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<document id="570E4422432B400C291BA66E8E9E969E" ID-DOI="10.5252/g2009n4a893" ID-ISSN="1638-9395" ID-Zenodo-Dep="5380423" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="donat" checkinTime="1630289815293" checkinUser="marcus" docAuthor="Geraads, Denis" docDate="2009" docId="03EF879FFFBDFFB7C4ADFB2F6D4AFAAF" docLanguage="fr" docName="Geodiversitas.2009.31.4.893-908.pdf" docOrigin="Geodiversitas 31 (4)" docSource="http://www.bioone.org/doi/abs/10.5252/g2009n4a893" docStyle="DocumentStyle:06460E2025300062676A2DBCBE4CBDE8.3:Geodiversitas.2009-2010.journal_article.type1" docStyleId="06460E2025300062676A2DBCBE4CBDE8" docStyleName="Geodiversitas.2009-2010.journal_article.type1" docStyleVersion="3" docTitle="Helladotherium duvernoyi" docType="treatment" docVersion="5" lastPageNumber="901" masterDocId="FFD6FFE7FFBAFFBFC440FFE26F09FFE7" masterDocTitle="Giraffidae (Mammalia) de la fin du Néogène de la République de Macédoine (ARYM)" masterLastPageNumber="908" masterPageNumber="893" pageNumber="900" updateTime="1712934493062" updateUser="donat" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="346C820F95D7FD1AB9661E23B09B73C0">Giraffidae (Mammalia) de la fin du Néogène de la République de Macédoine (ARYM)</mods:title>
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<mods:namePart id="8210674D6A2540C1CBFE0FD808A65E35">Geraads, Denis</mods:namePart>
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<mods:date id="735BE9EF4AE8C9D597C82B2DD167A692">2009</mods:date>
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<treatment id="03EF879FFFBDFFB7C4ADFB2F6D4AFAAF" ID-DOI="http://doi.org/10.5281/zenodo.5465597" ID-Zenodo-Dep="5465597" LSID="urn:lsid:plazi:treatment:03EF879FFFBDFFB7C4ADFB2F6D4AFAAF" httpUri="http://treatment.plazi.org/id/03EF879FFFBDFFB7C4ADFB2F6D4AFAAF" lastPageId="8" lastPageNumber="901" pageId="7" pageNumber="900">
<subSubSection id="C35C6502FFBDFFB8C4ADFB2F6D0DFAEF" pageId="7" pageNumber="900" type="nomenclature">
<paragraph id="8BF93689FFBDFFB8C4ADFB2F6D0DFAEF" blockId="7.[237,524,1229,1288]" pageId="7" pageNumber="900">
<heading id="D0B181E5FFBDFFB8C4ADFB2F6D0DFAEF" centered="true" fontSize="11" level="2" pageId="7" pageNumber="900" reason="2">
<taxonomicName id="4C464D0AFFBDFFB8C4ADFB2F6D0DFAEF" authority="(Gaudry &amp; Lartet, 1856)" baseAuthorityName="Gaudry &amp; Lartet" baseAuthorityYear="1856" class="Mammalia" family="Giraffidae" genus="Helladotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="900" phylum="Chordata" rank="species" species="duvernoyi">
<emphasis id="B932EA9BFFBDFFB8C4ADFB2F6D05FB0F" bold="true" box="[237,524,1229,1256]" italics="true" pageId="7" pageNumber="900">Helladotherium duvernoyi</emphasis>
(
<bibRefCitation id="EFD74B78FFBDFFB8C4BBFB0F6EF5FAEF" author="GAUDRY A. &amp; LARTET E." box="[251,508,1261,1288]" pageId="7" pageNumber="900" pagination="271 - 274" refId="ref10135" refString="GAUDRY A. &amp; LARTET E. 1856. - Resultats des recherches paleontologiques entreprises dans l'Attique sous les auspices de l'Academie. Comptes-rendus hebdomadaires des seances de l'Academie des Sciences 43: 271 - 274." type="journal article" year="1856">Gaudry &amp; Lartet, 1856</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35C6502FFBDFFB8C431FACA6D8EFAA6" box="[113,647,1320,1345]" pageId="7" pageNumber="900" type="reference_group">
<paragraph id="8BF93689FFBDFFB8C431FACA6D8EFAA6" blockId="7.[113,647,1320,1345]" box="[113,647,1320,1345]" pageId="7" pageNumber="900">
<treatmentCitationGroup id="AB5611A7FFBDFFB8C431FACA6D8EFAA6" box="[113,647,1320,1345]" pageId="7" pageNumber="900">
<taxonomicName id="4C464D0AFFBDFFB8C431FACA6D8AFAA6" ID-CoL="3G3KN" authority="Gaudry &amp; Lartet, 1856: 274" authorityName="Gaudry &amp; Lartet" authorityPageNumber="274" authorityYear="1856" box="[113,643,1320,1345]" class="Mammalia" family="Giraffidae" genus="Camelopardalis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="900" phylum="Chordata" rank="species" species="duvernoyi">
<emphasis id="B932EA9BFFBDFFB8C431FACA6E08FAA7" box="[113,257,1320,1344]" italics="true" pageId="7" pageNumber="900">Camelopardalis</emphasis>
<treatmentCitation id="0AE71098FFBDFFB8C548FACA6D8AFAA6" author="GAUDRY A. &amp; LARTET E." box="[264,643,1320,1345]" page="274" pageId="7" pageNumber="900" year="1856">
