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<document id="E0DAA471E7416A0B70544CA775321DA2" ID-CLB-Dataset="26049" ID-DOI="10.11646/zootaxa.4654.1.1" ID-GBIF-Dataset="9ce1d103-9234-4fbe-9544-f1c294bb8783" ID-ISSN="1175-5326" ID-Zenodo-Dep="3365803" ID-ZooBank="CFD82CC0-3110-472E-972B-7ADC0C523A04" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1565592061838" checkinUser="plazi" docAuthor="Hansen, Thomas" docDate="2019" docId="BF18F633A98FFF752B9CC667FE83FAEC" docLanguage="en" docName="zootaxa.4654.1.1.pdf" docOrigin="Zootaxa 4654 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Spirancilla milthersii D. D. D. D. 1902" docType="treatment" docVersion="21" lastPageNumber="145" masterDocId="43218E4BA900FFE52B0BC14DFFA9FFE3" masterDocTitle="Gastropods from the Cretaceous-Palaeogene boundary in Denmark" masterLastPageNumber="196" masterPageNumber="1" pageNumber="144" updateTime="1698750564626" updateUser="ExternalLinkService">
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<mods:title id="150ACD3405324AB9C17D017DBF01F31E">Gastropods from the Cretaceous-Palaeogene boundary in Denmark</mods:title>
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<treatment id="BF18F633A98FFF752B9CC667FE83FAEC" ID-GBIF-Taxon="159074708" LSID="urn:lsid:plazi:treatment:BF18F633A98FFF752B9CC667FE83FAEC" httpUri="http://treatment.plazi.org/id/BF18F633A98FFF752B9CC667FE83FAEC" lastPageId="144" lastPageNumber="145" pageId="143" pageNumber="144">
<subSubSection id="7FAB14AEA98FFF6A2B9CC667FD96F8A7" box="[151,575,1834,1861]" pageId="143" pageNumber="144" type="nomenclature">
<paragraph id="370E4725A98FFF6A2B9CC667FD96F8A7" blockId="143.[151,575,1834,1896]" box="[151,575,1834,1861]" pageId="143" pageNumber="144">
<heading id="6C46F049A98FFF6A2B9CC667FD96F8A7" bold="true" box="[151,575,1834,1861]" fontSize="11" level="1" pageId="143" pageNumber="144" reason="1">
<taxonomicName id="F0B13CA6A98FFF6A2B9CC667FD96F8A7" authority="(Ravn, 1902)" authorityName="D. D. D. D." authorityYear="1902" baseAuthorityName="Ravn" baseAuthorityYear="1902" box="[151,575,1834,1861]" class="Gastropoda" family="Olividae" genus="Spirancilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="143" pageNumber="144" phylum="Mollusca" rank="species" species="milthersii">
<emphasis id="05C59B37A98FFF6A2B9CC667FD96F8A7" bold="true" box="[151,575,1834,1861]" pageId="143" pageNumber="144">
<emphasis id="05C59B37A98FFF6A2B9CC667FE31F8A6" bold="true" box="[151,408,1834,1861]" italics="true" pageId="143" pageNumber="144">Spirancilla milthersii</emphasis>
(
<bibRefCitation id="53203AD4A98FFF6A2AA2C666FD9FF8A7" author="Ravn, J. P. J." box="[425,566,1834,1861]" pageId="143" pageNumber="144" pagination="205 - 270" refId="ref138257" refString="Ravn, J. P. J. (1902) Molluskerne i Danmarks Kridtaflejringer. II, Scaphopoder, Gastropoder og Cephalopoder. Det Kongelige Danske Videnskabernes Selskab Skrifter, 6. Raekke, Naturvidenskabelig og Mathematisk Afdeling XI, 4, 205 - 270, pls. 1 - 5." type="journal article" year="1902">Ravn, 1902</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7FAB14AEA98FFF6A2B9CC603FD5CF848" pageId="143" pageNumber="144" type="description">
<paragraph id="370E4725A98FFF6A2B9CC603FEB6F88B" blockId="143.[151,575,1834,1896]" box="[151,287,1870,1896]" pageId="143" pageNumber="144">
