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<document id="02C3D99E61BFC1DEE8E5B813B74DC200" ID-CLB-Dataset="30294" ID-DOI="10.11646/zootaxa.4407.3.6" ID-GBIF-Dataset="d6a4aaf4-4045-4302-8220-413a7eca4f99" ID-ISSN="1175-5326" ID-Zenodo-Dep="1216616" ID-ZooBank="3FF82287-8A03-4FE6-803D-EC8C2A370B6D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1523435013489" checkinUser="plazi" docAuthor="Dubatolov, Vladimir V., Volynkin, Anton V. &amp; Kishida, Yasunori" docDate="2018" docId="03EF87BAFF9AFFFE9EA93F76FC02FBEE" docLanguage="en" docName="zootaxa.4407.3.6.pdf" docOrigin="Zootaxa 4407 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macohasa tetraspila Dubatolov &amp; Volynkin &amp; Kishida 2018, comb. nov." docType="treatment" docVersion="7" lastPageNumber="391" masterDocId="FFD6FFC2FF9DFFF69E3E382AFFEDFF92" masterDocTitle="Review of the Prabhasa Moore, 1878 genus-group (Lepidoptera, Erebidae, Arctiinae)" masterLastPageNumber="400" masterPageNumber="383" pageNumber="390" updateTime="1698536239552" updateUser="ExternalLinkService">
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<mods:title id="54CFEFC47C355D5E1ED49DF829635A3B">Review of the Prabhasa Moore, 1878 genus-group (Lepidoptera, Erebidae, Arctiinae)</mods:title>
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<mods:name id="DB700B6C3DDFD3D65C12B3E9C4159B8E" type="personal">
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<mods:namePart id="8DF7455662A428D6491A9259E49549C6">Dubatolov, Vladimir V.</mods:namePart>
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<mods:namePart id="1D225B6A83B6C1BDB09F0C61D7B8831A">Volynkin, Anton V.</mods:namePart>
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<mods:namePart id="6F9B5211356ED0E96734B7D3FEE33243">Kishida, Yasunori</mods:namePart>
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<mods:title id="A08E3D505B973C29FE5189C2B7351F29">Zootaxa</mods:title>
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<mods:part id="383E690F96E6B2A1728C1C19604E6D76">
<mods:date id="B83A4816D9E670CF59DC46292DB48659">2018</mods:date>
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<mods:number id="86EF4DCA40C124B949A0914EED63E122">2018-04-11</mods:number>
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<mods:number id="EF4EB61E47139F2166975B82EF01E3F6">4407</mods:number>
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<treatment id="03EF87BAFF9AFFFE9EA93F76FC02FBEE" ID-DOI="http://doi.org/10.5281/zenodo.5997304" ID-GBIF-Taxon="143536669" ID-Zenodo-Dep="5997304" LSID="urn:lsid:plazi:treatment:03EF87BAFF9AFFFE9EA93F76FC02FBEE" httpUri="http://treatment.plazi.org/id/03EF87BAFF9AFFFE9EA93F76FC02FBEE" lastPageId="8" lastPageNumber="391" pageId="7" pageNumber="390">
<subSubSection id="C35C6527FF9AFFF19EA93F76FE6FF805" pageId="7" pageNumber="390" type="nomenclature">
<paragraph id="8BF936ACFF9AFFF19EA93F76FD3EF8E4" blockId="7.[151,723,1884,1943]" box="[151,723,1884,1910]" pageId="7" pageNumber="390">
<heading id="D0B181C0FF9AFFF19EA93F76FD3EF8E4" bold="true" box="[151,723,1884,1910]" fontSize="11" level="1" pageId="7" pageNumber="390" reason="1">
<emphasis id="B932EABEFF9AFFF19EA93F76FD3EF8E4" bold="true" box="[151,723,1884,1910]" pageId="7" pageNumber="390">
<taxonomicName id="4C464D2FFF9AFFF19EA93F76FDD2F8E4" ID-CoL="BFSPL" authority="(Cerny, 2009)" authorityName="Dubatolov &amp; Volynkin &amp; Kishida" authorityYear="2018" baseAuthorityName="Cerny" baseAuthorityYear="2009" box="[151,575,1884,1910]" class="Insecta" family="Erebidae" genus="Macohasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="390" phylum="Arthropoda" rank="species" species="tetraspila" status="comb. nov.">
<emphasis id="B932EABEFF9AFFF19EA93F76FE61F8E4" bold="true" box="[151,396,1884,1910]" italics="true" pageId="7" pageNumber="390">Macohasa tetraspila</emphasis>
(Černý, 2009)
</taxonomicName>
,
<taxonomicNameLabel id="A20157C5FF9AFFF19C733F76FD3EF8E4" box="[589,723,1884,1910]" pageId="7" pageNumber="390">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF936ACFF9AFFF19EA93F54FE6FF805" blockId="7.[151,723,1884,1943]" box="[151,386,1918,1943]" pageId="7" pageNumber="390">
(