<emphasis id="B932EA9BFFBDFFB8C548FACA6E6AFAA7" box="[264,355,1320,1344]" italics="true" pageId="7" pageNumber="900">duvernoyi</emphasis>
<bibRefCitation id="EFD74B78FFBDFFB8C529FACB6D8AFAA6" author="GAUDRY A. &amp; LARTET E." box="[361,643,1321,1345]" pageId="7" pageNumber="900" pagination="271 - 274" refId="ref10135" refString="GAUDRY A. &amp; LARTET E. 1856. - Resultats des recherches paleontologiques entreprises dans l'Attique sous les auspices de l'Academie. Comptes-rendus hebdomadaires des seances de l'Academie des Sciences 43: 271 - 274." type="journal article" year="1856">Gaudry &amp; Lartet, 1856: 274</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35C6502FFBDFFB8C431FABB6D79F993" pageId="7" pageNumber="900" type="materials_examined">
<paragraph id="8BF93689FFBDFFB8C431FABB6FF2FA12" blockId="7.[112,649,1368,1525]" pageId="7" pageNumber="900">
<collectionCode id="ED57AE4CFFBDFFB8C431FABB6FAAFA89" box="[113,163,1369,1392]" pageId="7" pageNumber="900">TYPE</collectionCode>
. —
<bibRefCitation id="EFD74B78FFBDFFB8C494FABA6EC6FA97" author="GAUDRY A. &amp; LARTET E." box="[212,463,1368,1392]" pageId="7" pageNumber="900" pagination="271 - 274" refId="ref10135" refString="GAUDRY A. &amp; LARTET E. 1856. - Resultats des recherches paleontologiques entreprises dans l'Attique sous les auspices de l'Academie. Comptes-rendus hebdomadaires des seances de l'Academie des Sciences 43: 271 - 274." type="journal article" year="1856">Gaudry &amp; Lartet (1856)</bibRefCitation>
n’ont pas désigné d’holotype parmi la série type, ultérieurement mélangée à d’autres spécimens, de sorte qu’il est impossible de désigner un
<typeStatus id="54FD882BFFBDFFB8C4AEFA4A6E40FA27" box="[238,329,1448,1472]" pageId="7" pageNumber="900" type="lectotype">lectotype</typeStatus>
, mais ceci est sans conséquence, tous les grands Giraffi dae de la localité type appartenant à cette espèce.
</paragraph>
<paragraph id="8BF93689FFBDFFB8C431F9EF6D79F993" blockId="7.[112,647,1548,1652]" pageId="7" pageNumber="900">
MATÉRIEL ÉTUDIÉ. — Maxillaire de Karaslari (Ka-2752), base d’un crâne de Kirokucuk (KK-2737), maxillaire de Vozarci (Voz-1870/79) et métacarpien sans numéro du même site. Tout ce matériel est conservé au
<collectionCode id="ED57AE4CFFBDFFB8C65DF9BE6D65F993" box="[541,620,1628,1652]" pageId="7" pageNumber="900">MHNS</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C35C6502FFBDFFB7C431F96E6D4AFAAF" lastPageId="8" lastPageNumber="901" pageId="7" pageNumber="900" type="description">
<paragraph id="8BF93689FFBDFFB8C431F96E6EE7F944" blockId="7.[113,494,1675,1699]" box="[113,494,1675,1699]" pageId="7" pageNumber="900">
LOCALITÉ
<typeStatus id="54FD882BFFBDFFB8C496F9736E0AF946" box="[214,259,1681,1697]" pageId="7" pageNumber="900">TYPE</typeStatus>
. — Pikermi (Turolien).
</paragraph>
<paragraph id="8BF93689FFBDFFB8C6E5FDD76CA4FD51" blockId="7.[675,1211,564,694]" pageId="7" pageNumber="900">
EXTENSION STRATIGRAPHIQUE ET GÉOGRAPHIQUE. — Vallésien final au Turolien final de la province balkanoiranienne; le genre a aussi été signalé en
<collectingCountry id="F3517619FFBDFFB8C061FD8B6B6BFD66" box="[1057,1122,617,641]" name="France" pageId="7" pageNumber="900">France</collectingCountry>
(
<bibRefCitation id="EFD74B78FFBDFFB8C02DFD8B6DDFFD7B" author="GAUDRY A." pageId="7" pageNumber="900" refId="ref10111" refString="GAUDRY A. 1873. - Animaux fossiles du Mont Leberon (Vaucluse). F. Savy, Paris, 112 p." type="book" year="1873">Gaudry 1873</bibRefCitation>
) et en
<collectingCountry id="F3517619FFBDFFB8C754FD666C6CFD7B" box="[788,869,644,668]" name="Hungary" pageId="7" pageNumber="900">Hongrie</collectingCountry>
(
<bibRefCitation id="EFD74B78FFBDFFB8C72FFD666B17FD7B" author="BERNOR R. L. &amp; FEIBEL C. &amp; VIRANTA S." box="[879,1054,643,668]" pageId="7" pageNumber="900" pagination="105 - 111" refId="ref9289" refString="BERNOR R. L., FEIBEL C. &amp; VIRANTA S. 2003. - The vertebrate locality of Hatvan, late Miocene (MiddleTurolian, MN 12), Hungary, in PETCULESCU A. &amp; STIUCA E. (eds), Advances in Vertebrate Paleontology &quot; Hen to Panta &quot;. A tribute to Constantin Radulescu and Petre Mihai Samson. Academie roumaine, Bucarest: 105 - 111." type="book chapter" year="2003">
Bernor
<emphasis id="B932EA9BFFBDFFB8C7F8FD666CECFD7C" box="[952,997,643,667]" italics="true" pageId="7" pageNumber="900">et al.</emphasis>
2003
</bibRefCitation>
), mais ces déterminations sont incertaines.