<figureCitation id="AF8A5BA0A98FFF6A2B9CC603FF41F88B" box="[151,232,1870,1896]" captionStart="FIGURE 31" captionStartId="141.[151,250,188,213]" captionTargetId="figure@140.[151,1435,179,1918]" captionTargetPageId="140" captionText="FIGURE 31. AC. Vexillum tenestolidum n. sp. AB. Silicone cast of holotype MGUH 33259. Width of protoconch 0.7 mm. Width of last whorl 3.4 mm. C. Silicone cast of paratype MGUH 33260. Width 5.8 mm. DG. Vexillum? rosenkrantzi n. sp. DE. Silicone cast and inner mould of holotype MGUH 33261. Width 1.9 and 1.6 mm respectively. F. Silicone cast of paratype MGUH 33262. Width 3.2 mm. G. Silicone cast of paratype MGUH 33263. Width 2.9 mm. H. Vexillum? sp. Silicone cast of MGUH 33264. Width 3.5 mm. I. Columbarium heberti (Briart &amp; Cornet, 1880). Silicone cast of last whorl MGUH 33265. Width 7.7 mm. Photograph by S.L. Jakobsen. J. Columbarium sp. Silicone cast of MGUH 33266. Width 13.3 mm. KN. Conomitra glabra (Ravn, 1902). K. Silicone cast of MGUH 33267. Width 3.6 mm. LM. Silicone cast of MGUH 33268. Width 3.6 and 3.9 mm respectively. N. External cast of syntype MGUH 116. Height 8.2 mm. Photograph by S.L. Jakobsen. O. Volutomitra cf. quinqueplicata Ravn, 1902. Silicone cast of MGUH 33269. Width 11.6 mm. P. Volutomitra quinqueplicata Ravn, 1902. External cast representing last whorl of type specimen MGUH 117. Middle Danien Faxe Formation, Denmark. Height 38.6 mm. Photograph by S.L. Jakobsen. QR. Spirancilla milthersii (Ravn, 1902). Q. Silicone cast of holotype MGUH 33270. Width 3.3 mm. R. Silicone cast of MGUH 33271. Width 3.1 mm. ST. Exilia sp., Silicone cast of MGUH 33272. Width 1.6 mm. UW. Latiromitra? sp. External cast and inner mould of MGUH 33273. Width 4.6 mm for casts and 4.0 mm for mould." figureDoi="http://doi.org/10.5281/zenodo.3442118" httpUri="https://zenodo.org/record/3442118/files/figure.png" pageId="143" pageNumber="144">Figs 31</figureCitation>
QR
</paragraph>
<paragraph id="370E4725A98FFF6A2B9CC6DEFD5CF848" blockId="143.[151,757,1938,1963]" box="[151,757,1938,1963]" pageId="143" pageNumber="144">
1902
<taxonomicName id="F0B13CA6A98FFF6A2BDBC6DFFE2DF849" authority="Milthersii" authorityName="Milthersii" box="[208,388,1938,1962]" class="Gastropoda" family="Olividae" genus="Ancilla" kingdom="Animalia" order="Neogastropoda" pageId="143" pageNumber="144" phylum="Mollusca" rank="genus">
<emphasis id="05C59B37A98FFF6A2BDBC6DFFE2DF849" box="[208,388,1938,1962]" italics="true" pageId="143" pageNumber="144">Ancilla Milthersii</emphasis>
</taxonomicName>
n. sp. Ravn: 234235, pl. II, fig. 18.
</paragraph>
</subSubSection>
<subSubSection id="7FAB14AEA98FFF752B9CC695FD67FB40" lastPageId="144" lastPageNumber="145" pageId="143" pageNumber="144" type="materials_examined">
<paragraph id="370E4725A98FFF752B9CC695FB8BFF18" blockId="143.[151,1436,2008,2035]" lastBlockId="144.[151,1437,153,1295]" lastPageId="144" lastPageNumber="145" pageId="143" pageNumber="144">
<emphasis id="05C59B37A98FFF6A2B9CC695FEBBF810" bold="true" box="[151,274,2008,2035]" pageId="143" pageNumber="144">Diagnosis.</emphasis>
Protoconch conical with just over two convex whorls. Teleoconch shell elongated with weakly ex- pressed sutures and no spire furrows. Columella almost vertical, bearing four to six narrow, oblique plaits. Parietal callus very thin except in front of adapical part of aperture, forming weak tongue adapically of aperture. Aperture taking up slightly over half the shell height; its outer lip lacking a labral denticle.
