398 lines
62 KiB
XML
398 lines
62 KiB
XML
<document id="BDA298FA85CCFDCACB2636D99BD55431" ID-CLB-Dataset="4970" ID-DOI="10.11646/zootaxa.4984.1.23" ID-GBIF-Dataset="ebbdb3a5-8b13-45db-936a-57c3868e18b7" ID-ISSN="1175-5326" ID-Zenodo-Dep="4927365" ID-ZooBank="919D259F-C161-473A-8731-0C20A3EE3793" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623430582221" checkinUser="plazi" docAuthor="Huber, Bernhard A." docDate="2021" docId="03EF879798657D68FF0BFF4E2ADBFF70" docLanguage="en" docName="zootaxa.4984.1.23.pdf" docOrigin="Zootaxa 4984 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Artema martensi Huber 2021, sp.n." docType="treatment" docVersion="7" lastPageNumber="332" masterDocId="FFD6FFEF98677D60FF9CFFD92E45FFA5" masterDocTitle="First Northwest African species of the spider genus Artema, from caves in Morocco with notes on body size in pholcid spiders (Araneae: Pholcidae)" masterLastPageNumber="334" masterPageNumber="324" pageNumber="326" updateTime="1699010424832" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="08B60C80558E8449FED844E679B6581D">
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<mods:title id="855410D054C870212EA98C3E45196912">First Northwest African species of the spider genus Artema, from caves in Morocco with notes on body size in pholcid spiders (Araneae: Pholcidae)</mods:title>
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<mods:name id="C1407446B999F0AC983FBDDB5FBE9B9A" type="personal">
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<mods:roleTerm id="1066AAC661475DCFA5B4BFABECF29951">Author</mods:roleTerm>
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<mods:namePart id="E514E1A115404D4B57B1BD7E044F2B3E">Huber, Bernhard A.</mods:namePart>
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<mods:title id="DD96733D5F173A894F8EC04D03656612">Zootaxa</mods:title>
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<mods:part id="FFE467FE2CB0FD263DF119B8C19C1EEB">
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<mods:date id="34B9D0BDDD49F05B0B668AFABFBF0F56">2021</mods:date>
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<mods:number id="3BFFBE5C83A3A7192E8D8C0DE15AD858">2021-06-10</mods:number>
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<mods:number id="ED0F35D6541BA0D8A54A8D6E95933A16">4984</mods:number>
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<mods:start id="71090B016DE4A2DB64EA0BE31C555502">324</mods:start>
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<mods:classification id="CC837CF339144256F503E58B54831FA3">journal article</mods:classification>
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<mods:identifier id="09F75CF35330363BC5F4AD73AAC8B5B0" type="CLB-Dataset">4970</mods:identifier>
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<mods:identifier id="B29F729C9B1CCB4A35BF0FA8058A815C" type="DOI">10.11646/zootaxa.4984.1.23</mods:identifier>
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<mods:identifier id="963AFBD145CDC484A5BFD0E632CA7021" type="GBIF-Dataset">ebbdb3a5-8b13-45db-936a-57c3868e18b7</mods:identifier>
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<mods:identifier id="4B715C9788D0ACF2B7F4E3DE928E7716" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="A286CED5B4AB93F454E0B4FF1E50A949" type="Zenodo-Dep">4927365</mods:identifier>
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<mods:identifier id="37BBC4B0F869149EEEEE6DEEFB32EA5F" type="ZooBank">919D259F-C161-473A-8731-0C20A3EE3793</mods:identifier>
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</mods:mods>
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<treatment id="03EF879798657D68FF0BFF4E2ADBFF70" ID-DOI="http://doi.org/10.5281/zenodo.4927383" ID-GBIF-Taxon="184162248" ID-Zenodo-Dep="4927383" LSID="urn:lsid:plazi:treatment:03EF879798657D68FF0BFF4E2ADBFF70" httpUri="http://treatment.plazi.org/id/03EF879798657D68FF0BFF4E2ADBFF70" lastPageId="8" lastPageNumber="332" pageId="2" pageNumber="326">
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<subSubSection id="C35C650A98657D62FF0BFF4E2FE1FF14" box="[151,420,151,177]" pageId="2" pageNumber="326" type="nomenclature">
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<paragraph id="8BF9368198657D62FF0BFF4E2FE1FF14" blockId="2.[151,420,151,213]" box="[151,420,151,177]" pageId="2" pageNumber="326">
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<heading id="D0B181ED98657D62FF0BFF4E2FE1FF14" bold="true" box="[151,420,151,177]" fontSize="11" level="1" pageId="2" pageNumber="326" reason="1">
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<emphasis id="B932EA9398657D62FF0BFF4E2FE1FF14" bold="true" box="[151,420,151,177]" pageId="2" pageNumber="326">
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<taxonomicName id="4C464D0298657D62FF0BFF4E2F27FF14" ID-CoL="98YTP" authority="Huber, 2021" authorityName="Huber" authorityYear="2021" box="[151,354,151,177]" class="Arachnida" family="Pholcidae" genus="Artema" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="326" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
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<emphasis id="B932EA9398657D62FF0BFF4E2F27FF14" bold="true" box="[151,354,151,177]" italics="true" pageId="2" pageNumber="326">Artema martensi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20157E898657D62FEF5FF4E2FE1FF14" box="[361,420,151,177]" pageId="2" pageNumber="326" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35C650A98657D62FF0BFF622F42FF70" box="[151,263,187,213]" pageId="2" pageNumber="326" type="description">
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<paragraph id="8BF9368198657D62FF0BFF622F42FF70" blockId="2.[151,420,151,213]" box="[151,263,187,213]" pageId="2" pageNumber="326">
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<figureCitation id="137D2A0498657D62FF0BFF622F42FF70" box="[151,263,187,213]" captionStart-0="FIGURES 1–4" captionStart-1="FIGURES 5–7" captionStart-2="FIGURES 8–13" captionStart-3="FIGURES 14–18" captionStart-4="FIGURES 19–24" captionStartId-0="1.[151,264,1982,2006]" captionStartId-1="2.[151,264,1936,1960]" captionStartId-2="3.[151,264,1828,1852]" captionStartId-3="4.[151,264,1403,1427]" captionStartId-4="5.[151,264,1787,1811]" captionTargetBox-0="[151,1436,869,1958]" captionTargetBox-1="[152,1435,1240,1913]" captionTargetBox-2="[158,1423,278,1793]" captionTargetBox-3="[159,1424,187,1374]" captionTargetBox-4="[151,1436,181,1763]" captionTargetId-0="figure-240@1.[151,1436,869,1958]" captionTargetId-1="figure-425@2.[152,1435,1240,1913]" captionTargetId-2="figure-71@3.[151,1435,273,1805]" captionTargetId-3="figure-16@4.[151,1436,181,1379]" captionTargetId-4="figure-23@5.[151,1436,181,1763]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="5" captionText-0="FIGURES 1–4. Artema martensi sp. n., habitus of living spiders from Morocco, between Lakhssas and Bouizakarne (1–2 holotype male, 3–4 female)." captionText-1="FIGURES 5–7. Artema martensi sp. n. (holotype), left male palp (5 retrolateral, 6 dorsal, 7 prolateral). Scale line: 1 mm." captionText-2="FIGURES 8–13. Artema martensi sp. n. (holotype). 8–9 Male chelicerae (8 frontal, 9 lateral; arrows point at distinctive pair of rounded processes). 10–13 Left procursus (10 retrolateral, 11 dorsal, 12 prolateral, 13 ventral; arrows point at distinctive ventral process). dp-dorsal process, vp—ventral pocket. Scale lines: 1 mm." captionText-3="FIGURES 14–18. Artema martensi sp. n. 14–17 Left genital bulb (holotype) (14 retrolateral, 15 ventral, 16 prolateral, 17 dorsal). 18 Cleared female genitalia (female from “cave 1”), dorsal view; arrow points at cushion-like median structure. a, b, c, d—bulbal processes a–d; em—embolus; bps—proximal bulbal sclerite; pp—pore plate. Scale lines: 1 mm." captionText-4="FIGURES 19–24. Artema martensi sp. n., female from “cave 1”. 19–21 Untreated female genitalia (19 ventral, 20 anterior, 21 lateral; arrows point at distinctive pair of lateral processes on posterior epigynal plate). 22–23 Cleared female genitalia (22 ventral, 23 dorsal; arrow points at cushion-like median structure). 24 Cleared female genitalia, dorsal, with dorsal arc folded backwards. aep—anterior epigynal processes; ep—epigynum (main epigynal plate); dar—dorsal arc; pep—posterior epigynal plate; pp—pore plate; var—ventral arc. Scale lines: 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4927367" figureDoi-1="http://doi.org/10.5281/zenodo.4927369" figureDoi-2="http://doi.org/10.5281/zenodo.4927371" figureDoi-3="http://doi.org/10.5281/zenodo.4927373" figureDoi-4="http://doi.org/10.5281/zenodo.4927375" httpUri-0="https://zenodo.org/record/4927367/files/figure.png" httpUri-1="https://zenodo.org/record/4927369/files/figure.png" httpUri-2="https://zenodo.org/record/4927371/files/figure.png" httpUri-3="https://zenodo.org/record/4927373/files/figure.png" httpUri-4="https://zenodo.org/record/4927375/files/figure.png" pageId="2" pageNumber="326">Figs 1–24</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C35C650A98657D62FF0BFF272DA3FE69" pageId="2" pageNumber="326" type="materials_examined">
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<paragraph id="8BF9368198657D62FF0BFF272A3DFE98" blockId="2.[151,1437,253,1216]" pageId="2" pageNumber="326">
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<emphasis id="B932EA9398657D62FF0BFF272F07FEB2" bold="true" box="[151,322,253,280]" pageId="2" pageNumber="326">
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<collectingRegion id="4982F86398657D62FF0BFF272E97FEBD" box="[151,210,254,280]" pageId="2" pageNumber="326">Type</collectingRegion>
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material.
