treatments-xml/data/82/65/80/82658048FFE7FFADFF3FFBCC6285F864.xml

<document id="E310128457F468C683C4EB0BFEB8A5D7" ID-DOI="10.5281/zenodo.202485" ID-GBIF-Dataset="53aa7273-94c8-4bec-88bf-41af0631c7c5" ID-ISSN="1175-5326" ID-Zenodo-Dep="202485" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460236554809" checkinUser="plazi" docAuthor="Howlader, Mohammad Sajid Ali" docDate="2011" docId="82658048FFE7FFADFF3FFBCC6285F864" docLanguage="en" docName="zt02761p050.pdf" docOrigin="Zootaxa 2761" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Fejervarya asmati Howlader, 2011, sp. nov." docType="treatment" docVersion="9" lastPageNumber="48" masterDocId="7E5CF830FFE5FFAAFFA8FFD56118FA5B" masterDocTitle="A new species of Fejervarya (Anura: Dicroglossidae) from Bangladesh" masterLastPageNumber="50" masterPageNumber="41" pageNumber="43" updateTime="1698239698720" updateUser="plazi">
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<mods:title id="57888DACCE6B36D4541E635487987E05">A new species of Fejervarya (Anura: Dicroglossidae) from Bangladesh</mods:title>
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<mods:namePart id="9E601949C7B252353DCD24BF4310B6AD">Howlader, Mohammad Sajid Ali</mods:namePart>
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<mods:date id="24D36749785A7B408416B698B36F7A8E">2011</mods:date>
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<subSubSection id="42D662D5FFE7FFA8FF3FFBCC61E7FE0F" pageId="2" pageNumber="43" type="nomenclature">
<paragraph id="0A73315EFFE7FFA8FF3FFBCC60CFFE69" blockId="2.[151,471,1048,1108]" box="[151,471,1048,1075]" pageId="2" pageNumber="43">
<heading id="513B8632FFE7FFA8FF3FFBCC60CFFE69" bold="true" box="[151,471,1048,1075]" fontSize="11" level="1" pageId="2" pageNumber="43" reason="1">
<emphasis id="38B8ED4CFFE7FFA8FF3FFBCC60CFFE69" bold="true" box="[151,471,1048,1075]" pageId="2" pageNumber="43">
<taxonomicName id="CDCC4ADDFFE7FFA8FF3FFBCC606AFE68" ID-CoL="3DXDH" box="[151,370,1049,1075]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="2" pageNumber="43" phylum="Chordata" rank="species" species="asmati" status="sp. nov.">
<emphasis id="38B8ED4CFFE7FFA8FF3FFBCC606AFE68" bold="true" box="[151,370,1049,1075]" italics="true" pageId="2" pageNumber="43">Fejervarya asmati</emphasis>
</taxonomicName>
<taxonomicNameLabel id="238B5037FFE7FFA8FED1FBCD60CFFE69" box="[377,471,1048,1074]" pageId="2" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="0A73315EFFE7FFA8FF3FFBE961E7FE0F" blockId="2.[151,471,1048,1108]" box="[151,255,1084,1108]" pageId="2" pageNumber="43">
<figureCitation id="92F72DDBFFE7FFA8FF3FFBE961E7FE0F" box="[151,255,1084,1108]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="3.[151,250,1973,1996]" captionStartId-1="4.[151,250,997,1020]" captionTargetBox-0="[156,1430,179,1949]" captionTargetBox-1="[429,1155,483,973]" captionTargetId-0="figure@3.[156,1431,179,1949]" captionTargetId-1="figure@4.[428,1158,483,973]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 2. Photographs in life of Fejervarya asmati sp. nov. A: Calling male. B: Mating of adult individuals. C: Ventral view of male." captionText-1="FIGURE 3. Ventral view of hand (on the left) and foot (on the right) of Fejervarya asmati." httpUri-0="https://zenodo.org/record/202486/files/figure.png" httpUri-1="https://zenodo.org/record/202487/files/figure.png" pageId="2" pageNumber="43">Figs. 2–3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="42D662D5FFE7FFA8FF3FFB516351FF14" pageId="2" pageNumber="43" type="materials_examined">
<paragraph id="0A73315EFFE7FFA8FF3FFB5161CEFEBF" blockId="2.[151,1436,1156,1972]" pageId="2" pageNumber="43">
<emphasis id="38B8ED4CFFE7FFA8FF3FFB51601CFEC7" bold="true" box="[151,260,1156,1180]" pageId="2" pageNumber="43">
<typeStatus id="D5778FFCFFE7FFA8FF3FFB51601CFEC7" box="[151,260,1156,1180]" pageId="2" pageNumber="43" type="holotype">Holotype</typeStatus>
</emphasis>
.—Adult male, MZD/F-01, collected in front of Teachers Club and Guest House (
<geoCoordinate id="6FF85799FFE7FFA8FB36FB51640AFEC7" box="[1182,1298,1156,1180]" direction="north" orientation="latitude" pageId="2" pageNumber="43" precision="925" value="22.466667">22° 28´N</geoCoordinate>
, 
<geoCoordinate id="6FF85799FFE7FFA8FAB6FB516497FEC7" box="[1310,1423,1156,1180]" direction="east" orientation="longitude" pageId="2" pageNumber="43" precision="925" value="91.78333">91° 47´E</geoCoordinate>
), 
<collectingCountry id="72DB71CEFFE7FFA8FF3FFB72600BFEE4" box="[151,275,1191,1215]" name="Bangladesh" pageId="2" pageNumber="43">Chittagong</collectingCountry>
University campus, Hathazari, 
<collectingCountry id="72DB71CEFFE7FFA8FD20FB72621FFEE4" box="[648,775,1191,1215]" name="Bangladesh" pageId="2" pageNumber="43">Chittagong</collectingCountry>
, 
<collectingCountry id="72DB71CEFFE7FFA8FCBFFB726285FEE4" box="[791,925,1191,1215]" name="Bangladesh" pageId="2" pageNumber="43">Bangladesh</collectingCountry>
; collected by M. S. A. Howlader, 
<date id="7E72179EFFE7FFA8FA92FB7D61CBFEBF" pageId="2" pageNumber="43" value="2008-05-15">May 15, 2008</date>
.
