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<document id="271D47918CA7134D1FE0EE22A8A6E9E4" ID-CLB-Dataset="26049" ID-DOI="10.11646/zootaxa.4654.1.1" ID-GBIF-Dataset="9ce1d103-9234-4fbe-9544-f1c294bb8783" ID-ISSN="1175-5326" ID-Zenodo-Dep="3365803" ID-ZooBank="CFD82CC0-3110-472E-972B-7ADC0C523A04" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1565592061838" checkinUser="plazi" docAuthor="Hansen, Thomas" docDate="2019" docId="BF18F633A96AFF8F2B9CC004FDFEFA01" docLanguage="en" docName="zootaxa.4654.1.1.pdf" docOrigin="Zootaxa 4654 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Vermetus Daudin 1800" docType="treatment" docVersion="20" lastPageNumber="107" masterDocId="43218E4BA900FFE52B0BC14DFFA9FFE3" masterDocTitle="Gastropods from the Cretaceous-Palaeogene boundary in Denmark" masterLastPageNumber="196" masterPageNumber="1" pageNumber="107" updateTime="1698750564626" updateUser="ExternalLinkService">
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<mods:title id="F6C3F68A302F5C2438B22A472FF6FBD4">Gastropods from the Cretaceous-Palaeogene boundary in Denmark</mods:title>
</mods:titleInfo>
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<mods:namePart id="91C64848A45FAC39BBEFBDF43D1947C5">Hansen, Thomas</mods:namePart>
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<mods:typeOfResource id="1BF2055F9DAA7F3786F38ACBDB60F12E">text</mods:typeOfResource>
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<mods:title id="52958F23C6C8DF24040658E35F3D7C64">Zootaxa</mods:title>
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<mods:part id="5CCEEF00EF871228C9AFEF0CEA328610">
<mods:date id="17504E7C4DF7DB934503F82A61FF635C">2019</mods:date>
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<mods:number id="4040A94BED3A7EA48738FFD2F31F3705">2019-08-12</mods:number>
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<mods:number id="3CFDDEC9AEDE4996DF587BA1683C637C">4654</mods:number>
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<mods:classification id="3DE68A33776C4974C3AC357FD4260B38">journal article</mods:classification>
<mods:identifier id="EE4536149D90B16A7EE3A6E28346681A" type="CLB-Dataset">26049</mods:identifier>
<mods:identifier id="C27719FF4CD100B907F931A0940484B3" type="DOI">10.11646/zootaxa.4654.1.1</mods:identifier>
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<treatment id="BF18F633A96AFF8F2B9CC004FDFEFA01" ID-GBIF-Taxon="159074762" LSID="urn:lsid:plazi:treatment:BF18F633A96AFF8F2B9CC004FDFEFA01" httpUri="http://treatment.plazi.org/id/BF18F633A96AFF8F2B9CC004FDFEFA01" lastPageNumber="107" pageId="106" pageNumber="107">
<subSubSection id="7FAB14AEA96AFF8F2B9CC004FE9BFE81" box="[151,306,328,355]" pageId="106" pageNumber="107" type="nomenclature">
<paragraph id="370E4725A96AFF8F2B9CC004FE9BFE81" blockId="106.[151,306,328,390]" box="[151,306,328,355]" pageId="106" pageNumber="107">
<heading id="6C46F049A96AFF8F2B9CC004FE9BFE81" bold="true" box="[151,306,328,355]" fontSize="11" level="1" pageId="106" pageNumber="107" reason="1">
<emphasis id="05C59B37A96AFF8F2B9CC004FE9BFE81" bold="true" box="[151,306,328,355]" pageId="106" pageNumber="107">
