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<document id="341C12A3B408A02E4547404092C125F6" ID-DOI="10.11646/zootaxa.3925.1.6" ID-GBIF-Dataset="08e0d80e-3aae-411c-acb9-450bbb4ae133" ID-ISSN="1175-5326" ID-Zenodo-Dep="232510" ID-ZooBank="2249FA36-09F3-4A85-A79B-7564A187CCEE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461000639597" checkinUser="plazi" docAuthor="Salazar-Niño, Karen &amp; Serrão, José Eduardo" docDate="2015" docId="D64F5523FFDBFF831CD8FAFF4F21FB21" docLanguage="en" docName="zt03925p108.pdf" docOrigin="Zootaxa 3925 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Veturius" docType="treatment" docVersion="8" lastPageNumber="96" masterDocId="2A762D5BFFDAFF811C4FFC2C4D16FFE3" masterDocTitle="Description of the immature stages of nine species of Veturius (Coleoptera: Passalidae)" masterLastPageNumber="108" masterPageNumber="94" pageNumber="95" updateTime="1698564453905" updateUser="plazi">
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<mods:title id="7DB6906B82B6A4CC361C180BED739D3F">Description of the immature stages of nine species of Veturius (Coleoptera: Passalidae)</mods:title>
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<mods:namePart id="F1935C6456417B296F9E7C7F8A9DF5BC">Salazar-Niño, Karen</mods:namePart>
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<mods:namePart id="A6073DB521A8EC537DB842C40D6EA638">Serrão, José Eduardo</mods:namePart>
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<treatment id="D64F5523FFDBFF831CD8FAFF4F21FB21" ID-DOI="http://doi.org/10.5281/zenodo.5618094" ID-GBIF-Taxon="126095364" ID-Zenodo-Dep="5618094" LSID="urn:lsid:plazi:treatment:D64F5523FFDBFF831CD8FAFF4F21FB21" httpUri="http://treatment.plazi.org/id/D64F5523FFDBFF831CD8FAFF4F21FB21" lastPageId="2" lastPageNumber="96" pageId="1" pageNumber="95">
<subSubSection id="16FCB7BEFFDBFF831CD8FAFF497BFBE8" lastPageId="2" lastPageNumber="96" pageId="1" pageNumber="95" type="nomenclature">
<paragraph id="5E59E435FFDBFF801CD8FAFF4F92F90E" blockId="1.[151,644,1747,1807]" box="[151,644,1747,1773]" pageId="1" pageNumber="95">
<heading id="05115359FFDBFF801CD8FAFF4F92F90E" bold="true" box="[151,644,1747,1773]" fontSize="11" level="1" pageId="1" pageNumber="95" reason="1">
<emphasis id="6C923827FFDBFF801CD8FAFF4F92F90E" bold="true" box="[151,644,1747,1773]" pageId="1" pageNumber="95">
<taxonomicName id="99E69FB6FFDBFF801CD8FAFF4DEAF90E" box="[151,252,1747,1773]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="6C923827FFDBFF801CD8FAFF4DEAF90E" bold="true" box="[151,252,1747,1773]" italics="true" pageId="1" pageNumber="95">Veturius</emphasis>
</taxonomicName>
(
<emphasis id="6C923827FFDBFF801D42FAFF4C79F90E" bold="true" box="[269,367,1747,1773]" italics="true" pageId="1" pageNumber="95">Vetur ius</emphasis>
)
<taxonomicName id="99E69FB6FFDBFF801DCCFAFF4F92F90E" authority="Kuwert, 1898" authorityName="Kuwert" authorityYear="1898" box="[387,644,1747,1773]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="95" phylum="Arthropoda" rank="species" species="aspina">
<emphasis id="6C923827FFDBFF801DCCFAFF4CC5F90E" bold="true" box="[387,467,1747,1773]" italics="true" pageId="1" pageNumber="95">aspina</emphasis>
Kuwert, 1898
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="5E59E435FFDBFF801CD8FADA4CB2F8EC" blockId="1.[151,644,1747,1807]" box="[151,420,1782,1807]" pageId="1" pageNumber="95">
(
<figureCitation id="C6DDF8B0FFDBFF801CD0FADA4DFFF8EC" box="[159,233,1782,1807]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1970]" captionTargetBox="[316,1244,210,1934]" captionTargetId="figure@3.[310,1255,196,1934]" captionTargetPageId="3" captionText="FIGURE 1. Light micrographs of the head. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. crassus, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. Epicranial suture (ES) and frontal suture (FS) and post-antennal head setae (HPA). Scale 1 mm." httpUri="https://zenodo.org/record/232511/files/figure.png" pageId="1" pageNumber="95">Figs. 1</figureCitation>
a, 2a, 3a, 4a, 9a)
</paragraph>
<paragraph id="5E59E435FFDBFF801CD8FB12488AF804" blockId="1.[151,1436,1854,2023]" pageId="1" pageNumber="95">
<emphasis id="6C923827FFDBFF801CD8FB124C27F8B4" bold="true" box="[151,305,1854,1879]" pageId="1" pageNumber="95">Third instar.</emphasis>
<emphasis id="6C923827FFDBFF801D75FB124C62F8B4" box="[314,372,1854,1879]" italics="true" pageId="1" pageNumber="95">Body</emphasis>