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</emphasis>
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<materialsCitation id="3B2E3CDC98657D62FEDAFF242A31FE98" ID-GBIF-Occurrence="3334346301" collectingDate="2018-09-15" collectionCode="ZFMK" collectorName="B. A. Huber & H. Belhadj" country="Morocco" elevation="1000" latitude="29.2496" location="Lakhssas" longLatPrecision="6" longitude="-9.7435" pageId="2" pageNumber="326" specimenCount="1" stateProvince="Souss-Massa" typeStatus="holotype">
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<emphasis id="B932EA9398657D62FEDAFF242CC9FEBD" bold="true" box="[326,652,253,280]" pageId="2" pageNumber="326">
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<collectingCountry id="F351761198657D62FEDAFF242FA6FEB2" box="[326,483,253,279]" name="Morocco" pageId="2" pageNumber="326">MOROCCO</collectingCountry>
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:
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||
<emphasis id="B932EA9398657D62FE70FF272CC6FEBD" bold="true" box="[492,643,254,280]" italics="true" pageId="2" pageNumber="326">
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<collectingRegion id="4982F86398657D62FE70FF272CC6FEBD" box="[492,643,254,280]" pageId="2" pageNumber="326">Souss-Massa</collectingRegion>
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||
</emphasis>
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||
:
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</emphasis>
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<typeStatus id="54FD882398657D62FD0DFF272CB6FEBD" box="[657,755,254,280]" pageId="2" pageNumber="326" type="holotype">holotype</typeStatus>
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male (
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<collectionCode id="ED57AE4498657D62FCA5FF272DCEFEBD" box="[825,907,254,280]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="2" pageNumber="326" type="Museum">ZFMK</collectionCode>
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||
,Ar 22208), between
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||
<location id="8E99605A98657D62FBECFF272A92FEBD" LSID="urn:lsid:plazi:treatment:03EF879798657D68FF0BFF4E2ADBFF70:8E99605A98657D62FBECFF272A92FEBD" box="[1136,1239,254,280]" country="Morocco" latitude="29.2496" longLatPrecision="6" longitude="-9.7435" name="Lakhssas" pageId="2" pageNumber="326" stateProvince="Souss-Massa">Lakhssas</location>
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||
and Bouizakarne, “cave 2”,
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||
<geoCoordinate id="EE72504698657D62FE9AFEFB2FC1FE99" box="[262,388,290,316]" degrees="29.2496" direction="north" orientation="latitude" pageId="2" pageNumber="326" precision="5" value="29.2496">29.2496°N</geoCoordinate>
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||
,
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||
<geoCoordinate id="EE72504698657D62FE13FEFB2C47FE98" box="[399,514,290,317]" degrees="9.7435" direction="west" orientation="longitude" pageId="2" pageNumber="326" precision="5" value="-9.7435">9.7435°W</geoCoordinate>
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||
,
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||
<elevation id="006BD1B298657D62FD91FEFB2CDBFE99" box="[525,670,290,316]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="326" unit="m" value="1000.0">
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<quantity id="4CBE9B6498657D62FD91FEFB2C27FE99" box="[525,610,290,316]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="326" unit="m" value="1000.0">1000 m</quantity>
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||
a.s.l.
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||
</elevation>
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,
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||
<collectorName id="26B3535798657D62FD37FEFB2D68FE99" box="[683,813,290,316]" pageId="2" pageNumber="326">B.A. Huber</collectorName>
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&
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<collectorName id="26B3535798657D62FCCCFEFB2D8FFE99" box="[848,970,290,316]" pageId="2" pageNumber="326">H. Belhadj</collectorName>
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leg.
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<date id="FFF8104198657D62FB9DFEFB2A31FE98" box="[1025,1140,290,317]" pageId="2" pageNumber="326" value="2018-09-15">
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<collectingDate id="EFBCE9A998657D62FB9DFEFB2A31FE98" box="[1025,1140,290,317]" pageId="2" pageNumber="326" value="2018-09-15">15.9.2018</collectingDate>
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</date>
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</materialsCitation>
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.
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||
</paragraph>
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<paragraph id="8BF9368198657D62FF5BFE9C2DA3FE69" blockId="2.[151,1437,253,1216]" pageId="2" pageNumber="326">
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<materialsCitation id="3B2E3CDC98657D62FF5BFE9C2AB7FEC5" ID-GBIF-Occurrence="3334346303" box="[199,1266,325,352]" collectionCode="ZFMK" country="Morocco" location="Souss-Massa" pageId="2" pageNumber="326" specimenCount="2" specimenCount-female="2" stateProvince="Souss-Massa">
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<emphasis id="B932EA9398657D62FF5BFE9C2DCAFEC5" bold="true" box="[199,911,325,352]" pageId="2" pageNumber="326">
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||
Additional material examined.
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||
<collectingCountry id="F351761198657D62FDDFFE9C2CA5FEFA" box="[579,736,325,351]" name="Morocco" pageId="2" pageNumber="326">MOROCCO</collectingCountry>
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:
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<emphasis id="B932EA9398657D62FD73FE9F2DC3FEC5" bold="true" box="[751,902,326,352]" italics="true" pageId="2" pageNumber="326">
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<collectingRegion id="4982F86398657D62FD73FE9F2DC3FEC5" box="[751,902,326,352]" pageId="2" pageNumber="326">Souss-Massa</collectingRegion>
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</emphasis>
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:
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||
</emphasis>
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||
<specimenCount id="9D40FD0898657D62FC05FE9F2A4DFEC5" box="[921,1032,326,352]" pageId="2" pageNumber="326" type="female">2 females</specimenCount>
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(
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||
<collectionCode id="ED57AE4498657D62FB85FE9F2A2EFEC5" box="[1049,1131,326,352]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="2" pageNumber="326" type="Museum">ZFMK</collectionCode>
|
||
, Ar 22209)
|
||
</materialsCitation>
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||
,
|
||
<materialsCitation id="3B2E3CDC98657D62FA9CFE9F2C71FE0C" ID-GBIF-Occurrence="3334346304" collectingDate="2018-09-15" collectionCode="ZFMK" collectorName="B. A. Huber & H. Belhadj" country="Morocco" elevation="1000" latitude="29.2488" location="Lakhssas" longLatPrecision="6" longitude="-9.7435" pageId="2" pageNumber="326" specimenCount="3" specimenCount-juvenile="3" stateProvince="Souss-Massa">
|
||
<specimenCount id="9D40FD0898657D62FA9CFE9F2B38FEC5" box="[1280,1405,326,352]" pageId="2" pageNumber="326" type="juvenile">3 juveniles</specimenCount>
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||
in pure ethanol (
|
||
<collectionCode id="ED57AE4498657D62FEACFEB32FC7FE21" box="[304,386,362,388]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="2" pageNumber="326" type="Museum">ZFMK</collectionCode>
|
||
, Mor82), with same data as for holotype, but: “cave 1” (probably another entrance to the same cave system),
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||
<geoCoordinate id="EE72504698657D62FEABFE572FF1FE0D" box="[311,436,398,424]" degrees="29.2488" direction="north" orientation="latitude" pageId="2" pageNumber="326" precision="5" value="29.2488">29.2488°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE72504698657D62FE5CFE572C71FE0C" box="[448,564,398,425]" degrees="9.7435" direction="west" orientation="longitude" pageId="2" pageNumber="326" precision="5" value="-9.7435">9.7435°W</geoCoordinate>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B2E3CDC98657D62FDA2FE572DA7FE69" ID-GBIF-Occurrence="3334346302" collectingDate="2018-09-16" collectionCode="ZFMK" collectorName="B. A. Huber & H. Belhadj" country="Morocco" elevation="1380" latitude="29.8348" longLatPrecision="6" longitude="-9.0247" pageId="2" pageNumber="326" specimenCount="1" specimenCount-female="1" stateProvince="Souss-Massa">
|
||
<specimenCount id="9D40FD0898657D62FDA2FE572CE5FE0D" box="[574,672,398,424]" pageId="2" pageNumber="326" type="female">1 female</specimenCount>
|
||
(
|
||
<collectionCode id="ED57AE4498657D62FD33FE572D44FE0D" box="[687,769,398,424]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="2" pageNumber="326" type="Museum">ZFMK</collectionCode>
|
||
, Ar 22210),
|
||
<collectorName id="26B3535798657D62FC12FE572A17FE0D" box="[910,1106,398,424]" pageId="2" pageNumber="326">SW Ida Ougnidif</collectorName>
|
||
,
|
||
<geoCoordinate id="EE72504698657D62FBC1FE572A9FFE0D" box="[1117,1242,398,424]" degrees="29.8348" direction="north" orientation="latitude" pageId="2" pageNumber="326" precision="5" value="29.8348">29.8348°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE72504698657D62FB7AFE572B1DFE0D" box="[1254,1368,398,424]" degrees="9.0247" direction="west" orientation="longitude" pageId="2" pageNumber="326" precision="5" value="-9.0247">9.0247°W</geoCoordinate>
|
||
,
|
||
<elevation id="006BD1B298657D62FAF8FE572EAFFE69" metricMagnitude="3" metricUnit="m" metricValue="1.38" pageId="2" pageNumber="326" unit="m" value="1380.0">
|
||
<quantity id="4CBE9B6498657D62FAF8FE572EE8FE69" metricMagnitude="3" metricUnit="m" metricValue="1.38" pageId="2" pageNumber="326" unit="m" value="1380.0">1380 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
, small cave (rock shelter),
|
||
<collectorName id="26B3535798657D62FD84FE6B2CDFFE69" box="[536,666,434,460]" pageId="2" pageNumber="326">B.A. Huber</collectorName>
|
||
&
|
||
<collectorName id="26B3535798657D62FD22FE6B2D72FE69" box="[702,823,434,460]" pageId="2" pageNumber="326">H. Belhadj</collectorName>
|
||
leg.
