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<document id="B2DFA757452193045340EB67BAE67181" ID-DOI="10.1080/00222930701529273" ID-ISSN="1464-5262" ID-Zenodo-Dep="5232664" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629503227275" checkinUser="felipe" docAuthor="Arconada, Beatriz, Delicado, Diana &amp; Ramos, M. Ángeles" docDate="2007" docId="987BEB16FFA59E5BFE45D0185239B191" docLanguage="en" docName="JNatHist.41.29-32.2007-2035.pdf" docOrigin="Journal of Natural History 41 (29 - 32)" docSource="http://dx.doi.org/10.1080/00222930701529273" docStyle="DocumentStyle:5A9EB2ED5589A523F0236C66DE2A6BA8.4:JNatHist.2005-2007.journal_article.1cover" docStyleId="5A9EB2ED5589A523F0236C66DE2A6BA8" docStyleName="JNatHist.2005-2007.journal_article.1cover" docStyleVersion="4" docTitle="Iberhoratia gatoa Arconada &amp; Delicado &amp; Ramos 2007, n. comb." docType="treatment" docVersion="3" lastPageNumber="2028" masterDocId="6442936EFFB09E4CFFD1D2155620B704" masterDocTitle="A new genus and two new species of Hydrobiidae (Mollusca, Caenogastropoda) from the Iberian Peninsula" masterLastPageNumber="2035" masterPageNumber="2007" pageNumber="2027" updateTime="1699075173358" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="5FF7A8FADC38EE5C53B65B0293E628DA">A new genus and two new species of Hydrobiidae (Mollusca, Caenogastropoda) from the Iberian Peninsula</mods:title>
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<mods:namePart id="F071570CA2F26D572216F4C7AFFD6652">Arconada, Beatriz</mods:namePart>
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<mods:namePart id="FAAD6F881732C9EF97FA2D10E7890A06">Delicado, Diana</mods:namePart>
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<mods:namePart id="0142771971396C36A99C6B9156E62C8D">Ramos, M. Ángeles</mods:namePart>
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<mods:title id="CD580FC70EACE83F1474B3C688901293">Journal of Natural History</mods:title>
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<mods:date id="F727B38FF7F8AA4A93C7E9A2D850A68E">2007</mods:date>
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<mods:number id="02B5FD378EB1512FE9DF6B53F8E25F0B">2010-12-02</mods:number>
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<mods:number id="18B02EC284262EF97AA5C8ABA8EB902B">41</mods:number>
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<mods:number id="80C571F203077D53B6CDF968FECB4664">29 - 32</mods:number>
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<mods:start id="0CE03DB21B1305DB1AA7770D2E675B31">2007</mods:start>
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<mods:identifier id="E020B4540634778E0DC2E8EDA36EC8FA" type="ISSN">1464-5262</mods:identifier>
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<treatment id="987BEB16FFA59E5BFE45D0185239B191" LSID="urn:lsid:plazi:treatment:987BEB16FFA59E5BFE45D0185239B191" httpUri="http://treatment.plazi.org/id/987BEB16FFA59E5BFE45D0185239B191" lastPageId="23" lastPageNumber="2028" pageId="21" pageNumber="2027">
<subSubSection id="58C8098BFFA59E59FE45D018558BB521" box="[404,939,525,549]" pageId="21" pageNumber="2027" type="nomenclature">
<paragraph id="106D5A00FFA59E59FE45D018558BB521" blockId="21.[140,939,525,615]" box="[404,939,525,549]" pageId="21" pageNumber="2027">
<heading id="4B25ED6CFFA59E59FE45D018558BB521" box="[404,939,525,549]" centered="true" fontSize="10" level="2" pageId="21" pageNumber="2027" reason="2">
<taxonomicName id="D7D22183FFA59E59FE45D0185519B521" ID-CoL="3P78N" authority="(Boeters, 1980)" authorityName="Arconada &amp; Delicado &amp; Ramos" authorityYear="2007" baseAuthorityName="Boeters" baseAuthorityYear="1980" box="[404,825,525,549]" class="Gastropoda" family="Hydrobiidae" genus="Iberhoratia" kingdom="Animalia" order="Littorinimorpha" pageId="21" pageNumber="2027" phylum="Mollusca" rank="species" species="gatoa" status="comb. nov.">
<emphasis id="22A68612FFA59E59FE45D0185459B521" bold="true" box="[404,633,525,549]" italics="true" pageId="21" pageNumber="2027">Iberhoratia gatoa</emphasis>
(
<bibRefCitation id="744327F1FFA59E59FD5AD0185512B521" author="Boeters HD" box="[651,818,525,549]" pageId="21" pageNumber="2027" pagination="61 - 64" refId="ref12950" refString="Boeters HD. 1980. Unbekannte westeuropaische Prosobranchia, 3. Basteria 44: 61 - 64." type="journal article" year="1980">Boeters, 1980</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="39953B69FFA59E59FC93D018558BB521" box="[834,939,525,549]" pageId="21" pageNumber="2027" rank="species">n. comb.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="58C8098BFFA59E59FD87D03B54D3B542" box="[598,755,558,582]" pageId="21" pageNumber="2027" type="description">
<paragraph id="106D5A00FFA59E59FD87D03B54D3B542" blockId="21.[140,939,525,615]" box="[598,755,558,582]" pageId="21" pageNumber="2027">
(
<figureCitation id="88E94685FFA59E59FD8FD03B54EBB542" box="[606,715,558,582]" captionStart="Figure 8" captionStartId="22.[259,319,534,553]" captionTargetBox="[125,1173,133,498]" captionTargetId="figure-126@22.[125,1173,133,498]" captionTargetPageId="22" captionText="Figure 8. Holotype of Iberhoratia gatoa (RMNH 55467). Scale bar: 500 Mm (A, B)." figureDoi="http://doi.org/10.5281/zenodo.5232678" httpUri="https://zenodo.org/record/5232678/files/figure.png" pageId="21" pageNumber="2027">Figures 8</figureCitation>