: length 60.063.0 mm, width (prothorax) 12.0 mm; covered with uniform pile of fine, short setae (
<figureCitation id="C6DDF8B0FFDBFF801C90FB4E4C35F898" box="[223,291,1890,1915]" captionStart="FIGURE 9" captionStartId="12.[151,250,1377,1399]" captionTargetBox="[151,1436,193,1355]" captionTargetId="figure@12.[151,1436,193,1356]" captionTargetPageId="12" captionText="FIGURE 9. Light micrographs of the abdominal segment V (laterodorsal view), showing the setae of the body. (a) Uniform pile of fine short setae (Veturius aspina), with detail of the setae. (b) Tiny, stout setae (V. simillimus), with detail of the setae. (c) Tiny, stout setae (V. impressus), with detail of the setae. Scale 0.1 mm." httpUri="https://zenodo.org/record/232519/files/figure.png" pageId="1" pageNumber="95">Fig. 9</figureCitation>
a); prosternal setae absent. Primary setal pattern: 45 HPA, 1 PD, 4 PSL, 1 MSD, 23 MSL, 1 MTD, 23 MTL, 1 TM, 1 TSM, 1 TL, 1 AV9 (
<tableCitation id="1364D18EFFDBFF801E1DFBAA4FBFF87C" box="[594,681,1926,1951]" captionStart="TABLE 2" captionStartId="2.[151,239,332,355]" captionTargetBox="[159,1407,414,850]" captionTargetPageId="2" captionText="TABLE 2. Number of primary setae of larvae of Veturius species (Coleoptera: Passalidae). Numbers in parentheses refer to character states less frequent." httpUri="http://table.plazi.org/id/0A99B4BDFFD8FF831CD8FD604CFDFE60" pageId="1" pageNumber="95" tableUuid="0A99B4BDFFD8FF831CD8FD604CFDFE60">Table 2</tableCitation>
).
<emphasis id="6C923827FFDBFF801EF2FBAA4FECF87C" box="[701,762,1926,1951]" italics="true" pageId="1" pageNumber="95">Head</emphasis>
: covered with uniform pile of short, golden setae; epicranial suture short; frontal sutures extended to the base of the antennae; width of head capsule 7.08.0 mm (
<figureCitation id="C6DDF8B0FFDBFF801975FB86486AF820" box="[1338,1404,1962,1987]" captionStart="FIGURE 1" captionStartId="3.[151,250,1948,1970]" captionTargetBox="[316,1244,210,1934]" captionTargetId="figure@3.[310,1255,196,1934]" captionTargetPageId="3" captionText="FIGURE 1. Light micrographs of the head. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. crassus, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. Epicranial suture (ES) and frontal suture (FS) and post-antennal head setae (HPA). Scale 1 mm." httpUri="https://zenodo.org/record/232511/files/figure.png" pageId="1" pageNumber="95">Fig. 1</figureCitation>
a). Mandibles without micro-asperities on the ventral base. Maxillary stipes with 13 micro-conical projections (
<figureCitation id="C6DDF8B0FFDBFF801920FBE2488AF804" box="[1391,1436,1998,2023]" captionStart="FIGURE 2" captionStartId="4.[151,250,1651,1673]" captionTargetBox="[151,1436,193,1628]" captionTargetId="figure@4.[151,1436,193,1629]" captionTargetPageId="4" captionText="FIGURE 2. Scanning electronic micrographs of maxillae. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. crassus, (g) V. impressus, (h) high magnification of encircled area of Fig. 2 g of the microconical projections (arrow) on the stipes (i) V. oberthuri, (j) V. negroensis. Scale 200 µm." httpUri="https://zenodo.org/record/232512/files/figure.png" pageId="1" pageNumber="95">Fig.</figureCitation>
</paragraph>
<paragraph id="5E59E435FFD8FF831CD8FCB44F2BFEFF" blockId="2.[151,1436,151,285]" pageId="2" pageNumber="96">
2a).
<emphasis id="6C923827FFD8FF831C83FCB54C17FF53" box="[204,257,153,176]" italics="true" pageId="2" pageNumber="96">Legs</emphasis>
: pars stridens (mesocoxae) length 1097.80 µm, width 737.89 µm, distance into striate 20.12 µm, 41 striae; mesocoxae and metacoxae with 2 internal setae. Metathoracic legs covered with short setae and microasperities in medial and posterior areas; anterior margin with 5 well developed teeth, I is the largest tooth, IIV gradually decreasing in size (
<figureCitation id="C6DDF8B0FFD8FF831D91FD284F36FEFF" box="[478,544,260,285]" captionStart="FIGURE 3" captionStartId="5.[151,250,1952,1974]" captionTargetBox="[315,1253,196,1928]" captionTargetId="figure@5.[315,1255,196,1929]" captionTargetPageId="5" captionText="FIGURE 3. Scanning electronic micrographs of metathoracic legs. (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. c r a s s u s, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. I VI indicate the number of teeth. Scale 100 µm." httpUri="https://zenodo.org/record/232513/files/figure.png" pageId="2" pageNumber="96">Fig. 3</figureCitation>
a).