</paragraph>
<paragraph id="0A73315EFFE7FFA8FF6FFB256351FF14" blockId="2.[151,1436,1156,1972]" pageId="2" pageNumber="43">
<emphasis id="38B8ED4CFFE7FFA8FF6FFB25606BFF53" bold="true" box="[199,371,1264,1288]" pageId="2" pageNumber="43">Paratopotypes</emphasis>
.—MZD/F-02 (adult female), MZD/F-03 (adult male), MZD/F-04 (adult male), MZD/F-05 (adult male), and MZD/F-06 (adult male) collected from the same locality as the 
<typeStatus id="D5778FFCFFE7FFA8FB9DFAC16583FF77" box="[1077,1179,1300,1324]" pageId="2" pageNumber="43" type="holotype">holotype</typeStatus>
; collected by M. S. A. Howlader &amp; J. I. Taher, 
<date id="7E72179EFFE7FFA8FE0EFAED635EFF14" box="[422,582,1335,1360]" pageId="2" pageNumber="43" value="2008-05-16">May 16, 2008</date>
.
</paragraph>
</subSubSection>
<subSubSection id="42D662D5FFE7FFA8FF6FFA8E65C5FC73" pageId="2" pageNumber="43" type="diagnosis">
<paragraph id="0A73315EFFE7FFA8FF6FFA8E65C5FC73" blockId="2.[151,1436,1156,1972]" pageId="2" pageNumber="43">
<emphasis id="38B8ED4CFFE7FFA8FF6FFA8E6022FF2F" bold="true" box="[199,314,1371,1396]" pageId="2" pageNumber="43">Diagnosis</emphasis>
.—A species assigned to the genus 
<taxonomicName id="CDCC4ADDFFE7FFA8FD64FA886250FF2F" box="[716,840,1373,1396]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="2" pageNumber="43" phylum="Chordata" rank="genus">
<emphasis id="38B8ED4CFFE7FFA8FD64FA886250FF2F" box="[716,840,1373,1396]" italics="true" pageId="2" pageNumber="43">Fejervarya</emphasis>
</taxonomicName>
by having “Fejervaryan” line on both sides of belly, absence of rictal gland at the mouth commissure, smooth skin with minute warts or folds, lips more or less barred, fingers free of webbing, toes not fully webbed. The new species is characterized by a male SVL of 
<quantity id="CD349CBBFFE7FFA8FB43FA716480FFE7" box="[1259,1432,1444,1468]" metricMagnitude="-2" metricUnit="m" metricValue="3.125" metricValueMax="3.34" metricValueMin="2.91" pageId="2" pageNumber="43" unit="mm" value="31.25" valueMax="33.4" valueMin="29.1">29.1– 33.4 mm</quantity>
; butterfly shaped vocal marking present in male; forearm length 70% of hand length; relative length of fingers, shortest to longest: 2 &lt;4 &lt;1&lt;3; nostril much closer to snout tip than eye, nostril–snout length 57% of distance from front of eyes to nostril; nostril–snout length 0.67% of internarial distance; MBE 18% of HL.