<figureCitation id="137D2A29FF9AFFF19EA13F54FECBF805" box="[159,294,1918,1943]" captionStart="FIGURES 1119" captionStartId="4.[151,264,1844,1866]" captionTargetBox="[200,1361,194,1789]" captionTargetId="figure@4.[191,1396,193,1823]" captionTargetPageId="4" captionText="FIGURES 1119. Macohasa and Chinasa spp.: adults. 11, M. orientalis, ♂, Malaysia, Malacca Peninsula (MWM/ZSM); 12, M. orientalis, ♀, Malaysia, Malacca Peninsula (MWM/ZSM); 13, M. tetraspila, holotype ♂, Thailand, Chantaburi prov. (MWM/ZSM); 14, M. tetraspila, paratype ♀, Thailand, Chantaburi prov. (MWM/ZSM); 15, M. dimorpha, syntype ♂, Philippines, Luzon (©NHMUK); 16, M. dimorpha, syntype ♀, Philippines, Luzon (©NHMUK); 17, Ch. costalis costalis, holotype ♂, N China (©NHMUK); 18, Ch. costalis costalis, ♂, China, Zhejiang (MWM/ZSM); 19, Ch. costalis costalis, ♀, China, Guangdong (SZMN)." httpUri="https://zenodo.org/record/1216620/files/figure.png" pageId="7" pageNumber="390">Figs. 1314</figureCitation>
,
<figureCitation id="137D2A29FF9AFFF19F0C3F55FEBCF805" box="[306,337,1919,1943]" captionStart="FIGURES 2729" captionStartId="12.[151,264,1953,1975]" captionTargetBox="[180,1380,198,1932]" captionTargetId="figure@12.[151,1436,193,1932]" captionTargetPageId="12" captionText="FIGURES 2729. Macohasa spp.: male genitalia (genital capsules in ventral vieW). 27, M. orientalis, Malaysia, slide MWM 31775 Volynkin; 28, M. tetraspila, holotype, Thailand, Chantaburi prov., slide MWM 33812 Volynkin; 29, M. dimorpha, Philippines, PalaWan, slide MWM 31777 Volynkin." httpUri="https://zenodo.org/record/1216626/files/figure.png" pageId="7" pageNumber="390">28</figureCitation>
,
<figureCitation id="137D2A29FF9AFFF19F633F55FE97F805" box="[349,378,1919,1943]" captionStart="FIGURES 3641" captionStartId="15.[151,264,1891,1913]" captionTargetBox="[169,1399,193,1854]" captionTargetId="figure@15.[164,1400,193,1865]" captionTargetPageId="15" captionText="FIGURES 3641. Lithosiina spp.: female genitalia (ventral vieW). 36, Prabhasa venosa, [India, Meghalaya] Khasis, slide BMNH(E) Arct-4557f HolloWay (©NHMUK); 37, Macohasa cernyi, paratype, N Vietnam, slide MWM 31774 Volynkin; 38, Macohasa orientalis, Malaysia, slide MWM 31776 Volynkin; 39, Macohasa tetraspila, paratype, Thailand, Chantaburi prov., slide AV4186m Volynkin; 40, Macohasa dimorpha, Philippines, PalaWan, slide MWM 31778 Volynkin; 41, Macotasa biplagella, slide BMNH(E) Arct-2849 (©NHMUK)." httpUri="https://zenodo.org/record/1216634/files/figure.png" pageId="7" pageNumber="390">39</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35C6527FF9AFFF19EA93FE8FD08F864" pageId="7" pageNumber="390" type="description">
<paragraph id="8BF936ACFF9AFFF19EA93FE8FD08F864" blockId="7.[151,1436,1985,2038]" pageId="7" pageNumber="390">
<taxonomicName id="4C464D2FFF9AFFF19EA93FE8FE05F845" authority="Cerny, 2009" authorityName="Cerny" authorityYear="2009" box="[151,488,1985,2007]" class="Insecta" family="Erebidae" genus="Macotasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="390" phylum="Arthropoda" rank="species" species="tetraspila">
<emphasis id="B932EABEFF9AFFF19EA93FE8FE8FF845" box="[151,354,1985,2007]" italics="true" pageId="7" pageNumber="390">Macotasa tetraspila</emphasis>
Černý, 2009
</taxonomicName>
,
<emphasis id="B932EABEFF9AFFF19FCC3FEBFD40F845" box="[498,685,1985,2007]" italics="true" pageId="7" pageNumber="390">
Moths of
<collectingCountry id="F351763CFF9AFFF19C6F3FEBFD40F845" box="[593,685,1985,2007]" name="Thailand" pageId="7" pageNumber="390">Thailand</collectingCountry>
</emphasis>
, 6: 139, pl. 28, figs. 278a, 278b (
<typeStatus id="54FD880EFF9AFFF19DC23FEBFBDDF845" box="[1020,1072,1985,2007]" pageId="7" pageNumber="390">Type</typeStatus>
locality: “SE
<collectingCountry id="F351763CFF9AFFF19A803FEBFAF6F845" box="[1214,1307,1985,2007]" name="Thailand" pageId="7" pageNumber="390">Thailand</collectingCountry>
,
<collectingRegion id="4982F84EFF9AFFF19B1B3FEBFA75F845" box="[1317,1432,1985,2007]" country="Thailand" name="Chanthaburi" pageId="7" pageNumber="390">Chantaburi</collectingRegion>
, Kaeng Hang MaeW Dist.,
<geoCoordinate id="EE72506BFF9AFFF19FF53FCAFDACF864" box="[459,577,2016,2038]" direction="north" orientation="latitude" pageId="7" pageNumber="390" precision="15" value="10.078055">10°441” N</geoCoordinate>
,
<geoCoordinate id="EE72506BFF9AFFF19C753FCAFD23F864" box="[587,718,2016,2038]" direction="east" orientation="longitude" pageId="7" pageNumber="390" precision="15" value="101.80139">101°485” E</geoCoordinate>
”).