</paragraph>
<caption id="0A99B4BDFFD8FF831CD8FD604CFDFE60" ID-Table-UUID="0A99B4BDFFD8FF831CD8FD604CFDFE60" httpUri="http://table.plazi.org/id/0A99B4BDFFD8FF831CD8FD604CFDFE60" pageId="2" pageNumber="96" targetBox="[159,1407,414,850]" targetIsTable="true" targetPageId="2">
<paragraph id="5E59E435FFD8FF831CD8FD604CFDFE60" blockId="2.[151,1435,332,387]" pageId="2" pageNumber="96">
<emphasis id="6C923827FFD8FF831CD8FD604C1FFE80" bold="true" box="[151,265,332,355]" pageId="2" pageNumber="96">TABLE 2.</emphasis>
Number of primary setae of larvae of
<taxonomicName id="99E69FB6FFD8FF831EEDFD604FEFFE80" box="[674,761,332,355]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="6C923827FFD8FF831EEDFD604FEFFE80" box="[674,761,332,355]" italics="true" pageId="2" pageNumber="96">Veturius</emphasis>
</taxonomicName>
species (
<taxonomicName id="99E69FB6FFD8FF831F15FD604EC3FE80" box="[858,981,332,355]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="96" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="99E69FB6FFD8FF831F91FD604944FE80" box="[990,1106,332,355]" class="Insecta" family="Passalidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="96" phylum="Arthropoda" rank="family">Passalidae</taxonomicName>
). Numbers in parentheses refer to character states less frequent.
</paragraph>
</caption>
<paragraph id="5E59E435FFD8FF831D20FDB24843FCB1" pageId="2" pageNumber="96">
<table id="2CE61695FFD8007E1CD0FDB24869FCB1" box="[159,1407,414,850]" gridcols="11" gridrows="11" pageId="2" pageNumber="96">
<tr id="E0D6E677FFD8007E1CD0FDB24869FE50" box="[159,1407,414,435]" gridrow="0" pageId="2" pageNumber="96" rowspan-0="1" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="A3078F0BFFD8007E1D20FDB24F55FE50" box="[367,579,414,435]" gridcol="1" gridrow="0" pageId="2" pageNumber="96">
<emphasis id="6C923827FFD8FF831D20FDB24F0EFE50" bold="true" box="[367,536,414,435]" pageId="2" pageNumber="96">SETAL GROUP</emphasis>
*
</th>
</tr>
<tr id="E0D6E677FFD8007E1CD0FDE44869FE3D" box="[159,1407,456,478]" gridrow="1" pageId="2" pageNumber="96">
<th id="A3078F0BFFD8007E1CD0FDE44C2FFE3D" box="[159,313,456,478]" gridcol="0" gridrow="1" pageId="2" pageNumber="96">SPECIES</th>
<td id="A3078F0BFFD8007E1D20FDE44F55FE3D" box="[367,579,456,478]" gridcol="1" gridrow="1" pageId="2" pageNumber="96">HPA PD PSL</td>
<td id="A3078F0BFFD8007E1E24FDE44FB4FE3D" box="[619,674,456,478]" gridcol="2" gridrow="1" pageId="2" pageNumber="96">MSD</td>
<td id="A3078F0BFFD8007E1E86FDE44E1AFE3D" box="[713,780,456,478]" gridcol="3" gridrow="1" pageId="2" pageNumber="96">MSL</td>
<td id="A3078F0BFFD8007E1F7BFDE44E7AFE3D" box="[820,876,456,478]" gridcol="4" gridrow="1" pageId="2" pageNumber="96">MTD</td>
<td id="A3078F0BFFD8007E1FDAFDE44EDCFE3D" box="[917,970,456,478]" gridcol="5" gridrow="1" pageId="2" pageNumber="96">MTL</td>
<td id="A3078F0BFFD8007E1FBAFDE44909FE3D" box="[1013,1055,456,478]" gridcol="6" gridrow="1" pageId="2" pageNumber="96">TM</td>
<td id="A3078F0BFFD8007E180CFDE44961FE3D" box="[1091,1143,456,478]" gridcol="7" gridrow="1" pageId="2" pageNumber="96">TSM</td>
<td id="A3078F0BFFD8007E18EFFDE449A8FE3D" box="[1184,1214,456,478]" gridcol="8" gridrow="1" pageId="2" pageNumber="96">TL</td>
<td id="A3078F0BFFD8007E18AAFDE44805FE3D" box="[1253,1299,456,478]" gridcol="9" gridrow="1" pageId="2" pageNumber="96">AV9</td>
<td id="A3078F0BFFD8007E1974FDE44869FE3D" box="[1339,1407,456,478]" gridcol="10" gridrow="1" pageId="2" pageNumber="96">AR</td>
</tr>
<tr id="E0D6E677FFD8007E1CD0FDDD4869FDE4" box="[159,1407,497,519]" gridrow="2" pageId="2" pageNumber="96">
<th id="A3078F0BFFD8007E1CD0FDDD4C2FFDE4" box="[159,313,497,519]" gridcol="0" gridrow="2" pageId="2" pageNumber="96">
<taxonomicName id="99E69FB6FFD8FF831CD0FDDE4DEAFDE4" box="[159,252,498,519]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="96" phylum="Arthropoda" rank="species" species="aspina">
<emphasis id="6C923827FFD8FF831CD0FDDE4DEAFDE4" box="[159,252,498,519]" italics="true" pageId="2" pageNumber="96">V. aspina</emphasis>
</taxonomicName>
</th>
<td id="A3078F0BFFD8007E1D20FDDD4F55FDE4" box="[367,579,497,519]" gridcol="1" gridrow="2" pageId="2" pageNumber="96">45 1(2) 4</td>