|
||
<date id="FFF8104198657D62FCF2FE6B2DA7FE69" box="[878,994,434,460]" pageId="2" pageNumber="326" value="2018-09-16">
|
||
<collectingDate id="EFBCE9A998657D62FCF2FE6B2DA7FE69" box="[878,994,434,460]" pageId="2" pageNumber="326" value="2018-09-16">16.9.2018</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35C650A98657D62FF5BFE0C2B4DFD25" pageId="2" pageNumber="326" type="diagnosis">
|
||
<paragraph id="8BF9368198657D62FF5BFE0C2B4DFD25" blockId="2.[151,1437,253,1216]" pageId="2" pageNumber="326">
|
||
<emphasis id="B932EA9398657D62FF5BFE0C2F07FE4A" bold="true" box="[199,322,469,495]" pageId="2" pageNumber="326">Diagnosis.</emphasis>
|
||
Easily distinguished from other described congeners by posterior epigynal plate with pair of processes (arrows in
|
||
<figureCitation id="137D2A0498657D62FE92FE232FC9FDB1" box="[270,396,506,532]" captionStart="FIGURES 19–24" captionStartId="5.[151,264,1787,1811]" captionTargetBox="[151,1436,181,1763]" captionTargetId="figure-23@5.[151,1436,181,1763]" captionTargetPageId="5" captionText="FIGURES 19–24. Artema martensi sp. n., female from “cave 1”. 19–21 Untreated female genitalia (19 ventral, 20 anterior, 21 lateral; arrows point at distinctive pair of lateral processes on posterior epigynal plate). 22–23 Cleared female genitalia (22 ventral, 23 dorsal; arrow points at cushion-like median structure). 24 Cleared female genitalia, dorsal, with dorsal arc folded backwards. aep—anterior epigynal processes; ep—epigynum (main epigynal plate); dar—dorsal arc; pep—posterior epigynal plate; pp—pore plate; var—ventral arc. Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4927375" httpUri="https://zenodo.org/record/4927375/files/figure.png" pageId="2" pageNumber="326">Figs 20–21</figureCitation>
|
||
) and by male chelicerae with pair of proximal rounded processes (arrows in
|
||
<figureCitation id="137D2A0498657D62FB4AFE232B7FFDB1" box="[1238,1338,506,532]" captionStart="FIGURES 8–13" captionStartId="3.[151,264,1828,1852]" captionTargetBox="[158,1423,278,1793]" captionTargetId="figure-71@3.[151,1435,273,1805]" captionTargetPageId="3" captionText="FIGURES 8–13. Artema martensi sp. n. (holotype). 8–9 Male chelicerae (8 frontal, 9 lateral; arrows point at distinctive pair of rounded processes). 10–13 Left procursus (10 retrolateral, 11 dorsal, 12 prolateral, 13 ventral; arrows point at distinctive ventral process). dp-dorsal process, vp—ventral pocket. Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4927371" httpUri="https://zenodo.org/record/4927371/files/figure.png" pageId="2" pageNumber="326">Figs 8–9</figureCitation>
|
||
); also by the combination of the following characters: procursus without dorsal notch (
|
||
<figureCitation id="137D2A0498657D62FC6FFDC72A01FD9D" box="[1011,1092,542,568]" captionStart="FIGURES 8–13" captionStartId="3.[151,264,1828,1852]" captionTargetBox="[158,1423,278,1793]" captionTargetId="figure-71@3.[151,1435,273,1805]" captionTargetPageId="3" captionText="FIGURES 8–13. Artema martensi sp. n. (holotype). 8–9 Male chelicerae (8 frontal, 9 lateral; arrows point at distinctive pair of rounded processes). 10–13 Left procursus (10 retrolateral, 11 dorsal, 12 prolateral, 13 ventral; arrows point at distinctive ventral process). dp-dorsal process, vp—ventral pocket. Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4927371" httpUri="https://zenodo.org/record/4927371/files/figure.png" pageId="2" pageNumber="326">Fig. 11</figureCitation>
|
||
; cf. figs 20, 40 in
|
||
<bibRefCitation id="EFD74B7098657D62FA96FDC72E91FDF9" author="Aharon, S. & Huber, B. A. & Gavish-Regev, E." pageId="2" pageNumber="326" pagination="1 - 57" refId="ref3688" refString="Aharon, S., Huber, B. A. & Gavish-Regev, E. (2017) Daddy-long-leg giants: revision of the spider genus Artema Walckenaer, 1837 (Araneae, Pholcidae). European Journal of Taxonomy, 376, 1 - 57. https: // doi. org / 10.5852 / ejt. 2017.376" type="journal article" year="2017">
|
||
Aharon
|
||
<emphasis id="B932EA9398657D62FAF9FDC62BD8FD9D" box="[1381,1437,542,568]" italics="true" pageId="2" pageNumber="326">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
); procursus with large ventral distal apophysis (arrow in
|
||
<figureCitation id="137D2A0498657D62FCD0FD9B2DD8FDF9" box="[844,925,578,604]" captionStart="FIGURES 8–13" captionStartId="3.[151,264,1828,1852]" captionTargetBox="[158,1423,278,1793]" captionTargetId="figure-71@3.[151,1435,273,1805]" captionTargetPageId="3" captionText="FIGURES 8–13. Artema martensi sp. n. (holotype). 8–9 Male chelicerae (8 frontal, 9 lateral; arrows point at distinctive pair of rounded processes). 10–13 Left procursus (10 retrolateral, 11 dorsal, 12 prolateral, 13 ventral; arrows point at distinctive ventral process). dp-dorsal process, vp—ventral pocket. Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4927371" httpUri="https://zenodo.org/record/4927371/files/figure.png" pageId="2" pageNumber="326">Fig. 13</figureCitation>
|
||
); bulbal process c distinct, without prominent scales or hooks (
|
||
<figureCitation id="137D2A0498657D62FECEFDBF2FE0FD25" box="[338,421,614,640]" captionStart="FIGURES 14–18" captionStartId="4.[151,264,1403,1427]" captionTargetBox="[159,1424,187,1374]" captionTargetId="figure-16@4.[151,1436,181,1379]" captionTargetPageId="4" captionText="FIGURES 14–18. Artema martensi sp. n. 14–17 Left genital bulb (holotype) (14 retrolateral, 15 ventral, 16 prolateral, 17 dorsal). 18 Cleared female genitalia (female from “cave 1”), dorsal view; arrow points at cushion-like median structure. a, b, c, d—bulbal processes a–d; em—embolus; bps—proximal bulbal sclerite; pp—pore plate. Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4927373" httpUri="https://zenodo.org/record/4927373/files/figure.png" pageId="2" pageNumber="326">Fig. 14</figureCitation>
|
||
); main epigynal plate bean-shaped, without distinct median sclerite (
|
||
<figureCitation id="137D2A0498657D62FB3BFDBF2ABEFD25" box="[1191,1275,614,640]" captionStart="FIGURES 19–24" captionStartId="5.[151,264,1787,1811]" captionTargetBox="[151,1436,181,1763]" captionTargetId="figure-23@5.[151,1436,181,1763]" captionTargetPageId="5" captionText="FIGURES 19–24. Artema martensi sp. n., female from “cave 1”. 19–21 Untreated female genitalia (19 ventral, 20 anterior, 21 lateral; arrows point at distinctive pair of lateral processes on posterior epigynal plate). 22–23 Cleared female genitalia (22 ventral, 23 dorsal; arrow points at cushion-like median structure). 24 Cleared female genitalia, dorsal, with dorsal arc folded backwards. aep—anterior epigynal processes; ep—epigynum (main epigynal plate); dar—dorsal arc; pep—posterior epigynal plate; pp—pore plate; var—ventral arc. Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4927375" httpUri="https://zenodo.org/record/4927375/files/figure.png" pageId="2" pageNumber="326">Fig. 19</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35C650A98657D62FF5BFD532A71FD6D" pageId="2" pageNumber="326" type="etymology">
|
||
<paragraph id="8BF9368198657D62FF5BFD532A71FD6D" blockId="2.[151,1437,253,1216]" pageId="2" pageNumber="326">
|
||
<emphasis id="B932EA9398657D62FF5BFD532F0BFD01" bold="true" box="[199,334,650,676]" pageId="2" pageNumber="326">Etymology.</emphasis>
|
||
Named in honor of Jochen Martens in recognition of his contributions to arachnology, in particular his diligent analysis of harvestmen genitalia beyond their species diagnostic value.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35C650A98657D67FF5BFD082A76F84A" lastPageId="7" lastPageNumber="331" pageId="2" pageNumber="326" type="description">