</paragraph>
<paragraph id="8BF93689FFBDFFB8C6E5FD2C6CEFFC8D" blockId="7.[675,1211,718,874]" pageId="7" pageNumber="900">
DIAGNOSE. — C’est à cette espèce que sont habituellement attribués les nombreuses dentitions et os des membres de
<taxonomicName id="4C464D0AFFBDFFB8C766FCE16B2BFCFC" authority="du Miocene" authorityName="du Miocene" box="[806,1058,771,795]" class="Mammalia" family="Giraffidae" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="900" phylum="Chordata" rank="subFamily" subFamily="Sivatheriinae">Sivatheriinae du Miocène</taxonomicName>
supérieur de la région balkano-iranienne, mais le crâne du mâle étant toujours inconnu, aucune diagnose ne peut être proposée (voir Discussion de
<taxonomicName id="4C464D0AFFBDFFB8C725FCB06CD1FC8D" authorityName="Falconer &amp; Cautley" authorityYear="1836" box="[869,984,850,874]" class="Mammalia" family="Giraffidae" genus="Sivatherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="900" phylum="Chordata" rank="genus">
<emphasis id="B932EA9BFFBDFFB8C725FCB06CD1FC8D" box="[869,984,850,874]" italics="true" pageId="7" pageNumber="900">Sivatherium</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BF93689FFBDFFB8C6E5FC736B09FC4E" blockId="7.[674,1211,913,1700]" box="[677,1024,913,939]" pageId="7" pageNumber="900">DESCRIPTION ET COMPARAISONS</paragraph>
<paragraph id="8BF93689FFBDFFB8C6E5FC526C6DFAEF" blockId="7.[674,1211,913,1700]" pageId="7" pageNumber="900">
Un maxillaire de Karaslari (Ka-2752) comporte toutes les jugales supérieures, mais la plus grande partie des murailles externes des molaires manque. Les prémolaires sont grandes relativement aux molaires (dimensions:
<tableCitation id="C6C40332FFBDFFB8C73AFBCD6CEFFBAD" box="[890,998,1071,1098]" captionStart="TABLEAU" captionStartId="7.[113,122,221,238]" captionText="TABLEAU 3. — Dimensions (en mm) de quelques séries dentaires de Helladotherium Gaudry,1860 de la République de Macédoine, par comparaison avec quelques autres spécimens du Turolien. Notes: 1, Geraads et al. 2005; 2, Bohlin 1926; 3, PIUW." pageId="7" pageNumber="900">Tableau 3</tableCitation>
), P4 est à peu près aussi large que M1; ces caractères se retrouvent non seulement chez les autres
<taxonomicName id="4C464D0AFFBDFFB8C7B1FB8C6B9CFB6F" authorityName="Gaudry" authorityYear="1860" box="[1009,1173,1134,1160]" class="Mammalia" family="Giraffidae" genus="Helladotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="900" phylum="Chordata" rank="genus">
<emphasis id="B932EA9BFFBDFFB8C7B1FB8C6B9CFB6F" box="[1009,1173,1134,1160]" italics="true" pageId="7" pageNumber="900">Helladotherium</emphasis>
</taxonomicName>
(et
<taxonomicName id="4C464D0AFFBDFFB8C6E5FB6C6C37FB4F" authorityName="Falconer" authorityYear="1845" box="[677,830,1166,1192]" class="Mammalia" family="Giraffidae" genus="Bramatherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="900" phylum="Chordata" rank="genus">
<emphasis id="B932EA9BFFBDFFB8C6E5FB6C6C37FB4F" box="[677,830,1166,1192]" italics="true" pageId="7" pageNumber="900">Bramatherium</emphasis>
</taxonomicName>
), mais aussi chez les autres
<taxonomicName id="4C464D0AFFBDFFB8C023FB6C6DEFFB2E" class="Mammalia" family="Giraffidae" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="900" phylum="Chordata" rank="subFamily" subFamily="Sivatheriinae">Sivatheriinae</taxonomicName>
. Ils permettent de les distinguer aisément du paléotraginé
<taxonomicName id="4C464D0AFFBDFFB8C76FFB2F6CB2FB00" authorityName="Forsyth-Major" authorityYear="1888" box="[815,955,1229,1255]" class="Mammalia" family="Giraffidae" genus="Samotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="900" phylum="Chordata" rank="genus">
<emphasis id="B932EA9BFFBDFFB8C76FFB2F6CB2FB00" box="[815,955,1229,1255]" italics="true" pageId="7" pageNumber="900">Samotherium</emphasis>
</taxonomicName>
, qui peut atteindre une taille comparable.