<td id="A3078F0BFFD8007E1E24FDDD4FB4FDE4" box="[619,674,497,519]" gridcol="2" gridrow="2" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1E86FDDD4E1AFDE4" box="[713,780,497,519]" gridcol="3" gridrow="2" pageId="2" pageNumber="96">23(4)</td>
<td id="A3078F0BFFD8007E1F7BFDDD4E7AFDE4" box="[820,876,497,519]" gridcol="4" gridrow="2" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1FDAFDDD4EDCFDE4" box="[917,970,497,519]" gridcol="5" gridrow="2" pageId="2" pageNumber="96">23</td>
<td id="A3078F0BFFD8007E1FBAFDDD4909FDE4" box="[1013,1055,497,519]" gridcol="6" gridrow="2" pageId="2" pageNumber="96">1(2)</td>
<td id="A3078F0BFFD8007E180CFDDD4961FDE4" box="[1091,1143,497,519]" gridcol="7" gridrow="2" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18EFFDDD49A8FDE4" box="[1184,1214,497,519]" gridcol="8" gridrow="2" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18AAFDDD4805FDE4" box="[1253,1299,497,519]" gridcol="9" gridrow="2" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1974FDDD4869FDE4" box="[1339,1407,497,519]" gridcol="10" gridrow="2" pageId="2" pageNumber="96">2022</td>
</tr>
<tr id="E0D6E677FFD8007E1CD0FE364869FDD3" box="[159,1407,538,560]" gridrow="3" pageId="2" pageNumber="96">
<th id="A3078F0BFFD8007E1CD0FE364C2FFDD3" box="[159,313,538,560]" gridcol="0" gridrow="3" pageId="2" pageNumber="96">
<taxonomicName id="99E69FB6FFD8FF831CD0FE374C07FDD3" box="[159,273,538,560]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="96" phylum="Arthropoda" rank="species" species="assimilis">
<emphasis id="6C923827FFD8FF831CD0FE374C07FDD3" box="[159,273,538,560]" italics="true" pageId="2" pageNumber="96">V. assimilis</emphasis>
</taxonomicName>
</th>
<td id="A3078F0BFFD8007E1D20FE364F55FDD3" box="[367,579,538,560]" gridcol="1" gridrow="3" pageId="2" pageNumber="96">3 0 3</td>
<td id="A3078F0BFFD8007E1E24FE364FB4FDD3" box="[619,674,538,560]" gridcol="2" gridrow="3" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E1E86FE364E1AFDD3" box="[713,780,538,560]" gridcol="3" gridrow="3" pageId="2" pageNumber="96">2</td>
<td id="A3078F0BFFD8007E1F7BFE364E7AFDD3" box="[820,876,538,560]" gridcol="4" gridrow="3" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E1FDAFE364EDCFDD3" box="[917,970,538,560]" gridcol="5" gridrow="3" pageId="2" pageNumber="96">2</td>
<td id="A3078F0BFFD8007E1FBAFE364909FDD3" box="[1013,1055,538,560]" gridcol="6" gridrow="3" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E180CFE364961FDD3" box="[1091,1143,538,560]" gridcol="7" gridrow="3" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E18EFFE3649A8FDD3" box="[1184,1214,538,560]" gridcol="8" gridrow="3" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18AAFE364805FDD3" box="[1253,1299,538,560]" gridcol="9" gridrow="3" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1974FE364869FDD3" box="[1339,1407,538,560]" gridcol="10" gridrow="3" pageId="2" pageNumber="96">18</td>
</tr>
<tr id="E0D6E677FFD8007E1CD0FE684869FDB9" box="[159,1407,580,602]" gridrow="4" pageId="2" pageNumber="96">
<th id="A3078F0BFFD8007E1CD0FE684C2FFDB9" box="[159,313,580,602]" gridcol="0" gridrow="4" pageId="2" pageNumber="96">
<taxonomicName id="99E69FB6FFD8FF831CD0FE694C36FDB9" box="[159,288,580,602]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="96" phylum="Arthropoda" rank="species" species="simillimus">
<emphasis id="6C923827FFD8FF831CD0FE694C36FDB9" box="[159,288,580,602]" italics="true" pageId="2" pageNumber="96">V. simillimus</emphasis>
</taxonomicName>
</th>
<td id="A3078F0BFFD8007E1D20FE684F55FDB9" box="[367,579,580,602]" gridcol="1" gridrow="4" pageId="2" pageNumber="96">2 1 3</td>
<td id="A3078F0BFFD8007E1E24FE684FB4FDB9" box="[619,674,580,602]" gridcol="2" gridrow="4" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1E86FE684E1AFDB9" box="[713,780,580,602]" gridcol="3" gridrow="4" pageId="2" pageNumber="96">2</td>
<td id="A3078F0BFFD8007E1F7BFE684E7AFDB9" box="[820,876,580,602]" gridcol="4" gridrow="4" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E1FDAFE684EDCFDB9" box="[917,970,580,602]" gridcol="5" gridrow="4" pageId="2" pageNumber="96">2</td>