</paragraph>
</subSubSection>
<subSubSection id="42D662D5FFE7FFAFFF6FF9E66093FD89" lastPageId="5" lastPageNumber="46" pageId="2" pageNumber="43" type="description">
<paragraph id="0A73315EFFE7FFA8FF6FF9E663DBFD7F" blockId="2.[151,1436,1156,1972]" pageId="2" pageNumber="43">
<emphasis id="38B8ED4CFFE7FFA8FF6FF9E66382FC17" bold="true" box="[199,666,1587,1612]" pageId="2" pageNumber="43">
Description of 
<typeStatus id="D5778FFCFFE7FFA8FEDFF9E160C6FC17" box="[375,478,1588,1612]" pageId="2" pageNumber="43" type="holotype">holotype</typeStatus>
(adult male).—
</emphasis>
Small sized frog (SVL 30.0 mm). Head large, triangular, longer than wide, HW 93% of HL, HW 32% of SVL, HL 34% of SVL, MFE 60% of HL, MBE 18% of HL. Snout nearly rounded in lateral view, SL 40% of HL; canthus rostralis indistinct, loreal region concave. Nostrils much closer to snout tip than to eyes, NS 57% of EN; NS 5% of SVL, EN 9% of SVL; nostrils rounded and very small, NS 0.67% of IN, MN 85% HL. Eye large, EL 41% of HL, EL 14% of SVL; maximum width of upper eyelid greater than interorbital distance, IOD 83% of UEW, UEW 56% EL, UEW 8% SVL. Interorbital space convex, IOD 88% of IN. Tympanum round, TD 52% of EL, TEL 34% TD.
</paragraph>
<paragraph id="0A73315EFFE7FFA8FF6FF8E56244FDEF" blockId="2.[151,1436,1156,1972]" pageId="2" pageNumber="43">Arm moderately long, robust, FAL 70% HAL, FAL 17% of SVL, HAL 24% SVL. Fingers small, free of webbing, tips rounded. Relative length of fingers, shortest to longest: 2 &lt;4 &lt;1 &lt;3; tips of fingers bluntly rounded; fingers lacking dermal ridge. Subarticular tubercles prominent, rounded, single tubercle per digit; supernumerary tubercles absent; two capsule shaped, distinct palmar tubercles.</paragraph>
<caption id="5EB361D6FFE6FFA9FF3FF86061F3FDB1" httpUri="https://zenodo.org/record/202486/files/figure.png" pageId="3" pageNumber="44" targetBox="[156,1430,179,1949]" targetPageId="3">
<paragraph id="0A73315EFFE6FFA9FF3FF86061F3FDB1" blockId="3.[151,1436,1973,2026]" pageId="3" pageNumber="44">
<emphasis id="38B8ED4CFFE6FFA9FF3FF860600AFD91" bold="true" box="[151,274,1973,1996]" pageId="3" pageNumber="44">FIGURE 2.</emphasis>
Photographs in life of 
<taxonomicName id="CDCC4ADDFFE6FFA9FE53F86363AEFD90" box="[507,694,1974,1995]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="3" pageNumber="44" phylum="Chordata" rank="species" species="asmati" status="sp. nov.">
<emphasis id="38B8ED4CFFE6FFA9FE53F86363AEFD90" box="[507,694,1974,1995]" italics="true" pageId="3" pageNumber="44">Fejervarya asmati</emphasis>
</taxonomicName>
<taxonomicNameLabel id="238B5037FFE6FFA9FD14F8636212FD90" box="[700,778,1974,1995]" pageId="3" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
A: Calling male. B: Mating of adult individuals. C: Ventral view of male.
</paragraph>
</caption>
<paragraph id="0A73315EFFE1FFAEFF6FFF4261E9FBF7" blockId="4.[151,1436,151,428]" pageId="4" pageNumber="45">
<collectingCountry id="72DB71CEFFE1FFAEFF6FFF4261E7FAF4" box="[199,255,151,175]" name="India" pageId="4" pageNumber="45">Hind</collectingCountry>
limbs relatively long, TL 50% of SVL, THIGHL 87% of TL; FOL 50% SVL and equal to TL, FOL 70% of TFOL. Toes long, thin, tips rounded; webbing between toes weakly developed [1(1), 
<date id="7E72179EFFE1FFAEFB37FF6965AFFA8F" box="[1183,1207,188,212]" pageId="4" pageNumber="45">2i</date>
(1.5), 2e(0.5), 
<date id="7E72179EFFE1FFAEFAF6FF69646DFA8F" box="[1374,1397,188,212]" pageId="4" pageNumber="45">3i</date>
(2), 3e(1), 
<date id="7E72179EFFE1FFAEFF4AFF0A61E2FAAC" box="[226,250,223,247]" pageId="4" pageNumber="45">4i</date>
(2), 4e(2.5), 5(0.75)]. Relative lengths of toes, shortest to longest: 1 &lt;2 &lt;5 &lt;3 &lt;4; a weak, indistinct fringe of skin on outer side of toe 5. Inner metatarsal tubercle elongated, present at base of Toe 1; Outer metatarsal tubercle is oval, minute, distinct; subarticular tubercles well-developed, nearly oval. Dorsal surface smooth, tubercles present, arranging in row or reticulated pattern; tiny granules on upper eyelids, loreal, and cloacal region. Dorsal surface of forelimb, thigh and tarsus glandular. Throat, chest, abdomen and ventral part of thigh and tibia smooth.