</paragraph>
<paragraph id="8BF93689FFBDFFB7C6FBFAEF6D4CFC6B" blockId="7.[674,1211,913,1700]" lastBlockId="8.[111,647,849,1352]" lastPageId="8" lastPageNumber="901" pageId="7" pageNumber="900">
Le spécimen le plus complet est une partie de crâne de Kirokuçuk (KK-2737,
<figureCitation id="137D2A0CFFBDFFB8C040FACF6B40FAAF" box="[1024,1097,1325,1352]" captionStart="FIG" captionStartId="9.[113,124,1567,1584]" captionTargetBox="[113,1201,219,1510]" captionTargetId="figure-12@9.[113,1202,219,1511]" captionTargetPageId="9" captionText="FIG. 3. — A-G, Sivatherium garevskii n. sp.,holotype,Stramer près de Delcevo; A-C, corne postérieure gauche; A, vue médio-dorsale; B, vue antérieure ou antéro-latérale; C, détail de la surface pour montrer les sillons vasculaires; D, nasal en vue dorsale; E, reconstruction de la région de l’ouverture nasale en vue latérale; F, G, corne antérieure; F, vue dorsale; G, vue latérale; H, Sivatherium maurusium (Pomel, 1893), Djibouti, vue médio-dorsale du moulage de la corne postérieure à la même échelle qu’A et B pour montrer la taille très inférieure et la torsion; I, Helladotherium duvernoyi, partie inférieure d’un crâne de Kirokuçuk. Échelle: A, B, D-H, 40 cm; C, 20 cm; I, 25 cm." figureDoi="http://doi.org/10.5281/zenodo.5380427" httpUri="https://zenodo.org/record/5380427/files/figure.png" pageId="7" pageNumber="900">Fig. 3I</figureCitation>
), comportant les dents supérieures et la base du neurocrâne, semblables à ceux du crâne de Pikermi figuré par
<bibRefCitation id="EFD74B78FFBDFFB8C6E5FA6E6C49FA40" author="GAUDRY A." box="[677,832,1420,1447]" pageId="7" pageNumber="900" refId="ref10080" refString="GAUDRY A. 1862 - 67. - Animaux fossiles et Geologie de l'Attique. F. Savy, Paris, 2 vols, 472 p., 75 pls." type="book" year="1862">Gaudry (1862</bibRefCitation>
-67). Les dents présentent les mêmes caractères que sur la pièce précédente, P4 est presque rectangulaire, les molaires ne possèdent que de faibles piliers correspondant au paracône et métacône. La bulle tympanique est très petite, comme chez les
<taxonomicName id="4C464D0AFFBDFFB8C6E6F9C86C39F9A2" box="[678,816,1578,1605]" class="Mammalia" family="Giraffidae" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="900" phylum="Chordata" rank="subFamily" subFamily="Sivatheriinae">Sivatheriinae</taxonomicName>
plio-pléistocènes (
<taxonomicName id="4C464D0AFFBDFFB8C7B4F9C86B7AF9A3" authorityName="Falconer &amp; Cautley" authorityYear="1836" box="[1012,1139,1578,1604]" class="Mammalia" family="Giraffidae" genus="Sivatherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="7" pageNumber="900" phylum="Chordata" rank="genus">
<emphasis id="B932EA9BFFBDFFB8C7B4F9C86B7AF9A3" box="[1012,1139,1578,1604]" italics="true" pageId="7" pageNumber="900">Sivatherium</emphasis>
</taxonomicName>
), mais la base du crâne est moins raccourcie que chez ces derniers et l’axe du conduit auditif reste oblique, et non transversal (
<bibRefCitation id="EFD74B78FFBDFFB8C72CF96B6B0FF943" author="GERAADS D." box="[876,1030,1673,1700]" pageId="7" pageNumber="900" pagination="311 - 321" refId="ref10370" refString="GERAADS D. 1985. - Sivatherium maurusium (Pomel) (Giraffidae, Mammalia) du Pleistocene de la Republique de Djibouti. Palaontologische Zeitschrift 59 (3 / 4): 311 - 321." type="journal article" year="1985">Geraads 1985</bibRefCitation>
). Le basioccipital se rétrécit rapidement vers l’avant; les tubérosités antérieures sont petites mais bien saillantes.
</paragraph>
<caption id="DF396601FFB2FFB7C431FF3E6FC9FEFD" pageId="8" pageNumber="901" startId="8.[113,122,220,237]">
<paragraph id="8BF93689FFB2FFB7C431FF3E6FC9FEFD" blockId="8.[113,645,220,282]" pageId="8" pageNumber="901">
TABLEAU 4. — Dimensions (en mm) des métacarpiens rapportés à
<taxonomicName id="4C464D0AFFB2FFB7C431FF106E60FEE4" authority="Gaudry, 1860" authorityName="Gaudry" authorityYear="1860" box="[113,361,242,259]" class="Mammalia" family="Giraffidae" genus="Helladotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="8" pageNumber="901" phylum="Chordata" rank="genus">
<emphasis id="B932EA9BFFB2FFB7C431FF106FE6FEE4" box="[113,239,242,259]" italics="true" pageId="8" pageNumber="901">Helladotherium</emphasis>
Gaudry, 1860
</taxonomicName>
. Abréviations:
<emphasis id="B932EA9BFFB2FFB7C5A7FF106D07FEE4" bold="true" box="[487,526,242,259]" pageId="8" pageNumber="901">dist.</emphasis>
, distal;
<emphasis id="B932EA9BFFB2FFB7C613FF106D88FEE4" bold="true" box="[595,641,242,259]" pageId="8" pageNumber="901">prox.</emphasis>
, proximal.
</paragraph>
</caption>
<paragraph id="8BF93689FFB2FFB7C6E5FEDE6CC4FE89" blockId="8.[677,1209,315,366]" pageId="8" pageNumber="901">
<tableNote id="76A03707FFB2FFB7C6E5FEDE6CC4FE89" pageId="8" pageNumber="901" targetBox="[120,639,340,775]" targetPageId="8">
ESPÈCE TYPE. —
<taxonomicName id="4C464D0AFFB2FFB7C716FED96C39FE89" authority="Falconer &amp; Cautley, 1836" authorityName="Falconer &amp; Cautley" authorityYear="1836" class="Mammalia" family="Giraffidae" genus="Sivatherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="8" pageNumber="901" phylum="Chordata" rank="species" species="giganteum">
<emphasis id="B932EA9BFFB2FFB7C716FED96B35FEB4" box="[854,1084,315,339]" italics="true" pageId="8" pageNumber="901">Sivatherium giganteum</emphasis>
<bibRefCitation id="EFD74B78FFB2FFB7C005FEDE6C39FE89" author="FALCONER H. &amp; CAUTLEY P. T." pageId="8" pageNumber="901" pagination="1 - 24" refId="ref9763" refString="FALCONER H. &amp; CAUTLEY P. T. 1836. - Sivatherium giganteum, a new fossil Ruminant genus from the valley of Markanda, in the Siwalik branch of the Sub- Himalayan mountains. Asiatic Researches 19: 1 - 24." type="journal article" year="1836">Falconer &amp; Cautley, 1836</bibRefCitation>
</taxonomicName>
par monotypie.