<td id="A3078F0BFFD8007E1FBAFE684909FDB9" box="[1013,1055,580,602]" gridcol="6" gridrow="4" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E180CFE684961FDB9" box="[1091,1143,580,602]" gridcol="7" gridrow="4" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18EFFE6849A8FDB9" box="[1184,1214,580,602]" gridcol="8" gridrow="4" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18AAFE684805FDB9" box="[1253,1299,580,602]" gridcol="9" gridrow="4" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1974FE684869FDB9" box="[1339,1407,580,602]" gridcol="10" gridrow="4" pageId="2" pageNumber="96">19</td>
</tr>
<tr id="E0D6E677FFD8007E1CD0FE414869FD60" box="[159,1407,621,643]" gridrow="5" pageId="2" pageNumber="96">
<th id="A3078F0BFFD8007E1CD0FE414C2FFD60" box="[159,313,621,643]" gridcol="0" gridrow="5" pageId="2" pageNumber="96">
<taxonomicName id="99E69FB6FFD8FF831CD0FE424C2FFD60" box="[159,313,621,643]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="96" phylum="Arthropoda" rank="species" species="sinuatocollis">
<emphasis id="6C923827FFD8FF831CD0FE424C2FFD60" box="[159,313,621,643]" italics="true" pageId="2" pageNumber="96">V. sinuatocollis</emphasis>
</taxonomicName>
</th>
<td id="A3078F0BFFD8007E1D20FE414F55FD60" box="[367,579,621,643]" gridcol="1" gridrow="5" pageId="2" pageNumber="96">45 1 3</td>
<td id="A3078F0BFFD8007E1E24FE414FB4FD60" box="[619,674,621,643]" gridcol="2" gridrow="5" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1E86FE414E1AFD60" box="[713,780,621,643]" gridcol="3" gridrow="5" pageId="2" pageNumber="96">23</td>
<td id="A3078F0BFFD8007E1F7BFE414E7AFD60" box="[820,876,621,643]" gridcol="4" gridrow="5" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1FDAFE414EDCFD60" box="[917,970,621,643]" gridcol="5" gridrow="5" pageId="2" pageNumber="96">2</td>
<td id="A3078F0BFFD8007E1FBAFE414909FD60" box="[1013,1055,621,643]" gridcol="6" gridrow="5" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E180CFE414961FD60" box="[1091,1143,621,643]" gridcol="7" gridrow="5" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18EFFE4149A8FD60" box="[1184,1214,621,643]" gridcol="8" gridrow="5" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18AAFE414805FD60" box="[1253,1299,621,643]" gridcol="9" gridrow="5" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1974FE414869FD60" box="[1339,1407,621,643]" gridcol="10" gridrow="5" pageId="2" pageNumber="96">1721</td>
</tr>
<tr id="E0D6E677FFD8007E1CD0FEBA4869FD4E" box="[159,1407,662,685]" gridrow="6" pageId="2" pageNumber="96">
<th id="A3078F0BFFD8007E1CD0FEBA4C2FFD4E" box="[159,313,662,685]" gridcol="0" gridrow="6" pageId="2" pageNumber="96">
<taxonomicName id="99E69FB6FFD8FF831CD0FEB44C1AFD4E" box="[159,268,664,685]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="96" phylum="Arthropoda" rank="species" species="sinuatus">
<emphasis id="6C923827FFD8FF831CD0FEB44C1AFD4E" box="[159,268,664,685]" italics="true" pageId="2" pageNumber="96">V. sinuatus</emphasis>
</taxonomicName>
</th>
<td id="A3078F0BFFD8007E1D20FEBA4F55FD4E" box="[367,579,662,685]" gridcol="1" gridrow="6" pageId="2" pageNumber="96">3 0 3</td>
<td id="A3078F0BFFD8007E1E24FEBA4FB4FD4E" box="[619,674,662,685]" gridcol="2" gridrow="6" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E1E86FEBA4E1AFD4E" box="[713,780,662,685]" gridcol="3" gridrow="6" pageId="2" pageNumber="96">2</td>
<td id="A3078F0BFFD8007E1F7BFEBA4E7AFD4E" box="[820,876,662,685]" gridcol="4" gridrow="6" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E1FDAFEBA4EDCFD4E" box="[917,970,662,685]" gridcol="5" gridrow="6" pageId="2" pageNumber="96">2</td>
<td id="A3078F0BFFD8007E1FBAFEBA4909FD4E" box="[1013,1055,662,685]" gridcol="6" gridrow="6" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E180CFEBA4961FD4E" box="[1091,1143,662,685]" gridcol="7" gridrow="6" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18EFFEBA49A8FD4E" box="[1184,1214,662,685]" gridcol="8" gridrow="6" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18AAFEBA4805FD4E" box="[1253,1299,662,685]" gridcol="9" gridrow="6" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1974FEBA4869FD4E" box="[1339,1407,662,685]" gridcol="10" gridrow="6" pageId="2" pageNumber="96">20</td>
</tr>
<tr id="E0D6E677FFD8007E1CD0FEEC4869FD35" box="[159,1407,704,726]" gridrow="7" pageId="2" pageNumber="96">
<th id="A3078F0BFFD8007E1CD0FEEC4C2FFD35" box="[159,313,704,726]" gridcol="0" gridrow="7" pageId="2" pageNumber="96">
