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<taxonomicName id="E9C2D0E9EF51D320FF0FFBB70F51FBB1" ID-CoL="6VDQD" authority="Pocock, 1902" authorityName="Pocock" authorityYear="1902" box="[151,574,1053,1080]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="operculatus">
<emphasis id="1CB67778EF51D320FF0FFBB70F51FBB1" bold="true" box="[151,574,1053,1080]" pageId="2" pageNumber="153">
<emphasis id="1CB67778EF51D320FF0FFBB70CFCFBBE" bold="true" box="[151,403,1053,1080]" italics="true" pageId="2" pageNumber="153">Phrynus operculatus</emphasis>
<bibRefCitation id="4A53D69BEF51D320FE02FBB70F51FBB1" author="Pocock, R. I." box="[410,574,1053,1079]" pageId="2" pageNumber="153" refId="ref21088" refString="Pocock, R. I. (1902) Arachnida: Scorpiones, Pedipalpi &amp; Solifugae. In: Godman, F. D. &amp; Salvin, O. (Eds.), Biologia Centrali Americana. Vol. 3. Taylor and Francis, London, pp. 1 - 71." type="book" year="1902">Pocock, 1902</bibRefCitation>
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<paragraph id="2E7DAB6AEF51D320FF0FFBEB0F6EFBDD" blockId="2.[151,574,1053,1116]" box="[151,513,1089,1116]" pageId="2" pageNumber="153">
<figureCitation id="B6F9B7EFEF51D320FF0FFBEB0C4AFBDD" box="[151,293,1089,1115]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1942,1966]" captionStartId-1="5.[151,250,1903,1927]" captionStartId-2="6.[151,250,1775,1799]" captionTargetBox-0="[161,1423,388,1908]" captionTargetBox-1="[160,1423,531,1867]" captionTargetBox-2="[156,1432,852,1747]" captionTargetId-0="figure-88@3.[151,1436,381,1919]" captionTargetId-1="figure-157@5.[151,1436,525,1880]" captionTargetId-2="figure-368@6.[151,1436,848,1751]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 1. Phrynus operculatus. Female (CNAN-Am000407): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." captionText-1="FIGURE 2. Phrynus operculatus. Female (CNAN-Am000407): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." captionText-2="FIGURE 3. Male genital: Phrynus operculatus (CASENT 9060512): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1 = lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. Colored lines illustrate the comparison between the LoD medial width (green line) and the width of the base of LoL 1 (red line)." figureDoi-0="http://doi.org/10.5281/zenodo.4620708" figureDoi-1="http://doi.org/10.5281/zenodo.4620710" figureDoi-2="http://doi.org/10.5281/zenodo.4620718" httpUri-0="https://zenodo.org/record/4620708/files/figure.png" httpUri-1="https://zenodo.org/record/4620710/files/figure.png" httpUri-2="https://zenodo.org/record/4620718/files/figure.png" pageId="2" pageNumber="153">Figures. 13</figureCitation>
,
<figureCitation id="B6F9B7EFEF51D320FEA9FBEB0C15FBDD" box="[305,378,1089,1116]" captionStart-0="FIGURE 17" captionStart-1="FIGURE 18" captionStartId-0="27.[151,250,1596,1620]" captionStartId-1="28.[151,250,1479,1503]" captionTargetBox-0="[158,1428,188,1562]" captionTargetBox-1="[151,1436,813,1456]" captionTargetId-0="figure-15@27.[151,1437,181,1572]" captionTargetId-1="figure-330@28.[151,1436,813,1456]" captionTargetPageId-0="27" captionTargetPageId-1="28" captionText-0="FIGURE 17. Female genitalia: A. Phrynus operculatus (CNAN-Am000407). B. Phrynus alfonsoi sp. nov. (CNAN-T01383). C. Phrynus aliciae sp. nov. (CNAN-T01447). D. Phrynus giseae sp. nov. (CNAN-T01450). E. Phrynus tresmarias sp. nov. (CASENT 9060565). F. Phrynus jalisco (CASENT 9060519). Sclerotization of the gonopod bases (black arrows); colored lines illustrate the position of the sclerite of the gonopod with respect to the gonopod base, the length comparison between the gonopod base (green line) and sclerite of the gonopod (red line), and the comparison between width of the base of the sclerite (purple line) and the width of the medial region of the gonopod base (blue line)." captionText-1="FIGURE 18. Distribution: Phrynus alfonsoi sp. nov. (blue circle), Phrynus aliciae sp. nov. (yellow circle), Phrynus giseae sp. nov. (purple circle), Phrynus jalisco (light blue circle), Phrynus operculatus (red circle), Phrynus tresmarias sp. nov. (orange circle). Circles with black rings: type localities." figureDoi-0="http://doi.org/10.5281/zenodo.4620748" figureDoi-1="http://doi.org/10.5281/zenodo.4620750" httpUri-0="https://zenodo.org/record/4620748/files/figure.png" httpUri-1="https://zenodo.org/record/4620750/files/figure.png" pageId="2" pageNumber="153">1718</figureCitation>
.
<tableCitation id="63409ED1EF51D320FE1DFBEB0CB1FBDD" box="[389,478,1089,1115]" captionStart="TABLE 1" captionStartId="7.[151,239,151,175]" captionText="TABLE 1. Measurements (mm) of body, pedipalps segments, and femur of legs in Phrynus operculatus. When the segment is missing, the cell is empty." pageId="2" pageNumber="153">Table. 1</tableCitation>
,
<tableCitation id="63409ED1EF51D320FE74FBEB0C92FBDD" box="[492,509,1089,1115]" captionStart="TABLE 6" captionStartId="26.[151,239,368,392]" captionText="TABLE 6. Size proportions in the female genital: Gonopod base (Gb), Sclerite (Scl), and the male genital: lobus dorsalis (LoD), lobus lateralis primus (LoL1). Comparing their total length (l), medial width (mw), and base width (bw)." pageId="2" pageNumber="153">6</tableCitation>
.
</paragraph>
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<subSubSection id="66D8F8E1EF51D320FF0FFB2C0939F962" pageId="2" pageNumber="153" type="reference_group">
<paragraph id="2E7DAB6AEF51D320FF0FFB2C0E98FB18" blockId="2.[151,1437,1158,1795]" box="[151,1015,1158,1182]" pageId="2" pageNumber="153">
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<taxonomicName id="E9C2D0E9EF51D320FF0FFB2C0CA4FB18" authority="Herbst" authorityName="Herbst" authorityYear="1797" box="[151,459,1158,1182]" class="Arachnida" family="Phrynidae" genus="Phalangium" higherTaxonomySource="CoL" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="palmatum">
<emphasis id="1CB67778EF51D320FF0FFB2C0C11FB18" box="[151,382,1158,1182]" italics="true" pageId="2" pageNumber="153">Phalangium palmatum</emphasis>
Herbst
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, in
<treatmentCitation id="AF638D7BEF51D320FE77FB2C0E36FB18" author="Lichtenstein, A. A. H. &amp; Herbst, J. F. W." box="[495,857,1158,1182]" page="82" pageId="2" pageNumber="153" year="1797">
<bibRefCitation id="4A53D69BEF51D320FE77FB2C0E36FB18" author="Lichtenstein, A. A. H. &amp; Herbst, J. F. W." box="[495,857,1158,1182]" pageId="2" pageNumber="153" refId="ref20601" refString="Lichtenstein, A. A. H. &amp; Herbst, J. F. W. (1797) Naturgeschichte der Insekten-Gattungen Solpuga und Phalangium. In: Herbst, J. F. W. (Ed.), Natursystem der Ungeflugelten Insekten. Vol. 1. G. A. Lange, Berlin, pp. 1 - 88." type="book" year="1797">Lichtenstein &amp; Herbst 1797: 8284</bibRefCitation>
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, plate IV fig. 2.
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</paragraph>
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<taxonomicName id="E9C2D0E9EF51D320FF0FFB0E0CDFFB3A" authority="(Herbst)" baseAuthorityName="Herbst" box="[151,432,1188,1212]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="palmatus">
<emphasis id="1CB67778EF51D320FF0FFB0E0C3FFB3A" box="[151,336,1188,1212]" italics="true" pageId="2" pageNumber="153">Phrynus palmatus</emphasis>
(Herbst)
</taxonomicName>
:
<treatmentCitation id="AF638D7BEF51D320FE23FB0E0FD9FB3A" author="Latreille, P. A." box="[443,694,1188,1213]" page="136" pageId="2" pageNumber="153" year="1804">
<bibRefCitation id="4A53D69BEF51D320FE23FB0E0FD9FB3A" author="Latreille, P. A." box="[443,694,1188,1213]" pageId="2" pageNumber="153" refId="ref20566" refString="Latreille, P. A. (1804) Histoire naturelle, generale et particuliere, des Crustaces et des Insectes. Vol. 7. F. Dufart, Paris, 866 pp." type="book" year="1804">Latreille 1804: 136137</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="AF638D7BEF51D320FD5AFB0E0E06FB3A" author="Gervais, P." box="[706,873,1188,1213]" page="6" pageId="2" pageNumber="153" year="1844">
<bibRefCitation id="4A53D69BEF51D320FD5AFB0E0E06FB3A" author="Gervais, P." box="[706,873,1188,1213]" pageId="2" pageNumber="153" refId="ref19883" refString="Gervais, P. (1844) Histoire Naturelle des Insectes. Apteres. Vol. 3. Librairie Encyclopedique de Roret, Paris, 494 pp." type="book" year="1844">Gervais 1844: 6</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="4A53D69BEF51D320FCEDFB0E0E80FB3A" author="Wood, H. C." box="[885,1007,1188,1212]" pageId="2" pageNumber="153" pagination="107 - 112" refId="ref21903" refString="Wood, H. C. (1863) Description of new species of North American Pedipalpi. Proceedings of the Academy of Natural Sciences of Philadelphia, 15, 107 - 112." type="journal article" year="1863">Wood 1863</bibRefCitation>
: unnumbered fig. on p. 680;
<treatmentCitation id="AF638D7BEF51D320FA83FB0E0D81FB5D" author="Butler, A. G." page="118" pageId="2" pageNumber="153" year="1873">
<bibRefCitation id="4A53D69BEF51D320FA83FB0E0D81FB5D" author="Butler, A. G." pageId="2" pageNumber="153" pagination="415 - 425" refId="ref19114" refString="Butler, A. G. (1873) List of the species of Galeodides, with description of a new species in the collec- tion of the British Museum. Transactions of the Entomological Society of London, 1873, 415 - 425" type="journal article" year="1873">Butler 1873: 118</bibRefCitation>
</treatmentCitation>
, plate 6 fig. 5;
<treatmentCitation id="AF638D7BEF51D320FE1EFB690F5BFB5D" author="Butler, A. G." box="[390,564,1219,1243]" page="314" pageId="2" pageNumber="153" year="1879">
<bibRefCitation id="4A53D69BEF51D320FE1EFB690F5BFB5D" author="Butler, A. G." box="[390,564,1219,1243]" pageId="2" pageNumber="153" pagination="313 - 316" refId="ref19158" refString="Butler, A. G. (1879) Respecting a new distinction between the species of the genus Phrynus of authors. Annals and Magazine of Natural History, Series 5, 4, 313 - 316. https: // doi. org / 10.1080 / 00222937908679834" type="journal article" year="1879">Butler 1879: 314</bibRefCitation>
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, 315; Mello-Leit„o 1931: 424.
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</paragraph>
<paragraph id="2E7DAB6AEF51D320FF0FFB480E3FFB7C" blockId="2.[151,1437,1158,1795]" box="[151,848,1250,1274]" pageId="2" pageNumber="153">
<treatmentCitationGroup id="0ED28C44EF51D320FF0FFB480E3FFB7C" box="[151,848,1250,1274]" pageId="2" pageNumber="153">
<taxonomicName id="E9C2D0E9EF51D320FF0FFB480CDFFB7C" authority="(Herbst)" baseAuthorityName="Herbst" box="[151,432,1250,1274]" class="Arachnida" family="Phrynidae" genus="Admetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="palmatus">
<emphasis id="1CB67778EF51D320FF0FFB480C3DFB7C" box="[151,338,1250,1274]" italics="true" pageId="2" pageNumber="153">Admetus palmatus</emphasis>
(Herbst)
</taxonomicName>
: C.L.
<treatmentCitation id="AF638D7BEF51D320FE75FB480FE9FB7C" author="Koch, C. L." box="[493,646,1250,1274]" page="81" pageId="2" pageNumber="153" year="1850">
<bibRefCitation id="4A53D69BEF51D320FE75FB480FE9FB7C" author="Koch, C. L." box="[493,646,1250,1274]" pageId="2" pageNumber="153" refId="ref20363" refString="Koch, C. L. (1850) Ubersicht des Arachnidensystems. Vol. 5. J. L. Lotzbeck, N ʾ rnberg, 552 pp." type="book" year="1850">Koch 1850: 81</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="AF638D7BEF51D320FD09FB480E23FB7C" author="Hansen, H. J." box="[657,844,1250,1274]" page="161" pageId="2" pageNumber="153" year="1893">
<bibRefCitation id="4A53D69BEF51D320FD09FB480E23FB7C" author="Hansen, H. J." box="[657,844,1250,1274]" pageId="2" pageNumber="153" pagination="135 - 251" refId="ref20030" refString="Hansen, H. J. (1893) Organs and characters in different orders of arachnids. Entomologiske Meddelelser, 4, 135 - 251." type="journal article" year="1893">Hansen 1893: 161</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="2E7DAB6AEF51D320FF0FFAAA0972FA9F" blockId="2.[151,1437,1158,1795]" box="[151,1053,1280,1305]" pageId="2" pageNumber="153">
<treatmentCitationGroup id="0ED28C44EF51D320FF0FFAAA0972FA9F" box="[151,1053,1280,1305]" pageId="2" pageNumber="153">
<taxonomicName id="E9C2D0E9EF51D320FF0FFAAA0CBCFA9E" authority="(Herbst)" baseAuthorityName="Herbst" box="[151,467,1280,1304]" class="Arachnida" family="Phrynidae" genus="Neophrynus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="palmatus">
<emphasis id="1CB67778EF51D320FF0FFAAA0C1AFA9E" box="[151,373,1280,1304]" italics="true" pageId="2" pageNumber="153">Neophrynus palmatus</emphasis>
(Herbst)
</taxonomicName>
:
<treatmentCitation id="AF638D7BEF51D320FE45FAAA0FA6FA9F" author="Kraepelin, K." box="[477,713,1280,1305]" page="30" pageId="2" pageNumber="153" year="1895">
<bibRefCitation id="4A53D69BEF51D320FE45FAAA0FA6FA9F" author="Kraepelin, K." box="[477,713,1280,1305]" pageId="2" pageNumber="153" pagination="3 - 53" refId="ref20393" refString="Kraepelin, K. (1895) Revision der Tarantuliden Fabr. Verhandlungen des Naturwissenschaftlichen Vereins in Hamburg, 13 (3), 3 - 53." type="journal article" year="1895">Kraepelin 1895: 3034</bibRefCitation>
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, figs 3, 12, 15, 19, 25, 2728, 30.