</tableNote>
</paragraph>
<paragraph id="8BF93689FFB2FFB7C51EFEB66D6DFCE0" pageId="8" pageNumber="901">
<table id="F946C429FFB20040C438FEB66D76FCE0" box="[120,639,340,775]" gridcols="4" gridrows="18" pageId="8" pageNumber="901">
<tr id="357634CBFFB20040C438FEB66D76FE8F" box="[120,639,340,360]" gridrow="0" pageId="8" pageNumber="901" rowspan-0="1">
<th id="76A75DB7FFB20040C51EFEB66EBCFE8F" box="[350,437,340,360]" gridcol="1" gridrow="0" pageId="8" pageNumber="901">
<emphasis id="B932EA9BFFB2FFB7C51EFEB66EBCFE8F" bold="true" box="[350,437,340,360]" pageId="8" pageNumber="901">DT prox.</emphasis>
</th>
<th id="76A75DB7FFB20040C587FEB66D1CFE8F" box="[455,533,340,360]" gridcol="2" gridrow="0" pageId="8" pageNumber="901">
<emphasis id="B932EA9BFFB2FFB7C587FEB66D1CFE8F" bold="true" box="[455,533,340,360]" pageId="8" pageNumber="901">DT dist.</emphasis>
</th>
<th id="76A75DB7FFB20040C665FEB66D76FE8F" box="[549,639,340,360]" gridcol="3" gridrow="0" pageId="8" pageNumber="901">
<emphasis id="B932EA9BFFB2FFB7C665FEB66D76FE8F" bold="true" box="[549,639,340,360]" pageId="8" pageNumber="901">longueur</emphasis>
</th>
</tr>
<tr id="357634CBFFB20040C438FE906D76FE62" box="[120,639,370,389]" gridrow="1" pageId="8" pageNumber="901">
<th id="76A75DB7FFB20040C438FE906E5CFE62" box="[120,341,370,389]" gridcol="0" gridrow="1" pageId="8" pageNumber="901">Karaslari</th>
<td id="76A75DB7FFB20040C51EFE906EBCFE62" box="[350,437,370,389]" gridcol="1" gridrow="1" pageId="8" pageNumber="901">114</td>
<td id="76A75DB7FFB20040C587FE906D1CFE62" box="[455,533,370,389]" gridcol="2" gridrow="1" pageId="8" pageNumber="901">111</td>
<td id="76A75DB7FFB20040C665FE906D76FE62" box="[549,639,370,389]" gridcol="3" gridrow="1" pageId="8" pageNumber="901">436</td>
</tr>
<tr id="357634CBFFB20040C438FE686D76FE7A" box="[120,639,394,413]" gridrow="2" pageId="8" pageNumber="901">
<th id="76A75DB7FFB20040C438FE686E5CFE7A" box="[120,341,394,413]" gridcol="0" gridrow="2" pageId="8" pageNumber="901">Vozarci</th>
<td id="76A75DB7FFB20040C51EFE686EBCFE7A" box="[350,437,394,413]" gridcol="1" gridrow="2" pageId="8" pageNumber="901">102</td>
<td id="76A75DB7FFB20040C587FE686D1CFE7A" box="[455,533,394,413]" gridcol="2" gridrow="2" pageId="8" pageNumber="901">100</td>
<td id="76A75DB7FFB20040C665FE686D76FE7A" box="[549,639,394,413]" gridcol="3" gridrow="2" pageId="8" pageNumber="901">408</td>
</tr>
<tr id="357634CBFFB20040C438FE416D76FE51" box="[120,639,419,438]" gridrow="3" pageId="8" pageNumber="901">
<th id="76A75DB7FFB20040C438FE416D76FE51" box="[120,639,419,438]" colspan="4" colspanRight="3" gridcol="0" gridrow="3" pageId="8" pageNumber="901">Pikermi, Grèce</th>
</tr>
<tr id="357634CBFFB20040C438FE596D76FE29" box="[120,639,443,462]" gridrow="4" pageId="8" pageNumber="901">
<th id="76A75DB7FFB20040C438FE596E5CFE29" box="[120,341,443,462]" gridcol="0" gridrow="4" pageId="8" pageNumber="901">BMNH M 11382</th>
<td id="76A75DB7FFB20040C51EFE596EBCFE29" box="[350,437,443,462]" gridcol="1" gridrow="4" pageId="8" pageNumber="901">96</td>
<td id="76A75DB7FFB20040C587FE596D1CFE29" box="[455,533,443,462]" gridcol="2" gridrow="4" pageId="8" pageNumber="901">92</td>
<td id="76A75DB7FFB20040C665FE596D76FE29" box="[549,639,443,462]" gridcol="3" gridrow="4" pageId="8" pageNumber="901">418</td>
</tr>
<tr id="357634CBFFB20040C438FE316D76FE01" box="[120,639,467,486]" gridrow="5" pageId="8" pageNumber="901">
<th id="76A75DB7FFB20040C438FE316E5CFE01" box="[120,341,467,486]" gridcol="0" gridrow="5" pageId="8" pageNumber="901">BMNH M 11377</th>
<td id="76A75DB7FFB20040C51EFE316EBCFE01" box="[350,437,467,486]" gridcol="1" gridrow="5" pageId="8" pageNumber="901">105</td>
<td id="76A75DB7FFB20040C587FE316D1CFE01" box="[455,533,467,486]" gridcol="2" gridrow="5" pageId="8" pageNumber="901">103</td>
<td id="76A75DB7FFB20040C665FE316D76FE01" box="[549,639,467,486]" gridcol="3" gridrow="5" pageId="8" pageNumber="901">426</td>
</tr>
<tr id="357634CBFFB20040C438FE096D76FE19" box="[120,639,491,510]" gridrow="6" pageId="8" pageNumber="901">
<th id="76A75DB7FFB20040C438FE096E5CFE19" box="[120,341,491,510]" gridcol="0" gridrow="6" pageId="8" pageNumber="901">MNHN PIK-1526</th>