<taxonomicName id="99E69FB6FFD8FF831CD0FEED4C12FD35" box="[159,260,705,726]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="96" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="6C923827FFD8FF831CD0FEED4C12FD35" box="[159,260,705,726]" italics="true" pageId="2" pageNumber="96">V. crassus</emphasis>
</taxonomicName>
</th>
<td id="A3078F0BFFD8007E1D20FEEC4F55FD35" box="[367,579,704,726]" gridcol="1" gridrow="7" pageId="2" pageNumber="96">3(4) 0 3</td>
<td id="A3078F0BFFD8007E1E24FEEC4FB4FD35" box="[619,674,704,726]" gridcol="2" gridrow="7" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E1E86FEEC4E1AFD35" box="[713,780,704,726]" gridcol="3" gridrow="7" pageId="2" pageNumber="96">2</td>
<td id="A3078F0BFFD8007E1F7BFEEC4E7AFD35" box="[820,876,704,726]" gridcol="4" gridrow="7" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E1FDAFEEC4EDCFD35" box="[917,970,704,726]" gridcol="5" gridrow="7" pageId="2" pageNumber="96">2</td>
<td id="A3078F0BFFD8007E1FBAFEEC4909FD35" box="[1013,1055,704,726]" gridcol="6" gridrow="7" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E180CFEEC4961FD35" box="[1091,1143,704,726]" gridcol="7" gridrow="7" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18EFFEEC49A8FD35" box="[1184,1214,704,726]" gridcol="8" gridrow="7" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18AAFEEC4805FD35" box="[1253,1299,704,726]" gridcol="9" gridrow="7" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1974FEEC4869FD35" box="[1339,1407,704,726]" gridcol="10" gridrow="7" pageId="2" pageNumber="96">18(16)</td>
</tr>
<tr id="E0D6E677FFD8007E1CD0FEC54869FD1C" box="[159,1407,745,767]" gridrow="8" pageId="2" pageNumber="96">
<th id="A3078F0BFFD8007E1CD0FEC54C2FFD1C" box="[159,313,745,767]" gridcol="0" gridrow="8" pageId="2" pageNumber="96">
<taxonomicName id="99E69FB6FFD8FF831CD0FEC64C0BFD1C" box="[159,285,746,767]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="96" phylum="Arthropoda" rank="species" species="impressus">
<emphasis id="6C923827FFD8FF831CD0FEC64C0BFD1C" box="[159,285,746,767]" italics="true" pageId="2" pageNumber="96">V. impressus</emphasis>
</taxonomicName>
</th>
<td id="A3078F0BFFD8007E1D20FEC54F55FD1C" box="[367,579,745,767]" gridcol="1" gridrow="8" pageId="2" pageNumber="96">3 0 3</td>
<td id="A3078F0BFFD8007E1E24FEC54FB4FD1C" box="[619,674,745,767]" gridcol="2" gridrow="8" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E1E86FEC54E1AFD1C" box="[713,780,745,767]" gridcol="3" gridrow="8" pageId="2" pageNumber="96">2</td>
<td id="A3078F0BFFD8007E1F7BFEC54E7AFD1C" box="[820,876,745,767]" gridcol="4" gridrow="8" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E1FDAFEC54EDCFD1C" box="[917,970,745,767]" gridcol="5" gridrow="8" pageId="2" pageNumber="96">12</td>
<td id="A3078F0BFFD8007E1FBAFEC54909FD1C" box="[1013,1055,745,767]" gridcol="6" gridrow="8" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E180CFEC54961FD1C" box="[1091,1143,745,767]" gridcol="7" gridrow="8" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18EFFEC549A8FD1C" box="[1184,1214,745,767]" gridcol="8" gridrow="8" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18AAFEC54805FD1C" box="[1253,1299,745,767]" gridcol="9" gridrow="8" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1974FEC54869FD1C" box="[1339,1407,745,767]" gridcol="10" gridrow="8" pageId="2" pageNumber="96">1516</td>
</tr>
<tr id="E0D6E677FFD8007E1CD0FF3E4869FCCB" box="[159,1407,786,808]" gridrow="9" pageId="2" pageNumber="96">
<th id="A3078F0BFFD8007E1CD0FF3E4C2FFCCB" box="[159,313,786,808]" gridcol="0" gridrow="9" pageId="2" pageNumber="96">
<taxonomicName id="99E69FB6FFD8FF831CD0FF3F4C0EFCCB" box="[159,280,786,808]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="96" phylum="Arthropoda" rank="species" species="oberthuri">
<emphasis id="6C923827FFD8FF831CD0FF3F4C0EFCCB" box="[159,280,786,808]" italics="true" pageId="2" pageNumber="96">V. oberthuri</emphasis>
</taxonomicName>
</th>
<td id="A3078F0BFFD8007E1D20FF3E4F55FCCB" box="[367,579,786,808]" gridcol="1" gridrow="9" pageId="2" pageNumber="96">10 0 3</td>
<td id="A3078F0BFFD8007E1E24FF3E4FB4FCCB" box="[619,674,786,808]" gridcol="2" gridrow="9" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E1E86FF3E4E1AFCCB" box="[713,780,786,808]" gridcol="3" gridrow="9" pageId="2" pageNumber="96">2</td>