</paragraph>
<paragraph id="370E4725A990FF752BCCC049FA25FEA0" blockId="144.[151,1437,153,1295]" pageId="144" pageNumber="145">
<emphasis id="05C59B37A990FF752BCCC049FE9FFEFC" bold="true" box="[199,310,260,287]" pageId="144" pageNumber="145">Material.</emphasis>
<materialsCitation id="87D94D78A990FF752A35C048FA25FEA0" ID-GBIF-Occurrence="2331574421" collectionCode="MGUH" pageId="144" pageNumber="145" specimenCode="MGUH 33270, MGUH 33271" specimenCount="1" typeStatus="holotype">
<typeStatus id="E80AF987A990FF752A35C048FE0FFEFC" box="[318,422,261,287]" pageId="144" pageNumber="145" type="holotype">Holotype</typeStatus>
<specimenCode id="6717EF5EA990FF752AA4C048FDFBFEFC" box="[431,594,261,287]" collectionCode="MGUH" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="144" pageNumber="145">MGUH 33270</specimenCode>
(cast),
<specimenCode id="6717EF5EA990FF7529A3C048FCE7FEFC" box="[680,846,261,287]" collectionCode="MGUH" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="144" pageNumber="145">MGUH 33271</specimenCode>
, a mould with the informal sample number SH.192. AB, and two uncatalogued specimens deposited in the collections of the Natural History Museum, Copenhagen.
</materialsCitation>
</paragraph>
<paragraph id="370E4725A990FF752BCCC000FB87FE14" blockId="144.[151,1437,153,1295]" pageId="144" pageNumber="145">
<emphasis id="05C59B37A990FF752BCCC000FD92FE84" bold="true" box="[199,571,332,359]" pageId="144" pageNumber="145">Type stratum and type locality.</emphasis>
The single specimen of
<bibRefCitation id="53203AD4A990FF752844C000FC77FE84" author="Ravn, J. P. J." box="[847,990,333,359]" pageId="144" pageNumber="145" pagination="205 - 270" refId="ref138257" refString="Ravn, J. P. J. (1902) Molluskerne i Danmarks Kridtaflejringer. II, Scaphopoder, Gastropoder og Cephalopoder. Det Kongelige Danske Videnskabernes Selskab Skrifter, 6. Raekke, Naturvidenskabelig og Mathematisk Afdeling XI, 4, 205 - 270, pls. 1 - 5." type="journal article" year="1902">Ravn (1902)</bibRefCitation>
,
<collectionCode id="51A0DFE0A990FF7528E1C000FBE9FE84" box="[1002,1088,333,359]" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="144" pageNumber="145">MGUH</collectionCode>
33270, and therefore the
<typeStatus id="E80AF987A990FF752E6AC000FF63FE68" pageId="144" pageNumber="145" type="holotype">holotype</typeStatus>
, is from the
<taxonomicName id="F0B13CA6A990FF752A50C03CFD1AFE68" authority="Limestone Member" authorityName="Gilleaudeau et al." authorityYear="2018" box="[347,691,369,395]" class="Gastropoda" family="Cerithiidae" genus="Cerithium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="144" pageNumber="145" phylum="Mollusca" rank="genus">Cerithium Limestone Member</taxonomicName>
at Stevns Klint. The last known location for the specimen was in the collections of DGU [= Danmarks Geologiske Undersøgelse], which is now part of the semi-private institution
<collectionCode id="51A0DFE0A990FF752B9CC0F4FF49FE30" box="[151,224,441,467]" pageId="144" pageNumber="145">GEUS</collectionCode>
in
<collectingCountry id="4FA607B5A990FF752A0CC0F4FEDAFE30" box="[263,371,441,467]" name="Denmark" pageId="144" pageNumber="145">Denmark</collectingCountry>
. In spite of several attempts at contacting
<collectionCode id="51A0DFE0A990FF752858C0F4FC35FE30" box="[851,924,441,467]" pageId="144" pageNumber="145">GEUS</collectionCode>
for more information on its whereabouts, no information has come to hand, and the specimen may very well have become lost.
</paragraph>
<paragraph id="370E4725A990FF752BCCC34DFC45FDF8" blockId="144.[151,1437,153,1295]" box="[199,1004,512,539]" pageId="144" pageNumber="145">
<emphasis id="05C59B37A990FF752BCCC34DFEF3FDF8" bold="true" box="[199,346,512,539]" pageId="144" pageNumber="145">Occurrence.</emphasis>
Rare in the
<taxonomicName id="F0B13CA6A990FF752AE9C34CFC9EFDF8" authority="Limestone Member" authorityName="Gilleaudeau et al." authorityYear="2018" box="[482,823,513,539]" class="Gastropoda" family="Cerithiidae" genus="Cerithium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="144" pageNumber="145" phylum="Mollusca" rank="genus">Cerithium Limestone Member</taxonomicName>
at Stevns Klint.
</paragraph>
<paragraph id="370E4725A990FF752BCCC369FE4CFC28" blockId="144.[151,1437,153,1295]" pageId="144" pageNumber="145">
<emphasis id="05C59B37A990FF752BCCC369FEF0FDDC" bold="true" box="[199,345,548,575]" pageId="144" pageNumber="145">Description.</emphasis>
Shell small, narrowly ovate with bluntly pointed apex. Protoconch low conical with just over two convex whorls, about
<quantity id="F049EAC0A990FF752A9BC305FE5DFD80" box="[400,500,584,611]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="144" pageNumber="145" unit="mm" value="0.45">0.45 mm</quantity>
high and
<quantity id="F049EAC0A990FF75296FC305FD61FD80" box="[612,712,584,611]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="144" pageNumber="145" unit="mm" value="0.65">0.65 mm</quantity>
wide. Teleoconch spire weakly convex with flat whorls covered by thin callus obscuring sutures. Last whorl contributing around 78 % of shell height, moderately convex, evenly contracted anteriorly, with broad uncallused zone between abapical edge of spire callus and posterior fasciole.Aperture elongate-ovate, strongly notched anteriorly, taking up about 54 % of shell height. Outer lip with evenly curved anterior margin lacking labral denticle. Columella thickened, gently concave, with four to six narrow plaits continuing into the aperture. Columellar callus bordered adapically by shallow anterior fasciole furrow, furrow terminating at aperture. Middle fasciole slightly wider, terminating just adapical to columella, and divided from the former by more or less distinct groove. Posterior fasciole widest, often indistinct due to effacement of adapical groove, groove terminating at adapical aperture angle. Spire grooves, including ancillid groove and band absent. Parietal callus very thin but for weak, rather narrow and nearly straight-sided tongue reaching up from aperture to spire callus, widening just before reaching the latter.
</paragraph>
<paragraph id="370E4725A990FF752BCCC298FE08FBF0" blockId="144.[151,1437,153,1295]" pageId="144" pageNumber="145">
<emphasis id="05C59B37A990FF752BCCC298FED7FC0C" bold="true" box="[199,382,981,1007]" pageId="144" pageNumber="145">Measurements.</emphasis>
<typeStatus id="E80AF987A990FF752A8FC298FE45FC0C" box="[388,492,981,1007]" pageId="144" pageNumber="145" type="holotype">Holotype</typeStatus>
<collectionCode id="51A0DFE0A990FF752AF9C298FDE1FC0C" box="[498,584,981,1007]" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="144" pageNumber="145">MGUH</collectionCode>
33270 is
<quantity id="F049EAC0A990FF7529BFC298FCA0FC0C" box="[692,777,981,1007]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="144" pageNumber="145" unit="mm" value="3.3">3.3 mm</quantity>
wide and
<quantity id="F049EAC0A990FF752870C298FC79FC0C" box="[891,976,981,1007]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="144" pageNumber="145" unit="mm" value="7.8">7.8 mm</quantity>
high, composed of protoconch and about four teleoconch whorls.
</paragraph>
<paragraph id="370E4725A990FF752BCCC550FAEFFBB8" blockId="144.[151,1437,153,1295]" pageId="144" pageNumber="145">
<emphasis id="05C59B37A990FF752BCCC550FE92FBD4" bold="true" box="[199,315,1053,1079]" pageId="144" pageNumber="145">Remarks.</emphasis>
<bibRefCitation id="53203AD4A990FF752A4AC550FE67FBD4" author="Ravn, J. P. J." box="[321,462,1053,1079]" pageId="144" pageNumber="145" pagination="205 - 270" refId="ref138257" refString="Ravn, J. P. J. (1902) Molluskerne i Danmarks Kridtaflejringer. II, Scaphopoder, Gastropoder og Cephalopoder. Det Kongelige Danske Videnskabernes Selskab Skrifter, 6. Raekke, Naturvidenskabelig og Mathematisk Afdeling XI, 4, 205 - 270, pls. 1 - 5." type="journal article" year="1902">Ravn (1902)</bibRefCitation>
did not explain the reason for the taxon name, but it was clearly in honour of the geologist and collector of the specimen, Poul Christian Vilhelm Madsen Milthers, who found it in the year 1900.
</paragraph>
<paragraph id="370E4725A990FF752BCCC528FD67FB40" blockId="144.[151,1437,153,1295]" pageId="144" pageNumber="145">
In spite of what was indicated by
<bibRefCitation id="53203AD4A990FF752931C528FD77FB9C" author="Vermeij, G. J." box="[570,734,1125,1151]" pageId="144" pageNumber="145" pagination="461 - 508" refId="ref142778" refString="Vermeij, G. J. (2001) Innovation and evolution at the edge: origins and fates of gastropods with a labral tooth. Biological Journal of the Linnean Society, 72, 461 - 508. https: // doi. org / 10.1111 / j. 1095 - 8312.2001. tb 01333. x" type="journal article" year="2001">Vermeij (2001</bibRefCitation>
: p. 508)
<taxonomicName id="F0B13CA6A990FF752836C528FB82FB9C" authorityName="D. D. D. D." baseAuthorityName="Ravn" baseAuthorityYear="1902" box="[829,1067,1125,1151]" class="Gastropoda" family="Olividae" genus="Spirancilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="144" pageNumber="145" phylum="Mollusca" rank="species" species="milthersii">
<emphasis id="05C59B37A990FF752836C528FB82FB9C" box="[829,1067,1125,1151]" italics="true" pageId="144" pageNumber="145">Spirancilla milthersii</emphasis>
</taxonomicName>
does not posses a labral denticle, but has an evenly curved anterior outer lip margin.
</paragraph>
</subSubSection>
<subSubSection id="7FAB14AEA990FF752BCCC5E0FE83FAEC" pageId="144" pageNumber="145" type="reference_group">
<paragraph id="370E4725A990FF752BCCC5E0FE83FAEC" blockId="144.[151,1437,153,1295]" pageId="144" pageNumber="145">
<taxonomicName id="F0B13CA6A990FF752BCCC5E0FDF2FB24" authority="(Ravn, 1902)" authorityName="D. D. D. D." authorityYear="1902" baseAuthorityName="Ravn" baseAuthorityYear="1902" box="[199,603,1197,1223]" class="Gastropoda" family="Olividae" genus="Spirancilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="144" pageNumber="145" phylum="Mollusca" rank="species" species="milthersii">
<emphasis id="05C59B37A990FF752BCCC5E0FE10FB24" box="[199,441,1197,1223]" italics="true" pageId="144" pageNumber="145">Spirancilla milthersii</emphasis>
(
<bibRefCitation id="53203AD4A990FF752AC7C5E0FDFAFB24" author="Ravn, J. P. J." box="[460,595,1197,1223]" pageId="144" pageNumber="145" pagination="205 - 270" refId="ref138257" refString="Ravn, J. P. J. (1902) Molluskerne i Danmarks Kridtaflejringer. II, Scaphopoder, Gastropoder og Cephalopoder. Det Kongelige Danske Videnskabernes Selskab Skrifter, 6. Raekke, Naturvidenskabelig og Mathematisk Afdeling XI, 4, 205 - 270, pls. 1 - 5." type="journal article" year="1902">Ravn, 1902</bibRefCitation>
)
</taxonomicName>
resembles the Selandien
<taxonomicName id="F0B13CA6A990FF75288EC5E0FAF2FB24" authority="(von Koenen, 1885)" baseAuthorityName="von Koenen" baseAuthorityYear="1885" box="[901,1371,1196,1223]" class="Gastropoda" family="Olividae" genus="Spirancilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="144" pageNumber="145" phylum="Mollusca" rank="species" species="flexuosa">
<emphasis id="05C59B37A990FF75288EC5E0FBC0FB24" box="[901,1129,1197,1223]" italics="true" pageId="144" pageNumber="145">Spirancilla flexuosa</emphasis>
(
<bibRefCitation id="53203AD4A990FF752F77C5E0FAFAFB24" author="von Koenen, A." box="[1148,1363,1196,1223]" pageId="144" pageNumber="145" pagination="1 - 128" refId="ref143230" refString="von Koenen, A. (1885) Ueber eine Paleocane Fauna von Kopenhagen. Abhandlungen der Koniglichen Gesellschaft der Wissenschaften zu Gottingen, 32, 1 - 128, pls. 1 - 5." type="journal article" year="1885">von Koenen, 1885</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="4FA607B5A990FF752B9CC59CFEABFB08" box="[151,258,1233,1259]" name="Denmark" pageId="144" pageNumber="145">Denmark</collectingCountry>
, but it appears smaller with a more blunt spire, a deeper anterior fasciole furrow and with stronger columellar plaits.
</paragraph>
</subSubSection>
</treatment>
</document>