<td id="76A75DB7FFB20040C51EFE096EBCFE19" box="[350,437,491,510]" gridcol="1" gridrow="6" pageId="8" pageNumber="901">92</td>
<td id="76A75DB7FFB20040C587FE096D1CFE19" box="[455,533,491,510]" gridcol="2" gridrow="6" pageId="8" pageNumber="901">103</td>
<td id="76A75DB7FFB20040C665FE096D76FE19" box="[549,639,491,510]" gridcol="3" gridrow="6" pageId="8" pageNumber="901">410</td>
</tr>
<tr id="357634CBFFB20040C438FDE16D76FDF1" box="[120,639,515,534]" gridrow="7" pageId="8" pageNumber="901">
<th id="76A75DB7FFB20040C438FDE16E5CFDF1" box="[120,341,515,534]" gridcol="0" gridrow="7" pageId="8" pageNumber="901">MNHN PIK-1527</th>
<td id="76A75DB7FFB20040C51EFDE16EBCFDF1" box="[350,437,515,534]" gridcol="1" gridrow="7" pageId="8" pageNumber="901">97</td>
<td id="76A75DB7FFB20040C587FDE16D1CFDF1" box="[455,533,515,534]" gridcol="2" gridrow="7" pageId="8" pageNumber="901">102</td>
<td id="76A75DB7FFB20040C665FDE16D76FDF1" box="[549,639,515,534]" gridcol="3" gridrow="7" pageId="8" pageNumber="901">407</td>
</tr>
<tr id="357634CBFFB20040C438FDF96D76FDC9" box="[120,639,539,558]" gridrow="8" pageId="8" pageNumber="901">
<th id="76A75DB7FFB20040C438FDF96E5CFDC9" box="[120,341,539,558]" gridcol="0" gridrow="8" pageId="8" pageNumber="901">MNHN PIK-1525</th>
<td id="76A75DB7FFB20040C51EFDF96EBCFDC9" box="[350,437,539,558]" gridcol="1" gridrow="8" pageId="8" pageNumber="901">87</td>
<td id="76A75DB7FFB20040C587FDF96D1CFDC9" box="[455,533,539,558]" gridcol="2" gridrow="8" pageId="8" pageNumber="901">97</td>
<td id="76A75DB7FFB20040C665FDF96D76FDC9" box="[549,639,539,558]" gridcol="3" gridrow="8" pageId="8" pageNumber="901">427</td>
</tr>
<tr id="357634CBFFB20040C438FDD16D76FDA1" box="[120,639,563,582]" gridrow="9" pageId="8" pageNumber="901">
<th id="76A75DB7FFB20040C438FDD16D76FDA1" box="[120,639,563,582]" colspan="4" colspanRight="3" gridcol="0" gridrow="9" pageId="8" pageNumber="901">Hadjidimovo, Bulgarie</th>
</tr>
<tr id="357634CBFFB20040C438FDA96D76FDB9" box="[120,639,587,606]" gridrow="10" pageId="8" pageNumber="901">
<th id="76A75DB7FFB20040C438FDA96E5CFDB9" box="[120,341,587,606]" gridcol="0" gridrow="10" pageId="8" pageNumber="901">sans numero</th>
<td id="76A75DB7FFB20040C51EFDA96EBCFDB9" box="[350,437,587,606]" gridcol="1" gridrow="10" pageId="8" pageNumber="901">118</td>
<td id="76A75DB7FFB20040C587FDA96D1CFDB9" box="[455,533,587,606]" gridcol="2" gridrow="10" pageId="8" pageNumber="901">109</td>
<td id="76A75DB7FFB20040C665FDA96D76FDB9" box="[549,639,587,606]" gridcol="3" gridrow="10" pageId="8" pageNumber="901">418</td>
</tr>
<tr id="357634CBFFB20040C438FD816D76FD91" box="[120,639,611,630]" gridrow="11" pageId="8" pageNumber="901">
<th id="76A75DB7FFB20040C438FD816E5CFD91" box="[120,341,611,630]" gridcol="0" gridrow="11" pageId="8" pageNumber="901">HD-5492</th>
<td id="76A75DB7FFB20040C51EFD816EBCFD91" box="[350,437,611,630]" gridcol="1" gridrow="11" pageId="8" pageNumber="901">115</td>
<td id="76A75DB7FFB20040C587FD816D1CFD91" box="[455,533,611,630]" gridcol="2" gridrow="11" pageId="8" pageNumber="901">102</td>
<td id="76A75DB7FFB20040C665FD816D76FD91" box="[549,639,611,630]" gridcol="3" gridrow="11" pageId="8" pageNumber="901">430</td>
</tr>
<tr id="357634CBFFB20040C438FD996D76FD69" box="[120,639,635,654]" gridrow="12" pageId="8" pageNumber="901">
<th id="76A75DB7FFB20040C438FD996E5CFD69" box="[120,341,635,654]" gridcol="0" gridrow="12" pageId="8" pageNumber="901">HD-5504</th>
<td id="76A75DB7FFB20040C51EFD996EBCFD69" box="[350,437,635,654]" gridcol="1" gridrow="12" pageId="8" pageNumber="901">110</td>
<td id="76A75DB7FFB20040C587FD996D1CFD69" box="[455,533,635,654]" gridcol="2" gridrow="12" pageId="8" pageNumber="901">104</td>
<td id="76A75DB7FFB20040C665FD996D76FD69" box="[549,639,635,654]" gridcol="3" gridrow="12" pageId="8" pageNumber="901">435</td>
</tr>
<tr id="357634CBFFB20040C438FD716D76FD41" box="[120,639,659,678]" gridrow="13" pageId="8" pageNumber="901">
<th id="76A75DB7FFB20040C438FD716D76FD41" box="[120,639,659,678]" colspan="4" colspanRight="3" gridcol="0" gridrow="13" pageId="8" pageNumber="901">Kalimantsi, Bulgarie</th>