,
<figureCitation id="88E94685FFA59E59FD0BD03B54CAB542" box="[730,746,558,582]" captionStart="Figure 9" captionStartId="22.[125,185,1611,1630]" captionTargetBox="[127,1170,897,1573]" captionTargetId="figure-84@22.[125,1173,897,1575]" captionTargetPageId="22" captionText="Figure 9. Original illustrations of Iberhoratia gatoa (Boeters 1980, p 63). (A, B) Holotype (RMNH 55467), shell width 1.5 mm; (C, D) paratype (BOE 948), shell width 1.2 mm; (E) penis (paratype BOE 948); (F) anterior female genitalia (paratype BOE 948)." figureDoi="http://doi.org/10.5281/zenodo.5232680" httpUri="https://zenodo.org/record/5232680/files/figure.png" pageId="21" pageNumber="2027">9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="58C8098BFFA59E59FF5DD05A54D5B563" box="[140,757,591,615]" pageId="21" pageNumber="2027" type="reference_group">
<paragraph id="106D5A00FFA59E59FF5DD05A54D5B563" blockId="21.[140,939,525,615]" box="[140,757,591,615]" pageId="21" pageNumber="2027">
<treatmentCitationGroup id="30C27D2EFFA59E59FF5DD05A54D5B563" box="[140,757,591,615]" pageId="21" pageNumber="2027">
<treatmentCitation id="91737C11FFA59E59FF5DD05A542BB563" author="Boeters HD" box="[140,523,591,615]" page="62" pageId="21" pageNumber="2027" year="1980">
<taxonomicName id="D7D22183FFA59E59FF5DD05A542BB563" ID-CoL="7RPRQ" authority="Boeters 1980, p 62" authorityName="Boeters" authorityPageNumber="62" authorityYear="1980" box="[140,523,591,615]" class="Gastropoda" family="Hydrobiidae" genus="Horatia" kingdom="Animalia" order="Littorinimorpha" pageId="21" pageNumber="2027" phylum="Mollusca" rank="species" species="gatoa">
<emphasis id="22A68612FFA59E59FF5DD05A5705B563" box="[140,293,591,615]" italics="true" pageId="21" pageNumber="2027">Horatia gatoa</emphasis>
<bibRefCitation id="744327F1FFA59E59FEFFD05A57ECB563" author="Boeters HD" box="[302,460,591,615]" pageId="21" pageNumber="2027" pagination="61 - 64" refId="ref12950" refString="Boeters HD. 1980. Unbekannte westeuropaische Prosobranchia, 3. Basteria 44: 61 - 64." type="journal article" year="1980">Boeters 1980</bibRefCitation>
, p 62
</taxonomicName>
</treatmentCitation>
64,
<figureCitation id="88E94685FFA59E59FD97D05A5491B563" box="[582,689,591,615]" captionStart="Figure 5" captionStartId="18.[125,185,1313,1332]" captionTargetBox="[125,1173,133,1277]" captionTargetId="figure-98@18.[125,1173,133,1277]" captionTargetPageId="18" captionText="Figure 5. Shells of Iberhoratia aurorae. (AD, G) Shells from Robladillo de Gata; (E, I) shells from Roturas; (F, H, J, K) shell (holotype) from population between Hervás and Jerte (type locality). (HK) Protoconch and microsculpture. Scale bar: 500 Mm (AG)." figureDoi="http://doi.org/10.5281/zenodo.5232672" httpUri="https://zenodo.org/record/5232672/files/figure.png" pageId="21" pageNumber="2027">Figures 5</figureCitation>
,
<figureCitation id="88E94685FFA59E59FD11D05A54F1B563" box="[704,721,591,615]" captionStart="Figure 7" captionStartId="20.[125,185,1507,1526]" captionTargetBox="[125,1173,133,1471]" captionTargetId="figure-0@20.[125,1173,133,1471]" captionTargetPageId="20" captionText="Figure 7. Anatomy of Iberhoratia aurorae from type locality. (A, B) Partial nervous system; (C) osphradium and ctenidium; (D) rectum curve; (E) stomach; (F) head of a male and penis; (G) anterior female genitalia; (H) bursa copulatrix and seminal receptacles. Abbreviations in text. Scale bar: 500 Mm (AH)." figureDoi="http://doi.org/10.5281/zenodo.5232676" httpUri="https://zenodo.org/record/5232676/files/figure.png" pageId="21" pageNumber="2027">7</figureCitation>
,
<figureCitation id="88E94685FFA59E59FD0ED05A54D0B563" box="[735,752,591,615]" captionStart="Figure 8" captionStartId="22.[259,319,534,553]" captionTargetBox="[125,1173,133,498]" captionTargetId="figure-126@22.[125,1173,133,498]" captionTargetPageId="22" captionText="Figure 8. Holotype of Iberhoratia gatoa (RMNH 55467). Scale bar: 500 Mm (A, B)." figureDoi="http://doi.org/10.5281/zenodo.5232678" httpUri="https://zenodo.org/record/5232678/files/figure.png" pageId="21" pageNumber="2027">8</figureCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="58C8098BFFA59E5BFF5BD0B45239B191" lastPageId="23" lastPageNumber="2029" pageId="21" pageNumber="2027" type="materials_examined">
<paragraph id="106D5A00FFA59E59FF5BD0B45736B5BC" blockId="21.[138,278,672,696]" box="[138,278,672,696]" pageId="21" pageNumber="2027">
<emphasis id="22A68612FFA59E59FF5BD0B45736B5BC" box="[138,278,672,696]" italics="true" pageId="21" pageNumber="2027">
<typeStatus id="CF69E4A2FFA59E59FF5BD0B456E0B5BC" box="[138,192,673,696]" pageId="21" pageNumber="2027">Type</typeStatus>
locality
</emphasis>
</paragraph>
<paragraph id="106D5A00FFA59E59FF5BD0C45203B5ED" blockId="21.[138,1059,721,745]" box="[138,1059,721,745]" pageId="21" pageNumber="2027">
<materialsCitation id="A0BA505DFFA59E59FF5BD0C4523FB5ED" box="[138,1055,721,745]" location="Hundidero del Gato" pageId="21" pageNumber="2027" specimenCount="1" typeStatus="holotype">
A cave named
<location id="150D0CDBFFA59E59FEE8D0C4540AB5ED" LSID="urn:lsid:plazi:treatment:987BEB16FFA59E5BFE45D0185239B191:150D0CDBFFA59E59FEE8D0C4540AB5ED" box="[313,554,721,745]" name="Hundidero del Gato" pageId="21" pageNumber="2027">Hundidero del Gato</location>
, Benaoján, Málaga, UTM: 30SVF003673
</materialsCitation>
.