</paragraph>
<caption id="5EB361D6FFE1FFAEFF3FFC30652BF9A0" box="[151,1075,997,1020]" httpUri="https://zenodo.org/record/202487/files/figure.png" pageId="4" pageNumber="45" targetBox="[429,1155,483,973]" targetPageId="4">
<paragraph id="0A73315EFFE1FFAEFF3FFC30652BF9A0" blockId="4.[151,1075,997,1020]" box="[151,1075,997,1020]" pageId="4" pageNumber="45">
<emphasis id="38B8ED4CFFE1FFAEFF3FFC30600AF9A1" bold="true" box="[151,274,997,1020]" pageId="4" pageNumber="45">FIGURE 3.</emphasis>
Ventral view of hand (on the left) and foot (on the right) of 
<taxonomicName id="CDCC4ADDFFE1FFAEFCDAFC336528F9A0" box="[882,1072,998,1019]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="4" pageNumber="45" phylum="Chordata" rank="species" species="asmati">Fejervarya asmati</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="0A73315EFFE1FFAEFF6FFBF2654DFE8B" blockId="4.[151,1437,1063,2025]" pageId="4" pageNumber="45">
<emphasis id="38B8ED4CFFE1FFAEFF6FFBF26086FE1B" bold="true" box="[199,414,1063,1088]" pageId="4" pageNumber="45">Coloration in life.</emphasis>
—Basic color varies from olive green to greenish brown, with a few dark irregular spots particularly fused into transverse bands; spots are formed by tubercle arrangement on dorsal sides; sides of body marbled. A narrow middorsal line usually present up to the posterior of eyes. Two red spot found on the middorsal line and another two red spot on each of the forelimbs. Toes webbing faintly marbled. Forelimbs with dark transverse bands; hind limbs dotted on thighs and strips on lower legs. Venter immaculate.
</paragraph>
<paragraph id="0A73315EFFE1FFAEFF6FFB0965E8FF43" blockId="4.[151,1437,1063,2025]" pageId="4" pageNumber="45">
<emphasis id="38B8ED4CFFE1FFAEFF6FFB096312FEAE" bold="true" box="[199,522,1244,1269]" pageId="4" pageNumber="45">Coloration in preservative.</emphasis>
—In preservative olive green surface become whitish brown. Middorsal line and forelimbs lost red spots. Color arrangement of the total body become fade to their original position.
</paragraph>
<paragraph id="0A73315EFFE1FFAEFF6FFAF063B8FF97" blockId="4.[151,1437,1063,2025]" pageId="4" pageNumber="45">
<emphasis id="38B8ED4CFFE1FFAEFF6FFAF060FEFF66" bold="true" box="[199,486,1316,1341]" pageId="4" pageNumber="45">Measurements (in mm).</emphasis>
—Male (
<typeStatus id="D5778FFCFFE1FFAEFDE3FAF163ACFF67" box="[587,692,1316,1340]" pageId="4" pageNumber="45" type="holotype">holotype</typeStatus>
): SVL 30.0; HL 10.3; HW 9.6; MN 8.7; SL 4.1; MFE 6.2; MBE 1.9; IN 2.3; IOD 2.0; EN 2.7; NS 1.5; EL 4.3; UEW 2.4; TD 2.2; TEL 0.8; HAL 7.2; FAL 5.1; THIGHL 13.1; TL 15.1; TFOL 22.3; FOL 15.7; IMTL 1.4. Female (paratopotype): SVL 33.4; HL 11.4; HW 10.5; MN 9.5; SL 4.6; MFE 6.8; MBE 2.2; IN 2.4; IOD 2.1; EN 2.8; NS 1.7; EL 4.7; UEW 2.6; TD 2.3; TEL 0.8; HAL 7.6; FAL 5.8; THIGHL 13.7; TL 16.1; FOL 16.7; IMTL 1.7.
</paragraph>
<paragraph id="0A73315EFFE1FFAEFF6FFA02656FFC4F" blockId="4.[151,1437,1063,2025]" pageId="4" pageNumber="45">
<emphasis id="38B8ED4CFFE1FFAEFF6FFA026023FFAB" bold="true" box="[199,315,1495,1520]" pageId="4" pageNumber="45">Variation.</emphasis>
—Morphometric variation is shown on the table 1. Two specimens lack mid-dorsal lines (males MZD/F-05 and MZD/F-06), female (MZD/F-02) has no vocal marking and no red spots.