</paragraph>
</subSubSection>
<subSubSection id="C35C6527FF95FFFE9EA938BDFDA6FED2" pageId="8" pageNumber="391" type="materials_examined">
<paragraph id="8BF936ACFF95FFFE9EA938BDFDA6FED2" blockId="8.[151,1437,151,1149]" pageId="8" pageNumber="391">
<materialsCitation id="3B2E3CF1FF95FFFE9EA938BDFA74FF46" ID-GBIF-Occurrence="1837804148" collectingDate="2005-06-02" collectingDateMax="2005-06-03" collectingDateMin="2005-06-02" collectionCode="ZSM" collectorName="K. Cerny &amp; Volynkin &amp; Coll." country="Thailand" latitude="10.078055" location="Kaeng Hang Maew Dist." longLatPrecision="21" longitude="101.80139" pageId="8" pageNumber="391" specimenCount="1" specimenCount-male="1" stateProvince="Chantaburi" typeStatus="holotype">
<emphasis id="B932EABEFF95FFFE9EA938BDFE29FF22" bold="true" box="[151,452,151,176]" pageId="8" pageNumber="391">
<typeStatus id="54FD880EFF95FFFE9EA938BDFF3FFF22" box="[151,210,151,176]" pageId="8" pageNumber="391">Type</typeStatus>
material examined
</emphasis>
.
<typeStatus id="54FD880EFF95FFFE9FE738BDFDABFF22" box="[473,582,151,176]" pageId="8" pageNumber="391" type="holotype">
<emphasis id="B932EABEFF95FFFE9FE738BDFDABFF22" bold="true" box="[473,582,151,176]" pageId="8" pageNumber="391">Holotype</emphasis>
</typeStatus>
:
<specimenCount id="9D40FD25FF95FFFE9C6538B2FD9DFF3D" box="[603,624,152,175]" pageId="8" pageNumber="391" type="male"></specimenCount>
(
<figureCitation id="137D2A29FF95FFFE9CBB38BDFD04FF22" box="[645,745,151,176]" captionStart="FIGURES 1119" captionStartId="4.[151,264,1844,1866]" captionTargetBox="[200,1361,194,1789]" captionTargetId="figure@4.[191,1396,193,1823]" captionTargetPageId="4" captionText="FIGURES 1119. Macohasa and Chinasa spp.: adults. 11, M. orientalis, ♂, Malaysia, Malacca Peninsula (MWM/ZSM); 12, M. orientalis, ♀, Malaysia, Malacca Peninsula (MWM/ZSM); 13, M. tetraspila, holotype ♂, Thailand, Chantaburi prov. (MWM/ZSM); 14, M. tetraspila, paratype ♀, Thailand, Chantaburi prov. (MWM/ZSM); 15, M. dimorpha, syntype ♂, Philippines, Luzon (©NHMUK); 16, M. dimorpha, syntype ♀, Philippines, Luzon (©NHMUK); 17, Ch. costalis costalis, holotype ♂, N China (©NHMUK); 18, Ch. costalis costalis, ♂, China, Zhejiang (MWM/ZSM); 19, Ch. costalis costalis, ♀, China, Guangdong (SZMN)." httpUri="https://zenodo.org/record/1216620/files/figure.png" pageId="8" pageNumber="391">Figs. 13</figureCitation>
,
<figureCitation id="137D2A29FF95FFFE9CC538B2FCF7FF22" box="[763,794,152,176]" captionStart="FIGURES 2729" captionStartId="12.[151,264,1953,1975]" captionTargetBox="[180,1380,198,1932]" captionTargetId="figure@12.[151,1436,193,1932]" captionTargetPageId="12" captionText="FIGURES 2729. Macohasa spp.: male genitalia (genital capsules in ventral vieW). 27, M. orientalis, Malaysia, slide MWM 31775 Volynkin; 28, M. tetraspila, holotype, Thailand, Chantaburi prov., slide MWM 33812 Volynkin; 29, M. dimorpha, Philippines, PalaWan, slide MWM 31777 Volynkin." httpUri="https://zenodo.org/record/1216626/files/figure.png" pageId="8" pageNumber="391">28</figureCitation>
), SE
<collectingCountry id="F351763CFF95FFFE9D5D38BDFC24FF22" box="[867,969,151,176]" name="Thailand" pageId="8" pageNumber="391">Thailand</collectingCountry>
,
<collectingRegion id="4982F84EFF95FFFE9DE538BDFBB7FF22" box="[987,1114,151,176]" country="Thailand" name="Chanthaburi" pageId="8" pageNumber="391">Chantaburi</collectingRegion>
,