<taxonomicName id="F0B13CA6A96AFF8F2B9CC004FEAEFE80" ID-CoL="7PKR2" authorityName="Daudin" authorityYear="1800" box="[151,263,329,355]" class="Gastropoda" family="Vermetidae" genus="Vermetus" kingdom="Animalia" order="Littorinimorpha" pageId="106" pageNumber="107" phylum="Mollusca" rank="genus">
<emphasis id="05C59B37A96AFF8F2B9CC004FEAEFE80" bold="true" box="[151,263,329,355]" italics="true" pageId="106" pageNumber="107">Vermetus</emphasis>
</taxonomicName>
sp.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7FAB14AEA96AFF8F2B9CC021FEB5FE65" box="[151,284,364,390]" pageId="106" pageNumber="107" type="description">
<paragraph id="370E4725A96AFF8F2B9CC021FEB5FE65" blockId="106.[151,306,328,390]" box="[151,284,364,390]" pageId="106" pageNumber="107">
<figureCitation id="AF8A5BA0A96AFF8F2B9CC021FF40FE65" box="[151,233,364,390]" captionStart="FIGURE 27" captionStartId="109.[151,250,188,213]" captionTargetId="figure@108.[151,1435,179,1918]" captionTargetPageId="108" captionText="FIGURE 27. A–B. Melanella? sp., Silicone cast of MGUH 33183. Width 1.4 mm. C–E. Vermetus sp., C. Silicone cast of attached tube MGUH 33184. Largest diameter 6.6 mm. D. Inner mould of protoconch and initial teleoconch inside bryozoan ball. MGUH 33185. Protoconch 0.5 mm wide. E. Silicone cast of MGUH 33186. Width 3.5 mm. F–J. Thylacodes contendis n. sp. F. Attached holotype mould MGUH 33187. Width 2.4 mm. G–H. Silicone cast of paratype MGUH 33188. Width 2.9 mm. I. Silicone cast of paratype MGUH 33189. Width 3.0 mm. J. Silicone cast of free whorl paratype MGUH 33190. Tube diameter 1.8 mm. K–M. Thylacodes? sp., Silicone cast of MGUH 33191. Width 2.4 mm. N. Dendropoma sp., Protoconch and initial teleoconch tube of MGUH 33192. Width 1.7 mm. O. Laxispira sp., Silicone cast of MGUH 33193. Width 3.6 mm. P–R. Epetrium s.l. skeldervigensis n. sp. P. Silicone cast of holotype MGUH 33194. Width 2.4 mm. Q. Silicone cast of paratype MGUH 33195. Width 4.0 mm. R. Silicone cast of paratype MGUH 33196. Width 3.6 mm. S. Triphoridae genus et sp. indet. Sinistrally coiled protoconch MGUH 33197. Width 0.7 mm. T–W. Cerithiopsis restemlongis n. sp. T. Silicone cast of holotype MGUH 33198. Width 1.0 mm. U. Silicone cast of paratype MGUH 33199. Width 1.5 mm. V. Silicone cast of paratype protoconch MGUH 33200. Width 0.5 mm. W. Silicone cast of paratype protoconch MGUH 33201. Width 0.6 mm and height 1.1 mm. X. Cerithiopsis? sp., Silicone cast of MGUH 33202. Width 0.6 mm. Y–AE. Thereitis carlsbergi n. sp. Y–Z. Silicone cast of holotype specimen MGUH 33203. Width of protoconch 0.6 mm and teleoconch 1.4 mm. AE. Silicone cast of paratype MGUH 33204. Width 0.7 mm." figureDoi="http://doi.org/10.5281/zenodo.3442110" httpUri="https://zenodo.org/record/3442110/files/figure.png" pageId="106" pageNumber="107">Figs 27</figureCitation>
C–E
</paragraph>
</subSubSection>
<subSubSection id="7FAB14AEA96AFF8F2B9CC0FEFB7DFAB1" pageId="106" pageNumber="107" type="materials_examined">
<paragraph id="370E4725A96AFF8F2B9CC0FEFCBBFCB8" blockId="106.[151,1437,435,1507]" pageId="106" pageNumber="107">