</tr>
<tr id="357634CBFFB20040C438FD4E6D76FD58" box="[120,639,684,703]" gridrow="14" pageId="8" pageNumber="901">
<th id="76A75DB7FFB20040C438FD4E6E5CFD58" box="[120,341,684,703]" gridcol="0" gridrow="14" pageId="8" pageNumber="901">sans numero</th>
<td id="76A75DB7FFB20040C51EFD4E6EBCFD58" box="[350,437,684,703]" gridcol="1" gridrow="14" pageId="8" pageNumber="901">125</td>
<td id="76A75DB7FFB20040C587FD4E6D1CFD58" box="[455,533,684,703]" gridcol="2" gridrow="14" pageId="8" pageNumber="901">100</td>
<td id="76A75DB7FFB20040C665FD4E6D76FD58" box="[549,639,684,703]" gridcol="3" gridrow="14" pageId="8" pageNumber="901">470</td>
</tr>
<tr id="357634CBFFB20040C438FD266D76FD30" box="[120,639,708,727]" gridrow="15" pageId="8" pageNumber="901">
<th id="76A75DB7FFB20040C438FD266E5CFD30" box="[120,341,708,727]" gridcol="0" gridrow="15" pageId="8" pageNumber="901">sans numero</th>
<td id="76A75DB7FFB20040C51EFD266EBCFD30" box="[350,437,708,727]" gridcol="1" gridrow="15" pageId="8" pageNumber="901">115</td>
<td id="76A75DB7FFB20040C587FD266D1CFD30" box="[455,533,708,727]" gridcol="2" gridrow="15" pageId="8" pageNumber="901">110</td>
<td id="76A75DB7FFB20040C665FD266D76FD30" box="[549,639,708,727]" gridcol="3" gridrow="15" pageId="8" pageNumber="901">435</td>
</tr>
<tr id="357634CBFFB20040C438FD3E6D76FD08" box="[120,639,732,751]" gridrow="16" pageId="8" pageNumber="901">
<th id="76A75DB7FFB20040C438FD3E6D76FD08" box="[120,639,732,751]" colspan="4" colspanRight="3" gridcol="0" gridrow="16" pageId="8" pageNumber="901">Maragha, Iran</th>
</tr>
<tr id="357634CBFFB20040C438FD166D76FCE0" box="[120,639,756,775]" gridrow="17" pageId="8" pageNumber="901">
<th id="76A75DB7FFB20040C438FD166E5CFCE0" box="[120,341,756,775]" gridcol="0" gridrow="17" pageId="8" pageNumber="901">MNHN, sans numero</th>
<td id="76A75DB7FFB20040C51EFD166EBCFCE0" box="[350,437,756,775]" gridcol="1" gridrow="17" pageId="8" pageNumber="901">92</td>
<td id="76A75DB7FFB20040C587FD166D1CFCE0" box="[455,533,756,775]" gridcol="2" gridrow="17" pageId="8" pageNumber="901">&gt; 99</td>
<td id="76A75DB7FFB20040C665FD166D76FCE0" box="[549,639,756,775]" gridcol="3" gridrow="17" pageId="8" pageNumber="901">465</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF93689FFB2FFB7C6E5FD0F6CC7FBCF" blockId="8.[675,1211,749,1064]" pageId="8" pageNumber="901">
<tableNote id="76A03707FFB2FFB7C6E5FD0F6CC7FBCF" pageId="8" pageNumber="901" targetBox="[120,639,340,775]" targetPageId="8">
LOCALITÉ TYPE. — Stamer près de Delcevo, nord-est de la République de Macédoine. Le sédiment adhérant aux fossiles ne permet pas de choisir entre la Formation de Pancerevo, attribuée au Miocène supérieur, et celle de Solnje, attribuée au Pliocène (
<bibRefCitation id="EFD74B78FFB2FFB7C7B2FCB46DD3FC6E" author="DUMURDZANOV N. &amp; SERAFIMOVSKI T. &amp; BURCHFIEL B. C." pageId="8" pageNumber="901" pagination="1 - 20" refId="ref9680" refString="DUMURDZANOV N., SERAFIMOVSKI T. &amp; BURCHFIEL B. C. 2004. - Evolution of the Neogene-Pleistocene basins of Macedonia. Geological Society of America Digital Maps and Chart Series 1: 1 - 20." type="book chapter" year="2004">
Dumurdzanov
<emphasis id="B932EA9BFFB2FFB7C0C9FCB56BB3FC89" box="[1161,1210,854,878]" italics="true" pageId="8" pageNumber="901">et al.</emphasis>
2004
</bibRefCitation>
); cependant, ce site a livré trois autres fossiles, une P3 référable à
<taxonomicName id="4C464D0AFFB2FFB7C765FC696CC0FC44" box="[805,969,907,931]" class="Mammalia" family="Suidae" genus="Propotamochoerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="8" pageNumber="901" phylum="Chordata" rank="genus">
<emphasis id="B932EA9BFFB2FFB7C765FC696CC0FC44" box="[805,969,907,931]" italics="true" pageId="8" pageNumber="901">Propotamochoerus</emphasis>
</taxonomicName>
? sp., genre mio-pliocène, et deux fragments de bois de cervidés n’appartenant sans doute pas à des genres modernes et, bien que très pauvre, un tel assemblage s’accorde mieux avec le Pliocène qu’avec le Miocène supérieur, où les cervidés sont rares et les bovidés dominants.
</tableNote>
</paragraph>
<paragraph id="8BF93689FFB2FFB7C4C8FC736D4AFAAF" blockId="8.[111,647,849,1352]" pageId="8" pageNumber="901">
De Vozarci proviennent un maxillaire avec P3-M2, Voz-1870/79, ainsi qu’un métacarpien semblable par ses proportions à ceux de Pikermi. Un spécimen de Karaslari est un peu plus gros, quoique dans les limites de variation de ceux de Perivolaki (
<bibRefCitation id="EFD74B78FFB2FFB7C5A1FBF26FF2FBAD" author="KOSTOPOULOS D. S. &amp; KOUFOS G. D." pageId="8" pageNumber="901" pagination="135 - 149" refId="ref10976" refString="KOSTOPOULOS D. S. &amp; KOUFOS G. D. 2006. - The late Miocene vertebrate locality of Perivolaki, Thessaly, Greece. 8. Giraffidae. Palaeontographica 276 (1 - 6): 135 - 149." type="journal article" year="2006">Kostopoulos &amp; Koufos 2006</bibRefCitation>
), ou de ceux de
<collectingCountry id="F3517619FFB2FFB7C5DBFBCD6EF8FBAD" box="[411,497,1071,1098]" name="Bulgaria" pageId="8" pageNumber="901">Bulgarie</collectingCountry>
(
<bibRefCitation id="EFD74B78FFB2FFB7C5BCFBCD6FA2FB8D" author="GERAADS D. &amp; SPASSOV N. &amp; KOVACHEV D." pageId="8" pageNumber="901" pagination="11 - 18" refId="ref10589" refString="GERAADS D., SPASSOV N. &amp; KOVACHEV D. 2005. - Giraffidae (Artiodactyla, Mammalia) from the late Miocene of Kalimantsi and Hadjidimovo, South-Western Bulgaria. Geologica Balcanica 35 (1 - 2): 11 - 18." type="journal article" year="2005">
Geraads
<emphasis id="B932EA9BFFB2FFB7C617FBD26D8EFBAE" box="[599,647,1071,1097]" italics="true" pageId="8" pageNumber="901">et al.</emphasis>
2005
</bibRefCitation>
), mais plus court et plus massif qu’un métacarpien de Maragha en
<collectingCountry id="F3517619FFB2FFB7C579FB8D6E6AFB6D" box="[313,355,1135,1162]" name="Iran" pageId="8" pageNumber="901">Iran</collectingCountry>
(dimensions:
<tableCitation id="C6C40332FFB2FFB7C5ADFB8D6D59FB6D" box="[493,592,1135,1162]" captionStart="TABLEAU" captionStartId="8.[113,122,220,237]" captionText="TABLEAU 4. — Dimensions (en mm) des métacarpiens rapportés à Helladotherium Gaudry, 1860. Abréviations: dist., distal; prox., proximal." pageId="8" pageNumber="901">Tableau 4</tableCitation>
).Ces différences de proportions s’accordent aussi bien avec la variabilité intraspécifique qu’avec une distinction taxonomique (la forme de Maragha a été distinguée sous le nom de
<taxonomicName id="4C464D0AFFB2FFB7C54AFB0F6FAEFACF" authority="Mecquenem, 1924" authorityName="Mecquenem" authorityYear="1924" class="Mammalia" family="Giraffidae" genus="Helladotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="8" pageNumber="901" phylum="Chordata" rank="species" species="gaudryi">
<emphasis id="B932EA9BFFB2FFB7C54AFB0F6EF1FAE0" box="[266,504,1261,1287]" italics="true" pageId="8" pageNumber="901">Helladotherium gaudryi</emphasis>
Mecquenem, 1924
</taxonomicName>
, et rien ne montre même que le
<taxonomicName id="4C464D0AFFB2FFB7C5A2FAEF6D6DFACF" box="[482,612,1293,1320]" class="Mammalia" family="Giraffidae" kingdom="Animalia" order="Artiodactyla" pageId="8" pageNumber="901" phylum="Chordata" rank="subFamily" subFamily="Sivatheriinae">Sivatheriinae</taxonomicName>
du site iranien appartienne vraiment à ce genre).
</paragraph>
</subSubSection>
</treatment>
</document>
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