<location id="8E996077FF95FFFE9A5238B2FA75FF22" LSID="urn:lsid:plazi:treatment:03EF87BAFF9AFFFE9EA93F76FC02FBEE:8E996077FF95FFFE9A5238B2FA75FF22" box="[1132,1432,151,176]" country="Thailand" latitude="10.078055" longLatPrecision="21" longitude="101.80139" name="Kaeng Hang Maew Dist." pageId="8" pageNumber="391" stateProvince="Chantaburi">Kaeng Hang Maew Dist.</location>
,
<geoCoordinate id="EE72506BFF95FFFE9EA93896FEC9FF46" box="[151,292,188,212]" direction="north" orientation="latitude" pageId="8" pageNumber="391" precision="15" value="10.078055">10°441” N</geoCoordinate>
,
<geoCoordinate id="EE72506BFF95FFFE9F063896FE3CFF46" box="[312,465,188,213]" direction="east" orientation="longitude" pageId="8" pageNumber="391" precision="15" value="101.80139">101°485” E</geoCoordinate>
,
<collectingDate id="EFBCE984FF95FFFE9FDA3896FD82FF47" box="[484,623,188,213]" pageId="8" pageNumber="391" value="2005-06-02" valueMax="2005-06-03" valueMin="2005-06-02">
<date id="FFF8106CFF95FFFE9FDA3896FD82FF47" box="[484,623,188,213]" pageId="8" pageNumber="391" value="2005-06-02" valueMax="2005-06-03" valueMin="2005-06-02">2-3.VI.2005</date>
</collectingDate>
, leg.
<collectorName id="26B3537AFF95FFFE9C833897FCC3FF47" box="[701,814,188,213]" pageId="8" pageNumber="391">K. Černý</collectorName>
, slide MWM 33812
<collectorName id="26B3537AFF95FFFE9A063896FB73FF47" box="[1080,1182,188,213]" pageId="8" pageNumber="391">Volynkin</collectorName>
(
<collectorName id="26B3537AFF95FFFE9A883896FB02FF47" box="[1206,1263,188,213]" pageId="8" pageNumber="391">Coll.</collectorName>
MWM/
<collectionCode id="ED57AE69FF95FFFE9B683896FA7CFF46" box="[1366,1425,188,212]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="8" pageNumber="391">ZSM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B2E3CF1FF95FFFE9EA938F5FAD9FF6A" ID-GBIF-Occurrence="1837804129" box="[151,1332,223,248]" collectingDate="2005-06-02" collectingDateMax="2005-06-03" collectingDateMin="2005-06-02" collectionCode="ZSM" collectorName="K. Cerny &amp; Volynkin &amp; Coll." country="Thailand" latitude="10.078055" location="Kaeng Hang Maew Dist." longLatPrecision="21" longitude="101.80139" pageId="8" pageNumber="391" specimenCount="19" specimenCount-female="11" specimenCount-male="8" stateProvince="Chantaburi" typeStatus="paratype">
<typeStatus id="54FD880EFF95FFFE9EA938F5FEE2FF6A" box="[151,271,223,248]" pageId="8" pageNumber="391" type="paratype">
<emphasis id="B932EABEFF95FFFE9EA938F5FEE2FF6A" bold="true" box="[151,271,223,248]" pageId="8" pageNumber="391">Paratypes</emphasis>
</typeStatus>
:
<specimenCount id="9D40FD25FF95FFFE9F2038CAFEA1FF65" box="[286,332,224,247]" pageId="8" pageNumber="391" type="male">7 ♂</specimenCount>
,
<specimenCount id="9D40FD25FF95FFFE9F6738CAFE7FFF65" box="[345,402,224,248]" pageId="8" pageNumber="391" type="female">10 ♀</specimenCount>
, same data as in the
<typeStatus id="54FD880EFF95FFFE9CBC38F5FD0AFF6A" box="[642,743,223,248]" pageId="8" pageNumber="391" type="holotype">holotype</typeStatus>
, slides AV4183
<specimenCount id="9D40FD25FF95FFFE9DA038CAFC5DFF6A" box="[926,944,224,248]" pageId="8" pageNumber="391" type="male"></specimenCount>
, AV4184
<specimenCount id="9D40FD25FF95FFFE9A1C38CAFBC2FF6A" box="[1058,1071,224,248]" pageId="8" pageNumber="391" type="female"></specimenCount>
Volynkin (Coll. CKC)
</materialsCitation>
;