|
||
<paragraph id="8BF9368198657D62FF5BFD082CF3FCFD" blockId="2.[151,1437,253,1216]" pageId="2" pageNumber="326">
|
||
<emphasis id="B932EA9398657D62FF5BFD082FDBFD49" bold="true" box="[199,414,721,748]" pageId="2" pageNumber="326">Description. Male</emphasis>
|
||
(
|
||
<typeStatus id="54FD882398657D62FE32FD0B2C52FD48" box="[430,535,722,749]" pageId="2" pageNumber="326" type="holotype">holotype</typeStatus>
|
||
):
|
||
<emphasis id="B932EA9398657D62FDB7FD0A2C88FD49" box="[555,717,723,748]" italics="true" pageId="2" pageNumber="326">Measurements</emphasis>
|
||
. Total length 9.3, carapace width 4.5. Distance PME-PME
|
||
<quantity id="4CBE9B6498657D62FAEEFD0B2E85FCB5" metricMagnitude="-1" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="326" unit="mm" value="220.0">220 µm</quantity>
|
||
; diameter PME 210x
|
||
<quantity id="4CBE9B6498657D62FE36FD2F2C46FCB5" box="[426,515,758,784]" metricMagnitude="-1" metricUnit="m" metricValue="2.3" pageId="2" pageNumber="326" unit="mm" value="230.0">230 µm</quantity>
|
||
; distance PME-ALE
|
||
<quantity id="4CBE9B6498657D62FD73FD2F2D0DFCB5" box="[751,840,758,784]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="326" unit="mm" value="100.0">100 µm</quantity>
|
||
; diameter AME
|
||
<quantity id="4CBE9B6498657D62FC62FD2F2A12FCB5" box="[1022,1111,758,784]" metricMagnitude="-1" metricUnit="m" metricValue="2.3" pageId="2" pageNumber="326" unit="mm" value="230.0">230 µm</quantity>
|
||
; distance AME-AME
|
||
<quantity id="4CBE9B6498657D62FAD1FD2F2BDDFCB5" box="[1357,1432,758,784]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="326" unit="mm" value="60.0">60 µm</quantity>
|
||
. Leg 1: 68.0 (17.1 + 2.1 + 19.2 + 24.9 + 4.7), tibia 2: 14.9, tibia 3: 11.2, tibia 4: 14.3; tibia 1 L/d: 42; femora 1–4 diameters (at half length): 0.70, 0.72, 0.72, 0.76.
|
||
</paragraph>
|
||
<paragraph id="8BF9368198657D62FF5BFCBB2F08FB95" blockId="2.[151,1437,253,1216]" pageId="2" pageNumber="326">
|
||
<emphasis id="B932EA9398657D62FF5BFCBB2F4DFCD9" box="[199,264,866,892]" italics="true" pageId="2" pageNumber="326">Color</emphasis>
|
||
(in ethanol). Carapace pale ochre-yellow, with brown median mark and three pairs of lateral marks (anterior pair very small); ocular area laterally light brown; clypeus with brown median band; sternum ochre-yellow, with dark brown margins and dark ochre median mark; legs ochre-yellow, metatarsi and tarsi light brown; femur 1 proximally red-brown; femora and tibiae with light tips and dark subdistal rings, additional dark ring on patellae + tibiae proximally; abdomen pale grey, dorsally and laterally with distinct pattern of large dark marks; ventrally monochromous.
|
||
</paragraph>
|
||
<paragraph id="8BF9368198657D62FF5BFBE32F9BFB65" blockId="2.[151,1437,253,1216]" pageId="2" pageNumber="326">
|
||
<emphasis id="B932EA9398657D62FF5BFBE32F44FBF1" box="[199,257,1082,1108]" italics="true" pageId="2" pageNumber="326">Body</emphasis>
|
||
. Habitus as in
|
||
<figureCitation id="137D2A0498657D62FE2FFBE32C5FFBF1" box="[435,538,1082,1108]" captionStart="FIGURES 1–4" captionStartId="1.[151,264,1982,2006]" captionTargetBox="[151,1436,869,1958]" captionTargetId="figure-240@1.[151,1436,869,1958]" captionTargetPageId="1" captionText="FIGURES 1–4. Artema martensi sp. n., habitus of living spiders from Morocco, between Lakhssas and Bouizakarne (1–2 holotype male, 3–4 female)." figureDoi="http://doi.org/10.5281/zenodo.4927367" httpUri="https://zenodo.org/record/4927367/files/figure.png" pageId="2" pageNumber="326">Figs 1–2</figureCitation>
|
||
. Ocular area slightly elevated. Deep thoracic pit transforming posteriorly into shallow median furrow. Clypeus unmodified but rim more sclerotized than in female. Sternum wider than long (2.6/2.1), unmodified. Abdomen globular, slightly higher than long and laterally slightly compressed; gonopore with four epiandrous spigots.
|
||
</paragraph>
|
||
<caption id="DF39660998657D62FF0BF8492B20F80D" ID-DOI="http://doi.org/10.5281/zenodo.4927369" ID-Zenodo-Dep="4927369" box="[151,1381,1936,1961]" httpUri="https://zenodo.org/record/4927369/files/figure.png" pageId="2" pageNumber="326" startId="2.[151,264,1936,1960]" targetBox="[152,1435,1240,1913]" targetPageId="2">
|
||
<paragraph id="8BF9368198657D62FF0BF8492B20F80D" blockId="2.[151,1381,1936,1961]" box="[151,1381,1936,1961]" pageId="2" pageNumber="326">
|
||
<emphasis id="B932EA9398657D62FF0BF8492F7EF80C" bold="true" box="[151,315,1936,1961]" pageId="2" pageNumber="326">FIGURES 5–7.</emphasis>
|
||
<taxonomicName id="4C464D0298657D62FEDDF8482FAEF80D" authority="Huber, 2021" authorityName="Huber" authorityYear="2021" box="[321,491,1937,1960]" class="Arachnida" family="Pholcidae" genus="Artema" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="326" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
|
||
<emphasis id="B932EA9398657D62FEDDF8482FAEF80D" box="[321,491,1937,1960]" italics="true" pageId="2" pageNumber="326">Artema martensi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20157E898657D62FE6DF8482C62F80D" box="[497,551,1937,1960]" pageId="2" pageNumber="326" rank="species">sp. n.</taxonomicNameLabel>
|
||
(holotype), left male palp (5 retrolateral, 6 dorsal, 7 prolateral). Scale line: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF9368198647D63FF5BFF4E2A81FF5C" blockId="3.[151,1436,151,249]" pageId="3" pageNumber="327">
|
||
<emphasis id="B932EA9398647D63FF5BFF4E2F07FF14" box="[199,322,151,177]" italics="true" pageId="3" pageNumber="327">Chelicerae</emphasis>
|
||
. As in
|
||
<figureCitation id="137D2A0498647D63FE09FF4E2FBCFF14" box="[405,505,151,177]" captionStart="FIGURES 8–13" captionStartId="3.[151,264,1828,1852]" captionTargetBox="[158,1423,278,1793]" captionTargetId="figure-71@3.[151,1435,273,1805]" captionTargetPageId="3" captionText="FIGURES 8–13. Artema martensi sp. n. (holotype). 8–9 Male chelicerae (8 frontal, 9 lateral; arrows point at distinctive pair of rounded processes). 10–13 Left procursus (10 retrolateral, 11 dorsal, 12 prolateral, 13 ventral; arrows point at distinctive ventral process). dp-dorsal process, vp—ventral pocket. Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4927371" httpUri="https://zenodo.org/record/4927371/files/figure.png" pageId="3" pageNumber="327">Figs 8–9</figureCitation>
|
||
, distally with pair of strongly protruding frontal processes set with large modified (cone-shaped) hairs (25 and 27, respectively); with additional pair of rounded processes more proximally (arrows in
|
||
<figureCitation id="137D2A0498647D63FF0BFF062F66FF5C" box="[151,291,223,249]" captionStart="FIGURES 8–13" captionStartId="3.[151,264,1828,1852]" captionTargetBox="[158,1423,278,1793]" captionTargetId="figure-71@3.[151,1435,273,1805]" captionTargetPageId="3" captionText="FIGURES 8–13. Artema martensi sp. n. (holotype). 8–9 Male chelicerae (8 frontal, 9 lateral; arrows point at distinctive pair of rounded processes). 10–13 Left procursus (10 retrolateral, 11 dorsal, 12 prolateral, 13 ventral; arrows point at distinctive ventral process). dp-dorsal process, vp—ventral pocket. Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4927371" httpUri="https://zenodo.org/record/4927371/files/figure.png" pageId="3" pageNumber="327">Figs 8 and 9</figureCitation>
|
||
); lateral stridulatory ridges very indistinct, barely visible in dissecting microscope.
