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<document id="FED2165720F39808494EB3D96C4E2094" ID-CLB-Dataset="26049" ID-DOI="10.11646/zootaxa.4654.1.1" ID-GBIF-Dataset="9ce1d103-9234-4fbe-9544-f1c294bb8783" ID-ISSN="1175-5326" ID-Zenodo-Dep="3365803" ID-ZooBank="CFD82CC0-3110-472E-972B-7ADC0C523A04" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1565592061838" checkinUser="plazi" docAuthor="Hansen, Thomas" docDate="2019" docId="BF18F633A94CFFA82B9CC5C9FDFFFE84" docLanguage="en" docName="zootaxa.4654.1.1.pdf" docOrigin="Zootaxa 4654 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Arene jakobseni Hansen 2019, n. sp." docType="treatment" docUuid="CECD3AEE-2619-4933-A2F8-084BFC442056" docUuidSource="ZooBank" docVersion="22" lastPageNumber="78" masterDocId="43218E4BA900FFE52B0BC14DFFA9FFE3" masterDocTitle="Gastropods from the Cretaceous-Palaeogene boundary in Denmark" masterLastPageNumber="196" masterPageNumber="1" pageNumber="77" updateTime="1698750564626" updateUser="ExternalLinkService">
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<mods:title id="C30FA03B33A8954BEEC04868D73F02C0">Gastropods from the Cretaceous-Palaeogene boundary in Denmark</mods:title>
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<mods:namePart id="AF2192AF41F553D11DF88787BDD72A2C">Hansen, Thomas</mods:namePart>
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<mods:title id="897DF682D8830490A4E58499D5EE68DB">Zootaxa</mods:title>
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<mods:date id="39FDCF36D5499553033715F044D5F16C">2019</mods:date>
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<mods:number id="0E9645A24EDF4CF0C948DC6A425AF177">2019-08-12</mods:number>
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<treatment id="BF18F633A94CFFA82B9CC5C9FDFFFE84" ID-GBIF-Taxon="159074878" LSID="urn:lsid:zoobank.org:act:CECD3AEE-2619-4933-A2F8-084BFC442056" httpUri="http://treatment.plazi.org/id/BF18F633A94CFFA82B9CC5C9FDFFFE84" lastPageId="77" lastPageNumber="78" pageId="76" pageNumber="77">
<subSubSection id="7FAB14AEA94CFFA92B9CC5C9FE0CFB7D" box="[151,421,1156,1183]" pageId="76" pageNumber="77" type="nomenclature">
<paragraph id="370E4725A94CFFA92B9CC5C9FE0CFB7D" blockId="76.[151,421,1156,1218]" box="[151,421,1156,1183]" pageId="76" pageNumber="77">
<heading id="6C46F049A94CFFA92B9CC5C9FE0CFB7D" bold="true" box="[151,421,1156,1183]" fontSize="11" level="1" pageId="76" pageNumber="77" reason="1">
<emphasis id="05C59B37A94CFFA92B9CC5C9FE0CFB7D" bold="true" box="[151,421,1156,1183]" pageId="76" pageNumber="77">
<taxonomicName id="F0B13CA6A94CFFA92B9CC5C9FEF5FB7C" authority="Hansen, 2019" authorityName="Hansen" authorityYear="2019" box="[151,348,1156,1183]" class="Gastropoda" family="Areneidae" genus="Arene" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="76" pageNumber="77" phylum="Mollusca" rank="species" species="jakobseni" status="sp. nov.">
<emphasis id="05C59B37A94CFFA92B9CC5C9FEF5FB7C" bold="true" box="[151,348,1156,1183]" italics="true" pageId="76" pageNumber="77">Arene jakobseni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1EF6264CA94CFFA92A68C5C9FE0CFB7D" box="[355,421,1156,1182]" pageId="76" pageNumber="77" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7FAB14AEA94CFFA92B9CC5E5FEF7FB21" box="[151,350,1192,1218]" pageId="76" pageNumber="77" type="description">
<paragraph id="370E4725A94CFFA92B9CC5E5FEF7FB21" blockId="76.[151,421,1156,1218]" box="[151,350,1192,1218]" pageId="76" pageNumber="77">
<figureCitation id="AF8A5BA0A94CFFA92B9CC5E5FF43FB21" box="[151,234,1192,1218]" captionStart="FIGURE 22" captionStartId="73.[151,250,188,213]" captionTargetId="figure@72.[151,1435,181,1920]" captionTargetPageId="72" captionText="FIGURE 22. A. Phorcus sp., Silicone cast of MGUH 33109. Exposed width 2.2 mm. BC. Stomatella sp., External mould of MGUH 33110. Figure B has been colour inverted and mirrored to show it in positive. Width 1.5 mm. DF. Solariella sp. DE. Silicone cast of MGUH 33111. Width 6.2 mm. F. Silicone cast of aperture of MGUH 33112. Width 5.95 mm. GI. Skenea sp. GH. Inner mould of MGUH 33113. Width 5.2 mm. I. Inner mould of MGUH 33114. Width 10.2 mm. JK. Leucorhynchia sp., Silicone cast of MGUH 33115. Width 2.4 mm. LN. Tectus? indecorus n. sp., L. Umbilical view of inner mould of paratype ØSM.10042-327-b. Width 19.7 mm. MN. Silicone cast of holotype MGUH 33116. Width 1.7 mm. OP. Homalopoma sp., Silicone cast and inner mould of MGUH 33117. Width 9.6, 6.9 mm respectively. Q. Calliomphalus (Calliomphalus) sp., Silicone cast of MGUH 33118. Width 3.1 mm. RU. Arene jakobseni n. sp. RT. Silicone cast of holotype MGUH 33119. Width 8.6 mm. U. Silicone cast of paratype MGUH 33120. Width of aperture 4.8 mm." figureDoi="http://doi.org/10.5281/zenodo.3442100" httpUri="https://zenodo.org/record/3442100/files/figure.png" pageId="76" pageNumber="77">Figs 22</figureCitation>
RU, 23A
</paragraph>
</subSubSection>
<subSubSection id="7FAB14AEA94CFFA82B9CC5A2FDFFFE84" lastPageId="77" lastPageNumber="78" pageId="76" pageNumber="77" type="materials_examined">
<paragraph id="370E4725A94CFFA92B9CC5A2FCAAFACD" blockId="76.[151,1437,1263,2011]" pageId="76" pageNumber="77">
<emphasis id="05C59B37A94CFFA92B9CC5A2FEBBFAE9" bold="true" box="[151,274,1263,1290]" pageId="76" pageNumber="77">Diagnosis.</emphasis>
Shell relatively high; planispirally coiled protoconch featuring convex whorls; teleoconch with three to four strong, beaded spiral cords above whorl periphery.
</paragraph>
<paragraph id="370E4725A94CFFA92BCCC47AFD4AFA95" blockId="76.[151,1437,1263,2011]" pageId="76" pageNumber="77">
<emphasis id="05C59B37A94CFFA92BCCC47AFE1DFAB1" bold="true" box="[199,436,1335,1362]" pageId="76" pageNumber="77">Derivation of name.</emphasis>
In honour of Sten Lennart Jakobsen, conservator at the Geological Museum of Copenhagen, who has been a tremendous help to this project.
</paragraph>
<paragraph id="370E4725A94CFFA92BCCC4CDFE5DF9E5" blockId="76.[151,1437,1263,2011]" pageId="76" pageNumber="77">
<emphasis id="05C59B37A94CFFA92BCCC4CDFEDCFA79" bold="true" box="[199,373,1407,1434]" pageId="76" pageNumber="77">Type material.</emphasis>
<materialsCitation id="87D94D78A94CFFA92A77C4CDFB7AFA79" ID-GBIF-Occurrence="2331574434" box="[380,1235,1408,1435]" collectionCode="MGUH" pageId="76" pageNumber="77" specimenCode="MGUH 33119" specimenCount="1" typeStatus="holotype">
<typeStatus id="E80AF987A94CFFA92A77C4CDFE4DFA79" box="[380,484,1408,1434]" pageId="76" pageNumber="77" type="holotype">Holotype</typeStatus>
<specimenCode id="6717EF5EA94CFFA92AE0C4CDFD24FA79" box="[491,653,1408,1434]" collectionCode="MGUH" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="76" pageNumber="77">MGUH 33119</specimenCode>
is a natural external and internal mould of the spire
</materialsCitation>
.