<materialsCitation id="3B2E3CF1FF95FFFE9B7F38CAFDA6FED2" ID-GBIF-Occurrence="1837804124" collectingDate="2005-11-21" collectorName="K. Cerny &amp; Volynkin &amp; Coll." country="Thailand" elevation="265" latitude="13.061111" location="Soi Dao distr." longLatPrecision="21" longitude="102.24583" municipality="Bar Ta Moon" pageId="8" pageNumber="391" specimenCount="4" specimenCount-male="4" stateProvince="Chantaburi" typeStatus="paratype">
<specimenCount id="9D40FD25FF95FFFE9B7F38CAFA83FF65" box="[1345,1390,224,248]" pageId="8" pageNumber="391" type="male">3 ♂</specimenCount>
, SE
<collectingCountry id="F351763CFF95FFFE9EA9392EFF10FE8F" box="[151,253,260,285]" name="Thailand" pageId="8" pageNumber="391">Thailand</collectingCountry>
,
<collectingRegion id="4982F84EFF95FFFE9F37392EFE65FE8F" box="[265,392,260,285]" country="Thailand" name="Chanthaburi" pageId="8" pageNumber="391">Chantaburi</collectingRegion>
,
<elevation id="006BD19FFF95FFFE9FAA392EFE33FE8E" box="[404,478,260,285]" metricMagnitude="2" metricUnit="m" metricValue="2.65" pageId="8" pageNumber="391" unit="m" value="265.0">
<quantity id="4CBE9B49FF95FFFE9FAA392EFE33FE8E" box="[404,478,260,285]" metricMagnitude="2" metricUnit="m" metricValue="2.65" pageId="8" pageNumber="391" unit="m" value="265.0">265 m</quantity>
</elevation>
,
<collectingMunicipality id="6B9DACD6FF95FFFE9FD5392FFD69FE8E" box="[491,644,261,284]" pageId="8" pageNumber="391">Bar Ta Moon</collectingMunicipality>
,
<location id="8E996077FF95FFFE9CAE392EFCC3FE8F" LSID="urn:lsid:plazi:treatment:03EF87BAFF9AFFFE9EA93F76FC02FBEE:8E996077FF95FFFE9CAE392EFCC3FE8F" box="[656,814,260,285]" country="Thailand" latitude="13.061111" longLatPrecision="21" longitude="102.24583" municipality="Bar Ta Moon" name="Soi Dao distr." pageId="8" pageNumber="391" stateProvince="Chantaburi">Soi Dao distr.</location>
,
<geoCoordinate id="EE72506BFF95FFFE9D07392EFC56FE8E" box="[825,955,260,284]" direction="north" orientation="latitude" pageId="8" pageNumber="391" precision="15" value="13.061111">13°340N</geoCoordinate>
,
<geoCoordinate id="EE72506BFF95FFFE9DF6392EFB8FFE8F" box="[968,1122,260,285]" direction="east" orientation="longitude" pageId="8" pageNumber="391" precision="15" value="102.24583">102°1445E</geoCoordinate>
,
<collectingDate id="EFBCE984FF95FFFE9A50392EFB1DFE8F" box="[1134,1264,260,285]" pageId="8" pageNumber="391" value="2005-11-21">
<date id="FFF8106CFF95FFFE9A50392EFB1DFE8F" box="[1134,1264,260,285]" pageId="8" pageNumber="391" value="2005-11-21">21.XI.2005</date>
</collectingDate>
, leg.
<collectorName id="26B3537AFF95FFFE9B10392FFA75FE8F" box="[1326,1432,260,285]" pageId="8" pageNumber="391">K. Černý</collectorName>
, slide AV4184
<specimenCount id="9D40FD25FF95FFFE9F0D3902FEA8FED2" box="[307,325,296,320]" pageId="8" pageNumber="391" type="male"></specimenCount>
<collectorName id="26B3537AFF95FFFE9F72390DFE5FFED2" box="[332,434,295,320]" pageId="8" pageNumber="391">Volynkin</collectorName>
(
<collectorName id="26B3537AFF95FFFE9FFF390DFE17FED2" box="[449,506,295,320]" pageId="8" pageNumber="391">Coll.</collectorName>
CKC).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35C6527FF95FFFE9EF93966FD87FCCF" pageId="8" pageNumber="391" type="description">
<paragraph id="8BF936ACFF95FFFE9EF93966FD87FCCF" blockId="8.[151,1437,151,1149]" pageId="8" pageNumber="391">
<emphasis id="B932EABEFF95FFFE9EF93966FE80FEF7" bold="true" box="[199,365,332,357]" pageId="8" pageNumber="391">Redescription</emphasis>
.
