmas I; I, sigmas II. Scales: A, C, E = 100 µm; B, D, H = 10 µm; F G, I = 5 µm." httpUri="https://zenodo.org/record/266128/files/figure.png" pageId="15" pageNumber="84">Fig. 13</figureCitation>
CD), and the smaller category of acanthostyles may bear secondary microspines over the main spines seen at the base of the spicule (
<figureCitation id="137D2A2DFFC2FFB24463F930FFEE75EA" captionStart="FIGURE 13" captionStartId="16.[151,250,1427,1449]" captionTargetBox="[188,1398,195,1405]" captionTargetId="figure@16.[188,1398,195,1405]" captionTargetPageId="16" captionText="FIGURE 13. Lissodendoryx (Ectyodoryx) anacantha (Hentschel, 1914), electron micrographs. A I, holotype (ZMH S 2327): A, aniso-subtylotes; B, extremities of aniso-subtylotes; C, acanthostyles I, and sigma I; D, bases of acanthostyles I; E, acanthostyles II, and sigma II; F, bases of acanthostyles II; G, arcuate isochelae; H, sigmas I; I, sigmas II. Scales: A, C, E = 100 µm; B, D, H = 10 µm; F G, I = 5 µm." httpUri="https://zenodo.org/record/266128/files/figure.png" pageId="15" pageNumber="84">Fig. 13</figureCitation>
EF). These characteristics of the acanthostyles have not been spotted in the
<typeStatus id="54FD880AFFC2FFB245ABF90CFA5675EA" box="[1191,1289,1713,1738]" pageId="15" pageNumber="84" type="holotype">holotype</typeStatus>
of
<emphasis id="B932EABAFFC2FFB2444BF90FFA0175E9" box="[1351,1374,1714,1737]" italics="true" pageId="15" pageNumber="84">L.</emphasis>
(
<emphasis id="B932EABAFFC2FFB24476F90FFACC75E9" box="[1402,1427,1714,1737]" italics="true" pageId="15" pageNumber="84">E.</emphasis>
)
<taxonomicName id="4C464D2BFFC2FFB2419BF968FE3C75CE" box="[151,355,1749,1774]" class="Demospongiae" family="Coelosphaeridae" genus="Lissodendoryx" kingdom="Animalia" order="Poecilosclerida" pageId="15" pageNumber="84" phylum="Porifera" rank="species" species="diegoramirezensis" status="sp. nov.">
<emphasis id="B932EABAFFC2FFB2419BF968FE3C75CE" box="[151,355,1749,1774]" italics="true" pageId="15" pageNumber="84">diegoramirezensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20157C1FFC2FFB24060F96BFE9875CD" box="[364,455,1750,1773]" pageId="15" pageNumber="84" rank="species">
<emphasis id="B932EABAFFC2FFB24060F96BFE9875CD" bold="true" box="[364,455,1750,1773]" pageId="15" pageNumber="84">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="137D2A2DFFC2FFB240D4F968FD7875CE" box="[472,551,1749,1774]" captionStart="FIGURE 11" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[188,1398,1251,1856]" captionTargetId="figure@14.[188,1398,1251,1856]" captionTargetPageId="14" captionText="FIGURE 11. Lissodendoryx (Ectyodoryx) diegoramirezensis sp. nov., skeleton. A D, holotype (IZUA POR 169): A, overview of skeletal architecture; B, choanosomal reticulation in detail; C, section of a main tract echinated by smaller acanthostyles, i = larger acanthostyle, ii = intermediate acanthostyle, iii = smaller acanthostyle; D, detail of surface with tylotes scattered in disorder. Scales: A = 500 µm; B D = 200 µm." httpUri="https://zenodo.org/record/266126/files/figure.png" pageId="15" pageNumber="84">Fig. 11</figureCitation>
CH). The re-examination of the
<typeStatus id="54FD880AFFC2FFB242A5F96BFC8675CD" box="[937,985,1750,1773]" pageId="15" pageNumber="84">type</typeStatus>
material of
<emphasis id="B932EABAFFC2FFB2456AF96BFB2275CD" box="[1126,1149,1750,1773]" italics="true" pageId="15" pageNumber="84">L.</emphasis>
(
<emphasis id="B932EABAFFC2FFB24583F96BFBF775CD" box="[1167,1192,1750,1773]" italics="true" pageId="15" pageNumber="84">E.</emphasis>
)
<taxonomicName id="4C464D2BFFC2FFB245B5F968FA7075CE" box="[1209,1327,1749,1774]" class="Demospongiae" family="Coelosphaeridae" genus="Lissodendoryx" kingdom="Animalia" order="Poecilosclerida" pageId="15" pageNumber="84" phylum="Porifera" rank="species" species="anacantha">
<emphasis id="B932EABAFFC2FFB245B5F968FA7075CE" box="[1209,1327,1749,1774]" italics="true" pageId="15" pageNumber="84">anacantha</emphasis>
</taxonomicName>
(
<figureCitation id="137D2A2DFFC2FFB2444DF968FACD75CE" box="[1345,1426,1749,1774]" captionStart="FIGURE 13" captionStartId="16.[151,250,1427,1449]" captionTargetBox="[188,1398,195,1405]" captionTargetId="figure@16.[188,1398,195,1405]" captionTargetPageId="16" captionText="FIGURE 13. Lissodendoryx (Ectyodoryx) anacantha (Hentschel, 1914), electron micrographs. A I, holotype (ZMH S 2327): A, aniso-subtylotes; B, extremities of aniso-subtylotes; C, acanthostyles I, and sigma I; D, bases of acanthostyles I; E, acanthostyles II, and sigma II; F, bases of acanthostyles II; G, arcuate isochelae; H, sigmas I; I, sigmas II. Scales: A, C, E = 100 µm; B, D, H = 10 µm; F G, I = 5 µm." httpUri="https://zenodo.org/record/266128/files/figure.png" pageId="15" pageNumber="84">Fig. 13</figureCitation>
) confirmed the spicule set originally reported by Hentschel (1914), and subsequently recognized by Koltun (1964). Nevertheless, new measurements obtained from the
<typeStatus id="54FD880AFFC2FFB243FAF8A0FC077416" box="[758,856,1821,1846]" pageId="15" pageNumber="84" type="holotype">holotype</typeStatus>
(
<tableCitation id="C6C40313FFC2FFB24266F8A0FCEA7416" box="[874,949,1821,1846]" captionStart="TABLE 2" captionStartId="5.[151,242,152,175]" captionTargetBox="[160,1427,266,1950]" captionTargetPageId="5" captionText="TABLE 2. Comparative table for the spicule micrometries (µm), distributions and depth of occurrence of Lissodendoryx (Ectyodoryx) Lundbeck, 1909 known from the Antarctic, the Subantarctic region, the SW Atlantic and SE Pacific; * (absent),? (no information)." httpUri="http://table.plazi.org/id/DF396620FFC8FFB8419BFF25FE9A73CF" pageId="15" pageNumber="84" tableUuid="DF396620FFC8FFB8419BFF25FE9A73CF">Tab. 2</tableCitation>
) revealed that acanthostyles can be much thicker than reported originally (Hentschel, 1914), matching more closely the measurements reported by Koltun (1964).
</paragraph>
<paragraph id="8BF936A8FFC2FFB241CBF834FA2574CA" blockId="15.[151,1437,1281,2026]" pageId="15" pageNumber="84">
Given the data at hand,
<emphasis id="B932EABAFFC2FFB240D9F837FEB37481" box="[469,492,1930,1953]" italics="true" pageId="15" pageNumber="84">L.</emphasis>
(
<emphasis id="B932EABAFFC2FFB240F1F837FD497481" box="[509,534,1930,1953]" italics="true" pageId="15" pageNumber="84">E.</emphasis>
)
<taxonomicName id="4C464D2BFFC2FFB2432BF834FDAB7482" box="[551,756,1929,1954]" class="Demospongiae" family="Coelosphaeridae" genus="Lissodendoryx" kingdom="Animalia" order="Poecilosclerida" pageId="15" pageNumber="84" phylum="Porifera" rank="species" species="diegoramirezensis" status="sp. nov.">
<emphasis id="B932EABAFFC2FFB2432BF834FDAB7482" box="[551,756,1929,1954]" italics="true" pageId="15" pageNumber="84">diegoramirezensis</emphasis>
</taxonomicName>
<emphasis id="B932EABAFFC2FFB243F7F837FC097481" bold="true" box="[763,854,1930,1953]" pageId="15" pageNumber="84">
<taxonomicNameLabel id="A20157C1FFC2FFB243F7F837FC097481" box="[763,854,1930,1953]" pageId="15" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is considered well distinguished from allied forms, but it is suggested that study of additional samples of
<emphasis id="B932EABAFFC2FFB243E6F813FC5E74E5" box="[746,769,1966,1989]" italics="true" pageId="15" pageNumber="84">L.</emphasis>
(
<emphasis id="B932EABAFFC2FFB2421DF813FC7674E5" box="[785,809,1966,1989]" italics="true" pageId="15" pageNumber="84">E.</emphasis>
)
<taxonomicName id="4C464D2BFFC2FFB24234F810FCF174E6" box="[824,942,1965,1990]" class="Demospongiae" family="Coelosphaeridae" genus="Lissodendoryx" kingdom="Animalia" order="Poecilosclerida" pageId="15" pageNumber="84" phylum="Porifera" rank="species" species="anacantha">
<emphasis id="B932EABAFFC2FFB24234F810FCF174E6" box="[824,942,1965,1990]" italics="true" pageId="15" pageNumber="84">anacantha</emphasis>
</taxonomicName>
, such as those registered (but not described) by Burton (1932) from
<collectingCountry id="F3517638FFC2FFB24092F86CFD1D74CA" box="[414,578,2001,2026]" name="South Georgia and the South Sandwich Islands" pageId="15" pageNumber="84">South Georgia</collectingCountry>
, is important to verify if these two taxa are distinct, as hypothesised here.
</paragraph>
</subSubSection>
</treatment>
</document>