</paragraph>
<paragraph id="106D5A00FFA59E59FF5BD1375778B43E" blockId="21.[138,344,802,826]" box="[138,344,802,826]" pageId="21" pageNumber="2027">
<emphasis id="22A68612FFA59E59FF5BD1375778B43E" box="[138,344,802,826]" italics="true" pageId="21" pageNumber="2027">Material examined</emphasis>
</paragraph>
<paragraph id="106D5A00FFA59E59FF5BD1465281B4E8" blockId="21.[138,1185,850,1004]" pageId="21" pageNumber="2027">
<emphasis id="22A68612FFA59E59FF5BD146570BB46E" box="[138,299,850,874]" italics="true" pageId="21" pageNumber="2027">Type material.</emphasis>
<materialsCitation id="A0BA505DFFA59E59FEE9D1475281B4E8" accessionNumber="NNM 55468" collectionCode="NNM" pageId="21" pageNumber="2027" specimenCode="NNM 9079" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="CF69E4A2FFA59E59FEE9D1475788B46E" box="[312,424,850,874]" pageId="21" pageNumber="2027" type="holotype">Holotype</typeStatus>
: (shell and operculum) in the
<collectionCode id="76C3C2C5FFA59E59FCDFD1475576B46E" box="[782,854,850,874]" country="Netherlands" httpUri="http://grbio.org/cool/wkmc-b7hm" name="National Museum of Natural History, Naturalis" pageId="21" pageNumber="2027" type="Museum">NNM</collectionCode>
collection (55467). Shell of the
<typeStatus id="CF69E4A2FFA59E59FF69D1665700B48F" box="[184,288,883,907]" pageId="21" pageNumber="2027" type="holotype">holotype</typeStatus>
, which has the peristome damaged, has been examined (
<figureCitation id="88E94685FFA59E59FC6ED166527BB48F" box="[959,1115,883,907]" captionStart="Figure 8" captionStartId="22.[259,319,534,553]" captionTargetBox="[125,1173,133,498]" captionTargetId="figure-126@22.[125,1173,133,498]" captionTargetPageId="22" captionText="Figure 8. Holotype of Iberhoratia gatoa (RMNH 55467). Scale bar: 500 Mm (A, B)." figureDoi="http://doi.org/10.5281/zenodo.5232678" httpUri="https://zenodo.org/record/5232678/files/figure.png" pageId="21" pageNumber="2027">Figure 8AC</figureCitation>
). The following
<typeStatus id="CF69E4A2FFA59E59FED3D1865750B4AF" box="[258,368,915,939]" pageId="21" pageNumber="2027" type="paratype">paratypes</typeStatus>
are deposited in different collections but they were not examined:
<collectionCode id="76C3C2C5FFA59E59FF5BD1A156F2B4C8" box="[138,210,948,972]" country="Netherlands" httpUri="http://grbio.org/cool/wkmc-b7hm" name="National Museum of Natural History, Naturalis" pageId="21" pageNumber="2027" type="Museum">NNM</collectionCode>
55468/1 (juvenile shell and operculum),
<specimenCode id="4074F27BFFA59E59FD7FD1A1551BB4C8" box="[686,827,948,972]" collectionCode="NNM" country="Netherlands" httpUri="http://grbio.org/cool/wkmc-b7hm" name="National Museum of Natural History, Naturalis" pageId="21" pageNumber="2027" type="Museum">NNM 9079</specimenCode>
/1 (juvenile in alcohol), Boeters 948/4 (damaged shell with operculum and three genitalia from
<specimenCount id="06D49189FFA59E59FCB2D1C155F8B4E8" box="[867,984,980,1004]" pageId="21" pageNumber="2027" type="male">two males</specimenCount>
and
<specimenCount id="06D49189FFA59E59FBC0D1C152B6B4E8" box="[1041,1174,980,1004]" pageId="21" pageNumber="2027" type="female">one female</specimenCount>
).
</materialsCitation>
</paragraph>
<paragraph id="106D5A00FFA59E59FF5BD60354E6B36A" blockId="21.[138,1185,1045,1134]" pageId="21" pageNumber="2027">
<emphasis id="22A68612FFA59E59FF5BD6035467B329" box="[138,583,1045,1069]" italics="true" pageId="21" pageNumber="2027">
<typeStatus id="CF69E4A2FFA59E59FF5BD60356E0B329" box="[138,192,1046,1069]" pageId="21" pageNumber="2027">Type</typeStatus>
material excluded from the species.