<emphasis id="05C59B37A96AFF8F2B9CC0FEFEAFFE2D" bold="true" box="[151,262,435,462]" pageId="106" pageNumber="107">Material.</emphasis>
<materialsCitation id="87D94D78A96AFF8F2A04C0F9FE13FE2D" ID-GBIF-Occurrence="2331574416" box="[271,442,436,462]" collectionCode="MGUH" pageId="106" pageNumber="107" specimenCode="MGUH 33184" specimenCount="1">
<specimenCode id="6717EF5EA96AFF8F2A04C0F9FE1FFE2D" box="[271,438,436,462]" collectionCode="MGUH" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="106" pageNumber="107">MGUH 33184</specimenCode>
,
</materialsCitation>
<materialsCitation id="87D94D78A96AFF8F2AC9C0F9FDC4FE2D" ID-GBIF-Occurrence="2331574417" box="[450,621,435,462]" collectionCode="MGUH" pageId="106" pageNumber="107" specimenCode="MGUH 33185" specimenCount="1">
<specimenCode id="6717EF5EA96AFF8F2AC9C0F9FDC0FE2D" box="[450,617,435,462]" collectionCode="MGUH" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="106" pageNumber="107">MGUH 33185</specimenCode>
,
</materialsCitation>
<materialsCitation id="87D94D78A96AFF8F297DC0F9FCB4FE2D" ID-GBIF-Occurrence="2331574418" box="[630,797,436,462]" collectionCode="MGUH" pageId="106" pageNumber="107" specimenCode="MGUH 33186" specimenCount="1">
<specimenCode id="6717EF5EA96AFF8F297DC0F9FCB4FE2D" box="[630,797,436,462]" collectionCode="MGUH" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="106" pageNumber="107">MGUH 33186</specimenCode>
</materialsCitation>
,
<collectionCode id="51A0DFE0A96AFF8F2822C0FEFCC3FE2D" box="[809,874,435,462]" country="USA" httpUri="http://grbio.org/cool/2e2p-880n" name="Ohio State University Museum" pageId="106" pageNumber="107">ØSM</collectionCode>
.10042-134-a,
<collectionCode id="51A0DFE0A96AFF8F2F07C0FEFBE4FE2D" box="[1036,1101,435,462]" country="USA" httpUri="http://grbio.org/cool/2e2p-880n" name="Ohio State University Museum" pageId="106" pageNumber="107">ØSM</collectionCode>
.10042-187,
<collectionCode id="51A0DFE0A96AFF8F2FD3C0FEFAB0FE2D" box="[1240,1305,435,462]" country="USA" httpUri="http://grbio.org/cool/2e2p-880n" name="Ohio State University Museum" pageId="106" pageNumber="107">ØSM</collectionCode>
.10042-188,
<collectionCode id="51A0DFE0A96AFF8F2B9CC09AFF7EFE11" box="[151,215,471,498]" country="USA" httpUri="http://grbio.org/cool/2e2p-880n" name="Ohio State University Museum" pageId="106" pageNumber="107">ØSM</collectionCode>
.10042-386,
<collectionCode id="51A0DFE0A96AFF8F2A69C09AFE0BFE11" box="[354,418,471,498]" country="USA" httpUri="http://grbio.org/cool/2e2p-880n" name="Ohio State University Museum" pageId="106" pageNumber="107">ØSM</collectionCode>
.10042-388-b and
<materialsCitation id="87D94D78A96AFF8F2966C09AFCA6FCB8" ID-GBIF-Occurrence="2331574412" collectionCode="OSM" collectorName="Natural History Museum" country="Denmark" county="Lower Danian" location="The" municipality="upper Maastrichtian Hojerup Member" pageId="106" pageNumber="107" specimenCount="4">
<collectionCode id="51A0DFE0A96AFF8F2966C09AFD07FE11" box="[621,686,471,498]" country="USA" httpUri="http://grbio.org/cool/2e2p-880n" name="Ohio State University Museum" pageId="106" pageNumber="107">ØSM</collectionCode>
.10042-393.
<location id="326E11FEA96AFF8F283CC095FCCAFE11" LSID="urn:lsid:plazi:treatment:BF18F633A96AFF8F2B9CC004FDFEFA01:326E11FEA96AFF8F283CC095FCCAFE11" box="[823,867,472,498]" country="Denmark" county="Lower Danian" municipality="upper Maastrichtian Hojerup Member" name="The" pageId="106" pageNumber="107">The</location>
collection of the
<location id="326E11FEA96AFF8F2F2CC095FA96FE10" LSID="urn:lsid:plazi:treatment:BF18F633A96AFF8F2B9CC004FDFEFA01:326E11FEA96AFF8F2F2CC095FA96FE10" box="[1063,1343,472,499]" country="Denmark" county="Lower Danian" municipality="upper Maastrichtian Hojerup Member" name="Natural History Museum" pageId="106" pageNumber="107">Natural History Museum</location>
of
<collectingCountry id="4FA607B5A96AFF8F2E6FC094FF78FDF5" name="Denmark" pageId="106" pageNumber="107">Denmark</collectingCountry>
furthermore contains several specimens lacking numbers.
<location id="326E11FEA96AFF8F2857C0B1FC36FDF5" LSID="urn:lsid:plazi:treatment:BF18F633A96AFF8F2B9CC004FDFEFA01:326E11FEA96AFF8F2857C0B1FC36FDF5" box="[860,927,508,534]" country="Denmark" county="Lower Danian" municipality="upper Maastrichtian Hojerup Member" name="These" pageId="106" pageNumber="107">These</location>
may be grouped into 14 very fragmentary external teleoconch moulds from the
<collectingMunicipality id="D76ADD5FA96AFF8F2910C36CFC6AFDD9" box="[539,963,544,571]" pageId="106" pageNumber="107">upper Maastrichtian Højerup Member</collectingMunicipality>
(samples: SH.4, SR.773, SR.914, SR.921, SR.924.A, SR.1039, SR.1067.A–B, SR.1069.A–B, SR.1086.A–B, SR.1091.B, SR.1092.A–B, SR.1105, SR.1107, SR.1114.A) and 52 fragmentary moulds from the
<collectingCounty id="DE6F3FA9A96AFF8F29C4C324FCDBFD61" box="[719,882,616,643]" pageId="106" pageNumber="107">Lower Danian</collectingCounty>
<taxonomicName id="F0B13CA6A96AFF8F2871C325FB7BFD61" authority="Limestone Member" authorityName="Gilleaudeau et al." authorityYear="2018" box="[890,1234,616,642]" class="Gastropoda" family="Cerithiidae" genus="Cerithium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="106" pageNumber="107" phylum="Mollusca" rank="genus">Cerithium Limestone Member</taxonomicName>
(informal sample numbers: SH.80.B, SH.84, SH.149.B–C, SH.169.A, SH.174, SH.191, SH.234, SH.254, SH.277, SH.297.A–B, SH.302.A, SH.415.B, SH.454, SH.455, SH.458.A–B, SH.468.A, B, SH.483.A–B, SH.483.B–C, SR.143, SR.146, SR.189, SR.211.A–B, SR.250, SR.253, SR.281 (
<specimenCount id="21B78CACA96AFF8F29B1C399FCE0FD0C" box="[698,841,724,751]" pageId="106" pageNumber="107" type="generic">2 specimens</specimenCount>
), SR.285, SR.291, SR.295, SR.312, SR.331, SR.372, SR.383, SR.384, SR.389.A–B, SR.400, SR.404.A–B, SR.450, SR.458, SR.586.A–B, SR.597, SR.616 (
<specimenCount id="21B78CACA96AFF8F2E4BC3B5FF7FFCD4" pageId="106" pageNumber="107" type="generic">2 specimens</specimenCount>
), SR.622, SR.632, SR.642.A–B, SR.648, SO.90.A–B, SO.101, SO.176, and two without number from the old collections at the
<collectorName id="9A4422F3A96AFF8F2A57C20DFDDDFCB8" box="[348,628,832,859]" pageId="106" pageNumber="107">Natural History Museum</collectorName>
of
<collectingCountry id="4FA607B5A96AFF8F2992C20DFCAEFCB8" box="[665,775,832,859]" name="Denmark" pageId="106" pageNumber="107">Denmark</collectingCountry>
)
</materialsCitation>
.