|
||
</paragraph>
|
||
<caption id="DF39660998647D63FF0BF8FD2D06F821" ID-DOI="http://doi.org/10.5281/zenodo.4927371" ID-Zenodo-Dep="4927371" httpUri="https://zenodo.org/record/4927371/files/figure.png" pageId="3" pageNumber="327" startId="3.[151,264,1828,1852]" targetBox="[158,1423,278,1793]" targetPageId="3">
|
||
<paragraph id="8BF9368198647D63FF0BF8FD2D06F821" blockId="3.[151,1436,1828,1925]" pageId="3" pageNumber="327">
|
||
<emphasis id="B932EA9398647D63FF0BF8FD2F02F899" bold="true" box="[151,327,1828,1852]" pageId="3" pageNumber="327">FIGURES 8–13.</emphasis>
|
||
<taxonomicName id="4C464D0298647D63FED1F8FC2FB2F899" authority="Huber, 2021" authorityName="Huber" authorityYear="2021" box="[333,503,1829,1852]" class="Arachnida" family="Pholcidae" genus="Artema" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="327" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
|
||
<emphasis id="B932EA9398647D63FED1F8FC2FB2F899" box="[333,503,1829,1852]" italics="true" pageId="3" pageNumber="327">Artema martensi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20157E898647D63FE61F8FC2C76F899" box="[509,563,1829,1852]" pageId="3" pageNumber="327" rank="species">sp. n.</taxonomicNameLabel>
|
||
(holotype). 8–9 Male chelicerae (8 frontal, 9 lateral; arrows point at distinctive pair of rounded processes). 10–13 Left procursus (10 retrolateral, 11 dorsal, 12 prolateral, 13 ventral; arrows point at distinctive ventral process). dp-dorsal process, vp—ventral pocket. Scale lines: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF39660998637D64FF0BFAA22A97FA7E" ID-DOI="http://doi.org/10.5281/zenodo.4927373" ID-Zenodo-Dep="4927373" httpUri="https://zenodo.org/record/4927373/files/figure.png" pageId="4" pageNumber="328" startId="4.[151,264,1403,1427]" targetBox="[159,1424,187,1374]" targetPageId="4">
|
||
<paragraph id="8BF9368198637D64FF0BFAA22A97FA7E" blockId="4.[151,1437,1403,1499]" pageId="4" pageNumber="328">
|
||
<emphasis id="B932EA9398637D64FF0BFAA22F12FA36" bold="true" box="[151,343,1403,1427]" pageId="4" pageNumber="328">FIGURES 14–18.</emphasis>
|
||
<taxonomicName id="4C464D0298637D64FEFCFAA22C48FA36" authority="Huber, 2021" authorityName="Huber" authorityYear="2021" box="[352,525,1403,1427]" class="Arachnida" family="Pholcidae" genus="Artema" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="328" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
|
||
<emphasis id="B932EA9398637D64FEFCFAA22C48FA36" box="[352,525,1403,1427]" italics="true" pageId="4" pageNumber="328">Artema martensi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20157E898637D64FD8AFAA52C15FA36" box="[534,592,1404,1427]" pageId="4" pageNumber="328" rank="species">sp. n.</taxonomicNameLabel>
|
||
14–17 Left genital bulb (holotype) (14 retrolateral, 15 ventral, 16 prolateral, 17 dorsal). 18 Cleared female genitalia (female from “cave 1”), dorsal view; arrow points at cushion-like median structure. a, b, c, d—bulbal processes a–d; em—embolus; bps—proximal bulbal sclerite; pp—pore plate. Scale lines: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF9368198637D64FF5BF9D02DA7F8E6" blockId="4.[151,1437,1545,2040]" pageId="4" pageNumber="328">
|
||
<emphasis id="B932EA9398637D64FF5BF9D02F42F986" box="[199,263,1545,1571]" italics="true" pageId="4" pageNumber="328">Palps</emphasis>
|
||
. As is
|
||
<figureCitation id="137D2A0498637D64FEC1F9D02F80F981" box="[349,453,1545,1572]" captionStart="FIGURES 5–7" captionStartId="2.[151,264,1936,1960]" captionTargetBox="[152,1435,1240,1913]" captionTargetId="figure-425@2.[152,1435,1240,1913]" captionTargetPageId="2" captionText="FIGURES 5–7. Artema martensi sp. n. (holotype), left male palp (5 retrolateral, 6 dorsal, 7 prolateral). Scale line: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4927369" httpUri="https://zenodo.org/record/4927369/files/figure.png" pageId="4" pageNumber="328">Figs 5–7</figureCitation>
|
||
; coxa unmodified; trochanter ventrally slightly protruding; femur very large, distally strongly widened, proximally with retrolateral ventral process, whitish ventral area, and very small prolateral stridulatory pick (modified hair); patella very short but ventrally closed; femur-patella joints in dorsal position (i.e., not shifted to one side), very close together; tibia-tarsus joints shifted toward retrolateral side; tarsus without macrotrichia; procursus (
|
||
<figureCitation id="137D2A0498637D64FE25F9402C7BF916" box="[441,574,1689,1715]" captionStart="FIGURES 8–13" captionStartId="3.[151,264,1828,1852]" captionTargetBox="[158,1423,278,1793]" captionTargetId="figure-71@3.[151,1435,273,1805]" captionTargetPageId="3" captionText="FIGURES 8–13. Artema martensi sp. n. (holotype). 8–9 Male chelicerae (8 frontal, 9 lateral; arrows point at distinctive pair of rounded processes). 10–13 Left procursus (10 retrolateral, 11 dorsal, 12 prolateral, 13 ventral; arrows point at distinctive ventral process). dp-dorsal process, vp—ventral pocket. Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4927371" httpUri="https://zenodo.org/record/4927371/files/figure.png" pageId="4" pageNumber="328">Figs 10–13</figureCitation>
|
||
) short, with distinct dorsal process and indistinct ventral pocket, distally strongly widened (in dorsal view), without notch, with whitish distal cushion-like structure; distally on ventral side with distinctive apophysis (arrows in
|
||
<figureCitation id="137D2A0498637D64FDDAF9382CB1F95E" box="[582,756,1761,1787]" captionStart="FIGURES 8–13" captionStartId="3.[151,264,1828,1852]" captionTargetBox="[158,1423,278,1793]" captionTargetId="figure-71@3.[151,1435,273,1805]" captionTargetPageId="3" captionText="FIGURES 8–13. Artema martensi sp. n. (holotype). 8–9 Male chelicerae (8 frontal, 9 lateral; arrows point at distinctive pair of rounded processes). 10–13 Left procursus (10 retrolateral, 11 dorsal, 12 prolateral, 13 ventral; arrows point at distinctive ventral process). dp-dorsal process, vp—ventral pocket. Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4927371" httpUri="https://zenodo.org/record/4927371/files/figure.png" pageId="4" pageNumber="328">Figs 10 and 13</figureCitation>
|
||
); genital bulb (
|
||
<figureCitation id="137D2A0498637D64FC38F9382A63F95E" box="[932,1062,1761,1787]" captionStart="FIGURES 14–18" captionStartId="4.[151,264,1403,1427]" captionTargetBox="[159,1424,187,1374]" captionTargetId="figure-16@4.[151,1436,181,1379]" captionTargetPageId="4" captionText="FIGURES 14–18. Artema martensi sp. n. 14–17 Left genital bulb (holotype) (14 retrolateral, 15 ventral, 16 prolateral, 17 dorsal). 18 Cleared female genitalia (female from “cave 1”), dorsal view; arrow points at cushion-like median structure. a, b, c, d—bulbal processes a–d; em—embolus; bps—proximal bulbal sclerite; pp—pore plate. Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4927373" httpUri="https://zenodo.org/record/4927373/files/figure.png" pageId="4" pageNumber="328">Figs 14–17</figureCitation>
|
||
) with spiraling proximal sclerite and several distal processes: processes a and b large, with whitish embolus in-between; process c distinct and sclerotized, with very small and indistinct scales; process d small and light.