</paragraph>
<paragraph id="0A73315EFFE1FFAEFF6FF9CA6484FCDB" blockId="4.[151,1437,1063,2025]" pageId="4" pageNumber="45">
<emphasis id="38B8ED4CFFE1FFAEFF6FF9CA60B6FC63" bold="true" box="[199,430,1567,1592]" pageId="4" pageNumber="45">Advertisement call.</emphasis>
—The advertisement call was recorded on 
<collectingCountry id="72DB71CEFFE1FFAEFC27F9F56513FC63" box="[911,1035,1568,1592]" name="Bangladesh" pageId="4" pageNumber="45">Chittagong</collectingCountry>
University campus at an air temperature of 24 ºC. The call consists of a very fast trill series of three note 
<typeStatus id="D5778FFCFFE1FFAEFC1BF99062EAFC06" box="[947,1010,1605,1629]" pageId="4" pageNumber="45">types</typeStatus>
: a group of similar pulsed trains (
<typeStatus id="D5778FFCFFE1FFAEFAC2F9916484FC07" box="[1386,1436,1604,1628]" pageId="4" pageNumber="45">type</typeStatus>
1) interspersed by short single notes (both 
<typeStatus id="D5778FFCFFE1FFAEFDC7F9BC6387FCDA" box="[623,671,1641,1665]" pageId="4" pageNumber="45">type</typeStatus>
2 and 3, or just 
<typeStatus id="D5778FFCFFE1FFAEFCF8F9BC6298FCDA" box="[848,896,1641,1665]" pageId="4" pageNumber="45">type</typeStatus>
3 notes) with 775 ms (N=1) of duration (
<figureCitation id="92F72DDBFFE1FFAEFAE2F9BD6497FCDB" box="[1354,1423,1640,1664]" captionStart="FIGURE 4" captionStartId="6.[151,250,821,844]" captionTargetBox="[351,1238,200,788]" captionTargetId="figure@6.[339,1243,187,800]" captionTargetPageId="6" captionText="FIGURE 4. Sonogram (top) and oscillogram (bottom) of the advertisement call of Fejervarya asmati." httpUri="https://zenodo.org/record/202488/files/figure.png" pageId="4" pageNumber="45">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="0A73315EFFE1FFAEFF6FF95963C0FD6F" blockId="4.[151,1437,1063,2025]" pageId="4" pageNumber="45">
Each pulsed note (train) of 
<typeStatus id="D5778FFCFFE1FFAEFE51F9586331FCFE" box="[505,553,1677,1701]" pageId="4" pageNumber="45">type</typeStatus>
1 has a duration of 92-134 ms (110±15.5, N=5); interval between two trains is about 43–71 ms (56.25±12.63, N=4). The number of pulses per train ranges from 15–18 (16.8±1.3, N=5). Pulse repetition rate within a train is 152.72/s. These pulses have a harmonious modulation with 5 recognizable bands: fundamental frequency around 0.2–1.0 kHz, a second band at 1.4–2.2 kHz, a formant band at 2.6–3.4 kHz; followed by further bands at 3.8–4.6 and 5.0–5.8 kHz.
</paragraph>
<paragraph id="0A73315EFFE1FFAEFF6FF89463C1FDFB" blockId="4.[151,1437,1063,2025]" pageId="4" pageNumber="45">
Between two trains of 
<typeStatus id="D5778FFCFFE1FFAEFE7CF894631CFD02" box="[468,516,1857,1881]" pageId="4" pageNumber="45">type</typeStatus>
1 can be emitted a single note of 
<typeStatus id="D5778FFCFFE1FFAEFC09F89462C9FD02" box="[929,977,1857,1881]" pageId="4" pageNumber="45">type</typeStatus>
2 (N=1) or 0–3 single notes of 
<typeStatus id="D5778FFCFFE1FFAEFAFBF894649BFD02" box="[1363,1411,1857,1881]" pageId="4" pageNumber="45">type</typeStatus>
3 (1.0±1.41, N=4). Occasionally, a single note of 
<typeStatus id="D5778FFCFFE1FFAEFD1FF8B063FFFD26" box="[695,743,1893,1917]" pageId="4" pageNumber="45">type</typeStatus>
3 can be also emitted between two pulses of 
<typeStatus id="D5778FFCFFE1FFAEFB58F8B06438FD26" box="[1264,1312,1893,1917]" pageId="4" pageNumber="45">type</typeStatus>
1 within a train with a frequency of occurrence of 20% (N=5).
</paragraph>
<paragraph id="0A73315EFFE1FFAEFF6FF87962DBFDB3" blockId="4.[151,1437,1063,2025]" pageId="4" pageNumber="45">
Single notes of 
<typeStatus id="D5778FFCFFE1FFAEFED0F87860B0FD9E" box="[376,424,1965,1989]" pageId="4" pageNumber="45">type</typeStatus>
2 have a duration of 17 ms (N=1) and are emitted 23 ms (N=1) after the last note of first 
<typeStatus id="D5778FFCFFE1FFAEFF3FF80461DFFDB2" box="[151,199,2001,2025]" pageId="4" pageNumber="45">type</typeStatus>
1 train, and 21ms (N=1) far from the following single note (
<typeStatus id="D5778FFCFFE1FFAEFCC5F8056287FDB3" box="[877,927,2000,2024]" pageId="4" pageNumber="45">type</typeStatus>
3).