</paragraph>
<paragraph id="370E4725A96AFF8F2BCCC22EFBD7FBED" blockId="106.[151,1437,435,1507]" pageId="106" pageNumber="107">
<emphasis id="05C59B37A96AFF8F2BCCC22EFEF3FC9D" bold="true" box="[199,346,867,894]" pageId="106" pageNumber="107">Occurrence.</emphasis>
Lithified top of the upper Maastrichtian Højerup Member at Rødvig and Højerup Church. Low- er Danian
<taxonomicName id="F0B13CA6A96AFF8F2A1BC2C5FDC3FC41" authority="Limestone Member" authorityName="Gilleaudeau et al." authorityYear="2018" box="[272,618,904,930]" class="Gastropoda" family="Cerithiidae" genus="Cerithium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="106" pageNumber="107" phylum="Mollusca" rank="genus">Cerithium Limestone Member</taxonomicName>
at Rødvig, Højerup Church, Skeldervig and Holtug, Stevns. The lower Danian ‘dead layer’ at Vokslev, Northern Jutland. The lower hardground (‘krabbelaget’) in the Korsnaeb Member at Korsnaeb, Stevns Klint. The best exposed protoconch possibly comes from the infilling of Korsnaeb Member into a burrow in the lower middle part of the
<taxonomicName id="F0B13CA6A96AFF8F2940C2B9FC36FBED" authority="Limestone Member" authorityName="Gilleaudeau et al." authorityYear="2018" box="[587,927,1012,1038]" class="Gastropoda" family="Cerithiidae" genus="Cerithium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="106" pageNumber="107" phylum="Mollusca" rank="genus">Cerithium Limestone Member</taxonomicName>
at Højerup Church.
</paragraph>
<paragraph id="370E4725A96AFF8F2BCCC55AFC0FFB7D" blockId="106.[151,1437,435,1507]" pageId="106" pageNumber="107">
<emphasis id="05C59B37A96AFF8F2BCCC55AFEF0FBD1" bold="true" box="[199,345,1047,1074]" pageId="106" pageNumber="107">Description.</emphasis>
Protoconch
<quantity id="F049EAC0A96AFF8F2AE3C55AFD3CFBD0" box="[488,661,1047,1075]" metricMagnitude="-4" metricUnit="m" metricValue="7.75" metricValueMax="9.0" metricValueMin="6.5" pageId="106" pageNumber="107" unit="mm" value="0.775" valueMax="0.9" valueMin="0.65">0.65 to 0.9 mm</quantity>
high and
<quantity id="F049EAC0A96AFF8F280DC55AFC1DFBD0" box="[774,948,1047,1075]" metricMagnitude="-4" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="106" pageNumber="107" unit="mm" value="0.525" valueMax="0.55" valueMin="0.5">0.5 to 0.55 mm</quantity>
wide and regularly coiled trochispiral with 3 ½ to 4 whorls. Surface without sculpture. Teleoconch uncoiled, irregular with few unevenly distributed and moderately sharp spiral ribs and fairly strong, irregularly distributed growth increments. Older specimens may have a denser spiral sculpture with more rounded ribs. Aperture tube-shaped.