<emphasis id="B932EABEFF95FFFE9F413966FE56FEF7" bold="true" box="[383,443,332,357]" pageId="8" pageNumber="391">Male</emphasis>
(
<figureCitation id="137D2A29FF95FFFE9FF13966FDCBFEF6" box="[463,550,332,357]" captionStart="FIGURES 1119" captionStartId="4.[151,264,1844,1866]" captionTargetBox="[200,1361,194,1789]" captionTargetId="figure@4.[191,1396,193,1823]" captionTargetPageId="4" captionText="FIGURES 1119. Macohasa and Chinasa spp.: adults. 11, M. orientalis, ♂, Malaysia, Malacca Peninsula (MWM/ZSM); 12, M. orientalis, ♀, Malaysia, Malacca Peninsula (MWM/ZSM); 13, M. tetraspila, holotype ♂, Thailand, Chantaburi prov. (MWM/ZSM); 14, M. tetraspila, paratype ♀, Thailand, Chantaburi prov. (MWM/ZSM); 15, M. dimorpha, syntype ♂, Philippines, Luzon (©NHMUK); 16, M. dimorpha, syntype ♀, Philippines, Luzon (©NHMUK); 17, Ch. costalis costalis, holotype ♂, N China (©NHMUK); 18, Ch. costalis costalis, ♂, China, Zhejiang (MWM/ZSM); 19, Ch. costalis costalis, ♀, China, Guangdong (SZMN)." httpUri="https://zenodo.org/record/1216620/files/figure.png" pageId="8" pageNumber="391">Fig. 13</figureCitation>
). Forewing length
<quantity id="4CBE9B49FF95FFFE9D3B3966FC9BFEF6" box="[773,886,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="8" pageNumber="391" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">910 mm</quantity>
. Forewing ground color yellowish brown, with costa slightly conVeX in subapical third and dorsum noticeably conVeX in basal half; costal margin with one broad, diffuse antemedial black spot; central cell with a long and Very narrow fold with androconial scales; posterior section of forewing posteriorly the costal spot with broad diffuse crossline presented as blackish suffusion. Hindwings ochreous.
<emphasis id="B932EABEFF95FFFE9FA739F6FDA7FE67" bold="true" box="[409,586,476,501]" pageId="8" pageNumber="391">Male genitalia</emphasis>
(
<figureCitation id="137D2A29FF95FFFE9C5E39F6FD56FE66" box="[608,699,476,501]" captionStart="FIGURES 2729" captionStartId="12.[151,264,1953,1975]" captionTargetBox="[180,1380,198,1932]" captionTargetId="figure@12.[151,1436,193,1932]" captionTargetPageId="12" captionText="FIGURES 2729. Macohasa spp.: male genitalia (genital capsules in ventral vieW). 27, M. orientalis, Malaysia, slide MWM 31775 Volynkin; 28, M. tetraspila, holotype, Thailand, Chantaburi prov., slide MWM 33812 Volynkin; 29, M. dimorpha, Philippines, PalaWan, slide MWM 31777 Volynkin." httpUri="https://zenodo.org/record/1216626/files/figure.png" pageId="8" pageNumber="391">Fig. 28</figureCitation>
). Uncus simple, long, basally broad, apically pointed, curVed downwards at apical half; scaphium broad, heaVily sclerotized; Vinculum elongated, more or less V-like, with tapered apeX; apeX of ValVe rounded; harpe-like fold small, pimple-like; distal saccular process short, broad, with apeX pointed and curVed dorsally. Aedeagus long, slightly curVed; Vesica broad, with two membranous subbasal diVerticula, one large globular distal diVerticulum with granulation, and one long and basally curVed distal diVerticulum with granulation; Vesica ejaculatorius broad, situated laterally.
<emphasis id="B932EABEFF95FFFE9DD93AA5FBD3FD3A" bold="true" box="[999,1086,655,680]" pageId="8" pageNumber="391">Female</emphasis>
(
<figureCitation id="137D2A29FF95FFFE9A733AA5FB4CFD3A" box="[1101,1185,655,680]" captionStart="FIGURES 1119" captionStartId="4.[151,264,1844,1866]" captionTargetBox="[200,1361,194,1789]" captionTargetId="figure@4.[191,1396,193,1823]" captionTargetPageId="4" captionText="FIGURES 1119. Macohasa and Chinasa spp.: adults. 11, M. orientalis, ♂, Malaysia, Malacca Peninsula (MWM/ZSM); 12, M. orientalis, ♀, Malaysia, Malacca Peninsula (MWM/ZSM); 13, M. tetraspila, holotype ♂, Thailand, Chantaburi prov. (MWM/ZSM); 14, M. tetraspila, paratype ♀, Thailand, Chantaburi prov. (MWM/ZSM); 15, M. dimorpha, syntype ♂, Philippines, Luzon (©NHMUK); 16, M. dimorpha, syntype ♀, Philippines, Luzon (©NHMUK); 17, Ch. costalis costalis, holotype ♂, N China (©NHMUK); 18, Ch. costalis costalis, ♂, China, Zhejiang (MWM/ZSM); 19, Ch. costalis costalis, ♀, China, Guangdong (SZMN)." httpUri="https://zenodo.org/record/1216620/files/figure.png" pageId="8" pageNumber="391">Fig. 14</figureCitation>
). Forewing length
<quantity id="4CBE9B49FF95FFFE9B4D3ABAFF01FD5E" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="8" pageNumber="391" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">11 12 mm</quantity>
. Forewing ground color pale yellow, with a row of 5 black spots, the middle one Very small, dot-like, sometimes absent; hindwing pale yellow.