<td id="A3078F0BFFD8007E1F7BFF3E4E7AFCCB" box="[820,876,786,808]" gridcol="4" gridrow="9" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E1FDAFF3E4EDCFCCB" box="[917,970,786,808]" gridcol="5" gridrow="9" pageId="2" pageNumber="96">2</td>
<td id="A3078F0BFFD8007E1FBAFF3E4909FCCB" box="[1013,1055,786,808]" gridcol="6" gridrow="9" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E180CFF3E4961FCCB" box="[1091,1143,786,808]" gridcol="7" gridrow="9" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18EFFF3E49A8FCCB" box="[1184,1214,786,808]" gridcol="8" gridrow="9" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18AAFF3E4805FCCB" box="[1253,1299,786,808]" gridcol="9" gridrow="9" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1974FF3E4869FCCB" box="[1339,1407,786,808]" gridcol="10" gridrow="9" pageId="2" pageNumber="96">18</td>
</tr>
<tr id="E0D6E677FFD8007E1CD0FF104869FCB1" box="[159,1407,828,850]" gridrow="10" pageId="2" pageNumber="96">
<th id="A3078F0BFFD8007E1CD0FF104C2FFCB1" box="[159,313,828,850]" gridcol="0" gridrow="10" pageId="2" pageNumber="96">
<taxonomicName id="99E69FB6FFD8FF831CD0FF114C33FCB1" box="[159,293,829,850]" class="Insecta" family="Passallidae" genus="Veturius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="96" phylum="Arthropoda" rank="species" species="negroensis">
<emphasis id="6C923827FFD8FF831CD0FF114C33FCB1" box="[159,293,829,850]" italics="true" pageId="2" pageNumber="96">V. negroensis</emphasis>
</taxonomicName>
</th>
<td id="A3078F0BFFD8007E1D20FF104F55FCB1" box="[367,579,828,850]" gridcol="1" gridrow="10" pageId="2" pageNumber="96">2 0 3</td>
<td id="A3078F0BFFD8007E1E24FF104FB4FCB1" box="[619,674,828,850]" gridcol="2" gridrow="10" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E1E86FF104E1AFCB1" box="[713,780,828,850]" gridcol="3" gridrow="10" pageId="2" pageNumber="96">2</td>
<td id="A3078F0BFFD8007E1F7BFF104E7AFCB1" box="[820,876,828,850]" gridcol="4" gridrow="10" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E1FDAFF104EDCFCB1" box="[917,970,828,850]" gridcol="5" gridrow="10" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1FBAFF104909FCB1" box="[1013,1055,828,850]" gridcol="6" gridrow="10" pageId="2" pageNumber="96">0</td>
<td id="A3078F0BFFD8007E180CFF104961FCB1" box="[1091,1143,828,850]" gridcol="7" gridrow="10" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18EFFF1049A8FCB1" box="[1184,1214,828,850]" gridcol="8" gridrow="10" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E18AAFF104805FCB1" box="[1253,1299,828,850]" gridcol="9" gridrow="10" pageId="2" pageNumber="96">1</td>
<td id="A3078F0BFFD8007E1974FF104869FCB1" box="[1339,1407,828,850]" gridcol="10" gridrow="10" pageId="2" pageNumber="96">14</td>
</tr>
</table>
</paragraph>
<paragraph id="5E59E435FFD8FF831CD8FF5E4898FC2A" blockId="2.[151,1435,882,969]" pageId="2" pageNumber="96">* Post-antennal head setae (HPA), dorsal pronotal setae (PD), lateral pronotal shield setae (PSL), dorsal mesonotal setae (MSD), lateral mesonotal setae (MSL), dorsal metanotal setae (MTD), lateral metanotal setae (MTL), median tergal setae (TM), submedial tergal setae (TSM), lateral tergal setae (TL), ventral abdominal setae (AV), anal ring setae (AR).</paragraph>
<paragraph id="5E59E435FFD8FF831C88FFDE497BFBE8" blockId="2.[151,1436,1010,1218]" box="[199,1133,1010,1035]" pageId="2" pageNumber="96">
<emphasis id="6C923827FFD8FF831C88FFDE4C27FBE8" box="[199,305,1010,1035]" italics="true" pageId="2" pageNumber="96">Abdomen</emphasis>
: raster with setae; anal ring with 2022 setae; anal slit T-shaped (
<figureCitation id="C6DDF8B0FFD8FF831841FFDE4944FBE8" box="[1038,1106,1010,1035]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1963]" captionTargetBox="[378,1203,192,1915]" captionTargetId="figure@6.[378,1205,182,1915]" captionTargetPageId="6" captionText="FIGURE 4. Light micrograph of the abdominal segment X, showing the anal opening (AP), anal ring setae (AR) and raster (R). (a) Veturius aspina, (b) V. assimilis, (c) V. simillimus, (d) V. sinuatocollis, (e) V. sinuatus, (f) V. c r a s s u s, (g) V. impressus, (h) V. oberthuri, (i) V. negroensis. Scale 1 mm." httpUri="https://zenodo.org/record/232514/files/figure.png" pageId="2" pageNumber="96">Fig. 4</figureCitation>
a).