</paragraph>
<paragraph id="370E4725A96AFF8F2BCCC5E5FF5DFB04" blockId="106.[151,1437,435,1507]" pageId="106" pageNumber="107">
<emphasis id="05C59B37A96AFF8F2BCCC5E5FED7FB21" bold="true" box="[199,382,1192,1218]" pageId="106" pageNumber="107">Measurements.</emphasis>
Largest specimen from Holtug with a largest whorl diameter of
<quantity id="F049EAC0A96AFF8F2F44C5E5FB0CFB20" box="[1103,1189,1192,1219]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="106" pageNumber="107" unit="mm" value="6.6">6.6 mm</quantity>
and aperture width of
<quantity id="F049EAC0A96AFF8F2B9CC581FF59FB04" box="[151,240,1228,1255]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="106" pageNumber="107" unit="mm" value="2.6">2.6 mm</quantity>
.
</paragraph>
<paragraph id="370E4725A96AFF8F2BCCC5BDFB7DFAB1" blockId="106.[151,1437,435,1507]" pageId="106" pageNumber="107">
<emphasis id="05C59B37A96AFF8F2BCCC5BDFE92FAE9" bold="true" box="[199,315,1264,1290]" pageId="106" pageNumber="107">Remarks.</emphasis>
Except for the protoconch specimen, which is located inside a bryozoan colony, none of the vermetid tubes studied is attached to a substrate. Most probably they have been associated with poriferans, which based on the numerous spicules present in the rock must have been quite abundant in the palaeo-environment.
</paragraph>
</subSubSection>
<subSubSection id="7FAB14AEA96AFF8F2BCCC410FDFEFA01" pageId="106" pageNumber="107" type="reference_group">
<paragraph id="370E4725A96AFF8F2BCCC410FDFEFA01" blockId="106.[151,1437,435,1507]" pageId="106" pageNumber="107">
<taxonomicName id="F0B13CA6A96AFF8F2BCCC410FE87FA95" ID-CoL="7PKR2" authorityName="Daudin" authorityYear="1800" box="[199,302,1373,1398]" class="Gastropoda" family="Vermetidae" genus="Vermetus" kingdom="Animalia" order="Littorinimorpha" pageId="106" pageNumber="107" phylum="Mollusca" rank="genus">
<emphasis id="05C59B37A96AFF8F2BCCC410FE87FA95" box="[199,302,1373,1398]" italics="true" pageId="106" pageNumber="107">Vermetus</emphasis>
</taxonomicName>
sp. differs from the Maastrichtian
<taxonomicName id="F0B13CA6A96AFF8F29A4C410FBF1FA95" authority="Kaunhowen, 1898" authorityName="Kaunhowen" authorityYear="1898" box="[687,1112,1372,1398]" class="Gastropoda" family="Vermetidae" genus="Vermetus" kingdom="Animalia" order="Littorinimorpha" pageId="106" pageNumber="107" phylum="Mollusca" rank="species" species="alternans">
<emphasis id="05C59B37A96AFF8F29A4C410FC2FFA95" box="[687,902,1372,1398]" italics="true" pageId="106" pageNumber="107">Vermetus alternans</emphasis>
<bibRefCitation id="53203AD4A96AFF8F2887C411FBF1FA95" author="Kaunhowen, F." box="[908,1112,1372,1398]" pageId="106" pageNumber="107" pagination="3 - 132" refId="ref133244" refString="Kaunhowen, F. (1898) Die Gastropoden der Maestrichter Kreide. Palaeontologische Abhandlungen, Neue Folge, 8 (1), 3 - 132, pls. 1 - 13. [for 1897]" type="journal article" year="1898">Kaunhowen, 1898</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="4FA607B5A96AFF8F2F90C411FB52FA95" box="[1179,1275,1372,1398]" name="Belgium" pageId="106" pageNumber="107">Belgium</collectingCountry>
by the sparser sculpture, which does not show any alternation between coarser and finer spiral ribs. Whether any of these diagnostic features separating the taxa could be solely ecophenetic is unclear, and at present it is deemed wisest to leave the Danish taxon under open nomenclature.
</paragraph>
</subSubSection>
</treatment>
</document>
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