</paragraph>
<caption id="5EB361D6FFE0FFAFFF3FFF4262A8FA89" ID-Table-UUID="5EB361D6FFE0FFAFFF3FFF4262A8FA89" httpUri="http://table.plazi.org/id/5EB361D6FFE0FFAFFF3FFF4262A8FA89" pageId="5" pageNumber="46" targetBox="[156,1432,244,1582]" targetIsTable="true" targetPageId="5">
<paragraph id="0A73315EFFE0FFAFFF3FFF4262A8FA89" blockId="5.[151,1435,151,210]" pageId="5" pageNumber="46">
<emphasis id="38B8ED4CFFE0FFAFFF3FFF426013FAF4" bold="true" box="[151,267,151,175]" pageId="5" pageNumber="46">TABLE 1.</emphasis>
Summary of qualitative diagnostic characters in 
<taxonomicName id="CDCC4ADDFFE0FFAFFC8CFF4D62F2FAF4" box="[804,1002,152,175]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="5" pageNumber="46" phylum="Chordata" rank="species" species="asmati" status="sp. nov.">
<emphasis id="38B8ED4CFFE0FFAFFC8CFF4D62F2FAF4" box="[804,1002,152,175]" italics="true" pageId="5" pageNumber="46">Fejervarya asmati</emphasis>
</taxonomicName>
<taxonomicNameLabel id="238B5037FFE0FFAFFC5BFF4D655FFAF4" box="[1011,1095,152,175]" pageId="5" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
and specimens of other congeners. Morphological ratios given as mean ± standard deviation over range.
</paragraph>
</caption>
<paragraph id="0A73315EFFE0FFAFFEC8FF206491FFF9" pageId="5" pageNumber="46">
<table id="78CCC3FEFFE00055FF34FF216480FC75" box="[156,1432,244,1582]" gridcols="6" gridrows="16" pageId="5" pageNumber="46">
<tr id="B4FC331CFFE00055FF34FF216480FB53" box="[156,1432,244,264]" gridrow="0" pageId="5" pageNumber="46" rowspan-0="1">
<th id="F72D5A60FFE00055FEB4FF2160F3FB53" box="[284,491,244,264]" gridcol="1" gridrow="0" pageId="5" pageNumber="46">
<taxonomicName id="CDCC4ADDFFE0FFAFFEC8FF2060AAFB5C" box="[352,434,245,263]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="5" pageNumber="46" phylum="Chordata" rank="species" species="asmati">
<emphasis id="38B8ED4CFFE0FFAFFEC8FF2060AAFB5C" box="[352,434,245,263]" italics="true" pageId="5" pageNumber="46">F. asmati</emphasis>
</taxonomicName>
</th>
<th id="F72D5A60FFE00055FDACFF2163C0FB53" box="[516,728,244,264]" gridcol="2" gridrow="0" pageId="5" pageNumber="46">
<taxonomicName id="CDCC4ADDFFE0FFAFFD9EFF2063B3FB5C" box="[566,683,244,263]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="5" pageNumber="46" phylum="Chordata" rank="species" species="nepalensis">
<emphasis id="38B8ED4CFFE0FFAFFD9EFF2063B3FB5C" box="[566,683,244,263]" italics="true" pageId="5" pageNumber="46">F. nepalensis</emphasis>
</taxonomicName>
</th>
<th id="F72D5A60FFE00055FD43FF2162D8FB53" box="[747,960,244,264]" gridcol="3" gridrow="0" pageId="5" pageNumber="46">
<taxonomicName id="CDCC4ADDFFE0FFAFFC86FF20629AFB5C" box="[814,898,245,263]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="5" pageNumber="46" phylum="Chordata" rank="species" species="pierrie">
<emphasis id="38B8ED4CFFE0FFAFFC86FF20629AFB5C" box="[814,898,245,263]" italics="true" pageId="5" pageNumber="46">F. pierrie</emphasis>
</taxonomicName>
</th>
<th id="F72D5A60FFE00055FC7CFF2165B3FB53" box="[980,1195,244,264]" gridcol="4" gridrow="0" pageId="5" pageNumber="46">
<taxonomicName id="CDCC4ADDFFE0FFAFFBAAFF20659AFB53" box="[1026,1154,244,264]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="5" pageNumber="46" phylum="Chordata" rank="species" species="syhadrensis">F. syhadrensis</taxonomicName>
</th>
<th id="F72D5A60FFE00055FB16FF216480FB53" box="[1214,1432,244,264]" gridcol="5" gridrow="0" pageId="5" pageNumber="46">