|
||
</paragraph>
|
||
<paragraph id="8BF9368198637D64FF5BF8972D98F852" blockId="4.[151,1437,1545,2040]" pageId="4" pageNumber="328">
|
||
<emphasis id="B932EA9398637D64FF5BF8972EB9F8C2" box="[199,252,1870,1895]" italics="true" pageId="4" pageNumber="328">Legs</emphasis>
|
||
. Without spines; with weakly curved hairs on all tibiae; few vertical hairs; with
|
||
<specimenCount id="9D40FD0898637D64FB0BF8942B57F8CD" box="[1175,1298,1869,1896]" pageId="4" pageNumber="328" type="generic" typeStatus="types">three types</specimenCount>
|
||
of ‘regular’ mechanoreceptors (i.e., except trichobothria and short vertical hairs): very long and strong hairs (up to
|
||
<quantity id="4CBE9B6498637D64FAD8F8A82BD8F829" box="[1348,1437,1905,1932]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="4" pageNumber="328" unit="mm" value="2.4">2.4 mm</quantity>
|
||
long, ~
|
||
<quantity id="4CBE9B6498637D64FF7AF84C2F74F815" box="[230,305,1941,1968]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="328" unit="mm" value="30.0">30 µm</quantity>
|
||
thick at basis), much shorter strong hairs (~
|
||
<quantity id="4CBE9B6498637D64FCB9F84C2DEAF815" box="[805,943,1941,1968]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="4" pageNumber="328" unit="mm" value="0.2" valueMax="0.3" valueMin="0.1">0.1–0.3 mm</quantity>
|
||
long,
|
||
<quantity id="4CBE9B6498637D64FC64F84C2A29F815" box="[1016,1132,1941,1968]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="4" pageNumber="328" unit="mm" value="15.0" valueMax="20.0" valueMin="10.0">10–20 µm</quantity>
|
||
thick), and very small and thin hairs, in high density (~
|
||
<quantity id="4CBE9B6498637D64FE7DF8602C7EF871" box="[481,571,1977,2004]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="328" unit="mm" value="0.1">0.1 mm</quantity>
|
||
long, ~
|
||
<quantity id="4CBE9B6498637D64FD0BF8602C91F871" box="[663,724,1977,2004]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="328" unit="mm" value="5.0">5 µm</quantity>
|
||
thick); retrolateral trichobothrium of tibia 1 at 8%; prolateral trichobothrium present on all tibiae; tarsal pseudosegmentation not visible.
|
||
</paragraph>
|
||
<caption id="DF39660998627D65FF0BF9222CACF806" ID-DOI="http://doi.org/10.5281/zenodo.4927375" ID-Zenodo-Dep="4927375" httpUri="https://zenodo.org/record/4927375/files/figure.png" pageId="5" pageNumber="329" startId="5.[151,264,1787,1811]" targetBox="[151,1436,181,1763]" targetPageId="5">
|
||
<paragraph id="8BF9368198627D65FF0BF9222CACF806" blockId="5.[151,1437,1787,1956]" pageId="5" pageNumber="329">
|
||
<emphasis id="B932EA9398627D65FF0BF9222F13F8B6" bold="true" box="[151,342,1787,1811]" pageId="5" pageNumber="329">FIGURES 19–24.</emphasis>
|
||
<taxonomicName id="4C464D0298627D65FEC2F9252C4FF8B6" authority="Huber, 2021" authorityName="Huber" authorityYear="2021" box="[350,522,1788,1811]" class="Arachnida" family="Pholcidae" genus="Artema" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="329" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
|
||
<emphasis id="B932EA9398627D65FEC2F9252C4FF8B6" box="[350,522,1788,1811]" italics="true" pageId="5" pageNumber="329">Artema martensi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20157E898627D65FD8EF9252C0EF8B6" box="[530,587,1788,1811]" pageId="5" pageNumber="329" rank="species">sp. n.</taxonomicNameLabel>
|
||
, female from “cave 1”. 19–21 Untreated female genitalia (19 ventral, 20 anterior, 21 lateral; arrows point at distinctive pair of lateral processes on posterior epigynal plate). 22–23 Cleared female genitalia (22 ventral, 23 dorsal; arrow points at cushion-like median structure). 24 Cleared female genitalia, dorsal, with dorsal arc folded backwards. aep—anterior epigynal processes; ep—epigynum (main epigynal plate); dar—dorsal arc; pep—posterior epigynal plate; pp—pore plate; var—ventral arc. Scale lines: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF9368198617D66FF5BFF4E2C55FE0B" blockId="6.[151,1437,151,430]" pageId="6" pageNumber="330">
|
||
<emphasis id="B932EA9398617D66FF5BFF4E2F6DFF14" bold="true" box="[199,296,151,177]" pageId="6" pageNumber="330">Female:</emphasis>
|
||
In general similar to male (
|
||
<figureCitation id="137D2A0498617D66FDFCFF4E2C81FF14" box="[608,708,151,177]" captionStart="FIGURES 1–4" captionStartId="1.[151,264,1982,2006]" captionTargetBox="[151,1436,869,1958]" captionTargetId="figure-240@1.[151,1436,869,1958]" captionTargetPageId="1" captionText="FIGURES 1–4. Artema martensi sp. n., habitus of living spiders from Morocco, between Lakhssas and Bouizakarne (1–2 holotype male, 3–4 female)." figureDoi="http://doi.org/10.5281/zenodo.4927367" httpUri="https://zenodo.org/record/4927367/files/figure.png" pageId="6" pageNumber="330">Figs 3–4</figureCitation>
|
||
) but chelicerae with very distinct stridulatory files and leg tibiae with additional (less distinct) dark ring at ~1/3 of length. Tibia
|
||
<quantity id="4CBE9B6498617D66FCC7FF622DC3FF70" box="[859,902,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="6" pageNumber="330" unit="in" value="1.0">1 in</quantity>
|
||
<specimenCount id="9D40FD0898617D66FC11FF622DBBFF70" box="[909,1022,187,213]" pageId="6" pageNumber="330" type="female">2 females</specimenCount>
|
||
: 14.0, 18.3 (missing in third female). Epigynum as in
|
||
<figureCitation id="137D2A0498617D66FED0FF062F8EFF5C" box="[332,459,223,249]" captionStart="FIGURES 19–24" captionStartId="5.[151,264,1787,1811]" captionTargetBox="[151,1436,181,1763]" captionTargetId="figure-23@5.[151,1436,181,1763]" captionTargetPageId="5" captionText="FIGURES 19–24. Artema martensi sp. n., female from “cave 1”. 19–21 Untreated female genitalia (19 ventral, 20 anterior, 21 lateral; arrows point at distinctive pair of lateral processes on posterior epigynal plate). 22–23 Cleared female genitalia (22 ventral, 23 dorsal; arrow points at cushion-like median structure). 24 Cleared female genitalia, dorsal, with dorsal arc folded backwards. aep—anterior epigynal processes; ep—epigynum (main epigynal plate); dar—dorsal arc; pep—posterior epigynal plate; pp—pore plate; var—ventral arc. Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4927375" httpUri="https://zenodo.org/record/4927375/files/figure.png" pageId="6" pageNumber="330">Figs 19–21</figureCitation>
|
||
, main epigynal plate semi-circular to bean-shaped, weakly protruding, with pair of dark areas laterally; posterior plate large, with distinctive pair of lateral processes (arrows in
|
||
<figureCitation id="137D2A0498617D66FB1EFEDA2B6BFEB8" box="[1154,1326,259,285]" captionStart="FIGURES 19–24" captionStartId="5.[151,264,1787,1811]" captionTargetBox="[151,1436,181,1763]" captionTargetId="figure-23@5.[151,1436,181,1763]" captionTargetPageId="5" captionText="FIGURES 19–24. Artema martensi sp. n., female from “cave 1”. 19–21 Untreated female genitalia (19 ventral, 20 anterior, 21 lateral; arrows point at distinctive pair of lateral processes on posterior epigynal plate). 22–23 Cleared female genitalia (22 ventral, 23 dorsal; arrow points at cushion-like median structure). 24 Cleared female genitalia, dorsal, with dorsal arc folded backwards. aep—anterior epigynal processes; ep—epigynum (main epigynal plate); dar—dorsal arc; pep—posterior epigynal plate; pp—pore plate; var—ventral arc. Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4927375" httpUri="https://zenodo.org/record/4927375/files/figure.png" pageId="6" pageNumber="330">Figs 20 and 21</figureCitation>
|
||
) (slightly smaller in female from SW Ida Ougnidif); with pair of distinct elevations (“anterior epigynal processes”) in front of epigynum. Internal genitalia (
|
||
<figureCitation id="137D2A0498617D66FE7EFE922C27FEC0" box="[482,610,331,357]" captionStart="FIGURES 19–24" captionStartId="5.[151,264,1787,1811]" captionTargetBox="[151,1436,181,1763]" captionTargetId="figure-23@5.[151,1436,181,1763]" captionTargetPageId="5" captionText="FIGURES 19–24. Artema martensi sp. n., female from “cave 1”. 19–21 Untreated female genitalia (19 ventral, 20 anterior, 21 lateral; arrows point at distinctive pair of lateral processes on posterior epigynal plate). 22–23 Cleared female genitalia (22 ventral, 23 dorsal; arrow points at cushion-like median structure). 24 Cleared female genitalia, dorsal, with dorsal arc folded backwards. aep—anterior epigynal processes; ep—epigynum (main epigynal plate); dar—dorsal arc; pep—posterior epigynal plate; pp—pore plate; var—ventral arc. Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4927375" httpUri="https://zenodo.org/record/4927375/files/figure.png" pageId="6" pageNumber="330">Figs 22–24</figureCitation>
|
||
) with pair of elongated brown pore plates, mostly membranous dorsal arc with cushion-like median structure (arrow in
|
||
<figureCitation id="137D2A0498617D66FD0BFEB62CAFFE2C" box="[663,746,367,393]" captionStart="FIGURES 19–24" captionStartId="5.[151,264,1787,1811]" captionTargetBox="[151,1436,181,1763]" captionTargetId="figure-23@5.[151,1436,181,1763]" captionTargetPageId="5" captionText="FIGURES 19–24. Artema martensi sp. n., female from “cave 1”. 19–21 Untreated female genitalia (19 ventral, 20 anterior, 21 lateral; arrows point at distinctive pair of lateral processes on posterior epigynal plate). 22–23 Cleared female genitalia (22 ventral, 23 dorsal; arrow points at cushion-like median structure). 24 Cleared female genitalia, dorsal, with dorsal arc folded backwards. aep—anterior epigynal processes; ep—epigynum (main epigynal plate); dar—dorsal arc; pep—posterior epigynal plate; pp—pore plate; var—ventral arc. Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4927375" httpUri="https://zenodo.org/record/4927375/files/figure.png" pageId="6" pageNumber="330">Fig. 23</figureCitation>
|
||
), ventral arc partly sclerotized, with pair of ventral processes (directed towards epigynal plate).