<emphasis id="B932EABEFF95FFFE9C523AFDFCDFFD62" bold="true" box="[620,818,727,752]" pageId="8" pageNumber="391">Female genitalia</emphasis>
(
<figureCitation id="137D2A29FF95FFFE9D7F3AFDFC7BFD62" box="[833,918,727,752]" captionStart="FIGURES 3641" captionStartId="15.[151,264,1891,1913]" captionTargetBox="[169,1399,193,1854]" captionTargetId="figure@15.[164,1400,193,1865]" captionTargetPageId="15" captionText="FIGURES 3641. Lithosiina spp.: female genitalia (ventral vieW). 36, Prabhasa venosa, [India, Meghalaya] Khasis, slide BMNH(E) Arct-4557f HolloWay (©NHMUK); 37, Macohasa cernyi, paratype, N Vietnam, slide MWM 31774 Volynkin; 38, Macohasa orientalis, Malaysia, slide MWM 31776 Volynkin; 39, Macohasa tetraspila, paratype, Thailand, Chantaburi prov., slide AV4186m Volynkin; 40, Macohasa dimorpha, Philippines, PalaWan, slide MWM 31778 Volynkin; 41, Macotasa biplagella, slide BMNH(E) Arct-2849 (©NHMUK)." httpUri="https://zenodo.org/record/1216634/files/figure.png" pageId="8" pageNumber="391">Fig. 39</figureCitation>
). OVipositor short, broad; apophyses long and thin, apophyses anteriores slightly longer than apophyses posteriores; ductus bursae short, narrow, membranous, with large, Ventral plate; corpus bursae broad, sack-like, membranous, with one elliptical signum; appendiX bursae short, bulbous, situated latero-posteriorly.
</paragraph>
</subSubSection>
<subSubSection id="C35C6527FF95FFFE9EF93B4DFC51FBCA" pageId="8" pageNumber="391" type="discussion">
<paragraph id="8BF936ACFF95FFFE9EF93B4DFC51FBCA" blockId="8.[151,1437,151,1149]" pageId="8" pageNumber="391">
<emphasis id="B932EABEFF95FFFE9EF93B4DFEC4FC12" bold="true" box="[199,297,871,896]" pageId="8" pageNumber="391">Remark</emphasis>
.
<taxonomicName id="4C464D2FFF95FFFE9F023B43FE3CFC12" authorityName="Cerny" authorityYear="2009" box="[316,465,871,896]" class="Insecta" family="Erebidae" genus="Macotasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="391" phylum="Arthropoda" rank="species" species="tetraspila">
<emphasis id="B932EABEFF95FFFE9F023B43FE3CFC12" box="[316,465,871,896]" italics="true" pageId="8" pageNumber="391">M. tetraspila</emphasis>
</taxonomicName>
is a closest relatiVe of
<taxonomicName id="4C464D2FFF95FFFE9CD33B43FC68FC12" authorityName="Dubatolov &amp; Volynkin &amp; Kishida" authorityYear="2018" baseAuthorityName="Hampson" baseAuthorityYear="1918" box="[749,901,871,896]" class="Insecta" family="Erebidae" genus="Macohasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="391" phylum="Arthropoda" rank="species" species="dimorpha">
<emphasis id="B932EABEFF95FFFE9CD33B43FC68FC12" box="[749,901,871,896]" italics="true" pageId="8" pageNumber="391">M. dimorpha</emphasis>
</taxonomicName>
. Both species haVe female genitalia differing significantly from those of two other congeners by the large Ventral plate of ductus bursae and elliptical signum (whereas in
<taxonomicName id="4C464D2FFF95FFFE9F1D3B9BFE7DFC5A" authority="Dubatolov &amp; Volynkin &amp; Kishida, 2018" authorityName="Dubatolov &amp; Volynkin &amp; Kishida" authorityYear="2018" box="[291,400,945,968]" class="Insecta" family="Erebidae" genus="Macohasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="391" phylum="Arthropoda" rank="species" species="cernyi" status="sp. nov.">
<emphasis id="B932EABEFF95FFFE9F1D3B9BFE7DFC5A" box="[291,400,945,968]" italics="true" pageId="8" pageNumber="391">M. cernyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20157C5FF95FFFE9FA73B85FE18FC5A" box="[409,501,943,968]" pageId="8" pageNumber="391" rank="species">
<emphasis id="B932EABEFF95FFFE9FA73B85FE18FC5A" bold="true" box="[409,501,943,968]" pageId="8" pageNumber="391">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C464D2FFF95FFFE9C103B9BFD2CFC5A" authorityName=": Hampson" authorityYear="1914" box="[558,705,943,968]" class="Insecta" family="Arctiidae" genus="Phaeosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="391" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B932EABEFF95FFFE9C103B9BFD2CFC5A" box="[558,705,943,968]" italics="true" pageId="8" pageNumber="391">M. orientalis</emphasis>