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</paragraph>
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<treatmentCitationGroup id="0ED28C44EF51D320FF0FFAB50CDBFA57" pageId="2" pageNumber="153">
<taxonomicName id="E9C2D0E9EF51D320FF0FFAB50CD9FAB1" authority="(Herbst)" baseAuthorityName="Herbst" box="[151,438,1311,1335]" class="Arachnida" family="Phrynidae" genus="Tarantula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="palmata">
<emphasis id="1CB67778EF51D320FF0FFAB50C3AFAB1" box="[151,341,1311,1335]" italics="true" pageId="2" pageNumber="153">Tarantula palmata</emphasis>
(Herbst)
</taxonomicName>
:
<treatmentCitation id="AF638D7BEF51D320FE59FAB50F88FAB1" author="Pereyaslawzewa, S." box="[449,743,1311,1335]" page="319" pageId="2" pageNumber="153" year="1897">
<bibRefCitation id="4A53D69BEF51D320FE59FAB50F88FAB1" author="Pereyaslawzewa, S." box="[449,743,1311,1335]" pageId="2" pageNumber="153" pagination="319 - 321" refId="ref20793" refString="Pereyaslawzewa, S. (1897 a) Les premiers stades du developpement de Pedipalpes. Comptes Rendus Hebdomadaires des Seances de l'Academie des Sciences, Paris, 125, 319 - 321." type="journal article" year="1897">Pereyaslawzewa 1897a: 319</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="AF638D7BEF51D320FD6BFAB50975FAB1" author="Pereyaslawzewa, S." box="[755,1050,1311,1335]" page="377" pageId="2" pageNumber="153" year="1897">
<bibRefCitation id="4A53D69BEF51D320FD6BFAB50975FAB1" author="Pereyaslawzewa, S." box="[755,1050,1311,1335]" pageId="2" pageNumber="153" pagination="377 - 380" refId="ref20827" refString="Pereyaslawzewa, S. (1897 b) Les derniers stades du developpement de Pedipalpes. Comptes Rendus Hebdomadaires des Seances de l'Academie des Sciences, Paris, 125, 377 - 380." type="journal article" year="1897">Pereyaslawzewa 1897b: 377</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="AF638D7BEF51D320FBBEFAB50992FAB1" author="Kraepelin, K." box="[1062,1277,1311,1335]" page="264" pageId="2" pageNumber="153" year="1901">
<bibRefCitation id="4A53D69BEF51D320FBBEFAB50992FAB1" author="Kraepelin, K." box="[1062,1277,1311,1335]" pageId="2" pageNumber="153" pagination="263 - 264" refId="ref20421" refString="Kraepelin, K. (1901) Catalogue des Pedipalpes des collections du Museum d'Histoire Naturelle de Paris. Bulletin du Museum National d'Histoire Naturelle, Paris, 7, 263 - 264." type="journal article" year="1901">Kraepelin 1901: 264</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="4A53D69BEF51D320FA91FA8A08F6FAB1" author="Borner, C." box="[1289,1433,1311,1335]" pageId="2" pageNumber="153" pagination="433 - 466" refId="ref18949" refString="Borner, C. (1902 a) Arachnologische Studien. (II und III). Zoologischer Anzeiger, 25, 433 - 466." type="journal article" year="1902">Börner 1902a</bibRefCitation>
: fig. 6;
<treatmentCitation id="AF638D7BEF51D320FE92FA940CA2FAD0" author="Borner, C." box="[266,461,1342,1366]" page="214" pageId="2" pageNumber="153" year="1902">
<bibRefCitation id="4A53D69BEF51D320FE92FA940CA2FAD0" author="Borner, C." box="[266,461,1342,1366]" pageId="2" pageNumber="153" pagination="214 - 215" refId="ref18975" refString="Borner, C. (1902 b) Koenenia mirabilis und andere Pedipalpen. Verhandlungen der Deutschen Zoologischen Gesellschaft, 12, 214 - 215." type="journal article" year="1902">Börner 1902b: 214</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="AF638D7BEF51D320FE4FFA940F1DFAD0" author="Borner, C." box="[471,626,1342,1366]" page="5" pageId="2" pageNumber="153" year="1904">
<bibRefCitation id="4A53D69BEF51D320FE4FFA940F1DFAD0" author="Borner, C." box="[471,626,1342,1366]" pageId="2" pageNumber="153" pagination="1 - 174" refId="ref19001" refString="Borner, C. (1904) Beitrage zur Morphologie der Arthropoden. I. Ein Beitrag zur Kenntnis der Pedipalpen. Zoologica, Stuttgart, 42, 1 - 174. https: // doi. org / 10.5962 / bhl. title. 59544" type="journal article" year="1904">Börner 1904: 5</bibRefCitation>
</treatmentCitation>
, figs 6, 18, 23, 26, 32, 57, 102114, figs 1011, 38, 46, 64, 85;
<treatmentCitation id="AF638D7BEF51D320FB6EFA940D8CFAF3" author="Comstock, J. H." page="20" pageId="2" pageNumber="153" year="1913">
<bibRefCitation id="4A53D69BEF51D320FB6EFA940D8CFAF3" author="Comstock, J. H." pageId="2" pageNumber="153" refId="ref19546" refString="Comstock, J. H. (1913) The spider book. Doubleday, Page and Co., New York, 746 pp." type="book" year="1913">Comstock 1913: 20</bibRefCitation>
</treatmentCitation>
, fig. 19;
<treatmentCitation id="AF638D7BEF51D320FEA5FAF60F43FAF2" author="Petrunkevitch, A." box="[317,556,1372,1396]" page="60" pageId="2" pageNumber="153" year="1925">
<bibRefCitation id="4A53D69BEF51D320FEA5FAF60F43FAF2" author="Petrunkevitch, A." box="[317,556,1372,1396]" pageId="2" pageNumber="153" pagination="51 - 248" refId="ref20861" refString="Petrunkevitch, A. (1925) Arachnida from Panama. Transactions of the Connecticut Academy of Arts and Sciences, 27, 51 - 248." type="journal article" year="1925">Petrunkevitch 1925: 60</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="4A53D69BEF51D320FDAEFAF70FAEFAF3" author="Kastner, A." box="[566,705,1373,1397]" pageId="2" pageNumber="153" pagination="1 - 76" refId="ref20269" refString="Kastner, A. (1932) Pedipalpi Latreille = Geissel Scorpione. In: K ʾ kenthal, W. (Ed.), Handbuch der Zoologie. Vol. 3 (2) (2). Walter de Gruyter &amp; Co., Berlin und Leipzig. pp. 1 - 76." type="book chapter" year="1932">Kästner 1932</bibRefCitation>
: figs 8, 23, 28, 31, 7374, 81;
<treatmentCitation id="AF638D7BEF51D320FC6DFAF709C2FAF2" author="Werner, F." box="[1013,1197,1372,1396]" page="475" pageId="2" pageNumber="153" year="1935">
<bibRefCitation id="4A53D69BEF51D320FC6DFAF709C2FAF2" author="Werner, F." box="[1013,1197,1372,1396]" pageId="2" pageNumber="153" pagination="317 - 490" refId="ref21772" refString="Werner, F. (1935) Klasse: Arachnoidea, Spinnentiere. Pedipalpen. In: Bronn, H. G. (Ed.), Klassen und Ordnungen des Tierreichs. Vol. 5 (IV) (8) (3). Akademische Verlagsgesellschaft, Leipzig, pp. 317 - 490." type="book chapter" year="1935">Werner 1935: 475</bibRefCitation>
</treatmentCitation>
, figs 25, 3435, 40, 47, 56, 59, 7273, 100, 110, 113, 141ab, 182b;
<treatmentCitation id="AF638D7BEF51D320FD0AFAD10E12FA15" author="Speijer, E. A. M." box="[658,893,1403,1427]" page="260" pageId="2" pageNumber="153" year="1936">
<bibRefCitation id="4A53D69BEF51D320FD0AFAD10E12FA15" author="Speijer, E. A. M." box="[658,893,1403,1427]" pageId="2" pageNumber="153" pagination="249 - 263" refId="ref21415" refString="Speijer, E. A. M. (1936) Die orientalischen Pedipalpi des Zoologischen Museums der Universitat Berlin. Mitteilung aus dem Zoologischen Museum in Berlin, 21, 249 - 263." type="journal article" year="1936">Speijer 1936: 260261</bibRefCitation>
</treatmentCitation>
, figs 1215;
<bibRefCitation id="4A53D69BEF51D320FB9AFAD109E5FA15" author="Millot, J." box="[1026,1162,1403,1427]" pageId="2" pageNumber="153" refId="ref20696" refString="Millot, J. (1949 a) Classe de Arachnides (Arachnida). I. Morphologie generale et anatomie interne. In: Grasse, P. P. (Ed.), Traite de zoologie. Vol. 6. Masson, Paris, pp. 263 - 319." type="book" year="1949">Millot 1949a</bibRefCitation>
: figs 68, 71;
<bibRefCitation id="4A53D69BEF51D320FA97FAD108F6FA15" author="Millot, J." box="[1295,1433,1403,1427]" pageId="2" pageNumber="153" refId="ref20750" refString="Millot, J. (1949 b) Ordre des Amblypyges. In: Grasse, P. P. (Ed.), Traite de zoologie. Vol. 6. Masson, Paris, pp. 563 - 588." type="book" year="1949">Millot 1949b</bibRefCitation>
: figs 326, 328, 333, 344;
<bibRefCitation id="4A53D69BEF51D320FE51FA300FFBFA34" author="Petrunkevitch, A." box="[457,660,1434,1458]" pageId="2" pageNumber="153" pagination="69 - 315" refId="ref20888" refString="Petrunkevitch, A. (1949) A study of Palaeozoic Arachnida. Transactions of the Connecticut Academy of Arts and Sciences, 37, 69 - 315." type="journal article" year="1949">Petrunkevitch 1949</bibRefCitation>
, fig. 17;
<treatmentCitation id="AF638D7BEF51D320FD6CFA300EA1FA34" author="Caporiacco, L. di" box="[756,974,1434,1458]" page="36" pageId="2" pageNumber="153" year="1951">
<bibRefCitation id="4A53D69BEF51D320FD6CFA300EA1FA34" author="Caporiacco, L. di" box="[756,974,1434,1458]" pageId="2" pageNumber="153" pagination="1 - 46" refId="ref19207" refString="Caporiacco, L. di (1951) Studi sugli Aracnidi del Venezuela raccolti dalla Sezione di Biologia (Universita Centrale del Venezuela). 1 Parte: Scorpiones, Opiliones, Solifuga y Chernetes. Acta Biologica Venezuelica, 1, 1 - 46." type="journal article" year="1951">Caporiacco 1951: 36</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="4A53D69BEF51D320FC42FA3009E3FA34" author="Zakhvatkin, A. A." box="[986,1164,1434,1458]" pageId="2" pageNumber="153" pagination="5 - 46" refId="ref21937" refString="Zakhvatkin, A. A. (1952) Subdivision of the mites (Acarina) into orders and the position of these in the system of the Chelicerata. Parazitologicheskii Sbornik Zoologicheskii Institut Akademii Nauk SSSR, 12, 5 - 46." type="journal article" year="1952">Zakhvatkin 1952</bibRefCitation>
: fig. 28b;
<treatmentCitation id="AF638D7BEF51D320FB60FA300C4CFA56" author="Weygoldt, P." page="273" pageId="2" pageNumber="153" year="1977">
<bibRefCitation id="4A53D69BEF51D320FB60FA300C4CFA56" author="Weygoldt, P." pageId="2" pageNumber="153" pagination="271 - 286" refId="ref21830" refString="Weygoldt, P. (1977) Kampf, Paarungsverhalten, Spermatophorenmorphologie und weibliche Genitalien bei neotropischen Geisselspinnen (Amblypygi, Arachnida). Zoomorphologie, 86, 271 - 286. https: // doi. org / 10.1007 / BF 00993669" type="journal article" year="1977">Weygoldt 1977: 273275</bibRefCitation>
</treatmentCitation>
, figs 1ac, 3a.
</treatmentCitationGroup>
</paragraph>
<paragraph id="2E7DAB6AEF51D320FF0FFA7D0F10F940" blockId="2.[151,1437,1158,1795]" pageId="2" pageNumber="153">
<treatmentCitationGroup id="0ED28C44EF51D320FF0FFA7D0F10F940" pageId="2" pageNumber="153">
<treatmentCitation id="AF638D7BEF51D320FF0FFA7D0F4FFA69" author="Pocock, R. I." box="[151,544,1495,1519]" page="52" pageId="2" pageNumber="153" year="1902">
<taxonomicName id="E9C2D0E9EF51D320FF0FFA7D0F4FFA69" ID-CoL="6VDQD" authority="Pocock, 1902: 52" authorityName="Pocock" authorityPageNumber="52" authorityYear="1902" box="[151,544,1495,1519]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="operculatus">
<emphasis id="1CB67778EF51D320FF0FFA7D0C07FA69" box="[151,360,1495,1519]" italics="true" pageId="2" pageNumber="153">Phrynus operculatus</emphasis>
<bibRefCitation id="4A53D69BEF51D320FEF6FA7D0F4FFA69" author="Pocock, R. I." box="[366,544,1495,1519]" pageId="2" pageNumber="153" refId="ref21088" refString="Pocock, R. I. (1902) Arachnida: Scorpiones, Pedipalpi &amp; Solifugae. In: Godman, F. D. &amp; Salvin, O. (Eds.), Biologia Centrali Americana. Vol. 3. Taylor and Francis, London, pp. 1 - 71." type="book" year="1902">Pocock, 1902: 52</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, plate 10 figs 8, 8ac;
<treatmentCitation id="AF638D7BEF51D320FC99FA7D0E94FA69" author="Franganillo, P." box="[769,1019,1495,1519]" page="66" pageId="2" pageNumber="153" year="1926">
<bibRefCitation id="4A53D69BEF51D320FC99FA7D0E94FA69" author="Franganillo, P." box="[769,1019,1495,1519]" pageId="2" pageNumber="153" pagination="42 - 68" refId="ref19780" refString="Franganillo, P. (1926) Aracnidos nuevos o poco conocidos de la Isla de Cuba. Boletin de la Sociedad Entomologica de Espana, 9, 42 - 68." type="journal article" year="1926">Franganillo 1926: 6667</bibRefCitation>
</treatmentCitation>
(probable misid);
<treatmentCitation id="AF638D7BEF51D320FB2DFA7D08F7FA69" author="Franganillo, P." box="[1205,1432,1495,1519]" page="120" pageId="2" pageNumber="153" year="1931">
<bibRefCitation id="4A53D69BEF51D320FB2DFA7D08F7FA69" author="Franganillo, P." box="[1205,1432,1495,1519]" pageId="2" pageNumber="153" pagination="44 - 49" refId="ref19812" refString="Franganillo, P. (1931) Excursiones aracnologicas durante el mes de Agosto de 1930. Revista Belen, La Habana, 24, 44 - 49 + 116 - 120 + 168 - 171 + 219 - 224 + 285 - 288." type="journal article" year="1931">Franganillo 1931: 120</bibRefCitation>
</treatmentCitation>
; Mello-Leit„o 1931: 42;
<treatmentCitation id="AF638D7BEF51D320FE20FA5C0FF3F988" author="Franganillo, P." box="[440,668,1526,1550]" page="149" pageId="2" pageNumber="153" year="1936">
<bibRefCitation id="4A53D69BEF51D320FE20FA5C0FF3F988" author="Franganillo, P." box="[440,668,1526,1550]" pageId="2" pageNumber="153" refId="ref19857" refString="Franganillo, P. (1936) Los Aracnidos de Cuba hasta 1936. Cultural S. A., La Habana, 178 pp." type="book" year="1936">Franganillo 1936: 149</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="AF638D7BEF51D320FD3EFA5C0ECFF988" author="Quintero, D." box="[678,928,1526,1550]" page="155" pageId="2" pageNumber="153" year="1981">
<bibRefCitation id="4A53D69BEF51D320FD3EFA5C0ECFF988" author="Quintero, D." box="[678,928,1526,1550]" pageId="2" pageNumber="153" pagination="117 - 166" refId="ref21143" refString="Quintero, D. (1981) The Amblypygid genus Phrynus in the Americas (Amblypygi, Phrynidae). Journal of Arachnology, 9, 117 - 166." type="journal article" year="1981">Quintero 1981: 155157</bibRefCitation>
</treatmentCitation>
, figs 102108, 154155, 157, 162, map;
<treatmentCitation id="AF638D7BEF51D320FAA4FA5C0C12F9AB" author="Coddington, J. A. &amp; Larcher, S. F. &amp; Cokendolpher, J. C." page="11" pageId="2" pageNumber="153" year="1990">
<bibRefCitation id="4A53D69BEF51D320FAA4FA5C0C12F9AB" author="Coddington, J. A. &amp; Larcher, S. F. &amp; Cokendolpher, J. C." pageId="2" pageNumber="153" pagination="5 - 20" refId="ref19415" refString="Coddington, J. A., Larcher, S. F. &amp; Cokendolpher, J. C. (1990) The systematic status of Arachnida, exclusive of Acari, in North America north of Mexico. In: Kosztarab, M. &amp; Schaefer, C. W. (Eds.), Systematics of the North American insects and arachnids: status and needs. Virginia Polytechnic Institute and State University, Blacksburg, pp. 5 - 20." type="book chapter" year="1990">
Coddington
<emphasis id="1CB67778EF51D320FF75F9BF0C70F9AA" box="[237,287,1556,1580]" italics="true" pageId="2" pageNumber="153">et al.</emphasis>
1990: 11
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="AF638D7BEF51D320FE1EF9BE0F11F9AB" author="Vazquez-Rojas, I." box="[390,638,1556,1581]" page="30" pageId="2" pageNumber="153" year="1995">
<bibRefCitation id="4A53D69BEF51D320FE1EF9BE0F11F9AB" author="Vazquez-Rojas, I." box="[390,638,1556,1581]" pageId="2" pageNumber="153" pagination="15 - 37" refId="ref21499" refString="Vazquez-Rojas, I. (1995) Los aracnidos de Mexico parte 1: Ricinulei, Amblypygi, Solifugae, Palpigradi, Schizomida, Uropygi. Dugesiana, 2, 15 - 37." type="journal article" year="1995">Vázquez-Rojas 1995: 30</bibRefCitation>
</treatmentCitation>
(as
<taxonomicName id="E9C2D0E9EF51D320FD3FF9BE0E1BF9AA" box="[679,884,1556,1580]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="opercularis">
<emphasis id="1CB67778EF51D320FD3FF9BE0E1BF9AA" box="[679,884,1556,1580]" italics="true" pageId="2" pageNumber="153">Phrynus opercularis</emphasis>
</taxonomicName>
[sic]);
<treatmentCitation id="AF638D7BEF51D320FC22F9BE09DAF9AB" author="Vazquez-Rojas, I." box="[954,1205,1556,1581]" page="72" pageId="2" pageNumber="153" year="1996">
<bibRefCitation id="4A53D69BEF51D320FC22F9BE09DAF9AB" author="Vazquez-Rojas, I." box="[954,1205,1556,1581]" pageId="2" pageNumber="153" pagination="71 - 73" refId="ref21533" refString="Vazquez-Rojas, I. (1996) Amblypygi. In: Llorente Bousquets, J. E., Garcia Aldrete, A. N. &amp; Gonzalez Soriano, E. (Eds.), Biodiversidad, taxonomia y biogeografia de artropodos de Mexico: hacia una sintesis de su conocimiento. Universidad Nacional Autonoma de Mexico, Mexico, D. F, pp. 71 - 73." type="book chapter" year="1996">Vázquez-Rojas 1996: 72</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="AF638D7BEF51D320FB27F9BC0C4CF9CD" author="Avila-Calvo, A. F. &amp; Armas, L. F. de" page="32" pageId="2" pageNumber="153" year="1997">
<bibRefCitation id="4A53D69BEF51D320FB27F9BC0C4CF9CD" author="Avila-Calvo, A. F. &amp; Armas, L. F. de" pageId="2" pageNumber="153" pagination="31 - 32" refId="ref18907" refString="Avila-Calvo, A. F. &amp; Armas, L. F. de (1997) Lista de los Amblipigidos (Arachnida: Amblypygi) de Mexico, Centroamerica y las Antillas. Cocuyo, 6, 31 - 32." type="journal article" year="1997">Ávila-Calvo &amp; Armas 1997: 32</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="AF638D7BEF51D320FEB5F9990CBFF9CD" author="Armas, L. F." box="[301,464,1587,1611]" page="29" pageId="2" pageNumber="153" year="1999">
<bibRefCitation id="4A53D69BEF51D320FEB5F9990CBFF9CD" author="Armas, L. F." box="[301,464,1587,1611]" pageId="2" pageNumber="153" pagination="29 - 30" refId="ref18521" refString="Armas, L. F. (1999) Ampliacion del area de distribucion de algunos Phrynus (Amblypygi: Phrynidae) y Centruroides (Scorpiones: Buthidae) de America Central y las Antillas. Cocuyo, 3, 29 - 30." type="journal article" year="1999">Armas 1999: 29</bibRefCitation>
</treatmentCitation>
(synonymized with
<taxonomicName id="E9C2D0E9EF51D320FD38F9990EE8F9CD" authorityName="Herbst" authorityYear="1797" box="[672,903,1587,1611]" class="Arachnida" family="Phrynidae" genus="Phalangium" higherTaxonomySource="CoL" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="palmatum">
<emphasis id="1CB67778EF51D320FD38F9990EE8F9CD" box="[672,903,1587,1611]" italics="true" pageId="2" pageNumber="153">Phalangium palmatum</emphasis>
</taxonomicName>
by
<bibRefCitation id="4A53D69BEF51D320FC35F999093BF9CD" author="Quintero, D." box="[941,1108,1587,1611]" pageId="2" pageNumber="153" pagination="117 - 166" refId="ref21143" refString="Quintero, D. (1981) The Amblypygid genus Phrynus in the Americas (Amblypygi, Phrynidae). Journal of Arachnology, 9, 117 - 166." type="journal article" year="1981">Quintero (1981)</bibRefCitation>
, but treated as senior synonym);
<treatmentCitation id="AF638D7BEF51D320FF5FF9F80CB4F9EC" author="Armas, L. F. &amp; Viquez, C." box="[199,475,1618,1642]" page="15" pageId="2" pageNumber="153" year="2001">
<bibRefCitation id="4A53D69BEF51D320FF5FF9F80CB4F9EC" author="Armas, L. F. &amp; Viquez, C." box="[199,475,1618,1642]" pageId="2" pageNumber="153" pagination="11 - 15" refId="ref18822" refString="Armas, L. F. &amp; Viquez, C. (2001) Nueva especie de Phrynus (Amblypygi: Phrynidae) de Costa Rica. Revista Iberica de Aracnologia, 4, 11 - 15." type="journal article" year="2001">Armas &amp; Víquez 2001: 15</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="AF638D7BEF51D320FE7EF9F80F85F9EC" author="Armas, L. F. &amp; Perez, A." box="[486,746,1618,1642]" page="48" pageId="2" pageNumber="153" year="2001">
<bibRefCitation id="4A53D69BEF51D320FE7EF9F80F85F9EC" author="Armas, L. F. &amp; Perez, A." box="[486,746,1618,1642]" pageId="2" pageNumber="153" pagination="47 - 66" refId="ref18786" refString="Armas, L. F. &amp; Perez, A. (2001) Los Amblipigidos de Republica Dominicana (Arachnida: Amblypygi). Revista Iberica de Aracnologia, 3, 47 - 66." type="journal article" year="2001">Armas &amp; Pérez 2001: 48</bibRefCitation>
</treatmentCitation>
; Jiménez &amp; Gutierrez 2004: 863;
<bibRefCitation id="4A53D69BEF51D320FBD1F9F809B8F9EC" author="Harvey, M. S." box="[1097,1239,1618,1642]" pageId="2" pageNumber="153" refId="ref20098" refString="Harvey, M. S. (2003) Catalogue of the smaller arachnid orders of the world: Amblypygi, Uropygi, Schizomida, Palpigradi, Ricinulei and Solifugae. CSIRO Publishing, Collingwood, Victoria, 385 pp. https: // doi. org / 10.1071 / 9780643090071" type="book" year="2003">Harvey, 2003</bibRefCitation>
: Order
<taxonomicName id="E9C2D0E9EF51D320FABAF9F808F7F9EC" ID-CoL="NV" authorityName="Thorell" authorityYear="1883" box="[1314,1432,1618,1642]" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="153" phylum="Arthropoda" rank="order">Amblypygi</taxonomicName>
, 2829;
<treatmentCitation id="AF638D7BEF51D320FE8CF9DB0F33F90E" author="Armas, L. F. &amp; Gadar, Y." box="[276,604,1648,1673]" page="134" pageId="2" pageNumber="153" year="2004">
<bibRefCitation id="4A53D69BEF51D320FE8CF9DB0F33F90E" author="Armas, L. F. &amp; Gadar, Y." box="[276,604,1648,1673]" pageId="2" pageNumber="153" pagination="136" refId="ref18701" refString="Armas, L. F. &amp; Gadar, Y. (2004) Nueva especie de Phrynus Lamarck, 1801 (Amblypygi: Phrynidae) de Chiapas, Mexico. Revista Iberica de Aracnologia, 10, 133 ‾ 136." type="journal article" year="2004">Armas &amp; Gadar 2004: 134136</bibRefCitation>
</treatmentCitation>
, fig. 8;
<treatmentCitation id="AF638D7BEF51D320FD35F9DB0963F90E" author="Viquez, C. &amp; Armas, L. F." box="[685,1036,1648,1673]" page="307" pageId="2" pageNumber="153" year="2006">
<bibRefCitation id="4A53D69BEF51D320FD35F9DB0963F90E" author="Viquez, C. &amp; Armas, L. F." box="[685,1036,1648,1673]" pageId="2" pageNumber="153" pagination="307 - 318" refId="ref21716" refString="Viquez, C. &amp; Armas, L. F. (2006) Los Amblipigidos (Arachnida: Amblypygi) de Guatemala. In: Cano, E. (Ed.), Biodiversidad de Guatemala. Vol 1. Universidad del Valle de Guatemala, Guatemala, pp. 307 - 318." type="book chapter" year="2006">Víquez &amp; Armas 2006: 3073016</bibRefCitation>
</treatmentCitation>
, figs. 9, 13;
<treatmentCitation id="AF638D7BEF51D320FB15F9DB0821F90F" author="Armas, L. F." box="[1165,1358,1648,1673]" page="232" pageId="2" pageNumber="153" year="2006">
<bibRefCitation id="4A53D69BEF51D320FB15F9DB0821F90F" author="Armas, L. F." box="[1165,1358,1648,1673]" pageId="2" pageNumber="153" pagination="223 - 245" refId="ref18565" refString="Armas, L. F. (2006 a) Sinopsis de los Amblipigidos Antillanos (Arachnida: Amblypygi). Boletin Sociedad Entomologica Aragonesa, 38, 223 - 245." type="journal article" year="2006">Armas 2006a: 232</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="AF638D7BEF51D320FAC1F9DB0C00F921" author="Armas, L. F." page="345" pageId="2" pageNumber="153" year="2006">
<bibRefCitation id="4A53D69BEF51D320FAC1F9DB0C00F921" author="Armas, L. F." pageId="2" pageNumber="153" pagination="345 - 359" refId="ref18597" refString="Armas, L. F. (2006 b) Los Amblypygidos Tendarapodos de Mexico (Arachnida: Amblypygi). Boletin Sociedad Entomologica Aragonesa, 39, 345 - 359." type="journal article" year="2006">Armas 2006b: 345358</bibRefCitation>
</treatmentCitation>
, fig. 18,
<tableCitation id="63409ED1EF51D320FE5EF9250F7EF921" box="[454,529,1679,1703]" captionStart="TABLE 1" captionStartId="7.[151,239,151,175]" captionText="TABLE 1. Measurements (mm) of body, pedipalps segments, and femur of legs in Phrynus operculatus. When the segment is missing, the cell is empty." pageId="2" pageNumber="153">Table 1</tableCitation>
;
<treatmentCitation id="AF638D7BEF51D320FD86F9250E58F921" author="Chirivi, D. &amp; Armas, L. F." box="[542,823,1679,1703]" page="396" pageId="2" pageNumber="153" year="2012">
<bibRefCitation id="4A53D69BEF51D320FD86F9250E58F921" author="Chirivi, D. &amp; Armas, L. F." box="[542,823,1679,1703]" pageId="2" pageNumber="153" pagination="395 - 402" refId="ref19317" refString="Chirivi, D. &amp; Armas, L. F. (2012) La subfamilia Phryninae (Amblypygi: Phrynidae) en Colombia. Boletin de la Sociedad Entomologica Aragonesa, 50, 395 - 402." type="journal article" year="2012">Chirivi &amp; Armas 2012: 396</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="AF638D7BEF51D320FCA7F925092CF921" author="Armas, L. F. &amp; Viquez, C. &amp; Trujillo, R." box="[831,1091,1679,1703]" page="25" pageId="2" pageNumber="153" year="2013">
<bibRefCitation id="4A53D69BEF51D320FCA7F925092CF921" author="Armas, L. F. &amp; Viquez, C. &amp; Trujillo, R." box="[831,1091,1679,1703]" pageId="2" pageNumber="153" pagination="25 - 31" refId="ref18860" refString="Armas, L. F., Viquez, C. &amp; Trujillo, R. (2013) Nueva especie de Phrynus Lamarck, 1801 (Amblypygi: Phrynidae) de Guatemala y Honduras. Revista Iberica de Aracnologia, 23, 25 - 31." type="journal article" year="2013">
Armas
<emphasis id="1CB67778EF51D320FC11F93A0ED3F921" box="[905,956,1679,1703]" italics="true" pageId="2" pageNumber="153">et al.</emphasis>
2013: 2530
</bibRefCitation>
</treatmentCitation>
; Armas 2014: 3035;
<treatmentCitation id="AF638D7BEF51D320FAB8F9250C40F940" author="Armas, L. F. &amp; Guzman, A. &amp; Francke, O." page="3" pageId="2" pageNumber="153" year="2014">
<bibRefCitation id="4A53D69BEF51D320FAB8F9250C40F940" author="Armas, L. F. &amp; Guzman, A. &amp; Francke, O." pageId="2" pageNumber="153" pagination="3 - 7" refId="ref18743" refString="Armas, L. F., Guzman, A. &amp; Francke, O. (2014) Una especie nueva de Phrynus (Amblypygi: Phrynidae) de Mexico. Revista Iberica de Aracnologia, 25, 3 - 7." type="journal article" year="2014">
Armas
<emphasis id="1CB67778EF51D320FAF2F93A08F2F921" box="[1386,1437,1679,1703]" italics="true" pageId="2" pageNumber="153">et al.</emphasis>
2014: 36
</bibRefCitation>
</treatmentCitation>
, fig. 3;
<treatmentCitation id="AF638D7BEF51D320FEE5F9040F14F940" author="Guzman, A. &amp; Chirivi, D. &amp; Francke, O." box="[381,635,1710,1734]" page="478" pageId="2" pageNumber="153" year="2015">
<bibRefCitation id="4A53D69BEF51D320FEE5F9040F14F940" author="Guzman, A. &amp; Chirivi, D. &amp; Francke, O." box="[381,635,1710,1734]" pageId="2" pageNumber="153" pagination="474 - 482" refId="ref19971" refString="Guzman, A., Chirivi, D. &amp; Francke, O. (2015) The first troglomorphic species of the genus Phrynus Lamarck, 1801 (Amblypygi: Phrynidae) from Mexico. Zootaxa, 3920 (3), 474 - 482. https: // doi. org / 10.11646 / zootaxa. 3920.3.6" type="journal article" year="2015">
Guzmán
<emphasis id="1CB67778EF51D320FE41F9050F63F940" box="[473,524,1710,1734]" italics="true" pageId="2" pageNumber="153">et al.</emphasis>
2015: 478
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="2E7DAB6AEF51D320FF0FF9670939F962" blockId="2.[151,1437,1158,1795]" box="[151,1110,1740,1765]" pageId="2" pageNumber="153">
<treatmentCitationGroup id="0ED28C44EF51D320FF0FF9670939F962" box="[151,1110,1740,1765]" pageId="2" pageNumber="153">
Not
<taxonomicName id="E9C2D0E9EF51D320FF5CF9660CB5F962" authority="(Herbst)" baseAuthorityName="Herbst" box="[196,474,1740,1764]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="palmatus">
<emphasis id="1CB67778EF51D320FF5CF9660C14F962" box="[196,379,1740,1764]" italics="true" pageId="2" pageNumber="153">Phrynus palmatus</emphasis>
(Herbst)
</taxonomicName>
:
<treatmentCitation id="AF638D7BEF51D320FE7BF9660FCDF962" author="Koch, C. L." box="[483,674,1740,1765]" page="13" pageId="2" pageNumber="153" year="1840">
<bibRefCitation id="4A53D69BEF51D320FE7BF9660FCDF962" author="Koch, C. L." box="[483,674,1740,1765]" pageId="2" pageNumber="153" refId="ref20326" refString="Koch, C. L. (1840) Die Arachniden. Getreu nach der Natur Abgebildet und Beschrieben. Vol. 8. C. H. Zeh'schen, N ʾ rnberg, 220 pp." type="book" year="1840">Koch 1840: 1315</bibRefCitation>
</treatmentCitation>
, fig. 601 (see
<taxonomicName id="E9C2D0E9EF51D320FCA9F966093DF962" ID-CoL="77DQ7" authority="(Pocock))" authorityName="" baseAuthorityName="Pocock" baseAuthorityYear="1894" box="[817,1106,1740,1764]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="gervaisii">
<emphasis id="1CB67778EF51D320FCA9F9660E8AF962" box="[817,997,1740,1764]" italics="true" pageId="2" pageNumber="153">Phrynus gervaisii</emphasis>
(Pocock))
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="66D8F8E1EF51D321FF0FF9410CB4FF53" lastPageId="3" lastPageNumber="154" pageId="2" pageNumber="153" type="materials_examined">
<paragraph id="2E7DAB6AEF51D320FF0FF9410CCAF885" blockId="2.[151,1437,1158,1795]" box="[151,421,1771,1795]" pageId="2" pageNumber="153">
<materialsCitation id="9EAAA137EF51D320FF0FF9410CCEF885" ID-GBIF-Occurrence="3064396310" box="[151,417,1771,1795]" country="Guatemala" location="Guatemala" pageId="2" pageNumber="153" specimenCount="1" typeStatus="holotype">
<emphasis id="1CB67778EF51D320FF0FF9410C45F885" bold="true" box="[151,298,1771,1795]" pageId="2" pageNumber="153">
<typeStatus id="F17915C8EF51D320FF0FF9410DA3F885" box="[151,204,1771,1795]" pageId="2" pageNumber="153">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="56D5EBFAEF51D320FEA8F9410CCEF885" box="[304,417,1771,1795]" name="Guatemala" pageId="2" pageNumber="153">Guatemala</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="2E7DAB6AEF51D321FF0FF89B0CB4FF53" blockId="2.[151,1436,1841,2012]" lastBlockId="3.[151,1436,150,357]" lastPageId="3" lastPageNumber="154" pageId="2" pageNumber="153">
<emphasis id="1CB67778EF51D320FF0FF89B0F3AF8CD" bold="true" box="[151,597,1841,1867]" pageId="2" pageNumber="153">
Revised material: (
<specimenCount id="38C460E3EF51D320FEE0F89B0C86F8CD" box="[376,489,1841,1867]" pageId="2" pageNumber="153" type="female">4 females</specimenCount>
,
<specimenCount id="38C460E3EF51D320FE6DF89B0F27F8CD" box="[501,584,1841,1867]" pageId="2" pageNumber="153" type="male">1 male</specimenCount>
):
</emphasis>
<materialsCitation id="9EAAA137EF51D320FDC4F89B0E81F8F6" ID-GBIF-Occurrence="3064396317" collectingDate="1972-08-23" collectionCode="CASENT" collectorName="C. Mullinex &amp; K. Lucas" country="Mexico" elevation="305" latitude="16.235832" location="Arriaga" longLatPrecision="1" longitude="-93.89972" pageId="2" pageNumber="153" specimenCode="CASENT 9060540" specimenCount="1" stateProvince="Chiapas">
<collectingCountry id="56D5EBFAEF51D320FDC4F89B0FA1F8CD" box="[604,718,1841,1867]" name="Mexico" pageId="2" pageNumber="153">MEXICO</collectingCountry>
.
<emphasis id="1CB67778EF51D320FD4FF89B0E5BF8CD" box="[727,820,1841,1867]" italics="true" pageId="2" pageNumber="153">
<collectingRegion id="EC066588EF51D320FD4FF89B0E5BF8CD" box="[727,820,1841,1867]" country="Mexico" name="Chiapas" pageId="2" pageNumber="153">Chiapas</collectingRegion>
</emphasis>
:
<location id="2B1DFDB1EF51D320FCD9F89B0EF8F8CD" LSID="urn:lsid:plazi:treatment:A66B1A7CEF51D324FF0FFBB70945FCBF:2B1DFDB1EF51D320FCD9F89B0EF8F8CD" box="[833,919,1841,1867]" country="Mexico" latitude="16.235832" longLatPrecision="1" longitude="-93.89972" name="Arriaga" pageId="2" pageNumber="153" stateProvince="Chiapas">Arriaga</location>
(
<geoCoordinate id="4BF6CDADEF51D320FC3DF89B094FF8CA" box="[933,1056,1841,1868]" degrees="16.235833" direction="north" orientation="latitude" pageId="2" pageNumber="153" precision="1" value="16.235832">16.235833</geoCoordinate>
,
<geoCoordinate id="4BF6CDADEF51D320FBB3F89B09DEF8CD" box="[1067,1201,1841,1867]" degrees="93.899722" direction="west" orientation="longitude" pageId="2" pageNumber="153" precision="1" value="-93.89972">-93.899722</geoCoordinate>
):
<quantity id="E93A068FEF51D320FB5BF89B0866F8CD" box="[1219,1289,1841,1867]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="153" unit="km" value="10.0">10 km</quantity>
at north,
<quantity id="E93A068FEF51D320FAEAF89B0DA0F8E9" metricMagnitude="2" metricUnit="m" metricValue="3.05" pageId="2" pageNumber="153" unit="m" value="305.0">
<elevation id="A5EF4C59EF51D320FAEAF89B0DA0F8E9" metricMagnitude="2" metricUnit="m" metricValue="3.05" pageId="2" pageNumber="153" unit="m" value="305.0">305 masl</elevation>
</quantity>
,
<date id="5A7C8DAAEF51D320FF43F8FF0CFDF8E9" box="[219,402,1877,1904]" pageId="2" pageNumber="153" value="1972-08-23">
<collectingDate id="4A387442EF51D320FF43F8FF0CFDF8E9" box="[219,402,1877,1904]" pageId="2" pageNumber="153" value="1972-08-23">23 August 1972</collectingDate>
</date>
,
<collectorName id="8337CEBCEF51D320FE07F8FF0F44F8E9" box="[415,555,1877,1903]" pageId="2" pageNumber="153">C. Mullinex</collectorName>
,
<collectorName id="8337CEBCEF51D320FDAFF8FC0FCFF8F6" box="[567,672,1878,1904]" pageId="2" pageNumber="153">K. Lucas</collectorName>
, female (
<specimenCode id="7E640311EF51D320FC90F8FF0E89F8F6" box="[776,998,1877,1904]" collectionCode="CASENT" pageId="2" pageNumber="153">CASENT 9060540</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="9EAAA137EF51D320FC63F8FF0E4DF812" ID-GBIF-Occurrence="3064396309" collectingDate="1972-08-23" collectionCode="CASENT" collectorName="C. Mullinex &amp; Lucas" country="Mexico" elevation="305" pageId="2" pageNumber="153" specimenCode="CASENT 9060527" specimenCount="1" stateProvince="Chiapas">
<quantity id="E93A068FEF51D320FC63F8FF092CF8E9" box="[1019,1091,1877,1903]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="153" unit="km" value="10.0">10 km</quantity>
at north,
<quantity id="E93A068FEF51D320FB2AF8FF0873F8E9" box="[1202,1308,1877,1904]" metricMagnitude="2" metricUnit="m" metricValue="3.05" pageId="2" pageNumber="153" unit="m" value="305.0">
<elevation id="A5EF4C59EF51D320FB2AF8FF0873F8E9" box="[1202,1308,1877,1904]" metricMagnitude="2" metricUnit="m" metricValue="3.05" pageId="2" pageNumber="153" unit="m" value="305.0">305 masl</elevation>
</quantity>
,
<date id="5A7C8DAAEF51D320FAB1F8FF0DBDF815" pageId="2" pageNumber="153" value="1972-08-23">
<collectingDate id="4A387442EF51D320FAB1F8FF0DBDF815" pageId="2" pageNumber="153" value="1972-08-23">23 August 1972</collectingDate>
</date>
,
<collectorName id="8337CEBCEF51D320FF45F8D30C08F815" box="[221,359,1913,1939]" pageId="2" pageNumber="153">C. Mullinex</collectorName>
, K.
<collectorName id="8337CEBCEF51D320FE0BF8D00CB6F812" box="[403,473,1914,1940]" pageId="2" pageNumber="153">Lucas</collectorName>
, female (
<specimenCode id="7E640311EF51D320FDA6F8D30E75F812" box="[574,794,1913,1940]" collectionCode="CASENT" pageId="2" pageNumber="153">CASENT 9060527</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="9EAAA137EF51D320FCB4F8D30F59F83E" ID-GBIF-Occurrence="3064396302" collectingDate="1972-08-23" collectionCode="CASENT" collectorName="C. Mullinex &amp; Lucas" country="Mexico" elevation="305" pageId="2" pageNumber="153" specimenCode="CASENT 9060512" specimenCount="1" stateProvince="Chiapas">
<quantity id="E93A068FEF51D320FCB4F8D30E1CF815" box="[812,883,1913,1939]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="153" unit="km" value="10.0">10 km</quantity>
at north,
<quantity id="E93A068FEF51D320FC45F8D30929F815" box="[989,1094,1913,1940]" metricMagnitude="2" metricUnit="m" metricValue="3.05" pageId="2" pageNumber="153" unit="m" value="305.0">
<elevation id="A5EF4C59EF51D320FC45F8D30929F815" box="[989,1094,1913,1940]" metricMagnitude="2" metricUnit="m" metricValue="3.05" pageId="2" pageNumber="153" unit="m" value="305.0">305 masl</elevation>
</quantity>
,
<date id="5A7C8DAAEF51D320FBC8F8D3086BF815" box="[1104,1284,1913,1940]" pageId="2" pageNumber="153" value="1972-08-23">
<collectingDate id="4A387442EF51D320FBC8F8D3086BF815" box="[1104,1284,1913,1940]" pageId="2" pageNumber="153" value="1972-08-23">23 August 1972</collectingDate>
</date>
,
<collectorName id="8337CEBCEF51D320FA96F8D308F6F815" box="[1294,1433,1913,1939]" pageId="2" pageNumber="153">C. Mullinex</collectorName>
, K.
<collectorName id="8337CEBCEF51D320FF22F8340C6FF83E" box="[186,256,1950,1976]" pageId="2" pageNumber="153">Lucas</collectorName>
, male (
<specimenCode id="7E640311EF51D320FEC9F8370F42F83E" box="[337,557,1949,1976]" collectionCode="CASENT" pageId="2" pageNumber="153">CASENT 9060512</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="9EAAA137EF51D320FDD8F8370926F85A" ID-GBIF-Occurrence="3064396311" collectingDate="1974-10-05" collectionCode="CASENT" collectorName="D. E. Breedlove &amp; J. A. Breedlove" country="Mexico" elevation="610" latitude="16.089722" location="Cerro Vernal" longLatPrecision="1" longitude="-93.75472" municipality="Tonala" pageId="2" pageNumber="153" specimenCode="CASENT 9060511" specimenCount="1" stateProvince="Chiapas">
<collectingMunicipality id="CE193110EF51D320FDD8F8370FE3F831" box="[576,652,1949,1975]" pageId="2" pageNumber="153">Tonalá</collectingMunicipality>
(
<geoCoordinate id="4BF6CDADEF51D320FD03F8370E79F831" box="[667,790,1949,1975]" degrees="16.089722" direction="north" orientation="latitude" pageId="2" pageNumber="153" precision="1" value="16.089722">16.089722</geoCoordinate>
,
<geoCoordinate id="4BF6CDADEF51D320FCBAF8370EC7F83E" box="[802,936,1949,1976]" degrees="93.754722" direction="west" orientation="longitude" pageId="2" pageNumber="153" precision="1" value="-93.75472">-93.754722</geoCoordinate>
):
<locationDeviation id="3FCFB248EF51D320FC23F8370909F831" box="[955,1126,1949,1975]" location="South" pageId="2" pageNumber="153">
<quantity id="E93A068FEF51D320FC23F8370974F831" box="[955,1051,1949,1975]" metricMagnitude="4" metricUnit="m" metricValue="2.0921472" pageId="2" pageNumber="153" unit="mi" value="13.0">13 miles</quantity>
South
</locationDeviation>
, slope of
<location id="2B1DFDB1EF51D320FB4CF8370808F831" LSID="urn:lsid:plazi:treatment:A66B1A7CEF51D324FF0FFBB70945FCBF:2B1DFDB1EF51D320FB4CF8370808F831" box="[1236,1383,1949,1975]" country="Mexico" latitude="16.089722" longLatPrecision="1" longitude="-93.75472" municipality="Tonala" name="Cerro Vernal" pageId="2" pageNumber="153" stateProvince="Chiapas">Cerro Vernal</location>
,
<quantity id="E93A068FEF51D320FAEAF8370DA0F85D" metricMagnitude="2" metricUnit="m" metricValue="6.1" pageId="2" pageNumber="153" unit="m" value="610.0">
<elevation id="A5EF4C59EF51D320FAEAF8370DA0F85D" metricMagnitude="2" metricUnit="m" metricValue="6.1" pageId="2" pageNumber="153" unit="m" value="610.0">610 masl</elevation>
</quantity>
,
<date id="5A7C8DAAEF51D320FF42F86B0CE3F85D" box="[218,396,1985,2012]" pageId="2" pageNumber="153" value="1974-10-05">
<collectingDate id="4A387442EF51D320FF42F86B0CE3F85D" box="[218,396,1985,2012]" pageId="2" pageNumber="153" value="1974-10-05">5 October 1974</collectingDate>
</date>
,
<collectorName id="8337CEBCEF51D320FE00F8680F26F85D" box="[408,585,1985,2012]" pageId="2" pageNumber="153">D.E. Breedlove</collectorName>
,
<collectorName id="8337CEBCEF51D320FDCCF8680F91F85D" box="[596,766,1985,2012]" pageId="2" pageNumber="153">J.A. Breedlove</collectorName>
, female (
<specimenCode id="7E640311EF51D320FCFDF86B092EF85A" box="[869,1089,1985,2012]" collectionCode="CASENT" pageId="2" pageNumber="153">CASENT 9060511</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="9EAAA137EF51D321FBCBF86B0CBCFF53" ID-GBIF-Occurrence="3064396304" collectingDate="2005-09-01" collectorName="O. Francke &amp; M. Jaimes &amp; Valdez &amp; Montano" country="Mexico" elevation="805" lastPageId="3" lastPageNumber="154" latitude="16.494444" location="Santa Isabel" longLatPrecision="1" longitude="-94.08195" municipality="Federal" pageId="2" pageNumber="153" specimenCount="1" stateProvince="Chiapas">
<collectingMunicipality id="CE193110EF51D320FBCBF86B09C8F85D" box="[1107,1191,1985,2011]" pageId="2" pageNumber="153">Federal</collectingMunicipality>
highway 190, near to
<location id="2B1DFDB1EF50D321FF0FFF3D0C71FF37" LSID="urn:lsid:plazi:treatment:A66B1A7CEF51D324FF0FFBB70945FCBF:2B1DFDB1EF50D321FF0FFF3D0C71FF37" box="[151,286,151,177]" country="Mexico" latitude="16.494444" longLatPrecision="1" longitude="-94.08195" municipality="Federal" name="Santa Isabel" pageId="3" pageNumber="154" stateProvince="Chiapas">Santa Isabel</location>
(
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,
<geoCoordinate id="4BF6CDADEF50D321FE29FF3D0F57FF37" box="[433,568,151,177]" degrees="94.081944" direction="west" orientation="longitude" pageId="3" pageNumber="154" precision="1" value="-94.08195">-94.081944</geoCoordinate>
):
<quantity id="E93A068FEF50D321FDD0FF3C0FDFFF37" box="[584,688,150,177]" metricMagnitude="2" metricUnit="m" metricValue="8.05" pageId="3" pageNumber="154" unit="m" value="805.0">
<elevation id="A5EF4C59EF50D321FDD0FF3C0FDFFF37" box="[584,688,150,177]" metricMagnitude="2" metricUnit="m" metricValue="8.05" pageId="3" pageNumber="154" unit="m" value="805.0">805 masl</elevation>
</quantity>
,
<date id="5A7C8DAAEF50D321FD22FF3D0EE9FF37" box="[698,902,150,177]" pageId="3" pageNumber="154" value="2005-09-01">
<collectingDate id="4A387442EF50D321FD22FF3D0EE9FF37" box="[698,902,150,177]" pageId="3" pageNumber="154" value="2005-09-01">1 September 2005</collectingDate>
</date>
,
<collectorName id="8337CEBCEF50D321FC08FF3D0961FF37" box="[912,1038,151,177]" pageId="3" pageNumber="154">O. Francke</collectorName>
,
<collectorName id="8337CEBCEF50D321FB80FF3D09E2FF37" box="[1048,1165,151,177]" pageId="3" pageNumber="154">M. Jaimes</collectorName>
, A.
<collectorName id="8337CEBCEF50D321FB2EFF3D086AFF37" box="[1206,1285,151,177]" pageId="3" pageNumber="154">Valdez</collectorName>
, H.
<collectorName id="8337CEBCEF50D321FAA8FF3D08F7FF37" box="[1328,1432,151,177]" pageId="3" pageNumber="154">Montaño</collectorName>
, female (CNAN-Am000407)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="66D8F8E1EF50D321FF5FFF740FBEFEE3" pageId="3" pageNumber="154" type="distribution">
<paragraph id="2E7DAB6AEF50D321FF5FFF740FBEFEE3" blockId="3.[151,1436,150,357]" pageId="3" pageNumber="154">
<emphasis id="1CB67778EF50D321FF5FFF740C0EFF7E" bold="true" box="[199,353,222,248]" pageId="3" pageNumber="154">Distribution.</emphasis>
<collectingCountry id="56D5EBFAEF50D321FEF0FF750CA3FF7F" box="[360,460,223,249]" name="Belize" pageId="3" pageNumber="154">BELIZE</collectingCountry>
.
<collectingCountry id="56D5EBFAEF50D321FE4FFF750FE6FF7F" box="[471,649,223,249]" name="Guatemala" pageId="3" pageNumber="154">GUATEMALA</collectingCountry>
.
<collectingCountry id="56D5EBFAEF50D321FD0CFF750E68FF7F" box="[660,775,223,249]" name="Mexico" pageId="3" pageNumber="154">MEXICO</collectingCountry>
: States of
<collectingRegion id="EC066588EF50D321FCE4FF750EB6FF7F" box="[892,985,223,249]" country="Mexico" name="Chiapas" pageId="3" pageNumber="154">Chiapas</collectingRegion>
,
<collectingRegion id="EC066588EF50D321FC7CFF750956FF7F" box="[996,1081,223,249]" country="Mexico" name="Colima" pageId="3" pageNumber="154">Colima</collectingRegion>
,
<collectingRegion id="EC066588EF50D321FBDEFF7509A5FF7F" box="[1094,1226,223,249]" country="Mexico" name="Guanajuato" pageId="3" pageNumber="154">Guanajuato</collectingRegion>
,
<collectingRegion id="EC066588EF50D321FB4DFF750852FF7F" box="[1237,1341,223,249]" country="Mexico" name="Guerrero" pageId="3" pageNumber="154">Guerrero</collectingRegion>
,
<collectingRegion id="EC066588EF50D321FAD1FF7508F7FF7F" box="[1353,1432,223,249]" country="Mexico" name="Jalisco" pageId="3" pageNumber="154">Jalisco</collectingRegion>
,
<collectingRegion id="EC066588EF50D321FF0FFEA90C79FE9B" box="[151,278,259,285]" country="Mexico" name="Michoacan de Ocampo" pageId="3" pageNumber="154">Michoacán</collectingRegion>
,
<collectingRegion id="EC066588EF50D321FEBBFEA90CECFE9B" box="[291,387,259,285]" country="Mexico" name="Morelos" pageId="3" pageNumber="154">Morelos</collectingRegion>
,
<collectingRegion id="EC066588EF50D321FE08FEA90C88FE9B" box="[400,487,259,285]" country="Mexico" name="Nayarit" pageId="3" pageNumber="154">Nayarit</collectingRegion>
,
<collectingRegion id="EC066588EF50D321FE6CFEA90FECFE9B" box="[500,643,259,285]" country="Mexico" name="Nuevo Leon" pageId="3" pageNumber="154">Nuevo León</collectingRegion>
,
<collectingRegion id="EC066588EF50D321FD08FEA90F88FE9B" box="[656,743,259,285]" country="Mexico" name="Oaxaca" pageId="3" pageNumber="154">Oaxaca</collectingRegion>
,
<collectingRegion id="EC066588EF50D321FD6CFEA90E23FE9B" box="[756,844,259,285]" country="Mexico" name="Sinaloa" pageId="3" pageNumber="154">Sinaloa</collectingRegion>
. UNITET STATES OF AMERICA: Texas. Many of these records are not supported with deposited material in collections and have no specific localities (
<bibRefCitation id="4A53D69BEF50D321FAA8FE8D0DBDFEE3" author="Quintero, D." pageId="3" pageNumber="154" pagination="117 - 166" refId="ref21143" refString="Quintero, D. (1981) The Amblypygid genus Phrynus in the Americas (Amblypygi, Phrynidae). Journal of Arachnology, 9, 117 - 166." type="journal article" year="1981">Quintero, 1981</bibRefCitation>
;
<bibRefCitation id="4A53D69BEF50D321FF44FEE10C18FEE3" author="Armas, L. F." box="[220,375,331,357]" pageId="3" pageNumber="154" pagination="345 - 359" refId="ref18597" refString="Armas, L. F. (2006 b) Los Amblypygidos Tendarapodos de Mexico (Arachnida: Amblypygi). Boletin Sociedad Entomologica Aragonesa, 39, 345 - 359." type="journal article" year="2006">Armas 2006b</bibRefCitation>
;
<bibRefCitation id="4A53D69BEF50D321FE19FEE10F39FEE3" author="Armas, L. F. &amp; Guzman, A. &amp; Francke, O." box="[385,598,331,357]" pageId="3" pageNumber="154" pagination="3 - 7" refId="ref18743" refString="Armas, L. F., Guzman, A. &amp; Francke, O. (2014) Una especie nueva de Phrynus (Amblypygi: Phrynidae) de Mexico. Revista Iberica de Aracnologia, 25, 3 - 7." type="journal article" year="2014">
Armas
<emphasis id="1CB67778EF50D321FE4BFEE10F7DFEE3" box="[467,530,331,357]" italics="true" pageId="3" pageNumber="154">et al.,</emphasis>
2014
</bibRefCitation>
). (
<figureCitation id="B6F9B7EFEF50D321FDE9FEE10FABFEE3" box="[625,708,331,357]" captionStart="FIGURE 18" captionStartId="28.[151,250,1479,1503]" captionTargetBox="[151,1436,813,1456]" captionTargetId="figure-330@28.[151,1436,813,1456]" captionTargetPageId="28" captionText="FIGURE 18. Distribution: Phrynus alfonsoi sp. nov. (blue circle), Phrynus aliciae sp. nov. (yellow circle), Phrynus giseae sp. nov. (purple circle), Phrynus jalisco (light blue circle), Phrynus operculatus (red circle), Phrynus tresmarias sp. nov. (orange circle). Circles with black rings: type localities." figureDoi="http://doi.org/10.5281/zenodo.4620750" httpUri="https://zenodo.org/record/4620750/files/figure.png" pageId="3" pageNumber="154">Fig. 18</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="7ABDFBE2EF50D321FF0FF83C0FA2F854" ID-DOI="http://doi.org/10.5281/zenodo.4620708" ID-Zenodo-Dep="4620708" httpUri="https://zenodo.org/record/4620708/files/figure.png" pageId="3" pageNumber="154" startId="3.[151,250,1942,1966]" targetBox="[161,1423,388,1908]" targetPageId="3">
<paragraph id="2E7DAB6AEF50D321FF0FF83C0FA2F854" blockId="3.[151,1437,1942,2002]" pageId="3" pageNumber="154">
<emphasis id="1CB67778EF50D321FF0FF83C0C7DF828" bold="true" box="[151,274,1942,1966]" pageId="3" pageNumber="154">FIGURE 1.</emphasis>
<taxonomicName id="E9C2D0E9EF50D321FE8FF83C0C87F828" authorityName="Pocock" authorityYear="1902" box="[279,488,1942,1966]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="operculatus">
<emphasis id="1CB67778EF50D321FE8FF83C0C87F828" box="[279,488,1942,1966]" italics="true" pageId="3" pageNumber="154">Phrynus operculatus</emphasis>
</taxonomicName>
. Female (CNAN-Am000407): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view.
</paragraph>
</caption>
<subSubSection id="66D8F8E1EF57D327FF5FFF3C0E22FE70" lastPageId="5" lastPageNumber="156" pageId="4" pageNumber="155" type="diagnosis">
<paragraph id="2E7DAB6AEF57D326FF5FFF3C0FC2FE0F" blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="155">
<emphasis id="1CB67778EF57D326FF5FFF3C0C2DFF36" bold="true" box="[199,322,150,176]" pageId="4" pageNumber="155">Diagnosis.</emphasis>
<taxonomicName id="E9C2D0E9EF57D326FEDFFF3C0F41FF36" authorityName="Pocock" authorityYear="1902" box="[327,558,150,176]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="operculatus">
<emphasis id="1CB67778EF57D326FEDFFF3C0F41FF36" box="[327,558,150,176]" italics="true" pageId="4" pageNumber="155">Phrynus operculatus</emphasis>
</taxonomicName>
can be distinguished from other species of the genus by the presence of the following set of characters: only one conspicuous tooth in the ectal row of the base of chelicerae, with an acuminated apex (
<figureCitation id="B6F9B7EFEF57D326FF42FF740C43FF7E" box="[218,300,222,248]" captionStart="FIGURE 1" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[161,1423,388,1908]" captionTargetId="figure-88@3.[151,1436,381,1919]" captionTargetPageId="3" captionText="FIGURE 1. Phrynus operculatus. Female (CNAN-Am000407): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.4620708" httpUri="https://zenodo.org/record/4620708/files/figure.png" pageId="4" pageNumber="155">Fig. 1F</figureCitation>
), does not present an inconspicuous spine in the dorsomedial area of the pedipalp tarsus, Pd2 is smaller than Pd4, Fv3 smaller than Fv5, Td1 is very small but longer than Td4 (
<figureCitation id="B6F9B7EFEF57D326FC5BFEA80966FE9A" box="[963,1033,258,284]" captionStart="FIGURE 2" captionStartId="5.[151,250,1903,1927]" captionTargetBox="[160,1423,531,1867]" captionTargetId="figure-157@5.[151,1436,525,1880]" captionTargetPageId="5" captionText="FIGURE 2. Phrynus operculatus. Female (CNAN-Am000407): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.4620710" httpUri="https://zenodo.org/record/4620710/files/figure.png" pageId="4" pageNumber="155">Fig. 2</figureCitation>
). Female genitalia have tubular and elongated gonopod bases, with claw-like sclerites of the gonopod, curved at the apical region (
<figureCitation id="B6F9B7EFEF57D326FB3FFE8C0860FEC6" box="[1191,1295,294,320]" captionStart="FIGURE 17" captionStartId="27.[151,250,1596,1620]" captionTargetBox="[158,1428,188,1562]" captionTargetId="figure-15@27.[151,1437,181,1572]" captionTargetPageId="27" captionText="FIGURE 17. Female genitalia: A. Phrynus operculatus (CNAN-Am000407). B. Phrynus alfonsoi sp. nov. (CNAN-T01383). C. Phrynus aliciae sp. nov. (CNAN-T01447). D. Phrynus giseae sp. nov. (CNAN-T01450). E. Phrynus tresmarias sp. nov. (CASENT 9060565). F. Phrynus jalisco (CASENT 9060519). Sclerotization of the gonopod bases (black arrows); colored lines illustrate the position of the sclerite of the gonopod with respect to the gonopod base, the length comparison between the gonopod base (green line) and sclerite of the gonopod (red line), and the comparison between width of the base of the sclerite (purple line) and the width of the medial region of the gonopod base (blue line)." figureDoi="http://doi.org/10.5281/zenodo.4620748" httpUri="https://zenodo.org/record/4620748/files/figure.png" pageId="4" pageNumber="155">Fig. 17A</figureCitation>
). In the male genitalia, the ornamentations over LoL 1 and 2 are tubular and long, and LoD is very thick and pronounced (
<figureCitation id="B6F9B7EFEF57D326FAD2FEE008E1FEE2" box="[1354,1422,330,356]" captionStart="FIGURE 3" captionStartId="6.[151,250,1775,1799]" captionTargetBox="[156,1432,852,1747]" captionTargetId="figure-368@6.[151,1436,848,1751]" captionTargetPageId="6" captionText="FIGURE 3. Male genital: Phrynus operculatus (CASENT 9060512): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1 = lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. Colored lines illustrate the comparison between the LoD medial width (green line) and the width of the base of LoL 1 (red line)." figureDoi="http://doi.org/10.5281/zenodo.4620718" httpUri="https://zenodo.org/record/4620718/files/figure.png" pageId="4" pageNumber="155">Fig. 3</figureCitation>
). Almost the wide of the base of LoL 1 (
<tableCitation id="63409ED1EF57D326FDD0FEC40FF0FE0F" box="[584,671,366,393]" captionStart="TABLE 6" captionStartId="26.[151,239,368,392]" captionText="TABLE 6. Size proportions in the female genital: Gonopod base (Gb), Sclerite (Scl), and the male genital: lobus dorsalis (LoD), lobus lateralis primus (LoL1). Comparing their total length (l), medial width (mw), and base width (bw)." pageId="4" pageNumber="155">Table 6</tableCitation>
).
</paragraph>
<paragraph id="2E7DAB6AEF57D326FF5FFE38086DFD2E" blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="155">
Species comparisons:
<taxonomicName id="E9C2D0E9EF57D326FE27FE380FCAFE2A" authorityName="Pocock" authorityYear="1902" box="[447,677,402,428]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="operculatus">
<emphasis id="1CB67778EF57D326FE27FE380FCAFE2A" box="[447,677,402,428]" italics="true" pageId="4" pageNumber="155">Phrynus operculatus</emphasis>
</taxonomicName>
is morphologically similar to
<taxonomicName id="E9C2D0E9EF57D326FC6CFE390935FE2A" authorityName="De Armas, Guzman &amp; Francke" authorityYear="2014" box="[1012,1114,402,428]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="jalisco">
<emphasis id="1CB67778EF57D326FC6CFE390935FE2A" box="[1012,1114,402,428]" italics="true" pageId="4" pageNumber="155">P. jalisco</emphasis>
</taxonomicName>
, however, the female genitalia are different, in
<taxonomicName id="E9C2D0E9EF57D326FEF5FE1D0CBCFE56" authorityName="De Armas, Guzman &amp; Francke" authorityYear="2014" box="[365,467,438,464]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="jalisco">
<emphasis id="1CB67778EF57D326FEF5FE1D0CBCFE56" box="[365,467,438,464]" italics="true" pageId="4" pageNumber="155">P. jalisco</emphasis>
</taxonomicName>
the gonopod bases are more elongated, the sclerites are thicker and have a pronounced inward curvature (
<figureCitation id="B6F9B7EFEF57D326FEFEFE700CA5FE72" box="[358,458,474,500]" captionStart="FIGURE 17" captionStartId="27.[151,250,1596,1620]" captionTargetBox="[158,1428,188,1562]" captionTargetId="figure-15@27.[151,1437,181,1572]" captionTargetPageId="27" captionText="FIGURE 17. Female genitalia: A. Phrynus operculatus (CNAN-Am000407). B. Phrynus alfonsoi sp. nov. (CNAN-T01383). C. Phrynus aliciae sp. nov. (CNAN-T01447). D. Phrynus giseae sp. nov. (CNAN-T01450). E. Phrynus tresmarias sp. nov. (CASENT 9060565). F. Phrynus jalisco (CASENT 9060519). Sclerotization of the gonopod bases (black arrows); colored lines illustrate the position of the sclerite of the gonopod with respect to the gonopod base, the length comparison between the gonopod base (green line) and sclerite of the gonopod (red line), and the comparison between width of the base of the sclerite (purple line) and the width of the medial region of the gonopod base (blue line)." figureDoi="http://doi.org/10.5281/zenodo.4620748" httpUri="https://zenodo.org/record/4620748/files/figure.png" pageId="4" pageNumber="155">Fig. 17F</figureCitation>
).
<taxonomicName id="E9C2D0E9EF57D326FE46FE700FA9FE72" authorityName="Pocock" authorityYear="1902" box="[478,710,474,500]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="operculatus">
<emphasis id="1CB67778EF57D326FE46FE700FA9FE72" box="[478,710,474,500]" italics="true" pageId="4" pageNumber="155">Phrynus operculatus</emphasis>
</taxonomicName>
is also similar to
<taxonomicName id="E9C2D0E9EF57D326FC17FE710E98FE72" authority="Joya, 2021" authorityName="Joya" authorityYear="2021" box="[911,1015,474,500]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="aliciae" status="sp. nov.">
<emphasis id="1CB67778EF57D326FC17FE710E98FE72" box="[911,1015,474,500]" italics="true" pageId="4" pageNumber="155">P. aliciae</emphasis>
</taxonomicName>
<emphasis id="1CB67778EF57D326FC66FE700937FE72" bold="true" box="[1022,1112,474,500]" pageId="4" pageNumber="155">
<taxonomicNameLabel id="0785CA03EF57D326FC66FE700937FE72" box="[1022,1112,474,500]" pageId="4" pageNumber="155" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, however, in this species the base of the sclerites of the female genitalia are straighter and surrounds the lateral margin of the base (
<figureCitation id="B6F9B7EFEF57D326FAB6FE5408FBFD9E" box="[1326,1428,510,536]" captionStart="FIGURE 17" captionStartId="27.[151,250,1596,1620]" captionTargetBox="[158,1428,188,1562]" captionTargetId="figure-15@27.[151,1437,181,1572]" captionTargetPageId="27" captionText="FIGURE 17. Female genitalia: A. Phrynus operculatus (CNAN-Am000407). B. Phrynus alfonsoi sp. nov. (CNAN-T01383). C. Phrynus aliciae sp. nov. (CNAN-T01447). D. Phrynus giseae sp. nov. (CNAN-T01450). E. Phrynus tresmarias sp. nov. (CASENT 9060565). F. Phrynus jalisco (CASENT 9060519). Sclerotization of the gonopod bases (black arrows); colored lines illustrate the position of the sclerite of the gonopod with respect to the gonopod base, the length comparison between the gonopod base (green line) and sclerite of the gonopod (red line), and the comparison between width of the base of the sclerite (purple line) and the width of the medial region of the gonopod base (blue line)." figureDoi="http://doi.org/10.5281/zenodo.4620748" httpUri="https://zenodo.org/record/4620748/files/figure.png" pageId="4" pageNumber="155">Fig. 17C</figureCitation>
) whereas in
<taxonomicName id="E9C2D0E9EF57D326FE80FD890CD7FDBA" authorityName="Pocock" authorityYear="1902" box="[280,440,546,572]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="operculatus">
<emphasis id="1CB67778EF57D326FE80FD890CD7FDBA" box="[280,440,546,572]" italics="true" pageId="4" pageNumber="155">P. operculatus</emphasis>
</taxonomicName>
the base is smaller. Is also similar to
<taxonomicName id="E9C2D0E9EF57D326FCF9FD890EB8FDBA" authority="Joya, 2021" authorityName="Joya" authorityYear="2021" box="[865,983,546,572]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="alfonsoi" status="sp. nov.">
<emphasis id="1CB67778EF57D326FCF9FD890EB8FDBA" box="[865,983,546,572]" italics="true" pageId="4" pageNumber="155">P. alfonsoi</emphasis>
</taxonomicName>
<emphasis id="1CB67778EF57D326FC47FD880955FDBA" bold="true" box="[991,1082,546,572]" pageId="4" pageNumber="155">
<taxonomicNameLabel id="0785CA03EF57D326FC47FD880955FDBA" box="[991,1082,546,572]" pageId="4" pageNumber="155" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but in this species, the sclerite of female gonopod is bigger with a more pronounced curvature in the apical region (17B). In the male genitalia, the projections on LoL 2 are completely tubular (
<figureCitation id="B6F9B7EFEF57D326FD15FDC00F8DFD02" box="[653,738,618,644]" captionStart="FIGURE 3" captionStartId="6.[151,250,1775,1799]" captionTargetBox="[156,1432,852,1747]" captionTargetId="figure-368@6.[151,1436,848,1751]" captionTargetPageId="6" captionText="FIGURE 3. Male genital: Phrynus operculatus (CASENT 9060512): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1 = lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. Colored lines illustrate the comparison between the LoD medial width (green line) and the width of the base of LoL 1 (red line)." figureDoi="http://doi.org/10.5281/zenodo.4620718" httpUri="https://zenodo.org/record/4620718/files/figure.png" pageId="4" pageNumber="155">Fig. 3B</figureCitation>
) whereas in
<taxonomicName id="E9C2D0E9EF57D326FCF6FDC10E8DFD02" authority="Joya, 2021" authorityName="Joya" authorityYear="2021" box="[878,994,618,644]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="alfonsoi" status="sp. nov.">
<emphasis id="1CB67778EF57D326FCF6FDC10E8DFD02" box="[878,994,618,644]" italics="true" pageId="4" pageNumber="155">P. alfonsoi</emphasis>
</taxonomicName>
<emphasis id="1CB67778EF57D326FC71FDC0092EFD02" bold="true" box="[1001,1089,618,644]" pageId="4" pageNumber="155">
<taxonomicNameLabel id="0785CA03EF57D326FC71FDC0092EFD02" box="[1001,1089,618,644]" pageId="4" pageNumber="155" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the projections bases are wider (
<figureCitation id="B6F9B7EFEF57D326FF38FD240D97FD2F" box="[160,248,654,681]" captionStart="FIGURE 6" captionStartId="11.[151,250,1779,1803]" captionTargetBox="[155,1432,855,1752]" captionTargetId="figure-323@11.[151,1437,849,1755]" captionTargetPageId="11" captionText="FIGURE 6. Male genitalia: Phrynus alfonsoi sp. nov. (CNAN-T01445): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1 = lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. Colored lines illustrate the comparison between the LoD medial width (green line) and the width of the base of LoL 1 (red line)." figureDoi="http://doi.org/10.5281/zenodo.4620724" httpUri="https://zenodo.org/record/4620724/files/figure.png" pageId="4" pageNumber="155">Fig. 6B</figureCitation>
). The LoD in
<taxonomicName id="E9C2D0E9EF57D326FE0DFD250F5CFD2E" authorityName="Pocock" authorityYear="1902" box="[405,563,654,680]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="operculatus">
<emphasis id="1CB67778EF57D326FE0DFD250F5CFD2E" box="[405,563,654,680]" italics="true" pageId="4" pageNumber="155">P. operculatus</emphasis>
</taxonomicName>
is thicker than in the other species compared (
<figureCitation id="B6F9B7EFEF57D326FBA4FD2409FCFD2E" box="[1084,1171,654,680]" captionStart="FIGURE 3" captionStartId="6.[151,250,1775,1799]" captionTargetBox="[156,1432,852,1747]" captionTargetId="figure-368@6.[151,1436,848,1751]" captionTargetPageId="6" captionText="FIGURE 3. Male genital: Phrynus operculatus (CASENT 9060512): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1 = lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. Colored lines illustrate the comparison between the LoD medial width (green line) and the width of the base of LoL 1 (red line)." figureDoi="http://doi.org/10.5281/zenodo.4620718" httpUri="https://zenodo.org/record/4620718/files/figure.png" pageId="4" pageNumber="155">Fig. 3C</figureCitation>
,
<tableCitation id="63409ED1EF57D326FB07FD24099CFD2F" box="[1183,1267,654,681]" captionStart="TABLE 6" captionStartId="26.[151,239,368,392]" captionText="TABLE 6. Size proportions in the female genital: Gonopod base (Gb), Sclerite (Scl), and the male genital: lobus dorsalis (LoD), lobus lateralis primus (LoL1). Comparing their total length (l), medial width (mw), and base width (bw)." pageId="4" pageNumber="155">Table 6</tableCitation>
).
</paragraph>
<paragraph id="2E7DAB6AEF57D326FF5FFD1809A2FCBE" blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="155">
The spine Pd1 is conspicuous (a quarter of the size of Pd2) (
<figureCitation id="B6F9B7EFEF57D326FCF6FD180EAAFD4A" box="[878,965,690,716]" captionStart="FIGURE 2" captionStartId="5.[151,250,1903,1927]" captionTargetBox="[160,1423,531,1867]" captionTargetId="figure-157@5.[151,1436,525,1880]" captionTargetPageId="5" captionText="FIGURE 2. Phrynus operculatus. Female (CNAN-Am000407): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.4620710" httpUri="https://zenodo.org/record/4620710/files/figure.png" pageId="4" pageNumber="155">Fig. 2D</figureCitation>
) similar to
<taxonomicName id="E9C2D0E9EF57D326FBDFFD1909D2FD4A" authority="Joya, 2021" authorityName="Joya" authorityYear="2021" box="[1095,1213,690,716]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="alfonsoi" status="sp. nov.">
<emphasis id="1CB67778EF57D326FBDFFD1909D2FD4A" box="[1095,1213,690,716]" italics="true" pageId="4" pageNumber="155">P. alfonsoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0785CA03EF57D326FB5DFD180870FD4B" box="[1221,1311,690,717]" pageId="4" pageNumber="155" rank="species">
<emphasis id="1CB67778EF57D326FB5DFD180875FD4A" bold="true" box="[1221,1306,690,716]" pageId="4" pageNumber="155">sp. nov</emphasis>
.
</taxonomicNameLabel>
,
<emphasis id="1CB67778EF57D326FAB7FD1908F3FD4A" box="[1327,1436,690,716]" italics="true" pageId="4" pageNumber="155">
<taxonomicName id="E9C2D0E9EF57D326FAB7FD1908F7FD4A" authorityName="De Armas, Guzman &amp; Francke" authorityYear="2014" box="[1327,1432,690,716]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="jalisco">P. jalisco</taxonomicName>
,
</emphasis>
and
<taxonomicName id="E9C2D0E9EF57D326FF5DFD7D0C43FD76" authority="Joya, 2021" authorityName="Joya" authorityYear="2021" box="[197,300,726,752]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="aliciae" status="sp. nov.">
<emphasis id="1CB67778EF57D326FF5DFD7D0C43FD76" box="[197,300,726,752]" italics="true" pageId="4" pageNumber="155">P. aliciae</emphasis>
</taxonomicName>
<emphasis id="1CB67778EF57D326FEAAFD7C0CFBFD76" bold="true" box="[306,404,726,752]" pageId="4" pageNumber="155">
<taxonomicNameLabel id="0785CA03EF57D326FEAAFD7C0CE2FD76" box="[306,397,726,752]" pageId="4" pageNumber="155" rank="species">sp. nov.</taxonomicNameLabel>
;
</emphasis>
Tv1 is shorter than Tv3 (
<figureCitation id="B6F9B7EFEF57D326FD30FD7C0F94FD76" box="[680,763,726,752]" captionStart="FIGURE 2" captionStartId="5.[151,250,1903,1927]" captionTargetBox="[160,1423,531,1867]" captionTargetId="figure-157@5.[151,1436,525,1880]" captionTargetPageId="5" captionText="FIGURE 2. Phrynus operculatus. Female (CNAN-Am000407): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.4620710" httpUri="https://zenodo.org/record/4620710/files/figure.png" pageId="4" pageNumber="155">Fig. 2E</figureCitation>
) similar to
<taxonomicName id="E9C2D0E9EF57D326FCE1FD7D0965FD76" authority="Joya, 2021" authorityName="Joya" authorityYear="2021" box="[889,1034,727,752]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="tresmarias" status="sp. nov.">
<emphasis id="1CB67778EF57D326FCE1FD7D0965FD76" box="[889,1034,727,752]" italics="true" pageId="4" pageNumber="155">P. tresmarias</emphasis>
</taxonomicName>
<emphasis id="1CB67778EF57D326FB88FD7C0907FD76" bold="true" box="[1040,1128,726,752]" pageId="4" pageNumber="155">
<taxonomicNameLabel id="0785CA03EF57D326FB88FD7C0907FD76" box="[1040,1128,726,752]" pageId="4" pageNumber="155" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="E9C2D0E9EF57D326FB04FD7D086DFD76" authority="Joya, 2021" authorityName="Joya" authorityYear="2021" box="[1180,1282,726,752]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="aliciae" status="sp. nov.">
<emphasis id="1CB67778EF57D326FB04FD7D086DFD76" box="[1180,1282,726,752]" italics="true" pageId="4" pageNumber="155">P. aliciae</emphasis>
</taxonomicName>
<emphasis id="1CB67778EF57D326FA90FD7C080EFD76" bold="true" box="[1288,1377,726,752]" pageId="4" pageNumber="155">
<taxonomicNameLabel id="0785CA03EF57D326FA90FD7C080EFD76" box="[1288,1377,726,752]" pageId="4" pageNumber="155" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="B6F9B7EFEF57D326FAF7FD7C0DD6FC92" captionStart="FIGURE 8" captionStartId="15.[151,250,1938,1962]" captionTargetBox="[163,1426,646,1903]" captionTargetId="figure-202@15.[151,1437,633,1914]" captionTargetPageId="15" captionText="FIGURE 8. Phrynus aliciae sp. nov. Female (CNAN-T01447): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.4620730" httpUri="https://zenodo.org/record/4620730/files/figure.png" pageId="4" pageNumber="155">Fig. 8E</figureCitation>
,
<figureCitation id="B6F9B7EFEF57D326FF5BFD500D9EFC92" box="[195,241,762,788]" captionStart="FIGURE 14" captionStartId="24.[151,250,1504,1528]" captionTargetBox="[167,1423,188,1480]" captionTargetId="figure-27@24.[151,1436,181,1481]" captionTargetPageId="24" captionText="FIGURE 14. Phrynus tresmarias sp. nov. Female (CASENT 9060565):A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.4620742" httpUri="https://zenodo.org/record/4620742/files/figure.png" pageId="4" pageNumber="155">14E</figureCitation>
) whereas, in
<taxonomicName id="E9C2D0E9EF57D326FE1BFD510C98FC92" authority="Joya, 2021" authorityName="Joya" authorityYear="2021" box="[387,503,762,788]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="alfonsoi" status="sp. nov.">
<emphasis id="1CB67778EF57D326FE1BFD510C98FC92" box="[387,503,762,788]" italics="true" pageId="4" pageNumber="155">P. alfonsoi</emphasis>
</taxonomicName>
<emphasis id="1CB67778EF57D326FE65FD500F3AFC92" bold="true" box="[509,597,762,788]" pageId="4" pageNumber="155">
<taxonomicNameLabel id="0785CA03EF57D326FE65FD500F3AFC92" box="[509,597,762,788]" pageId="4" pageNumber="155" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Tv1 is longer (
<figureCitation id="B6F9B7EFEF57D326FD63FD500E21FC93" box="[763,846,762,789]" captionStart="FIGURE 5" captionStartId="10.[151,250,1650,1674]" captionTargetBox="[162,1425,246,1606]" captionTargetId="figure-54@10.[151,1437,237,1627]" captionTargetPageId="10" captionText="FIGURE 5. Phrynus alfonsoi sp. nov. Female (CNAN-T01383): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.4620722" httpUri="https://zenodo.org/record/4620722/files/figure.png" pageId="4" pageNumber="155">Fig. 5E</figureCitation>
) and, in
<taxonomicName id="E9C2D0E9EF57D326FC34FD510960FC92" authority="Joya, 2021" authorityName="Joya" authorityYear="2021" box="[940,1039,763,788]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="giseae" status="sp. nov.">
<emphasis id="1CB67778EF57D326FC34FD510960FC92" box="[940,1039,763,788]" italics="true" pageId="4" pageNumber="155">P. giseae</emphasis>
</taxonomicName>
<emphasis id="1CB67778EF57D326FB8CFD500903FC92" bold="true" box="[1044,1132,762,788]" pageId="4" pageNumber="155">
<taxonomicNameLabel id="0785CA03EF57D326FB8CFD500903FC92" box="[1044,1132,762,788]" pageId="4" pageNumber="155" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<emphasis id="1CB67778EF57D326FB07FD510864FC92" box="[1183,1291,762,788]" italics="true" pageId="4" pageNumber="155">
<taxonomicName id="E9C2D0E9EF57D326FB07FD510868FC92" authorityName="De Armas, Guzman &amp; Francke" authorityYear="2014" box="[1183,1287,762,788]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="jalisco">P. jalisco</taxonomicName>
,
</emphasis>
are subequal (
<figureCitation id="B6F9B7EFEF57D326FF38FCB40C6CFCBE" box="[160,259,798,824]" captionStart="FIGURE 11" captionStartId="19.[151,250,1520,1544]" captionTargetBox="[159,1425,191,1484]" captionTargetId="figure-15@19.[151,1436,181,1496]" captionTargetPageId="19" captionText="FIGURE 11. Phrynus giseae sp. nov. Female (CNAN-T01450): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.4620736" httpUri="https://zenodo.org/record/4620736/files/figure.png" pageId="4" pageNumber="155">Fig. 11E</figureCitation>
). Td1 is a quarter of size Td2, so it differs from other species here compared (
<figureCitation id="B6F9B7EFEF57D326FBF2FCB409AFFCBE" box="[1130,1216,798,824]" captionStart="FIGURE 2" captionStartId="5.[151,250,1903,1927]" captionTargetBox="[160,1423,531,1867]" captionTargetId="figure-157@5.[151,1436,525,1880]" captionTargetPageId="5" captionText="FIGURE 2. Phrynus operculatus. Female (CNAN-Am000407): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.4620710" httpUri="https://zenodo.org/record/4620710/files/figure.png" pageId="4" pageNumber="155">Fig. 2F</figureCitation>
).
</paragraph>
<paragraph id="2E7DAB6AEF57D326FF5FFCE80FABFC22" blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="155">
<emphasis id="1CB67778EF57D326FF5FFCE80F12FCDA" bold="true" box="[199,637,834,860]" pageId="4" pageNumber="155">Complementary description. Female</emphasis>
(
<emphasis id="1CB67778EF57D326FD16FCE80E15FCDA" bold="true" box="[654,890,834,860]" pageId="4" pageNumber="155">CNAN-Am000407):</emphasis>
Total length
<quantity id="E93A068FEF57D326FB88FCE8090CFCDB" box="[1040,1123,834,861]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="155" unit="mm" value="16.0">16 mm</quantity>
. Carapace and opisthosoma color brown, pedipalps and legs are brown with red tonalities. Dimensions of prosoma, opisthosoma, pedipalps segments, and all leg femora are provided in
<tableCitation id="63409ED1EF57D326FDF2FC200FD1FC22" box="[618,702,906,932]" captionStart="TABLE 1" captionStartId="7.[151,239,151,175]" captionText="TABLE 1. Measurements (mm) of body, pedipalps segments, and femur of legs in Phrynus operculatus. When the segment is missing, the cell is empty." pageId="4" pageNumber="155">Table 1</tableCitation>
.
</paragraph>
<paragraph id="2E7DAB6AEF57D326FF5FFC040F3EFB96" blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="155">
<emphasis id="1CB67778EF57D326FF5FFC040C53FC4E" bold="true" box="[199,316,942,968]" italics="true" pageId="4" pageNumber="155">Carapace.</emphasis>
The frontal margin is almost straight. Carapace presents a group of granules widely spaced; anterior, lateral, and posterior margins with a white border, posterior margin concave. Lateral and anteromedial eyes clearly visible, ocular tubercle black (
<figureCitation id="B6F9B7EFEF57D326FE72FC5C0F2CFB96" box="[490,579,1014,1040]" captionStart="FIGURE 1" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[161,1423,388,1908]" captionTargetId="figure-88@3.[151,1436,381,1919]" captionTargetPageId="3" captionText="FIGURE 1. Phrynus operculatus. Female (CNAN-Am000407): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.4620708" httpUri="https://zenodo.org/record/4620708/files/figure.png" pageId="4" pageNumber="155">Fig. 1A</figureCitation>
).
</paragraph>
<paragraph id="2E7DAB6AEF57D326FF5FFBB00924FB42" blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="155">
<emphasis id="1CB67778EF57D326FF5FFBB00C5EFBB2" bold="true" box="[199,305,1050,1076]" italics="true" pageId="4" pageNumber="155">Sternum.</emphasis>
Tri-segmented, all segments poorly sclerotized, the area around segments is slightly less sclerotized. Tritosternum projected anteriorly, elongated, conical, with eight setae on the basal region, four on the medial region, and two more in the apex. Second segment (tetrasternum) oval, with two setae on the medial region. Third segment (pentasternum) oval, slightly smaller than the second segment, with two setae on the medial region. Metasternum longitudinally divided, with two setae on the posterior margin of each half (
<figureCitation id="B6F9B7EFEF57D326FC7EFB000952FB42" box="[998,1085,1194,1220]" captionStart="FIGURE 1" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[161,1423,388,1908]" captionTargetId="figure-88@3.[151,1436,381,1919]" captionTargetPageId="3" captionText="FIGURE 1. Phrynus operculatus. Female (CNAN-Am000407): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.4620708" httpUri="https://zenodo.org/record/4620708/files/figure.png" pageId="4" pageNumber="155">Fig. 1B</figureCitation>
).
</paragraph>
<paragraph id="2E7DAB6AEF57D326FF5FFB640FF3FA8A" blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="155">
<emphasis id="1CB67778EF57D326FF5FFB640C53FB6F" bold="true" box="[199,316,1230,1257]" italics="true" pageId="4" pageNumber="155">Abdomen.</emphasis>
Oblong, color dark brown. Presents patches of light coloration on both sides of each tergite. The carapace is wider than the abdomen (
<figureCitation id="B6F9B7EFEF57D326FDAFFB580FE1FA8A" box="[567,654,1266,1292]" captionStart="FIGURE 1" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[161,1423,388,1908]" captionTargetId="figure-88@3.[151,1436,381,1919]" captionTargetPageId="3" captionText="FIGURE 1. Phrynus operculatus. Female (CNAN-Am000407): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.4620708" httpUri="https://zenodo.org/record/4620708/files/figure.png" pageId="4" pageNumber="155">Fig. 1C</figureCitation>
).
</paragraph>
<paragraph id="2E7DAB6AEF57D326FF5FFABC0C51FA62" blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="155">
<emphasis id="1CB67778EF57D326FF5FFABC0C51FAB7" bold="true" box="[199,318,1302,1329]" italics="true" pageId="4" pageNumber="155">Chelicera.</emphasis>
The mesal row of the basal segment of chelicerae with three teeth, the first is bilobed, placed in the proximal portion, lobe 1b bigger than 1a, followed by one tooth shorter in medial portion, and the third, bigger than the others, placed in the distal region (
<figureCitation id="B6F9B7EFEF57D326FD1EFAF40F8EFAFE" box="[646,737,1374,1400]" captionStart="FIGURE 1" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[161,1423,388,1908]" captionTargetId="figure-88@3.[151,1436,381,1919]" captionTargetPageId="3" captionText="FIGURE 1. Phrynus operculatus. Female (CNAN-Am000407): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.4620708" httpUri="https://zenodo.org/record/4620708/files/figure.png" pageId="4" pageNumber="155">Fig. 1D</figureCitation>
). The ectal row with one conspicuous tooth (tooth 2) with an acuminated apex, placed in the medial region. There is a small keel on the most proximal region (tooth 1) (
<figureCitation id="B6F9B7EFEF57D326FAA2FA2808E1FA1A" box="[1338,1422,1410,1436]" captionStart="FIGURE 1" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[161,1423,388,1908]" captionTargetId="figure-88@3.[151,1436,381,1919]" captionTargetPageId="3" captionText="FIGURE 1. Phrynus operculatus. Female (CNAN-Am000407): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.4620708" httpUri="https://zenodo.org/record/4620708/files/figure.png" pageId="4" pageNumber="155">Fig. 1F</figureCitation>
). The mobile segment of chelicerae with four teeth, the first one is the biggest, followed by the third, and the fourth is the smallest.
</paragraph>
<paragraph id="2E7DAB6AEF57D327FF5FFA440CA6FF53" blockId="4.[151,1437,150,2013]" lastBlockId="5.[151,1437,151,502]" lastPageId="5" lastPageNumber="156" pageId="4" pageNumber="155">
<emphasis id="1CB67778EF57D326FF5FFA440CD9F98E" box="[199,438,1518,1545]" italics="true" pageId="4" pageNumber="155">
<emphasis id="1CB67778EF57D326FF5FFA440C5CF98F" bold="true" box="[199,307,1518,1545]" italics="true" pageId="4" pageNumber="155">Pedipalp.</emphasis>
Trochanter
</emphasis>
: Prolateral face with four spines; spines Tr1 and Tr3 placed in medial region, Tr2 placed near ventral margin. Tr3 and Tr4 subequal in length. Spine lengths: Tr2&gt;Tr1&gt;Tr4ŻTr3. Dorsal oblique series of five setiferous tubercles. Dorsomedial area without spines but with one big tubercle (
<figureCitation id="B6F9B7EFEF57D326FBC9F99C09C8F9D6" box="[1105,1191,1590,1616]" captionStart="FIGURE 1" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[161,1423,388,1908]" captionTargetId="figure-88@3.[151,1436,381,1919]" captionTargetPageId="3" captionText="FIGURE 1. Phrynus operculatus. Female (CNAN-Am000407): A. Carapace. B. Sternum. C. Dorsal opisthosoma. D. Chelicera, mesal view. E. Trochanter (Tr). F. Chelicera, ectal view." figureDoi="http://doi.org/10.5281/zenodo.4620708" httpUri="https://zenodo.org/record/4620708/files/figure.png" pageId="4" pageNumber="155">Fig. 1E</figureCitation>
).
<emphasis id="1CB67778EF57D326FB24F99D0868F9D6" box="[1212,1287,1591,1616]" italics="true" pageId="4" pageNumber="155">Femur</emphasis>
: Ventral face with five major spines, Fv4, Fv3a, Fv5a, and Fv7 are small. Spine lengths: Fv1&gt;Fv2&gt;Fv5&gt;Fv3&gt;Fv6&gt;Fv4&gt;Fv7&gt;F v5a&gt;Fv3a (
<figureCitation id="B6F9B7EFEF57D326FE8FF9D40C1DF91E" box="[279,370,1662,1688]" captionStart="FIGURE 2" captionStartId="5.[151,250,1903,1927]" captionTargetBox="[160,1423,531,1867]" captionTargetId="figure-157@5.[151,1436,525,1880]" captionTargetPageId="5" captionText="FIGURE 2. Phrynus operculatus. Female (CNAN-Am000407): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.4620710" httpUri="https://zenodo.org/record/4620710/files/figure.png" pageId="4" pageNumber="155">Fig. 2A</figureCitation>
). Dorsal face with five major spines; Fd4 small, with the spines Fd5a, Fd7, and Fd8 present as tubercles; Fd1 and Fd2 share the same base, Fd3 is separated from Fd2. Spine lengths: Fd2&gt;Fd3&gt;Fd5&gt;Fd1&gt;Fd6&gt;Fd 4=Fd5a&gt;Fd7&gt;Fd8 (
<figureCitation id="B6F9B7EFEF57D326FEEBF96C0CA5F966" box="[371,458,1734,1760]" captionStart="FIGURE 2" captionStartId="5.[151,250,1903,1927]" captionTargetBox="[160,1423,531,1867]" captionTargetId="figure-157@5.[151,1436,525,1880]" captionTargetPageId="5" captionText="FIGURE 2. Phrynus operculatus. Female (CNAN-Am000407): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.4620710" httpUri="https://zenodo.org/record/4620710/files/figure.png" pageId="4" pageNumber="155">Fig. 2B</figureCitation>
).
<taxonomicName id="E9C2D0E9EF57D326FE46F96C0FE1F966" authority=": Ventral" authorityName="Ventral" box="[478,654,1734,1761]" class="Gastropoda" family="Patellidae" genus="Patella" kingdom="Animalia" order="Pezizales" pageId="4" pageNumber="155" phylum="Mollusca" rank="genus">
<emphasis id="1CB67778EF57D326FE46F96C0F40F966" box="[478,559,1734,1760]" italics="true" pageId="4" pageNumber="155">Patella</emphasis>
: Ventral
</taxonomicName>
face with five major spines; with conspicuous spines Pv1-a and Pv1a, and one small spine between Pv2Pv3, Pv4Pv5, and Pv5Pv6. Spine lengths: Pv2&gt;Pv5&gt;Pv1&gt;Pv4&gt;Pv6&gt;Pv7&gt;Pv1- a=Pv1a&gt;Pv3 (
<figureCitation id="B6F9B7EFEF57D326FEAEF8A40CE0F8AE" box="[310,399,1806,1832]" captionStart="FIGURE 2" captionStartId="5.[151,250,1903,1927]" captionTargetBox="[160,1423,531,1867]" captionTargetId="figure-157@5.[151,1436,525,1880]" captionTargetPageId="5" captionText="FIGURE 2. Phrynus operculatus. Female (CNAN-Am000407): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.4620710" httpUri="https://zenodo.org/record/4620710/files/figure.png" pageId="4" pageNumber="155">Fig. 2C</figureCitation>
). Dorsal face with seven major spines, proximal to Pd1 there are two tubercles, Pd8 is small. Spine lengths: Pd5&gt;Pd3&gt;Pd4&gt;Pd2&gt;Pd6&gt;Pd7&gt;Pd1&gt;Pd8 (
<figureCitation id="B6F9B7EFEF57D326FC82F8980E1AF8CA" box="[794,885,1842,1868]" captionStart="FIGURE 2" captionStartId="5.[151,250,1903,1927]" captionTargetBox="[160,1423,531,1867]" captionTargetId="figure-157@5.[151,1436,525,1880]" captionTargetPageId="5" captionText="FIGURE 2. Phrynus operculatus. Female (CNAN-Am000407): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.4620710" httpUri="https://zenodo.org/record/4620710/files/figure.png" pageId="4" pageNumber="155">Fig. 2D</figureCitation>
).
<emphasis id="1CB67778EF57D326FC11F8980EACF8CA" box="[905,963,1842,1868]" italics="true" pageId="4" pageNumber="155">Tibia</emphasis>
: Ventral face with three major spines; has one tubercle and one small spine between Tv2Tv3. Spine lengths: Tv2&gt;Tv3&gt;Tv1 (
<figureCitation id="B6F9B7EFEF57D326FBA4F8FC09FCF8F6" box="[1084,1171,1878,1904]" captionStart="FIGURE 2" captionStartId="5.[151,250,1903,1927]" captionTargetBox="[160,1423,531,1867]" captionTargetId="figure-157@5.[151,1436,525,1880]" captionTargetPageId="5" captionText="FIGURE 2. Phrynus operculatus. Female (CNAN-Am000407): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.4620710" httpUri="https://zenodo.org/record/4620710/files/figure.png" pageId="4" pageNumber="155">Fig. 2E</figureCitation>
). Dorsal face with three major spines, one tubercle between Td2Td3, two conspicuous spines over Td3, distal to Td3 there are two small spines, the most distal spine is a little bigger and is here named Td4; Td1 is almost a half of Td3 length. Spine lengths: Td2&gt;Td3&gt;Td1&gt;Td4 (
<figureCitation id="B6F9B7EFEF57D326FE68F8680F26F85A" box="[496,585,1986,2012]" captionStart="FIGURE 2" captionStartId="5.[151,250,1903,1927]" captionTargetBox="[160,1423,531,1867]" captionTargetId="figure-157@5.[151,1436,525,1880]" captionTargetPageId="5" captionText="FIGURE 2. Phrynus operculatus. Female (CNAN-Am000407): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral patella (Pv). D. Dorsal patella (Pd). E. Ventral tibia (Tv). F. Dorsal tibia (Td)." figureDoi="http://doi.org/10.5281/zenodo.4620710" httpUri="https://zenodo.org/record/4620710/files/figure.png" pageId="4" pageNumber="155">Fig. 2F</figureCitation>
).
<emphasis id="1CB67778EF57D326FDF9F8690E57F85A" box="[609,824,1987,2012]" italics="true" pageId="4" pageNumber="155">Tarsus-metatarsus:</emphasis>
Internal face with two lines of dorsomedial bristles; suture between the tarsus and metatarsus is not visible. Tarsus without an inconspicuous spine on the proximal end of the dorsomedial surface.
</paragraph>
<paragraph id="2E7DAB6AEF56D327FF5FFF750E39FEC7" blockId="5.[151,1437,151,502]" pageId="5" pageNumber="156">
<emphasis id="1CB67778EF56D327FF5FFF750C6AFF7F" bold="true" box="[199,261,223,249]" pageId="5" pageNumber="156">
<emphasis id="1CB67778EF56D327FF5FFF750D92FF7F" bold="true" box="[199,253,223,249]" italics="true" pageId="5" pageNumber="156">Legs</emphasis>
.
</emphasis>
Femora brown. Femora lengths: I&gt;III&gt;II&gt;IV (
<tableCitation id="63409ED1EF56D327FC82FF750E1FFF7F" box="[794,880,223,249]" captionStart="TABLE 1" captionStartId="7.[151,239,151,175]" captionText="TABLE 1. Measurements (mm) of body, pedipalps segments, and femur of legs in Phrynus operculatus. When the segment is missing, the cell is empty." pageId="5" pageNumber="156">Table 1</tableCitation>
). Leg I: Tibia with 29 segments and tarsus with 62 segments in both legs. Leg IV: Basitibia with three segments. Basitibia-distitibia lengths: BT1&gt;DT&gt;BT3&gt;BT2. Basitarsus and Telotarsus subequal in size. Tarsus tetramerous.
</paragraph>
<paragraph id="2E7DAB6AEF56D327FF5FFEE00E22FE70" blockId="5.[151,1437,151,502]" pageId="5" pageNumber="156">
<emphasis id="1CB67778EF56D327FF5FFEE00CFDFEE3" bold="true" box="[199,402,330,357]" italics="true" pageId="5" pageNumber="156">Female genitalia.</emphasis>
Genital operculum pentagonal, it extends to the second segment of the opisthosoma; gonopod bases tubular, dorsal surface poorly sclerotized; on the most proximal area, there is a strongly sclerotized line joining both gonopod bases, coloration brown on the most proximal region, and white over entire the dorsal surface. Sclerites of the gonopods claw-like, the base is more than two times wider than medial and apical regions, apex slightly curved. Sclerite coloration brown (
<figureCitation id="B6F9B7EFEF56D327FDE0FE710FB0FE73" box="[632,735,475,501]" captionStart="FIGURE 17" captionStartId="27.[151,250,1596,1620]" captionTargetBox="[158,1428,188,1562]" captionTargetId="figure-15@27.[151,1437,181,1572]" captionTargetPageId="27" captionText="FIGURE 17. Female genitalia: A. Phrynus operculatus (CNAN-Am000407). B. Phrynus alfonsoi sp. nov. (CNAN-T01383). C. Phrynus aliciae sp. nov. (CNAN-T01447). D. Phrynus giseae sp. nov. (CNAN-T01450). E. Phrynus tresmarias sp. nov. (CASENT 9060565). F. Phrynus jalisco (CASENT 9060519). Sclerotization of the gonopod bases (black arrows); colored lines illustrate the position of the sclerite of the gonopod with respect to the gonopod base, the length comparison between the gonopod base (green line) and sclerite of the gonopod (red line), and the comparison between width of the base of the sclerite (purple line) and the width of the medial region of the gonopod base (blue line)." figureDoi="http://doi.org/10.5281/zenodo.4620748" httpUri="https://zenodo.org/record/4620748/files/figure.png" pageId="5" pageNumber="156">Fig. 17A</figureCitation>
,
<tableCitation id="63409ED1EF56D327FD72FE710E50FE70" box="[746,831,475,502]" captionStart="TABLE 6" captionStartId="26.[151,239,368,392]" captionText="TABLE 6. Size proportions in the female genital: Gonopod base (Gb), Sclerite (Scl), and the male genital: lobus dorsalis (LoD), lobus lateralis primus (LoL1). Comparing their total length (l), medial width (mw), and base width (bw)." pageId="5" pageNumber="156">Table 6</tableCitation>
).
</paragraph>
</subSubSection>
<caption id="7ABDFBE2EF56D327FF0FF8C50E32F82D" ID-DOI="http://doi.org/10.5281/zenodo.4620710" ID-Zenodo-Dep="4620710" httpUri="https://zenodo.org/record/4620710/files/figure.png" pageId="5" pageNumber="156" startId="5.[151,250,1903,1927]" targetBox="[160,1423,531,1867]" targetPageId="5">
<paragraph id="2E7DAB6AEF56D327FF0FF8C50E32F82D" blockId="5.[151,1437,1903,1963]" pageId="5" pageNumber="156">
<emphasis id="1CB67778EF56D327FF0FF8C50C7CF801" bold="true" box="[151,275,1903,1927]" pageId="5" pageNumber="156">FIGURE 2.</emphasis>
<taxonomicName id="E9C2D0E9EF56D327FE80F8C50C85F801" authorityName="Pocock" authorityYear="1902" box="[280,490,1903,1927]" class="Arachnida" family="Phrynidae" genus="Phrynus" kingdom="Animalia" order="Amblypygi" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="operculatus">
<emphasis id="1CB67778EF56D327FE80F8C50C85F801" box="[280,490,1903,1927]" italics="true" pageId="5" pageNumber="156">Phrynus operculatus</emphasis>
</taxonomicName>
. Female (CNAN-Am000407): A. Ventral femur (Fv). B. Dorsal femur (Fd). C. Ventral
<taxonomicName id="E9C2D0E9EF56D327FAC1F8C50DA9F82D" authority="(Pv)" baseAuthorityName="Pv" class="Gastropoda" family="Patellidae" genus="Patella" kingdom="Animalia" order="Pezizales" pageId="5" pageNumber="176" phylum="Mollusca" rank="species" species="patella">patella (Pv)</taxonomicName>
. D. Dorsal
<taxonomicName id="E9C2D0E9EF56D327FEA0F8390CDEF82D" authority="(Pd)" baseAuthorityName="Pd" box="[312,433,1939,1963]" class="Gastropoda" family="Patellidae" genus="Patella" kingdom="Animalia" order="Pezizales" pageId="5" pageNumber="176" phylum="Mollusca" rank="species" species="patella">patella (Pd)</taxonomicName>
. E. Ventral tibia (Tv). F. Dorsal tibia (Td).
</paragraph>
</caption>
<subSubSection id="66D8F8E1EF55D324FF5FFF3C0945FCBF" pageId="6" pageNumber="157" type="description">
<paragraph id="2E7DAB6AEF55D324FF5FFF3C0CA8FDE6" blockId="6.[151,1437,150,825]" pageId="6" pageNumber="157">
<emphasis id="1CB67778EF55D324FF5FFF3C0C50FF36" bold="true" box="[199,319,150,176]" pageId="6" pageNumber="157">Variation.</emphasis>
Measurements and the number of segments in the basitibia IV, Tibia I, and tarsus I, are summarized in
<tableCitation id="63409ED1EF55D324FF2DFF100C65FF52" box="[181,266,186,212]" captionStart="TABLE 1" captionStartId="7.[151,239,151,175]" captionText="TABLE 1. Measurements (mm) of body, pedipalps segments, and femur of legs in Phrynus operculatus. When the segment is missing, the cell is empty." pageId="6" pageNumber="157">Table 1</tableCitation>
.
<emphasis id="1CB67778EF55D324FE81FF100CE9FF52" bold="true" box="[281,390,186,212]" pageId="6" pageNumber="157">
<emphasis id="1CB67778EF55D324FE81FF100C12FF52" bold="true" box="[281,381,186,212]" italics="true" pageId="6" pageNumber="157">Sternum</emphasis>
:
</emphasis>
Setae over all segments of the sternum are variable in number and position.
<emphasis id="1CB67778EF55D324FB6EFF100D82FF7E" italics="true" pageId="6" pageNumber="157">
<emphasis id="1CB67778EF55D324FB6EFF10080BFF53" bold="true" box="[1270,1380,186,213]" italics="true" pageId="6" pageNumber="157">Pedipalp:</emphasis>
Trochanter
</emphasis>
: line of dorsal tubercles varies between four and six tubercles.
<emphasis id="1CB67778EF55D324FC20FF75096CFF7E" box="[952,1027,223,248]" italics="true" pageId="6" pageNumber="157">Femur</emphasis>
: Ventral face: tubercles are constant in most individuals, however, the size is variable, as well as the size of spines Fv4, Fv3a, Fv4a, and Fv5a, which are always smaller than major spines, but the relative sizes between them are variable. Dorsal face: the size and presence of tubercles mentioned in the complementary description are variable.
<taxonomicName id="E9C2D0E9EF55D324FB97FEE009D1FEE2" authority=": Ventral" authorityName="Ventral" box="[1039,1214,330,357]" class="Gastropoda" family="Patellidae" genus="Patella" kingdom="Animalia" order="Pezizales" pageId="6" pageNumber="157" phylum="Mollusca" rank="genus">
<emphasis id="1CB67778EF55D324FB97FEE0090FFEE2" box="[1039,1120,330,356]" italics="true" pageId="6" pageNumber="157">Patella</emphasis>
: Ventral
</taxonomicName>
face: Tubercles and small spines are more conspicuous in larger individuals. Dorsal face: tubercles can be more conspicuous, in some individuals the tubercles can be present between each spine, in larger specimens a small spine Pd8 can be present.
<emphasis id="1CB67778EF55D324FF0FFE1C0DBEFE56" box="[151,209,438,464]" italics="true" pageId="6" pageNumber="157">Tibia</emphasis>
: Ventral face: between Tv2Tv3, two or three small spines can be present. Dorsal face: tubercles between spines can be more conspicuous.
<emphasis id="1CB67778EF55D324FD93FE710F27FE73" bold="true" box="[523,584,475,501]" italics="true" pageId="6" pageNumber="157">Legs.</emphasis>
The number of segments of the tibia and tarsus of leg I, was variable among the individuals, the number of tibial segments varies between 29 and 42, and the tarsal segments between 62 and 69. In specimens studied, the number of segments did not depend on sex, frequently there are more tarsal segments in larger specimens (
<tableCitation id="63409ED1EF55D324FEFAFDEC0CD9FDE6" box="[354,438,582,608]" captionStart="TABLE 1" captionStartId="7.[151,239,151,175]" captionText="TABLE 1. Measurements (mm) of body, pedipalps segments, and femur of legs in Phrynus operculatus. When the segment is missing, the cell is empty." pageId="6" pageNumber="157">Table 1</tableCitation>
).
</paragraph>
<paragraph id="2E7DAB6AEF55D324FF5FFDC0094AFD2E" blockId="6.[151,1437,150,825]" pageId="6" pageNumber="157">
<emphasis id="1CB67778EF55D324FF5FFDC00C64FD03" bold="true" box="[199,267,618,645]" pageId="6" pageNumber="157">Male.</emphasis>
Males are similar in size to females (
<tableCitation id="63409ED1EF55D324FD33FDC00F92FD02" box="[683,765,618,644]" captionStart="TABLE 1" captionStartId="7.[151,239,151,175]" captionText="TABLE 1. Measurements (mm) of body, pedipalps segments, and femur of legs in Phrynus operculatus. When the segment is missing, the cell is empty." pageId="6" pageNumber="157">Table 1</tableCitation>
) spination pattern as in females, genital operculum with the posterior margin oval, and covering part of the third segment of the opisthosoma.
</paragraph>
<paragraph id="2E7DAB6AEF55D324FF5FFD180945FCBF" blockId="6.[151,1437,150,825]" pageId="6" pageNumber="157">
<emphasis id="1CB67778EF55D324FF5FFD180F12FD4A" bold="true" box="[199,637,690,717]" pageId="6" pageNumber="157">
<emphasis id="1CB67778EF55D324FF5FFD180C1DFD4B" bold="true" box="[199,370,690,717]" italics="true" pageId="6" pageNumber="157">Male genitalia</emphasis>
. (CASENT 9060512):
</emphasis>
LaM longer than Pi. LoL 1 a little longer than LoL 2, both densely covered with minute projections, in LoL 1, projections are tubular, thin, and elongated, placed close together, the insertion base is not visible; in LoL 2 projections are similar to LoL 1, the insertion base is also not visible. In dorsal view, the LoD is thick and elongated with a similar size to LoL1 (
<figureCitation id="B6F9B7EFEF55D324FCEFFCB40ED4FCBE" box="[887,955,798,824]" captionStart="FIGURE 3" captionStartId="6.[151,250,1775,1799]" captionTargetBox="[156,1432,852,1747]" captionTargetId="figure-368@6.[151,1436,848,1751]" captionTargetPageId="6" captionText="FIGURE 3. Male genital: Phrynus operculatus (CASENT 9060512): A. ventral view. B. Detail of ornamentation of LoL 1 and LoL 2. C. Dorsal view. Fi = fistula; LaM = lamina medialis; LoD = lobus dorsalis; LoL 1 = lobus lateralis primus; LoL 2 = lobus lateralis secundus; Pi = processus internus of gonopod. Colored lines illustrate the comparison between the LoD medial width (green line) and the width of the base of LoL 1 (red line)." figureDoi="http://doi.org/10.5281/zenodo.4620718" httpUri="https://zenodo.org/record/4620718/files/figure.png" pageId="6" pageNumber="157">Fig. 3</figureCitation>
,
<tableCitation id="63409ED1EF55D324FC5FFCB40973FCBF" box="[967,1052,798,825]" captionStart="TABLE 6" captionStartId="26.[151,239,368,392]" captionText="TABLE 6. Size proportions in the female genital: Gonopod base (Gb), Sclerite (Scl), and the male genital: lobus dorsalis (LoD), lobus lateralis primus (LoL1). Comparing their total length (l), medial width (mw), and base width (bw)." pageId="6" pageNumber="157">Table 6</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>