|
||
</paragraph>
|
||
<caption id="DF39660998617D66FF0BF93B2A34F8E7" ID-DOI="http://doi.org/10.5281/zenodo.4927377" ID-Zenodo-Dep="4927377" httpUri="https://zenodo.org/record/4927377/files/figure.png" pageId="6" pageNumber="330" startId="6.[151,264,1762,1786]" targetBox="[151,1436,453,1738]" targetPageId="6">
|
||
<paragraph id="8BF9368198617D66FF0BF93B2A34F8E7" blockId="6.[151,1436,1762,1858]" pageId="6" pageNumber="330">
|
||
<emphasis id="B932EA9398617D66FF0BF93B2F1DF95F" bold="true" box="[151,344,1762,1786]" pageId="6" pageNumber="330">FIGURES 25–30.</emphasis>
|
||
Collecting sites of
|
||
<taxonomicName id="4C464D0298617D66FDB5F93B2C93F95F" authority="Huber, 2021" authorityName="Huber" authorityYear="2021" box="[553,726,1762,1786]" class="Arachnida" family="Pholcidae" genus="Artema" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="330" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
|
||
<emphasis id="B932EA9398617D66FDB5F93B2C93F95F" box="[553,726,1762,1786]" italics="true" pageId="6" pageNumber="330">Artema martensi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20157E898617D66FD7CF93B2D5FF95C" box="[736,794,1762,1785]" pageId="6" pageNumber="330" rank="species">sp. n.</taxonomicNameLabel>
|
||
25–26 Between Lakhssas and Bouizakarne (25 “cave 1”, 26 surrounding area). 27–28 Between Lakhssas and Bouizakarne (27 “cave 2”, 28 surrounding area). 29–30 SW Ida Ougnidif (29 cave/rock shelter, 30 surrounding area; the cave is on the right side, where the road ‘disappears’).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF39660998607D67FF0BFCCF2C24FCD3" ID-DOI="http://doi.org/10.5281/zenodo.4927379" ID-Zenodo-Dep="4927379" httpUri="https://zenodo.org/record/4927379/files/figure.png" pageId="7" pageNumber="331" startId="7.[151,250,790,814]" targetBox="[151,1436,181,766]" targetPageId="7">
|
||
<paragraph id="8BF9368198607D67FF0BFCCF2C24FCD3" blockId="7.[151,1436,790,886]" pageId="7" pageNumber="331">
|
||
<emphasis id="B932EA9398607D67FF0BFCCF2F5AFC8B" bold="true" box="[151,287,790,814]" pageId="7" pageNumber="331">FIGURE 31.</emphasis>
|
||
Known distribution of
|
||
<taxonomicName id="4C464D0298607D67FD94FCCF2C16FC88" authorityName="Walckenaer" authorityYear="1837" box="[520,595,790,813]" class="Arachnida" family="Pholcidae" genus="Artema" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="331" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B932EA9398607D67FD94FCCF2C16FC88" box="[520,595,790,813]" italics="true" pageId="7" pageNumber="331">Artema</emphasis>
|
||
</taxonomicName>
|
||
, excluding the synanthropic
|
||
<taxonomicName id="4C464D0298607D67FCEFFCCF2D93FC8B" authorityName="Walckenaer" authorityYear="1837" box="[883,982,790,814]" class="Arachnida" family="Pholcidae" genus="Artema" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="331" phylum="Arthropoda" rank="species" species="atlanta">
|
||
<emphasis id="B932EA9398607D67FCEFFCCF2D93FC8B" box="[883,982,790,814]" italics="true" pageId="7" pageNumber="331">A. atlanta</emphasis>
|
||
</taxonomicName>
|
||
and undescribed putative species (cf.
|
||
<bibRefCitation id="EFD74B7098607D67FACCFCCF2F48FCF7" author="Aharon, S. & Huber, B. A. & Gavish-Regev, E." pageId="7" pageNumber="331" pagination="1 - 57" refId="ref3688" refString="Aharon, S., Huber, B. A. & Gavish-Regev, E. (2017) Daddy-long-leg giants: revision of the spider genus Artema Walckenaer, 1837 (Araneae, Pholcidae). European Journal of Taxonomy, 376, 1 - 57. https: // doi. org / 10.5852 / ejt. 2017.376" type="journal article" year="2017">
|
||
Aharon
|
||
<emphasis id="B932EA9398607D67FF0BFCE22E88FCF7" box="[151,205,826,850]" italics="true" pageId="7" pageNumber="331">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
). Coordinate data from
|
||
<bibRefCitation id="EFD74B7098607D67FD9BFCE32D58FCF7" author="Huber, B. A. & Kwapong, P." box="[519,797,826,850]" pageId="7" pageNumber="331" pagination="1 - 44" refId="ref4671" refString="Huber, B. A. & Kwapong, P. (2013) West African pholcid spiders: an overview, with descriptions of five new species (Araneae, Pholcidae). European Journal of Taxonomy, 59, 1 - 44. https: // doi. org / 10.5852 / ejt. 2013.59" type="journal article" year="2013">Huber & Kwapong (2013)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFD74B7098607D67FCB5FCE32A47FCF7" author="Aharon, S. & Huber, B. A. & Gavish-Regev, E." box="[809,1026,826,850]" pageId="7" pageNumber="331" pagination="1 - 57" refId="ref3688" refString="Aharon, S., Huber, B. A. & Gavish-Regev, E. (2017) Daddy-long-leg giants: revision of the spider genus Artema Walckenaer, 1837 (Araneae, Pholcidae). European Journal of Taxonomy, 376, 1 - 57. https: // doi. org / 10.5852 / ejt. 2017.376" type="journal article" year="2017">
|
||
Aharon
|
||
<emphasis id="B932EA9398607D67FCE3FCE22DF0FCF7" box="[895,949,826,850]" italics="true" pageId="7" pageNumber="331">et al.</emphasis>
|
||
(2017)
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFD74B7098607D67FB92FCE32B76FCF7" author="Al-Khazali, A. M. & Najim, S. A." box="[1038,1331,826,850]" pageId="7" pageNumber="331" pagination="179 - 187" refId="ref3748" refString="Al-Khazali, A. M. & Najim, S. A. (2018) First records of Pholcidae (Arachnida, Araneae) from Iraq. Bulletin of the Iraq Natural History Museum, 15, 179 - 187. https: // doi. org / 10.26842 / binhm. 7.2018.15.2.0179" type="journal article" year="2018">Al-Khazali & Najim (2018)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFD74B7098607D67FADCFCE32F7BFCD3" author="Huber, B. A. & Carvalho, L. S." pageId="7" pageNumber="331" pagination="1 - 96" refId="ref4504" refString="Huber, B. A. & Carvalho, L. S. (2019) Filling the gaps: descriptions of unnamed species included in the latest molecular phylogeny of Pholcidae (Araneae). Zootaxa, 4546 (1), 1 - 96. https: // doi. org / 10.11646 / zootaxa. 4546.1.1" type="journal article" year="2019">Huber & Carvalho (2019)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFD74B7098607D67FEF3FC872C18FCD3" author="Mammola, S. & Aharon, S. & Seifan, M. & Lubin, Y. & Gavish-Regev, E." box="[367,605,862,886]" pageId="7" pageNumber="331" pagination="119" refId="ref4811" refString="Mammola, S., Aharon, S., Seifan, M., Lubin, Y. & Gavish-Regev, E. (2019) Exploring the interplay between local and regional drivers of distribution of a subterranean organism. Diversity, 11, 119. https: // doi. org / 10.3390 / d 11080119" type="journal article" year="2019">
|
||
Mammola
|
||
<emphasis id="B932EA9398607D67FE42FC862C54FCD3" box="[478,529,862,886]" italics="true" pageId="7" pageNumber="331">et al.</emphasis>
|
||