</paragraph>
</subSubSection>
<subSubSection id="16FCB7BEFFD8FF831C88F83A4F21FB21" pageId="2" pageNumber="96" type="distribution">
<paragraph id="5E59E435FFD8FF831C88F83A4F21FB21" blockId="2.[151,1436,1010,1218]" pageId="2" pageNumber="96">
<emphasis id="6C923827FFD8FF831C88F83A4C76FBCC" bold="true" box="[199,352,1046,1071]" pageId="2" pageNumber="96">Distribution.</emphasis>
Species with geographical distribution from
<collectingCountry id="26F1A4A5FFD8FF831F15F83A4ED1FBCC" box="[858,967,1046,1071]" name="Honduras" pageId="2" pageNumber="96">Honduras</collectingCountry>
to
<collectingCountry id="26F1A4A5FFD8FF831FA5F83A4951FBCC" box="[1002,1095,1046,1071]" name="Ecuador" pageId="2" pageNumber="96">Ecuador</collectingCountry>
(
<bibRefCitation id="3A7799C4FFD8FF831819F83A49EDFBCC" author="Boucher" box="[1110,1275,1046,1071]" pageId="2" pageNumber="96" refString="Boucher, S. (2006) Evolution et phylogenie des Coleopteres Passalidae (Scarabaeoidea). Les taxons du groupe famille. La tribu neotropicale des Proculini et son complexe Veturius. Annales de la Societe Entomologique de France, 41, 239 - 604. http: // dx. doi. org / 10.1080 / 00379271.2005.10697444." type="journal article" year="2006">Boucher 2006</bibRefCitation>
).
<emphasis id="6C923827FFD8FF831C88F8174CA7FBB7" bold="true" box="[199,433,1083,1108]" pageId="2" pageNumber="96">Material examined.</emphasis>
<collectingCountry id="26F1A4A5FFD8FF831DF7F8104F46FBB7" box="[440,592,1084,1108]" name="Colombia" pageId="2" pageNumber="96">COLOMBIA</collectingCountry>
, TOLIMA, Municipio Ibagué, vereda Potrerillo, Sur el Totumo,
<geoCoordinate id="3BD282F2FFD8FF831968F810488AFBB7" box="[1319,1436,1082,1108]" direction="north" orientation="latitude" pageId="2" pageNumber="96" precision="15" value="4.3891664">4°23'21''N</geoCoordinate>
<geoCoordinate id="3BD282F2FFD8FF831CD8F84C4C23FB9A" box="[151,309,1119,1145]" direction="west" orientation="longitude" pageId="2" pageNumber="96" precision="1" value="-75.21547">75°12'55.7''W</geoCoordinate>
,
<quantity id="991E49D0FFD8FF831D07F84D4CB1FB9B" box="[328,423,1121,1145]" metricMagnitude="3" metricUnit="m" metricValue="1.04" pageId="2" pageNumber="96" unit="m" value="1040.0">1040 m</quantity>
,
<date id="2A58C2F5FFD8FF831DF5F84C4F31FB9A" box="[442,551,1120,1145]" pageId="2" pageNumber="96" value="2008-06-05">5.vi.2008</date>
. G. Amat, P. Reyes-Castillo, &amp; L. Jiménez (
<specimenCount id="48E02FBCFFD8FF831800F84C49F0FB9A" box="[1103,1254,1120,1145]" pageId="2" pageNumber="96" type="generic">5 specimens</specimenCount>
).
<collectingCountry id="26F1A4A5FFD8FF83194FF84D488EFB9A" box="[1280,1432,1121,1145]" name="Colombia" pageId="2" pageNumber="96">COLOMBIA</collectingCountry>
, ANTIOQUIA, Gómez Plata, vereda Vega Botero,
<quantity id="991E49D0FFD8FF831E98F8AA4E34FB7E" box="[727,802,1158,1182]" metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="2" pageNumber="96" unit="m" value="940.0">940 m</quantity>
,
<date id="2A58C2F5FFD8FF831F7FF8A94EA7FB7D" box="[816,945,1157,1182]" pageId="2" pageNumber="96" value="2011-07-12">12.vii.2011</date>
. Reyes-Castillo, Jiménez-Ferbans, Salazar- Niño, &amp; Amat-García (
<specimenCount id="48E02FBCFFD8FF831DD3F8854F3DFB21" box="[412,555,1193,1218]" pageId="2" pageNumber="96" type="generic">5 specimens</specimenCount>
).
</paragraph>
</subSubSection>
</treatment>
</document>