</taxonomicName>
the Ventral plates are round and bulbous, and the signa are barshaped). The male genitalia of both seXes are similar to those of
<taxonomicName id="4C464D2FFF95FFFE9DAD3BFFFBECFC7E" authority="Dubatolov &amp; Volynkin &amp; Kishida, 2018" authorityName="Dubatolov &amp; Volynkin &amp; Kishida" authorityYear="2018" box="[915,1025,981,1004]" class="Insecta" family="Erebidae" genus="Macohasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="391" phylum="Arthropoda" rank="species" species="cernyi" status="sp. nov.">
<emphasis id="B932EABEFF95FFFE9DAD3BFFFBECFC7E" box="[915,1025,981,1004]" italics="true" pageId="8" pageNumber="391">M. cernyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20157C5FF95FFFE9A323BFEFB84FC7F" box="[1036,1129,980,1005]" pageId="8" pageNumber="391" rank="species">
<emphasis id="B932EABEFF95FFFE9A323BFEFB84FC7F" bold="true" box="[1036,1129,980,1005]" pageId="8" pageNumber="391">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C464D2FFF95FFFE9A983BFFFAD7FC7F" authorityName=": Hampson" authorityYear="1914" box="[1190,1338,980,1005]" class="Insecta" family="Arctiidae" genus="Phaeosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="391" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B932EABEFF95FFFE9A983BFFFAD7FC7F" box="[1190,1338,980,1005]" italics="true" pageId="8" pageNumber="391">M. orientalis</emphasis>
</taxonomicName>
, but the scaphium is significantly broader, the harpe-like fold is significantly smaller, the distal saccular process is smaller, the Vinculum is significantly larger, and both distal diVerticula of aedeagus Vesica are granulated. Differences of
<taxonomicName id="4C464D2FFF95FFFE9B433C37FEEEFBCA" authorityName="Cerny" authorityYear="2009" class="Insecta" family="Erebidae" genus="Macotasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="391" phylum="Arthropoda" rank="species" species="tetraspila">
<emphasis id="B932EABEFF95FFFE9B433C37FEEEFBCA" italics="true" pageId="8" pageNumber="391">M. tetraspila</emphasis>
</taxonomicName>
from
<taxonomicName id="4C464D2FFF95FFFE9F783C6BFE34FBCA" authorityName="Dubatolov &amp; Volynkin &amp; Kishida" authorityYear="2018" baseAuthorityName="Hampson" baseAuthorityYear="1918" box="[326,473,1087,1112]" class="Insecta" family="Erebidae" genus="Macohasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="391" phylum="Arthropoda" rank="species" species="dimorpha">
<emphasis id="B932EABEFF95FFFE9F783C6BFE34FBCA" box="[326,473,1087,1112]" italics="true" pageId="8" pageNumber="391">M. dimorpha</emphasis>
</taxonomicName>
are listed in the diagnosis of
<emphasis id="B932EABEFF95FFFE9D1C3C6BFC51FBCA" box="[802,956,1087,1112]" italics="true" pageId="8" pageNumber="391">
<taxonomicName id="4C464D2FFF95FFFE9D1C3C6BFC55FBCA" authorityName="Dubatolov &amp; Volynkin &amp; Kishida" authorityYear="2018" baseAuthorityName="Hampson" baseAuthorityYear="1918" box="[802,952,1087,1112]" class="Insecta" family="Erebidae" genus="Macohasa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="391" phylum="Arthropoda" rank="species" species="dimorpha">M. dimorpha</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C35C6527FF95FFFE9EF93C4EFC02FBEE" box="[199,1007,1124,1149]" pageId="8" pageNumber="391" type="distribution">
<paragraph id="8BF936ACFF95FFFE9EF93C4EFC02FBEE" blockId="8.[151,1437,151,1149]" box="[199,1007,1124,1149]" pageId="8" pageNumber="391">
<emphasis id="B932EABEFF95FFFE9EF93C4EFEB4FBEF" bold="true" box="[199,345,1124,1149]" pageId="8" pageNumber="391">Distribution</emphasis>
. SE
<collectingCountry id="F351763CFF95FFFE9FB13C4EFE18FBEF" box="[399,501,1124,1149]" name="Thailand" pageId="8" pageNumber="391">Thailand</collectingCountry>
,
<collectingCountry id="F351763CFF95FFFE9C3E3C4EFD8BFBEF" box="[512,614,1124,1149]" name="Malaysia" pageId="8" pageNumber="391">Malaysia</collectingCountry>
(Malay Peninsula) (Bucsek 2014).
</paragraph>
</subSubSection>
</treatment>
</document>