(2019)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF39660998607D67FF0BF9B62FE4F96B" ID-DOI="http://doi.org/10.5281/zenodo.4927381" ID-Zenodo-Dep="4927381" httpUri="https://zenodo.org/record/4927381/files/figure.png" pageId="7" pageNumber="331" startId="7.[151,250,1647,1671]" targetBox="[157,1436,910,1618]" targetPageId="7">
|
||
<paragraph id="8BF9368198607D67FF0BF9B62FE4F96B" blockId="7.[151,1437,1647,1743]" pageId="7" pageNumber="331">
|
||
<emphasis id="B932EA9398607D67FF0BF9B62F52F922" bold="true" box="[151,279,1647,1671]" pageId="7" pageNumber="331">FIGURE 32</emphasis>
|
||
. Carapace width in 1632 species of
|
||
<taxonomicName id="4C464D0298607D67FDE0F9B62CA5F922" box="[636,736,1647,1671]" class="Arachnida" family="Pholcidae" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="331" phylum="Arthropoda" rank="family">Pholcidae</taxonomicName>
|
||
(1544 described, taken from the taxonomic literature; 88 undescribed, taken from specimens deposited in ZFMK). For raw data, see Supplementary file. The five highest values (>4.0) are all from representatives of
|
||
<taxonomicName id="4C464D0298607D67FECDF96E2FD9F96B" authorityName="Walckenaer" authorityYear="1837" box="[337,412,1719,1742]" class="Arachnida" family="Pholcidae" genus="Artema" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="331" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B932EA9398607D67FECDF96E2FD9F96B" box="[337,412,1719,1742]" italics="true" pageId="7" pageNumber="331">Artema</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF9368198607D67FF5BF9242A76F84A" blockId="7.[151,1436,1789,2032]" pageId="7" pageNumber="331">
|
||
<emphasis id="B932EA9398607D67FF5BF9242FC3F8B2" bold="true" box="[199,390,1789,1815]" pageId="7" pageNumber="331">Natural history.</emphasis>
|
||
This species does not show any troglomorphism and can be classified as troglophilic. At the
|
||
<typeStatus id="54FD882398607D67FF0BF8FB2E82F899" box="[151,199,1826,1852]" pageId="7" pageNumber="331">type</typeStatus>
|
||
locality, the surrounding area is flat and does not seem to provide suitable habitats (
|
||
<figureCitation id="137D2A0498607D67FBF5F8F82AAEF899" box="[1129,1259,1825,1852]" captionStart="FIGURES 25–30" captionStartId="6.[151,264,1762,1786]" captionTargetBox="[151,1436,453,1738]" captionTargetId="figure-145@6.[151,1436,453,1738]" captionTargetPageId="6" captionText="FIGURES 25–30. Collecting sites of Artema martensi sp. n. 25–26 Between Lakhssas and Bouizakarne (25 “cave 1”, 26 surrounding area). 27–28 Between Lakhssas and Bouizakarne (27 “cave 2”, 28 surrounding area). 29–30 SW Ida Ougnidif (29 cave/rock shelter, 30 surrounding area; the cave is on the right side, where the road ‘disappears’)." figureDoi="http://doi.org/10.5281/zenodo.4927377" httpUri="https://zenodo.org/record/4927377/files/figure.png" pageId="7" pageNumber="331">Figs 25–28</figureCitation>
|
||
). No webs were seen among larger rocks outside of two neighboring caves (or cave entrances). Within the caves, the spiders were found in the entrance area and slightly beyond (in the twilight area), with indistinct webs but sitting on the rock surface. They shared the caves with a much more abundant pholcid species, closely related to
|
||
<taxonomicName id="4C464D0298607D67FB2FF8542F12F86E" authority="Barrientos, 2019" authorityName="Barrientos" authorityYear="2019" class="Arachnida" family="Pholcidae" genus="Holocnemus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="331" phylum="Arthropoda" rank="species" species="aurouxi">
|
||
<emphasis id="B932EA9398607D67FB2FF8542BD9F802" box="[1203,1436,1933,1959]" italics="true" pageId="7" pageNumber="331">Holocnemus aurouxi</emphasis>
|
||
<bibRefCitation id="EFD74B7098607D67FF0BF8682F12F86E" author="Barrientos, J. A. & Branas, N. & Mederos, J." box="[151,343,1969,1995]" pageId="7" pageNumber="331" pagination="3 - 12" refId="ref3802" refString="Barrientos, J. A., Branas, N. & Mederos, J. (2019) Dos especies nuevas de Pholcidae (Araneae) de Marruecos. Revista Iberica de Aracnologia, 34, 3 - 12." type="journal article" year="2019">Barrientos, 2019</bibRefCitation>
|
||
</taxonomicName>
|
||
. The second locality near Ida Ougnidif is a cave with a large opening, i.e. a shelter rather than a cave (
|
||
<figureCitation id="137D2A0498607D67FF46F80C2F6EF84A" box="[218,299,2005,2031]" captionStart="FIGURES 25–30" captionStartId="6.[151,264,1762,1786]" captionTargetBox="[151,1436,453,1738]" captionTargetId="figure-145@6.[151,1436,453,1738]" captionTargetPageId="6" captionText="FIGURES 25–30. Collecting sites of Artema martensi sp. n. 25–26 Between Lakhssas and Bouizakarne (25 “cave 1”, 26 surrounding area). 27–28 Between Lakhssas and Bouizakarne (27 “cave 2”, 28 surrounding area). 29–30 SW Ida Ougnidif (29 cave/rock shelter, 30 surrounding area; the cave is on the right side, where the road ‘disappears’)." figureDoi="http://doi.org/10.5281/zenodo.4927377" httpUri="https://zenodo.org/record/4927377/files/figure.png" pageId="7" pageNumber="331">Fig. 29</figureCitation>
|
||
) but with crevices possibly leading into inaccessible deeper sections.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35C650A986F7D68FF5BFF4F2ADBFF70" pageId="8" pageNumber="332" type="distribution">
|
||
<paragraph id="8BF93681986F7D68FF5BFF4F2ADBFF70" blockId="8.[151,1437,150,213]" pageId="8" pageNumber="332">
|
||
<emphasis id="B932EA93986F7D68FF5BFF4F2F24FF15" bold="true" box="[199,353,150,176]" pageId="8" pageNumber="332">Distribution.</emphasis>
|
||
Known from two localities in the western Anti-Atlas of south-western
|
||
<collectingCountry id="F3517611986F7D68FB0CFF4E2AB0FF14" box="[1168,1269,151,177]" name="Morocco" pageId="8" pageNumber="332">Morocco</collectingCountry>
|
||
(
|
||
<figureCitation id="137D2A04986F7D68FA9AFF4F2B1EFF15" box="[1286,1371,150,176]" captionStart="FIGURE 31" captionStartId="7.[151,250,790,814]" captionTargetBox="[151,1436,181,766]" captionTargetId="figure-23@7.[151,1436,181,766]" captionTargetPageId="7" captionText="FIGURE 31. Known distribution of Artema, excluding the synanthropic A. atlanta and undescribed putative species (cf.Aharon et al. 2017). Coordinate data from Huber & Kwapong (2013), Aharon et al. (2017), Al-Khazali & Najim (2018), Huber & Carvalho (2019) and Mammola et al. (2019)." figureDoi="http://doi.org/10.5281/zenodo.4927379" httpUri="https://zenodo.org/record/4927379/files/figure.png" pageId="8" pageNumber="332">Fig. 31</figureCitation>
|
||
). The two localities are almost
|
||
<quantity id="4CBE9B64986F7D68FE33FF632C41FF71" box="[431,516,186,212]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="332" unit="km" value="100.0">100 km</quantity>
|
||
apart, suggesting that the species is widespread in the area.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |