treatments-xml/data/9C/54/87/9C5487F9FF8AFF9924D845B4FA00FBE7.xml
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<document id="453DAD97E3386BD9D924827460A88090" ID-DOI="10.5281/zenodo.188125" ID-GBIF-Dataset="c573d381-a5e4-4bb5-8c6b-5fd1eeeaf2ce" ID-ISSN="1175-5326" ID-Zenodo-Dep="188125" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461084044284" checkinUser="plazi" docAuthor="Mclay, Colin L." docDate="2009" docId="9C5487F9FF8AFF9924D845B4FA00FBE7" docLanguage="en" docName="zt02122p050.pdf" docOrigin="Zootaxa 2122" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Garthambrus" docType="key" docVersion="9" lastPageNumber="9" masterDocId="606DFF81FF82FF91244F4523FF9CFF9F" masterDocTitle="Revision of the crab genus Garthambrus Ng, 1996, with the description of two new genera and discussion of the status of Tutankhamen Rathbun, 1925 (Crustacea: Brachyura: Parthenopidae)" masterLastPageNumber="50" masterPageNumber="1" pageNumber="9" updateTime="1698587833449" updateUser="plazi">
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<mods:title id="8AB30EDC0BC320A484D99277F85BB7EF">Revision of the crab genus Garthambrus Ng, 1996, with the description of two new genera and discussion of the status of Tutankhamen Rathbun, 1925 (Crustacea: Brachyura: Parthenopidae)</mods:title>
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<mods:namePart id="68BA330A6AFB90E33B4FF44098129D65">Mclay, Colin L.</mods:namePart>
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<mods:date id="A5B86120D49C346500667F9EF4F9657B">2009</mods:date>
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<treatment id="9C5487F9FF8AFF9924D845B4FA00FBE7" ID-DOI="http://doi.org/10.5281/zenodo.6212854" ID-GBIF-Taxon="119618610" ID-Zenodo-Dep="6212854" LSID="urn:lsid:plazi:treatment:9C5487F9FF8AFF9924D845B4FA00FBE7" httpUri="http://treatment.plazi.org/id/9C5487F9FF8AFF9924D845B4FA00FBE7" lastPageNumber="9" pageId="8" pageNumber="9">
<subSubSection id="5CE76564FF8AFF9924D845B4FE6DFF2E" box="[151,497,151,177]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="144236EFFF8AFF9924D845B4FE6DFF2E" blockId="8.[151,497,151,177]" box="[151,497,151,177]" pageId="8" pageNumber="9">
<heading id="4F0A8183FF8AFF9924D845B4FE6DFF2E" bold="true" box="[151,497,151,177]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
<emphasis id="2689EAFDFF8AFF9924D845B4FE6DFF2E" bold="true" box="[151,497,151,177]" pageId="8" pageNumber="9">
Key to
<taxonomicName id="D3FD4D6CFF8AFF9924BF45B4FE0FFF2E" ID-CoL="7P36J" box="[240,403,151,177]" class="Malacostraca" family="Parthenopidae" genus="Garthambrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="2689EAFDFF8AFF9924BF45B4FE0FFF2E" bold="true" box="[240,403,151,177]" italics="true" pageId="8" pageNumber="9">Garthambrus</emphasis>
</taxonomicName>
species
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5CE76564FF8AFF9924D845C2FA00FBE7" pageId="8" pageNumber="9" type="key">
<keyStep id="AF092E4AFF8AFF9924D845C2FA07FEA7" pageId="8" pageNumber="9">
<paragraph id="144236EFFF8AFF9924D845C2FA00FE87" blockId="8.[151,1436,225,1144]" pageId="8" pageNumber="9">
<keyLead id="AF0C95DAFF8AFF9924D845C2FA00FE87" pageId="8" pageNumber="9">
1. Posterior edge of metabranchial region with a raised triangular patch and anterior epibranchial margin with well- separated evenly-spaced teeth each bearing minute spinules ..............................................................
<emphasis id="2689EAFDFF8AFF9920B94422FA00FE87" box="[1270,1436,257,280]" italics="true" pageId="8" pageNumber="9">
G.
<taxonomicName id="D3FD4D6CFF8AFF99215B4422FA00FE87" ID-CoL="3FBTC" box="[1300,1436,257,280]" class="Malacostraca" family="Parthenopidae" genus="Garthambrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="complanatus">complanatus</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="144236EFFF8AFF9924D84402FA07FEA7" blockId="8.[151,1436,225,1144]" box="[151,1435,289,312]" pageId="8" pageNumber="9">
<keyLead id="AF0C95DAFF8AFF9924D84402FA07FEA7" box="[151,1435,289,312]" pageId="8" pageNumber="9">- Posterior edge of metabranchial region without raised triangular patch and anterior epibranchial margin otherwise 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="AF092E4AFF8AFF9924D84462FA07FE07" pageId="8" pageNumber="9">
<paragraph id="144236EFFF8AFF9924D84462FA07FEE7" blockId="8.[151,1436,225,1144]" pageId="8" pageNumber="9">
<keyLead id="AF0C95DAFF8AFF9924D84462FA07FEE7" pageId="8" pageNumber="9">2. Posterior edge of metabranchial region with several tubercles, which can be stellate or granular leading to edge of posterior margin .......................................................................................................................................................... 3</keyLead>
</paragraph>
<paragraph id="144236EFFF8AFF9924D844A2FA07FE07" blockId="8.[151,1436,225,1144]" box="[151,1435,385,408]" pageId="8" pageNumber="9">
<keyLead id="AF0C95DAFF8AFF9924D844A2FA07FE07" box="[151,1435,385,408]" pageId="8" pageNumber="9">- Posterior edge of metabranchial region with either a spine or tubercle, not leading to edge of posterior margin ..... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="AF092E4AFF8AFF9924D84482FA00FD87" pageId="8" pageNumber="9">
<paragraph id="144236EFFF8AFF9924D84482FA00FE47" blockId="8.[151,1436,225,1144]" pageId="8" pageNumber="9">
<keyLead id="AF0C95DAFF8AFF9924D84482FA00FE47" pageId="8" pageNumber="9">
3. Dorsal surface of carapace with paxillate tubercles; tubercles on upper margin of ambulatory legs rounded at tip; tubercles on surfaces of ambulatory legs well spaced ..................................................................................
<emphasis id="2689EAFDFF8AFF99216244E2FA00FE47" box="[1325,1436,449,472]" italics="true" pageId="8" pageNumber="9">
G.
<taxonomicName id="D3FD4D6CFF8AFF99210444E2FA00FE47" ID-CoL="4DTWS" box="[1355,1436,449,472]" class="Malacostraca" family="Parthenopidae" genus="Parthenope" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="poupini">poupini</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="144236EFFF8AFF9924D844C2FA00FD87" blockId="8.[151,1436,225,1144]" pageId="8" pageNumber="9">
<keyLead id="AF0C95DAFF8AFF9924D844C2FA00FD87" pageId="8" pageNumber="9">
- Dorsal surface of carapace with stellate granules; sharp tipped tubercles on upper margin of ambulatory legs; tuber- cles on surfaces of ambulatory legs very close together ...............................................................................
<emphasis id="2689EAFDFF8AFF9921604722FA00FD87" box="[1327,1436,513,536]" italics="true" pageId="8" pageNumber="9">
G.
<taxonomicName id="D3FD4D6CFF8AFF9921034722FA00FD87" ID-CoL="6JYHW" box="[1356,1436,513,536]" class="Malacostraca" family="Parthenopidae" genus="Garthambrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="allisoni">allisoni</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="AF092E4AFF8AFF9924D84702FA07FDC7" pageId="8" pageNumber="9">
<paragraph id="144236EFFF8AFF9924D84702FA07FDA7" blockId="8.[151,1436,225,1144]" box="[151,1435,545,568]" pageId="8" pageNumber="9">
<keyLead id="AF0C95DAFF8AFF9924D84702FA07FDA7" box="[151,1435,545,568]" pageId="8" pageNumber="9">4. Upper margin of ambulatory legs cristate ................................................................................................................... 5</keyLead>
</paragraph>
<paragraph id="144236EFFF8AFF9924D84762FA07FDC7" blockId="8.[151,1436,225,1144]" box="[151,1435,577,600]" pageId="8" pageNumber="9">
<keyLead id="AF0C95DAFF8AFF9924D84762FA07FDC7" box="[151,1435,577,600]" pageId="8" pageNumber="9">- Upper margin of ambulatory legs tuberculate ............................................................................................................ 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="AF092E4AFF8AFF9924D84742FA07FD07" pageId="8" pageNumber="9">
<paragraph id="144236EFFF8AFF9924D84742FA00FDE7" blockId="8.[151,1436,225,1144]" box="[151,1436,609,632]" pageId="8" pageNumber="9">
<keyLead id="AF0C95DAFF8AFF9924D84742FA00FDE7" box="[151,1436,609,632]" pageId="8" pageNumber="9">
5. Subhepatic region not excavated ...............................................................................................................
<emphasis id="2689EAFDFF8AFF99215B4742FA00FDE7" box="[1300,1436,609,632]" italics="true" pageId="8" pageNumber="9">
G.
<taxonomicName id="D3FD4D6CFF8AFF99217E4742FA00FDE7" ID-CoL="6JYHN" box="[1329,1436,609,632]" class="Malacostraca" family="Parthenopidae" genus="Garthambrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lacunosus">lacunosus</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="144236EFFF8AFF9924D847A2FA07FD07" blockId="8.[151,1436,225,1144]" box="[151,1435,641,664]" pageId="8" pageNumber="9">
<keyLead id="AF0C95DAFF8AFF9924D847A2FA07FD07" box="[151,1435,641,664]" pageId="8" pageNumber="9">- Subhepatic region excavated ...................................................................................................................................... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="AF092E4AFF8AFF9924D84782FA00FD47" pageId="8" pageNumber="9">
<paragraph id="144236EFFF8AFF9924D84782FA00FD27" blockId="8.[151,1436,225,1144]" box="[151,1436,673,696]" pageId="8" pageNumber="9">
<keyLead id="AF0C95DAFF8AFF9924D84782FA00FD27" box="[151,1436,673,696]" pageId="8" pageNumber="9">
6. Hepatic tooth absent. Hepatic cavity broad and deep ............................................................................
<emphasis id="2689EAFDFF8AFF99214E4782FA00FD27" box="[1281,1436,673,696]" italics="true" pageId="8" pageNumber="9">
G.
<taxonomicName id="D3FD4D6CFF8AFF9921514782FA00FD27" ID-CoL="3FBTD" box="[1310,1436,673,696]" class="Malacostraca" family="Parthenopidae" genus="Garthambrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="darthvaderi">darthvaderi</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="144236EFFF8AFF9924D847E2FA00FD47" blockId="8.[151,1436,225,1144]" box="[151,1436,705,728]" pageId="8" pageNumber="9">
<keyLead id="AF0C95DAFF8AFF9924D847E2FA00FD47" box="[151,1436,705,728]" pageId="8" pageNumber="9">
- Hepatic tooth present. Hepatic cavity narrow and shallow ....................................................................
<emphasis id="2689EAFDFF8AFF99214947E2FA00FD47" box="[1286,1436,705,728]" italics="true" pageId="8" pageNumber="9">
G.
<taxonomicName id="D3FD4D6CFF8AFF99216B47E2FA00FD47" ID-CoL="3FBTJ" box="[1316,1436,705,728]" class="Malacostraca" family="Parthenopidae" genus="Garthambrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pteromerus">pteromerus</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="AF092E4AFF8AFF9924D847C2FA07FCA7" pageId="8" pageNumber="9">
<paragraph id="144236EFFF8AFF9924D847C2FA07FC87" blockId="8.[151,1436,225,1144]" pageId="8" pageNumber="9">
<keyLead id="AF0C95DAFF8AFF9924D847C2FA07FC87" pageId="8" pageNumber="9">
7. Upper margin of ambulatory legs with only one or two rounded tubercles on proximal and distal portion, middle portion smooth ..............................................................................................................................................
<emphasis id="2689EAFDFF8AFF9921654622FA07FC87" box="[1322,1435,769,792]" italics="true" pageId="8" pageNumber="9">
G.
<taxonomicName id="D3FD4D6CFF8AFF9921084622FA07FC87" ID-CoL="8YX8W" box="[1351,1435,769,792]" class="Malacostraca" family="Parthenopidae" genus="Garthambrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="posidon">posidon</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="144236EFFF8AFF9924D84602FA07FCA7" blockId="8.[151,1436,225,1144]" box="[151,1435,801,824]" pageId="8" pageNumber="9">
<keyLead id="AF0C95DAFF8AFF9924D84602FA07FCA7" box="[151,1435,801,824]" pageId="8" pageNumber="9">- Upper margin of ambulatory legs with well-spaced tubercles on entire upper margin .............................................. 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="AF092E4AFF8AFF9924D84662FA07FCE7" pageId="8" pageNumber="9">
<paragraph id="144236EFFF8AFF9924D84662FA00FCC7" blockId="8.[151,1436,225,1144]" box="[151,1436,833,856]" pageId="8" pageNumber="9">
<keyLead id="AF0C95DAFF8AFF9924D84662FA00FCC7" box="[151,1436,833,856]" pageId="8" pageNumber="9">
8. Dorsal surface of carapace densely covered with granulate tubercles, ambulatory legs with spines .........
<emphasis id="2689EAFDFF8AFF9921694662FA00FCC7" box="[1318,1436,833,856]" italics="true" pageId="8" pageNumber="9">
G.
<taxonomicName id="D3FD4D6CFF8AFF99210C4662FA00FCC7" ID-CoL="3FBTK" box="[1347,1436,833,856]" class="Malacostraca" family="Parthenopidae" genus="Garthambrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="stellatus">stellatus</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="144236EFFF8AFF9924D84642FA07FCE7" blockId="8.[151,1436,225,1144]" box="[151,1435,865,888]" pageId="8" pageNumber="9">
<keyLead id="AF0C95DAFF8AFF9924D84642FA07FCE7" box="[151,1435,865,888]" pageId="8" pageNumber="9">- Dorsal surface of carapace not densely tuberculate, ambulatory legs without spines ................................................ 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="AF092E4AFF8AFF9924D846A2FA07FC67" pageId="8" pageNumber="9">
<paragraph id="144236EFFF8AFF9924D846A2FA04FC27" blockId="8.[151,1436,225,1144]" pageId="8" pageNumber="9">
<keyLead id="AF0C95DAFF8AFF9924D846A2FA04FC27" pageId="8" pageNumber="9">
9. Distal tubercles of upper margin of ambulatory leg meri flattened laterally, sub-cristatiform. Hepatic tooth cristati- form. Teeth on anterior epibranchial margin narrow, well-spaced from each other ..........................................
<emphasis id="2689EAFDFF8AFF99211A4682FA04FC27" box="[1365,1432,929,952]" italics="true" pageId="8" pageNumber="9">G. tan i</emphasis>
</keyLead>
</paragraph>
<paragraph id="144236EFFF8AFF9924D846E2FA07FC67" blockId="8.[151,1436,225,1144]" pageId="8" pageNumber="9">
<keyLead id="AF0C95DAFF8AFF9924D846E2FA07FC67" pageId="8" pageNumber="9">- Distal tubercles of upper margin of ambulatory leg meri not flattened laterally. Hepatic shoulder forms right-angle not cristatiform. Teeth on anterior epibranchial margin broad, bases almost adjacent ............................................. 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="AF092E4AFF8AFF9924D84122FA00FBE7" pageId="8" pageNumber="9">
<paragraph id="144236EFFF8AFF9924D84122FA04FBA7" blockId="8.[151,1436,225,1144]" pageId="8" pageNumber="9">
<keyLead id="AF0C95DAFF8AFF9924D84122FA04FBA7" pageId="8" pageNumber="9">
10. Lateral tooth at epibranchial corner short, not elongated. Tubercles on dorsal surface of carapace paxilliform .......... ........................................................................................................................................................................
<emphasis id="2689EAFDFF8AFF99217B4102FA04FBA7" box="[1332,1432,1057,1080]" italics="true" pageId="8" pageNumber="9">G. c i d a r i s</emphasis>
</keyLead>
</paragraph>
<paragraph id="144236EFFF8AFF9924D84162FA00FBE7" blockId="8.[151,1436,225,1144]" pageId="8" pageNumber="9">
<keyLead id="AF0C95DAFF8AFF9924D84162FA00FBE7" pageId="8" pageNumber="9">
- Lateral tooth at epibranchial corner elongated, tip acute. Tubercles on dorsal surface of carapace not paxilliform ..... ...................................................................................................................................................................
<emphasis id="2689EAFDFF8AFF99215A4142FA00FBE7" box="[1301,1436,1121,1144]" italics="true" pageId="8" pageNumber="9">
G.
<taxonomicName id="D3FD4D6CFF8AFF99217C4142FA00FBE7" ID-CoL="3FBTM" box="[1331,1436,1121,1144]" class="Malacostraca" family="Parthenopidae" genus="Garthambrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="undulatus">undulatus</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
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