</emphasis>
Shell originally included by Boeters into
<taxonomicName id="D7D22183FFA59E59FBECD6035281B329" authorityName="Boeters" authorityYear="1980" box="[1085,1185,1046,1069]" class="Gastropoda" family="Hydrobiidae" genus="Horatia" kingdom="Animalia" order="Littorinimorpha" pageId="21" pageNumber="2027" phylum="Mollusca" rank="species" species="gatoa">
<emphasis id="22A68612FFA59E59FBECD6035281B329" box="[1085,1185,1046,1069]" italics="true" pageId="21" pageNumber="2027">H. gatoa</emphasis>
</taxonomicName>
(
<bibRefCitation id="744327F1FFA59E59FF43D6205712B349" author="Boeters HD" box="[146,306,1077,1101]" pageId="21" pageNumber="2027" pagination="61 - 64" refId="ref12950" refString="Boeters HD. 1980. Unbekannte westeuropaische Prosobranchia, 3. Basteria 44: 61 - 64." type="journal article" year="1980">Boeters 1980</bibRefCitation>
,
<figureCitation id="88E94685FFA59E59FEEED6205798B349" box="[319,440,1077,1101]" captionStart="Figure 6" captionStartId="19.[138,198,1425,1444]" captionTargetBox="[138,1186,133,1389]" captionTargetId="figure-42@19.[138,1186,133,1389]" captionTargetPageId="19" captionText="Figure 6. Operculum and radula of Iberhoratia aurorae from type locality. (A) Internal side of the operculum; (B) external side of the operculum; (C) radula; (D) central and lateral teeth; (E) central tooth; (F) lateral and internal marginal teeth. Scale bar: 200 Mm (A, B); 100 Mm (C)." figureDoi="http://doi.org/10.5281/zenodo.5232674" httpUri="https://zenodo.org/record/5232674/files/figure.png" pageId="21" pageNumber="2027">Figure 65</figureCitation>
<figureCitation id="88E94685FFA59E59FE69D6205476B349" box="[440,598,1077,1101]" captionStart="Figure 9" captionStartId="22.[125,185,1611,1630]" captionTargetBox="[127,1170,897,1573]" captionTargetId="figure-84@22.[125,1173,897,1575]" captionTargetPageId="22" captionText="Figure 9. Original illustrations of Iberhoratia gatoa (Boeters 1980, p 63). (A, B) Holotype (RMNH 55467), shell width 1.5 mm; (C, D) paratype (BOE 948), shell width 1.2 mm; (E) penis (paratype BOE 948); (F) anterior female genitalia (paratype BOE 948)." figureDoi="http://doi.org/10.5281/zenodo.5232680" httpUri="https://zenodo.org/record/5232680/files/figure.png" pageId="21" pageNumber="2027">Figure 9C, D</figureCitation>
in this paper), is re-identified as
<taxonomicName id="D7D22183FFA59E59FC0AD6205761B36A" authority="(Boeters, 1981)" baseAuthorityName="Boeters" baseAuthorityYear="1981" class="Gastropoda" family="Hydrobiidae" genus="Milesiana" kingdom="Animalia" order="Littorinimorpha" pageId="21" pageNumber="2027" phylum="Mollusca" rank="species" species="schuelei">
<emphasis id="22A68612FFA59E59FC0AD6205281B349" box="[987,1185,1077,1101]" italics="true" pageId="21" pageNumber="2027">Milesiana schuelei</emphasis>
(
<bibRefCitation id="744327F1FFA59E59FF43D6435718B36A" author="Boeters HD" box="[146,312,1110,1134]" pageId="21" pageNumber="2027" pagination="55 - 61" refId="ref12968" refString="Boeters HD. 1981. Unbekannte westeuropaische Prosobranchia, 2. Archiv fur Molluskenkunde (1980) 111 (1 / 3): 55 - 61." type="journal article" year="1981">Boeters, 1981</bibRefCitation>
)
</taxonomicName>
(in
<bibRefCitation id="744327F1FFA59E59FEA5D6435499B36A" author="Arconada B &amp; Ramos MA" box="[372,697,1110,1134]" pageId="21" pageNumber="2027" pagination="77 - 132" refId="ref12730" refString="Arconada B, Ramos MA. 2006. Revision of the genus Islamia Radoman, 1973 (Gastropoda, Prosobranchia, Hydrobiidae) on the Iberian Peninsula and description of two new genera and three new species. Malacologia 48 (1 / 2): 77 - 132." type="journal article" year="2006">Arconada and Ramos 2006</bibRefCitation>
).
</paragraph>
<paragraph id="106D5A00FFA59E59FF5BD6B35723B3BA" blockId="21.[138,259,1190,1214]" box="[138,259,1190,1214]" pageId="21" pageNumber="2027">
<emphasis id="22A68612FFA59E59FF5BD6B35723B3BA" box="[138,259,1190,1214]" italics="true" pageId="21" pageNumber="2027">Description</emphasis>
</paragraph>
<paragraph id="106D5A00FFA59E59FF5BD6C25480B254" blockId="21.[138,1185,1239,1523]" pageId="21" pageNumber="2027">
Original description in
<bibRefCitation id="744327F1FFA59E59FE71D6C25472B3EB" author="Boeters HD" box="[416,594,1239,1263]" pageId="21" pageNumber="2027" pagination="61 - 64" refId="ref12950" refString="Boeters HD. 1980. Unbekannte westeuropaische Prosobranchia, 3. Basteria 44: 61 - 64." type="journal article" year="1980">Boeters (1980)</bibRefCitation>
and redescription (
<bibRefCitation id="744327F1FFA59E59FCEDD6C255FCB3EB" author="Boeters HD" box="[828,988,1239,1263]" pageId="21" pageNumber="2027" pagination="181 - 261" refId="ref12996" refString="Boeters HD. 1988. Westeuropaische Moitessieriidae und Hydrobiidae in Spanien und Portugal (Gastropoda: Prosobranchia). Archiv fur Molluskenkunde 118: 181 - 261." type="journal article" year="1988">Boeters 1988</bibRefCitation>
) (
<figureCitation id="88E94685FFA59E59FC29D6C252BDB3EB" box="[1016,1181,1239,1263]" captionStart="Figure 8" captionStartId="22.[259,319,534,553]" captionTargetBox="[125,1173,133,498]" captionTargetId="figure-126@22.[125,1173,133,498]" captionTargetPageId="22" captionText="Figure 8. Holotype of Iberhoratia gatoa (RMNH 55467). Scale bar: 500 Mm (A, B)." figureDoi="http://doi.org/10.5281/zenodo.5232678" httpUri="https://zenodo.org/record/5232678/files/figure.png" pageId="21" pageNumber="2027">Figures 8AC</figureCitation>
,
<figureCitation id="88E94685FFA59E59FF5BD6ED5733B214" box="[138,275,1272,1296]" captionStart="Figure 9" captionStartId="22.[125,185,1611,1630]" captionTargetBox="[127,1170,897,1573]" captionTargetId="figure-84@22.[125,1173,897,1575]" captionTargetPageId="22" captionText="Figure 9. Original illustrations of Iberhoratia gatoa (Boeters 1980, p 63). (A, B) Holotype (RMNH 55467), shell width 1.5 mm; (C, D) paratype (BOE 948), shell width 1.2 mm; (E) penis (paratype BOE 948); (F) anterior female genitalia (paratype BOE 948)." figureDoi="http://doi.org/10.5281/zenodo.5232680" httpUri="https://zenodo.org/record/5232680/files/figure.png" pageId="21" pageNumber="2027">9A, B, E, F</figureCitation>
; Table
<collectionCode id="76C3C2C5FFA59E59FEB8D6ED5780B214" box="[361,416,1272,1296]" pageId="21" pageNumber="2027">VIII</collectionCode>
). Anatomical description was based on
<specimenCount id="06D49189FFA59E59FCBAD6ED55C0B214" box="[875,992,1272,1296]" pageId="21" pageNumber="2027" type="male">two males</specimenCount>
and
<specimenCount id="06D49189FFA59E59FBCAD6ED52BDB214" box="[1051,1181,1272,1296]" pageId="21" pageNumber="2027" type="female">one female</specimenCount>
. The following description is based on the original one excluding the shell re-identified as
<taxonomicName id="D7D22183FFA59E59FF5BD72C5726B254" baseAuthorityName="Boeters" baseAuthorityYear="1981" box="[138,262,1336,1360]" class="Gastropoda" family="Hydrobiidae" genus="Milesiana" kingdom="Animalia" order="Littorinimorpha" pageId="21" pageNumber="2027" phylum="Mollusca" rank="species" species="schuelei">
<emphasis id="22A68612FFA59E59FF5BD72C5726B254" box="[138,262,1336,1360]" italics="true" pageId="21" pageNumber="2027">M. schuelei</emphasis>
</taxonomicName>
, and on the study of the
<typeStatus id="CF69E4A2FFA59E59FDE2D72D54BBB254" box="[563,667,1336,1360]" pageId="21" pageNumber="2027" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="106D5A00FFA59E59FF74D74C5281B2F7" blockId="21.[138,1185,1239,1523]" pageId="21" pageNumber="2027">Shell shape valvatiform, with 3.5 whorls. The aperture does not ascend or descend near the last whorl. Shell aperture is rounded. Operculum pale yellowish, without any outgrowth. Rectum Z-shaped. Penis with a small lobe located in a medial position. Female genitalia has a well-developed bursa copulatrix, an elongated distal seminal receptacle, and a proximal receptacle lying tightly over a widened part of the renal oviduct.</paragraph>
<paragraph id="106D5A00FFA59E59FF5BD43956C9B140" blockId="21.[138,233,1580,1604]" box="[138,233,1580,1604]" pageId="21" pageNumber="2027">
<emphasis id="22A68612FFA59E59FF5BD43956C9B140" box="[138,233,1580,1604]" italics="true" pageId="21" pageNumber="2027">Remarks</emphasis>
</paragraph>
<paragraph id="106D5A00FFA59E5AFF5BD4485492B5A9" blockId="21.[138,1185,1629,1685]" lastBlockId="22.[125,1173,629,816]" lastPageId="22" lastPageNumber="2028" pageId="21" pageNumber="2027">
Original illustrations of female genitalia of
<taxonomicName id="D7D22183FFA59E59FD51D44855BAB171" authority="(Boeters 1980)" baseAuthorityName="Boeters" baseAuthorityYear="1980" box="[640,922,1629,1653]" class="Gastropoda" family="Hydrobiidae" genus="Horatia" kingdom="Animalia" order="Littorinimorpha" pageId="21" pageNumber="2027" phylum="Mollusca" rank="species" species="gatoa">
<emphasis id="22A68612FFA59E59FD51D44854C2B170" box="[640,738,1629,1653]" italics="true" pageId="21" pageNumber="2027">Ib. gatoa</emphasis>
(
<bibRefCitation id="744327F1FFA59E59FD22D44855B3B171" author="Boeters HD" box="[755,915,1629,1653]" pageId="21" pageNumber="2027" pagination="61 - 64" refId="ref12950" refString="Boeters HD. 1980. Unbekannte westeuropaische Prosobranchia, 3. Basteria 44: 61 - 64." type="journal article" year="1980">Boeters 1980</bibRefCitation>
)
</taxonomicName>
show a widened renal oviduct, which may have a seminal receptacle bent and adhered to the renal oviduct as in
<taxonomicName id="D7D22183FFA69E5AFFACD06056DCB589" authorityName="Arconada &amp; Delicado &amp; Ramos" authorityYear="2007" box="[125,252,629,653]" class="Gastropoda" family="Hydrobiidae" genus="Iberhoratia" kingdom="Animalia" order="Littorinimorpha" pageId="22" pageNumber="2028" phylum="Mollusca" rank="species" species="morenoi">
<emphasis id="22A68612FFA69E5AFFACD06056DCB589" box="[125,252,629,653]" italics="true" pageId="22" pageNumber="2028">Ib. morenoi</emphasis>
</taxonomicName>
and
<taxonomicName id="D7D22183FFA69E5AFEECD0605799B589" authorityName="Arconada &amp; Delicado &amp; Ramos" authorityYear="2007" box="[317,441,629,653]" class="Gastropoda" family="Hydrobiidae" genus="Iberhoratia" kingdom="Animalia" order="Littorinimorpha" pageId="22" pageNumber="2028" phylum="Mollusca" rank="species" species="aurorae">
<emphasis id="22A68612FFA69E5AFEECD0605799B589" box="[317,441,629,653]" italics="true" pageId="22" pageNumber="2028">Ib. aurorae</emphasis>
</taxonomicName>
, and a distal seminal receptacle (
<figureCitation id="88E94685FFA69E5AFC9DD06055E0B589" box="[844,960,629,653]" captionStart="Figure 9" captionStartId="22.[125,185,1611,1630]" captionTargetBox="[127,1170,897,1573]" captionTargetId="figure-84@22.[125,1173,897,1575]" captionTargetPageId="22" captionText="Figure 9. Original illustrations of Iberhoratia gatoa (Boeters 1980, p 63). (A, B) Holotype (RMNH 55467), shell width 1.5 mm; (C, D) paratype (BOE 948), shell width 1.2 mm; (E) penis (paratype BOE 948); (F) anterior female genitalia (paratype BOE 948)." figureDoi="http://doi.org/10.5281/zenodo.5232680" httpUri="https://zenodo.org/record/5232680/files/figure.png" pageId="22" pageNumber="2028">Figure 9F</figureCitation>
) as is the normal female genital disposition for
<taxonomicName id="D7D22183FFA69E5AFE08D0805492B5A9" box="[473,690,661,685]" class="Gastropoda" family="Hydrobiidae" genus="Iberhoratia" kingdom="Animalia" order="Littorinimorpha" pageId="22" pageNumber="2028" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="22A68612FFA69E5AFE08D0805470B5A9" box="[473,592,661,685]" italics="true" pageId="22" pageNumber="2028">Iberhoratia</emphasis>
species.
</taxonomicName>
</paragraph>
<caption id="44AD0A88FFA69E5AFED2D003522FB52D" ID-DOI="http://doi.org/10.5281/zenodo.5232678" ID-Zenodo-Dep="5232678" box="[259,1039,532,553]" httpUri="https://zenodo.org/record/5232678/files/figure.png" pageId="22" pageNumber="2028" startId="22.[259,319,534,553]" targetBox="[125,1173,133,498]" targetPageId="22">
<paragraph id="106D5A00FFA69E5AFED2D003522FB52D" blockId="22.[259,1039,532,553]" box="[259,1039,532,553]" pageId="22" pageNumber="2028">
Figure 8. Holotype of
<taxonomicName id="D7D22183FFA69E5AFE08D003544FB52D" authorityName="Arconada &amp; Delicado &amp; Ramos" authorityYear="2007" baseAuthorityName="Boeters" baseAuthorityYear="1980" box="[473,623,534,553]" class="Gastropoda" family="Hydrobiidae" genus="Iberhoratia" kingdom="Animalia" order="Littorinimorpha" pageId="22" pageNumber="2028" phylum="Mollusca" rank="species" species="gatoa">
<emphasis id="22A68612FFA69E5AFE08D003544FB52D" box="[473,623,534,553]" italics="true" pageId="22" pageNumber="2028">Iberhoratia gatoa</emphasis>
</taxonomicName>
(RMNH 55467). Scale bar: 500 Mm (A, B).
</paragraph>
</caption>
<paragraph id="106D5A00FFA69E5BFF49D0A35239B191" blockId="22.[125,1173,629,816]" lastBlockId="23.[138,1185,1245,1685]" lastPageId="23" lastPageNumber="2029" pageId="22" pageNumber="2028">
According to the measurements given by
<bibRefCitation id="744327F1FFA69E5AFDACD0A3550CB5CA" author="Boeters HD" box="[637,812,694,718]" pageId="22" pageNumber="2028" pagination="61 - 64" refId="ref12950" refString="Boeters HD. 1980. Unbekannte westeuropaische Prosobranchia, 3. Basteria 44: 61 - 64." type="journal article" year="1980">Boeters (1980)</bibRefCitation>
in the original description, the shell length varies from
<quantity id="D72AF7E5FFA69E5AFE48D0C25460B5EB" box="[409,576,727,751]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.5" metricValueMin="0.8" pageId="22" pageNumber="2028" unit="mm" value="1.15" valueMax="1.5" valueMin="0.8">0.8 to 1.5 mm</quantity>
and shell width from
<quantity id="D72AF7E5FFA69E5AFC98D0C255D4B5EB" box="[841,1012,727,751]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="22" pageNumber="2028" unit="mm" value="1.4" valueMax="1.5" valueMin="1.3">1.3 to 1.5 mm</quantity>
. In the figure caption the
<typeStatus id="CF69E4A2FFA69E5AFEDED0E25753B40B" box="[271,371,759,783]" pageId="22" pageNumber="2028" type="holotype">holotype</typeStatus>
width is
<quantity id="D72AF7E5FFA69E5AFE38D0E25461B40B" box="[489,577,759,783]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="22" pageNumber="2028" unit="mm" value="1.5">1.5 mm</quantity>
and length
<quantity id="D72AF7E5FFA69E5AFD06D0E25512B40B" box="[727,818,759,783]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="22" pageNumber="2028" unit="mm" value="1.5">1.5 mm</quantity>
. We cannot determine if this variability in shell values indicates that Boeters included in the species description measurements of the other
<specimenCount id="06D49189FFA79E5BFE18D6C854A3B3F1" box="[457,643,1245,1269]" pageId="23" pageNumber="2029" type="generic">three specimens</specimenCount>
for which anatomy was studied. Original shell illustrations from the
<typeStatus id="CF69E4A2FFA79E5BFE53D6E857C6B211" box="[386,486,1277,1301]" pageId="23" pageNumber="2029" type="holotype">holotype</typeStatus>
(
<figureCitation id="88E94685FFA79E5BFE24D6E854BAB211" box="[501,666,1277,1301]" captionStart="Figure 8" captionStartId="22.[259,319,534,553]" captionTargetBox="[125,1173,133,498]" captionTargetId="figure-126@22.[125,1173,133,498]" captionTargetPageId="22" captionText="Figure 8. Holotype of Iberhoratia gatoa (RMNH 55467). Scale bar: 500 Mm (A, B)." figureDoi="http://doi.org/10.5281/zenodo.5232678" httpUri="https://zenodo.org/record/5232678/files/figure.png" pageId="23" pageNumber="2029">Figures 8AC</figureCitation>
,
<figureCitation id="88E94685FFA79E5BFD74D6E854CAB211" box="[677,746,1277,1301]" captionStart="Figure 9" captionStartId="22.[125,185,1611,1630]" captionTargetBox="[127,1170,897,1573]" captionTargetId="figure-84@22.[125,1173,897,1575]" captionTargetPageId="22" captionText="Figure 9. Original illustrations of Iberhoratia gatoa (Boeters 1980, p 63). (A, B) Holotype (RMNH 55467), shell width 1.5 mm; (C, D) paratype (BOE 948), shell width 1.2 mm; (E) penis (paratype BOE 948); (F) anterior female genitalia (paratype BOE 948)." figureDoi="http://doi.org/10.5281/zenodo.5232680" httpUri="https://zenodo.org/record/5232680/files/figure.png" pageId="23" pageNumber="2029">9A, B</figureCitation>
) and
<typeStatus id="CF69E4A2FFA79E5BFCFDD6E855AFB211" box="[812,911,1277,1301]" pageId="23" pageNumber="2029" type="paratype">paratype</typeStatus>
of
<taxonomicName id="D7D22183FFA79E5BFC65D6E85234B210" authorityName="Arconada &amp; Delicado &amp; Ramos" authorityYear="2007" baseAuthorityName="Boeters" baseAuthorityYear="1980" box="[948,1044,1277,1300]" class="Gastropoda" family="Hydrobiidae" genus="Iberhoratia" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="2029" phylum="Mollusca" rank="species" species="gatoa">
<emphasis id="22A68612FFA79E5BFC65D6E85234B210" box="[948,1044,1277,1300]" italics="true" pageId="23" pageNumber="2029">H. gatoa</emphasis>
</taxonomicName>
(
<figureCitation id="88E94685FFA79E5BFBF2D6E85682B231" captionStart="Figure 9" captionStartId="22.[125,185,1611,1630]" captionTargetBox="[127,1170,897,1573]" captionTargetId="figure-84@22.[125,1173,897,1575]" captionTargetPageId="22" captionText="Figure 9. Original illustrations of Iberhoratia gatoa (Boeters 1980, p 63). (A, B) Holotype (RMNH 55467), shell width 1.5 mm; (C, D) paratype (BOE 948), shell width 1.2 mm; (E) penis (paratype BOE 948); (F) anterior female genitalia (paratype BOE 948)." figureDoi="http://doi.org/10.5281/zenodo.5232680" httpUri="https://zenodo.org/record/5232680/files/figure.png" pageId="23" pageNumber="2029">Figure 9C, D</figureCitation>
) correspond to
<specimenCount id="06D49189FFA79E5BFE8DD708542AB231" box="[348,522,1309,1333]" pageId="23" pageNumber="2029" type="generic">two specimens</specimenCount>
, which may not be conspecific. The
<typeStatus id="CF69E4A2FFA79E5BFC64D7085238B231" box="[949,1048,1309,1333]" pageId="23" pageNumber="2029" type="paratype">paratype</typeStatus>
of
<taxonomicName id="D7D22183FFA79E5BFBEED7085281B230" authorityName="Arconada &amp; Delicado &amp; Ramos" authorityYear="2007" baseAuthorityName="Boeters" baseAuthorityYear="1980" box="[1087,1185,1309,1332]" class="Gastropoda" family="Hydrobiidae" genus="Iberhoratia" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="2029" phylum="Mollusca" rank="species" species="gatoa">
<emphasis id="22A68612FFA79E5BFBEED7085281B230" box="[1087,1185,1309,1332]" italics="true" pageId="23" pageNumber="2029">H. gatoa</emphasis>
</taxonomicName>
(
<collectionCode id="76C3C2C5FFA79E5BFF43D72856EEB251" box="[146,206,1341,1365]" pageId="23" pageNumber="2029">BOE</collectionCode>
948) has a much flatter shell relative to the
<typeStatus id="CF69E4A2FFA79E5BFD20D7285579B251" box="[753,857,1341,1365]" pageId="23" pageNumber="2029" type="holotype">holotype</typeStatus>
, probably corresponding to
<taxonomicName id="D7D22183FFA79E5BFF5BD748542FB271" authority="(Boeters, 1981)" baseAuthorityName="Boeters" baseAuthorityYear="1981" box="[138,527,1373,1397]" class="Gastropoda" family="Hydrobiidae" genus="Milesiana" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="2029" phylum="Mollusca" rank="species" species="schuelei">
<emphasis id="22A68612FFA79E5BFF5BD7485770B271" box="[138,336,1373,1397]" italics="true" pageId="23" pageNumber="2029">Milesiana schuelei</emphasis>
(
<bibRefCitation id="744327F1FFA79E5BFEB1D7485426B271" author="Boeters HD" box="[352,518,1373,1397]" pageId="23" pageNumber="2029" pagination="55 - 61" refId="ref12968" refString="Boeters HD. 1981. Unbekannte westeuropaische Prosobranchia, 2. Archiv fur Molluskenkunde (1980) 111 (1 / 3): 55 - 61." type="journal article" year="1981">Boeters, 1981</bibRefCitation>
)
</taxonomicName>
(in
<bibRefCitation id="744327F1FFA79E5BFD90D74855A5B271" author="Arconada B &amp; Ramos MA" box="[577,901,1373,1397]" pageId="23" pageNumber="2029" pagination="77 - 132" refId="ref12730" refString="Arconada B, Ramos MA. 2006. Revision of the genus Islamia Radoman, 1973 (Gastropoda, Prosobranchia, Hydrobiidae) on the Iberian Peninsula and description of two new genera and three new species. Malacologia 48 (1 / 2): 77 - 132." type="journal article" year="2006">Arconada and Ramos 2006</bibRefCitation>
), a species which is very abundant in the type locality of
<taxonomicName id="D7D22183FFA79E5BFDD6D768544BB290" authorityName="Arconada &amp; Delicado &amp; Ramos" authorityYear="2007" baseAuthorityName="Boeters" baseAuthorityYear="1980" box="[519,619,1405,1429]" class="Gastropoda" family="Hydrobiidae" genus="Iberhoratia" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="2029" phylum="Mollusca" rank="species" species="gatoa">
<emphasis id="22A68612FFA79E5BFDD6D768544BB290" box="[519,619,1405,1429]" italics="true" pageId="23" pageNumber="2029">Ib. gatoa</emphasis>
</taxonomicName>
. In fact, shell measurements from this
<typeStatus id="CF69E4A2FFA79E5BFBECD7685280B291" box="[1085,1184,1405,1429]" pageId="23" pageNumber="2029" type="paratype">paratype</typeStatus>
(shell width
<quantity id="D72AF7E5FFA79E5BFECAD7885756B2B1" box="[283,374,1437,1461]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="23" pageNumber="2029" unit="mm" value="1.2">1.2 mm</quantity>
; length
<quantity id="D72AF7E5FFA79E5BFE02D788540DB2B1" box="[467,557,1437,1461]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="23" pageNumber="2029" unit="mm" value="0.8">0.8 mm</quantity>
) correspond to that described for
<taxonomicName id="D7D22183FFA79E5BFC6CD7885219B2B1" baseAuthorityName="Boeters" baseAuthorityYear="1981" box="[957,1081,1437,1461]" class="Gastropoda" family="Hydrobiidae" genus="Milesiana" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="2029" phylum="Mollusca" rank="species" species="schuelei">
<emphasis id="22A68612FFA79E5BFC6CD7885219B2B1" box="[957,1081,1437,1461]" italics="true" pageId="23" pageNumber="2029">M. schuelei</emphasis>
</taxonomicName>
. We can either consider that this
<typeStatus id="CF69E4A2FFA79E5BFE74D7A85428B2D1" box="[421,520,1469,1493]" pageId="23" pageNumber="2029" type="paratype">paratype</typeStatus>
is actually a juvenile of
<taxonomicName id="D7D22183FFA79E5BFCF1D7A855A1B2D0" authorityName="Arconada &amp; Delicado &amp; Ramos" authorityYear="2007" baseAuthorityName="Boeters" baseAuthorityYear="1980" box="[800,897,1469,1493]" class="Gastropoda" family="Hydrobiidae" genus="Iberhoratia" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="2029" phylum="Mollusca" rank="species" species="gatoa">
<emphasis id="22A68612FFA79E5BFCF1D7A855A1B2D0" box="[800,897,1469,1493]" italics="true" pageId="23" pageNumber="2029">Ib. gatoa</emphasis>
</taxonomicName>
, or that it belongs to
<taxonomicName id="D7D22183FFA79E5BFBAFD7A856FBB2F1" baseAuthorityName="Boeters" baseAuthorityYear="1981" class="Gastropoda" family="Hydrobiidae" genus="Milesiana" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="2029" phylum="Mollusca" rank="species" species="schuelei">
<emphasis id="22A68612FFA79E5BFBAFD7A856FBB2F1" italics="true" pageId="23" pageNumber="2029">M. schuelei</emphasis>
</taxonomicName>
. The second hypothesis seems to be more feasible because the growth rate of this shell is greater than that of the
<typeStatus id="CF69E4A2FFA79E5BFDC6D7E8545FB111" box="[535,639,1533,1557]" pageId="23" pageNumber="2029" type="holotype">holotype</typeStatus>
. For this reason we have proposed that this specimen might be excluded from the type material (
<bibRefCitation id="744327F1FFA79E5BFCE4D40852B5B131" author="Arconada B &amp; Ramos MA" box="[821,1173,1565,1589]" pageId="23" pageNumber="2029" pagination="77 - 132" refId="ref12730" refString="Arconada B, Ramos MA. 2006. Revision of the genus Islamia Radoman, 1973 (Gastropoda, Prosobranchia, Hydrobiidae) on the Iberian Peninsula and description of two new genera and three new species. Malacologia 48 (1 / 2): 77 - 132." type="journal article" year="2006">Arconada and Ramos 2006</bibRefCitation>
). Nevertheless, the genital illustrations from two other
<typeStatus id="CF69E4A2FFA79E5BFD25D4285542B151" box="[756,866,1597,1621]" pageId="23" pageNumber="2029" type="paratype">paratypes</typeStatus>
(
<specimenCount id="06D49189FFA79E5BFCA3D42855FDB151" box="[882,989,1597,1621]" pageId="23" pageNumber="2029" type="male">one male</specimenCount>
and
<specimenCount id="06D49189FFA79E5BFBC9D42852B9B151" box="[1048,1177,1597,1621]" pageId="23" pageNumber="2029" type="female">one female</specimenCount>
) with the same collection number (
<collectionCode id="76C3C2C5FFA79E5BFDF0D448547DB171" box="[545,605,1629,1653]" pageId="23" pageNumber="2029">BOE</collectionCode>
948) illustrated by
<bibRefCitation id="744327F1FFA79E5BFC9AD44855D3B171" author="Boeters HD" box="[843,1011,1629,1653]" pageId="23" pageNumber="2029" pagination="61 - 64" refId="ref12950" refString="Boeters HD. 1980. Unbekannte westeuropaische Prosobranchia, 3. Basteria 44: 61 - 64." type="journal article" year="1980">Boeters (1980</bibRefCitation>
,
<figureCitation id="88E94685FFA79E5BFBD2D44852B8B171" box="[1027,1176,1629,1653]" captionStart="Figure 9" captionStartId="22.[125,185,1611,1630]" captionTargetBox="[127,1170,897,1573]" captionTargetId="figure-84@22.[125,1173,897,1575]" captionTargetPageId="22" captionText="Figure 9. Original illustrations of Iberhoratia gatoa (Boeters 1980, p 63). (A, B) Holotype (RMNH 55467), shell width 1.5 mm; (C, D) paratype (BOE 948), shell width 1.2 mm; (E) penis (paratype BOE 948); (F) anterior female genitalia (paratype BOE 948)." figureDoi="http://doi.org/10.5281/zenodo.5232680" httpUri="https://zenodo.org/record/5232680/files/figure.png" pageId="23" pageNumber="2029">Figure 9E, F</figureCitation>
) actually correspond to the typical anatomical scheme of the genus
<emphasis id="22A68612FFA79E5BFC49D4685239B191" box="[920,1049,1661,1685]" italics="true" pageId="23" pageNumber="2029">
<taxonomicName id="D7D22183FFA79E5BFC49D4685235B191" authorityName="Arconada and Ramos" authorityYear="2007" box="[920,1045,1661,1685]" class="Gastropoda" family="Hydrobiidae" genus="Iberhoratia" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="2029" phylum="Mollusca" rank="genus">Iberhoratia</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>