<materialsCitation id="87D94D78A94CFFA92FD5C4CCFE59F9E5" ID-GBIF-Occurrence="2331574343" collectionCode="MGUH" pageId="76" pageNumber="77" specimenCode="MGUH 33120" specimenCount="1" typeStatus="paratype">
<typeStatus id="E80AF987A94CFFA92FD5C4CCFAE9FA78" box="[1246,1344,1409,1435]" pageId="76" pageNumber="77" type="paratype">Paratype</typeStatus>
<specimenCode id="6717EF5EA94CFFA92E4CC4CDFF74FA5D" collectionCode="MGUH" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="76" pageNumber="77">MGUH 33120</specimenCode>
is an external mould of the base and aperture from Holtug Quarry, possible from one of the Korsnaeb Member infilled
<emphasis id="05C59B37A94CFFA92BFBC485FE31FA01" box="[240,408,1480,1506]" italics="true" pageId="76" pageNumber="77">Thalassinoides</emphasis>
burrows in the
<taxonomicName id="F0B13CA6A94CFFA92947C485FC0CFA01" authority="Limestone Member" authorityName="Gilleaudeau et al." authorityYear="2018" box="[588,933,1480,1506]" class="Gastropoda" family="Cerithiidae" genus="Cerithium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="76" pageNumber="77" phylum="Mollusca" rank="genus">Cerithium Limestone Member</taxonomicName>
, but the gastropod is filled out with material from the
<taxonomicName id="F0B13CA6A94CFFA92BF5C4A1FE59F9E5" authority="Limestone" authorityName="Gilleaudeau et al." authorityYear="2018" box="[254,496,1516,1542]" class="Gastropoda" family="Cerithiidae" genus="Cerithium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="76" pageNumber="77" phylum="Mollusca" rank="genus">Cerithium Limestone</taxonomicName>
</materialsCitation>
.
</paragraph>
<paragraph id="370E4725A94CFFA92BCCC75DFF71F9AD" blockId="76.[151,1437,1263,2011]" pageId="76" pageNumber="77">
<materialsCitation id="87D94D78A94CFFA92BCCC75DFF7DF9AD" ID-GBIF-Occurrence="2331574457" county="Stevns Klint" location="Lower Danian" municipality="Holtug Quarry" pageId="76" pageNumber="77" specimenCount="2" typeStatus="holotype">
<emphasis id="05C59B37A94CFFA92BCCC75DFDE7F9C9" bold="true" box="[199,590,1551,1578]" pageId="76" pageNumber="77">
<typeStatus id="E80AF987A94CFFA92BCCC75DFEABF9C9" box="[199,258,1552,1578]" pageId="76" pageNumber="77">Type</typeStatus>
stratum and
<typeStatus id="E80AF987A94CFFA92AB9C75DFE4CF9C9" box="[434,485,1552,1578]" pageId="76" pageNumber="77">type</typeStatus>
locality.
</emphasis>
<location id="326E11FEA94CFFA92951C75CFCA9F9C9" LSID="urn:lsid:plazi:treatment:BF18F633A94CFFA82B9CC5C9FDFFFE84:326E11FEA94CFFA92951C75CFCA9F9C9" box="[602,768,1552,1579]" county="Stevns Klint" municipality="Holtug Quarry" name="Lower Danian" pageId="76" pageNumber="77">Lower Danian</location>
<taxonomicName id="F0B13CA6A94CFFA92807C75DFBC2F9C9" authority="Limestone Member" authorityName="Gilleaudeau et al." authorityYear="2018" box="[780,1131,1552,1578]" class="Gastropoda" family="Cerithiidae" genus="Cerithium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="76" pageNumber="77" phylum="Mollusca" rank="genus">Cerithium Limestone Member</taxonomicName>
at
<collectingMunicipality id="D76ADD5FA94CFFA92F9CC75DFAE8F9C9" box="[1175,1345,1552,1578]" pageId="76" pageNumber="77">Holtug Quarry</collectingMunicipality>
,
<collectingCounty id="DE6F3FA9A94CFFA92E59C75DFF7DF9AD" pageId="76" pageNumber="77">Stevns Klint</collectingCounty>
</materialsCitation>
.
</paragraph>
<paragraph id="370E4725A94CFFA92BCCC71AFF71F975" blockId="76.[151,1437,1263,2011]" pageId="76" pageNumber="77">
<emphasis id="05C59B37A94CFFA92BCCC71AFEF3F991" bold="true" box="[199,346,1623,1650]" pageId="76" pageNumber="77">Occurrence.</emphasis>
Lower Danian
<taxonomicName id="F0B13CA6A94CFFA92901C715FCF6F991" authority="Limestone Member" authorityName="Gilleaudeau et al." authorityYear="2018" box="[522,863,1624,1650]" class="Gastropoda" family="Cerithiidae" genus="Cerithium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="76" pageNumber="77" phylum="Mollusca" rank="genus">Cerithium Limestone Member</taxonomicName>
of the Rødvig Formation at Holtug Quarry, Stevns Klint.
</paragraph>
<paragraph id="370E4725A94CFFA92BCCC7D2FDADF839" blockId="76.[151,1437,1263,2011]" pageId="76" pageNumber="77">
<emphasis id="05C59B37A94CFFA92BCCC7D2FEF0F959" bold="true" box="[199,345,1695,1722]" pageId="76" pageNumber="77">Description.</emphasis>
Shell relatively high with planispirally coiled protoconch. Protoconch
<quantity id="F049EAC0A94CFFA92F8FC7EDFB75F958" box="[1156,1244,1696,1723]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="76" pageNumber="77" unit="mm" value="1.6">1.6 mm</quantity>
wide, consisting of 2 smooth or weakly transversely striated and slightly depressed whorls. Teleoconch turbiniform with deeply impressed sutures and strongly convex whorls; a weak keel may be present; base of shell weakly flattened with deep, but narrow umbilicus. Aperture circular with thick and smooth, flattened lip and low varix. Teleoconch surface heavily sculptured with eight to ten beaded or spiny spiral cords, three to four strong cords on adapical side, one at the periphery and the rest, which generally are much finer and more closely spaced, on the base. Spiny tubercles hollow in growth direction; restricted to primary three spiral ribs. Spiral cords on base decreasing in strength towards umbilicus, which is bounded by a single more strongly developed beaded spiral rib. Cords crossed by very fine and densely spaced transverse striae.
</paragraph>
<paragraph id="370E4725A94DFFA82BCCC1D4FBBCFF51" blockId="77.[151,1437,152,359]" box="[199,1045,152,179]" pageId="77" pageNumber="78">
<emphasis id="05C59B37A94DFFA82BCCC1D4FED7FF50" bold="true" box="[199,382,153,179]" pageId="77" pageNumber="78">Measurements.</emphasis>
<typeStatus id="E80AF987A94DFFA82A8FC1D5FE45FF51" box="[388,492,152,178]" pageId="77" pageNumber="78" type="holotype">Holotype</typeStatus>
<collectionCode id="51A0DFE0A94DFFA82AFFC1D4FDE3FF50" box="[500,586,153,179]" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="77" pageNumber="78">MGUH</collectionCode>
33119 is
<quantity id="F049EAC0A94DFFA829BDC1D4FCA5FF50" box="[694,780,153,179]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="77" pageNumber="78" unit="mm" value="9.2">9.2 mm</quantity>
high and
<quantity id="F049EAC0A94DFFA82870C1D5FC78FF50" box="[891,977,152,179]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="77" pageNumber="78" unit="mm" value="8.6">8.6 mm</quantity>
wide.
</paragraph>
<paragraph id="370E4725A94DFFA82BCCC1F0FDFFFE84" blockId="77.[151,1437,152,359]" pageId="77" pageNumber="78">
<emphasis id="05C59B37A94DFFA82BCCC1F0FE92FF34" bold="true" box="[199,315,189,215]" pageId="77" pageNumber="78">Remarks.</emphasis>
<taxonomicName id="F0B13CA6A94DFFA82A4FC1F0FE53FF35" authority="Hansen, 2019" authorityName="Hansen" authorityYear="2019" box="[324,506,188,215]" class="Gastropoda" family="Areneidae" genus="Arene" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="77" pageNumber="78" phylum="Mollusca" rank="species" species="jakobseni" status="sp. nov.">
<emphasis id="05C59B37A94DFFA82A4FC1F0FE53FF35" box="[324,506,188,215]" italics="true" pageId="77" pageNumber="78">Arene jakobseni</emphasis>
</taxonomicName>
<emphasis id="05C59B37A94DFFA8290FC1F0FDECFF34" bold="true" box="[516,581,189,215]" pageId="77" pageNumber="78">
<taxonomicNameLabel id="1EF6264CA94DFFA8290FC1F0FDECFF34" box="[516,581,189,215]" pageId="77" pageNumber="78" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs from the Campanian
<taxonomicName id="F0B13CA6A94DFFA8289FC1F0FABDFF34" authority="Kiel &amp; Bandel, 2002 a" authorityName="Kiel &amp; Bandel" authorityYear="2002" box="[916,1300,188,215]" class="Gastropoda" family="Areneidae" genus="Arene" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="77" pageNumber="78" phylum="Mollusca" rank="species" species="mcleani">
<emphasis id="05C59B37A94DFFA8289FC1F0FBA7FF35" box="[916,1038,188,215]" italics="true" pageId="77" pageNumber="78">A. mcleani</emphasis>
<bibRefCitation id="53203AD4A94DFFA82F1CC1F1FABDFF34" author="Kiel, S. &amp; Bandel, K." box="[1047,1300,188,215]" pageId="77" pageNumber="78" pagination="239 - 249" refId="ref133671" refString="Kiel, S. &amp; Bandel, K. (2002 a) Further Archaeogastropoda from the Campanian of Torallola, northern Spain. Acta Geologica Polonica, 52, 239 - 249, pls. 1 - 3." type="journal article" year="2002">Kiel &amp; Bandel, 2002a</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="4FA607B5A94DFFA82E57C1F1FA35FF35" box="[1372,1436,188,214]" name="Spain" pageId="77" pageNumber="78">Spain</collectingCountry>
and the Maastrichtian
<taxonomicName id="F0B13CA6A94DFFA82A9BC1ACFD73FF18" authority="Sohl, 1998" authorityName="Sohl" authorityYear="1998" box="[400,730,224,251]" class="Gastropoda" family="Areneidae" genus="Arene" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="77" pageNumber="78" phylum="Mollusca" rank="species" species="truncatospaera">
<emphasis id="05C59B37A94DFFA82A9BC1ACFDF3FF19" box="[400,602,225,251]" italics="true" pageId="77" pageNumber="78">A. truncatospaera</emphasis>
<bibRefCitation id="53203AD4A94DFFA8296AC1ADFD73FF18" author="Sohl, N. F." box="[609,730,224,251]" pageId="77" pageNumber="78" pagination="1 - 109" refId="ref140958" refString="Sohl, N. F. (1998) Upper Cretaceous Trochacean Gastropods from Puerto Rico and Jamaica. Palaeontographica Americana, 60, 1 - 109." type="journal article" year="1998">Sohl, 1998</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="4FA607B5A94DFFA82815C1ADFCD1FF19" box="[798,888,224,250]" name="Jamaica" pageId="77" pageNumber="78">Jamaica</collectingCountry>
in the sparser and spinier sculpture and the more strongly convex whorls. It closely resembles the middle Danian
<taxonomicName id="F0B13CA6A94DFFA8287AC048FB13FEFC" authority="(Ravn, 1933)" baseAuthorityName="Ravn" baseAuthorityYear="1933" box="[881,1210,260,287]" class="Gastropoda" family="Areneidae" genus="Arene" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="77" pageNumber="78" phylum="Mollusca" rank="species" species="depressa">
<emphasis id="05C59B37A94DFFA8287AC048FBB7FEFD" box="[881,1054,260,287]" italics="true" pageId="77" pageNumber="78">Arene depressa</emphasis>
(
<bibRefCitation id="53203AD4A94DFFA82F25C048FB1BFEFC" author="Ravn, J. P. J." box="[1070,1202,261,287]" pageId="77" pageNumber="78" pagination="1 - 74" refId="ref138358" refString="Ravn, J. P. J. (1933) Etudes sur les Pelecypodes et Gastropodes Daniens du Calcaire de Faxe. Det Kongelige Danske Videnskabernes Selskabs Skrifter, 9. raekke, Naturvidenskabelig og mathematisk Afdeling, 2, 1 - 74, pls. 1 - 7." type="journal article" year="1933">Ravn, 1933</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="4FA607B5A94DFFA82E0AC049FAC5FEFD" box="[1281,1388,260,286]" name="Denmark" pageId="77" pageNumber="78">Denmark</collectingCountry>
, but differs by the distinctly larger size, the coarser and sparser teleoconch sculpture and possible by a narrower umbilicus, though this may relate to ontogeny.
</paragraph>
</subSubSection>
</treatment>
</document>