<taxonomicName id="CDCC4ADDFFE0FFAFFB5FFF20647BFB53" box="[1271,1379,244,264]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="5" pageNumber="46" phylum="Chordata" rank="species" species="teraiensis">F. teraiensis</taxonomicName>
</th>
</tr>
<tr id="B4FC331CFFE00055FF34FEF16480FB0E" box="[156,1432,292,341]" gridrow="1" pageId="5" pageNumber="46" rowspan-0="1">
<td id="F72D5A60FFE00055FEB4FEF160F3FB0E" box="[284,491,292,341]" gridcol="1" gridrow="1" pageId="5" pageNumber="46">Male Female n=5 n=1</td>
<td id="F72D5A60FFE00055FDACFEF163C0FB0E" box="[516,728,292,341]" gridcol="2" gridrow="1" pageId="5" pageNumber="46">Male Female n=5 n=6</td>
<td id="F72D5A60FFE00055FD43FEF162D8FB0E" box="[747,960,292,341]" gridcol="3" gridrow="1" pageId="5" pageNumber="46">Male Female n=2 n=3</td>
<td id="F72D5A60FFE00055FC7CFEF165B3FB0E" box="[980,1195,292,341]" gridcol="4" gridrow="1" pageId="5" pageNumber="46">Male Female n=6 n=3</td>
<td id="F72D5A60FFE00055FB16FEF16480FB0E" box="[1214,1432,292,341]" gridcol="5" gridrow="1" pageId="5" pageNumber="46">Male Female n=10 n=11</td>
</tr>
<tr id="B4FC331CFFE00055FF34FEA46480FBC4" box="[156,1432,369,415]" gridrow="2" pageId="5" pageNumber="46">
<th id="F72D5A60FFE00055FF34FEA461E2FBC4" box="[156,250,369,415]" gridcol="0" gridrow="2" pageId="5" pageNumber="46">HL:SVL</th>
<td id="F72D5A60FFE00055FEB4FEA460F3FBC4" box="[284,491,369,415]" gridcol="1" gridrow="2" pageId="5" pageNumber="46">0.34±0.01 0.34 (0.32–0.36)</td>
<td id="F72D5A60FFE00055FDACFEA463C0FBC4" box="[516,728,369,415]" gridcol="2" gridrow="2" pageId="5" pageNumber="46">0.34±0.01 0.34±0.01 (0.32–0.36) (0.33–0.37)</td>
<td id="F72D5A60FFE00055FD43FEA462D8FBC4" box="[747,960,369,415]" gridcol="3" gridrow="2" pageId="5" pageNumber="46">0.31 0.33±0.00 (0.30–0.33) (0.32–0.33)</td>
<td id="F72D5A60FFE00055FC7CFEA465B3FBC4" box="[980,1195,369,415]" gridcol="4" gridrow="2" pageId="5" pageNumber="46">0.32±0.00 0.33±0.01 (0.32–0.36) (0.32–0.35)</td>
<td id="F72D5A60FFE00055FB16FEA46480FBC4" box="[1214,1432,369,415]" gridcol="5" gridrow="2" pageId="5" pageNumber="46">0.33±0.01 0.33±0.01 (0.31–0.36) (0.31–0.34)</td>
</tr>
<tr id="B4FC331CFFE00055FF34FE6E6480FBB2" box="[156,1432,443,489]" gridrow="3" pageId="5" pageNumber="46">
<th id="F72D5A60FFE00055FF34FE6E61E2FBB2" box="[156,250,443,489]" gridcol="0" gridrow="3" pageId="5" pageNumber="46">HL:HW</th>
<td id="F72D5A60FFE00055FEB4FE6E60F3FBB2" box="[284,491,443,489]" gridcol="1" gridrow="3" pageId="5" pageNumber="46">0.93±0.01 0.92 (0.90–0.95)</td>
<td id="F72D5A60FFE00055FDACFE6E63C0FBB2" box="[516,728,443,489]" gridcol="2" gridrow="3" pageId="5" pageNumber="46">1.06±0.02 1.02±0.03 (0.98–1.02) (0.99–1.05)</td>
<td id="F72D5A60FFE00055FD43FE6E62D8FBB2" box="[747,960,443,489]" gridcol="3" gridrow="3" pageId="5" pageNumber="46">1.02 1.06±0.02 (1.01–1.03) (1.03–1.08)</td>
<td id="F72D5A60FFE00055FC7CFE6E65B3FBB2" box="[980,1195,443,489]" gridcol="4" gridrow="3" pageId="5" pageNumber="46">1.03±0.03 1.04±0.03 (0.99–1.09) (1.03–1.08)</td>
<td id="F72D5A60FFE00055FB16FE6E6480FBB2" box="[1214,1432,443,489]" gridcol="5" gridrow="3" pageId="5" pageNumber="46">1.00±0.01 1.03±0.03 (0.96–1.03) (0.95–1.06)</td>
</tr>
<tr id="B4FC331CFFE00055FF34FDD36480F868" box="[156,1432,518,563]" gridrow="4" pageId="5" pageNumber="46">
<th id="F72D5A60FFE00055FF34FDD361E2F868" box="[156,250,518,563]" gridcol="0" gridrow="4" pageId="5" pageNumber="46">SL:HL</th>
<td id="F72D5A60FFE00055FEB4FDD360F3F868" box="[284,491,518,563]" gridcol="1" gridrow="4" pageId="5" pageNumber="46">0.38±0.01 0.40 (0.37–0.40)</td>
<td id="F72D5A60FFE00055FDACFDD363C0F868" box="[516,728,518,563]" gridcol="2" gridrow="4" pageId="5" pageNumber="46">0.45±0.04 0.39±0.04 (0.39–0.49) (0.34–0.44)</td>
<td id="F72D5A60FFE00055FD43FDD362D8F868" box="[747,960,518,563]" gridcol="3" gridrow="4" pageId="5" pageNumber="46">0.43 0.43±0.03 (0.42–0.44) (0.40–0.47)</td>
<td id="F72D5A60FFE00055FC7CFDD365B3F868" box="[980,1195,518,563]" gridcol="4" gridrow="4" pageId="5" pageNumber="46">0.47±0.03 0.45±0.06 (0.42–0.51) (0.40–0.51)</td>
<td id="F72D5A60FFE00055FB16FDD36480F868" box="[1214,1432,518,563]" gridcol="5" gridrow="4" pageId="5" pageNumber="46">0.47±0.02 0.43±0.02 (0.44–0.49) (0.40–0.46)</td>
</tr>
<tr id="B4FC331CFFE00055FF34FD9A6480F826" box="[156,1432,591,637]" gridrow="5" pageId="5" pageNumber="46">
<th id="F72D5A60FFE00055FF34FD9A61E2F826" box="[156,250,591,637]" gridcol="0" gridrow="5" pageId="5" pageNumber="46">EN:NS</th>
<td id="F72D5A60FFE00055FEB4FD9A60F3F826" box="[284,491,591,637]" gridcol="1" gridrow="5" pageId="5" pageNumber="46">1.77±0.04 1.66 (1.74–1.80)</td>
<td id="F72D5A60FFE00055FDACFD9A63C0F826" box="[516,728,591,637]" gridcol="2" gridrow="5" pageId="5" pageNumber="46">1.24±0.12 1.45±0.19 (1.05–1.35) (1.13–1.64)</td>
<td id="F72D5A60FFE00055FD43FD9A62D8F826" box="[747,960,591,637]" gridcol="3" gridrow="5" pageId="5" pageNumber="46">1.18 1.13±0.07 (1.13–1.24) (1.05–1.20)</td>
<td id="F72D5A60FFE00055FC7CFD9A65B3F826" box="[980,1195,591,637]" gridcol="4" gridrow="5" pageId="5" pageNumber="46">1.14±0.16 1.22±0.15 (0.92–1.41) (1.05–1.35)</td>
<td id="F72D5A60FFE00055FB16FD9A6480F826" box="[1214,1432,591,637]" gridcol="5" gridrow="5" pageId="5" pageNumber="46">1.03±0.07 1.06±0.11 (0.91–1.13) (0.93–1.32)</td>
</tr>
<tr id="B4FC331CFFE00055FF34FD4F6480F89C" box="[156,1432,666,711]" gridrow="6" pageId="5" pageNumber="46">
<th id="F72D5A60FFE00055FF34FD4F61E2F89C" box="[156,250,666,711]" gridcol="0" gridrow="6" pageId="5" pageNumber="46">IN:NS</th>
<td id="F72D5A60FFE00055FEB4FD4F60F3F89C" box="[284,491,666,711]" gridcol="1" gridrow="6" pageId="5" pageNumber="46">1.49±0.03 1.43 (1.45–1.51)</td>
<td id="F72D5A60FFE00055FDACFD4F63C0F89C" box="[516,728,666,711]" gridcol="2" gridrow="6" pageId="5" pageNumber="46">1.33±0.10 1.54±0.14 (1.15–1.39) (1.31–1.73)</td>
<td id="F72D5A60FFE00055FD43FD4F62D8F89C" box="[747,960,666,711]" gridcol="3" gridrow="6" pageId="5" pageNumber="46">1.18 1.23±0.09 (1.18–1.19) (1.15–1.33)</td>
<td id="F72D5A60FFE00055FC7CFD4F65B3F89C" box="[980,1195,666,711]" gridcol="4" gridrow="6" pageId="5" pageNumber="46">1.38±0.21 1.44±0.12 (1.18–1.76) (1.30–1.52)</td>
<td id="F72D5A60FFE00055FB16FD4F6480F89C" box="[1214,1432,666,711]" gridcol="5" gridrow="6" pageId="5" pageNumber="46">0.99±0.09 1.10±0.09 (0.76–1.09) (0.95–1.31)</td>
</tr>
<tr id="B4FC331CFFE00055FF34FD316480F94A" box="[156,1432,740,785]" gridrow="7" pageId="5" pageNumber="46">
<th id="F72D5A60FFE00055FF34FD3161E2F94A" box="[156,250,740,785]" gridcol="0" gridrow="7" pageId="5" pageNumber="46">EL:HL</th>
<td id="F72D5A60FFE00055FEB4FD3160F3F94A" box="[284,491,740,785]" gridcol="1" gridrow="7" pageId="5" pageNumber="46">0.42±0.01 0.41 (0.40–0.43)</td>
<td id="F72D5A60FFE00055FDACFD3163C0F94A" box="[516,728,740,785]" gridcol="2" gridrow="7" pageId="5" pageNumber="46">0.37±0.03 0.35±0.03 (0.32–0.40) (0.30–0.39)</td>
<td id="F72D5A60FFE00055FD43FD3162D8F94A" box="[747,960,740,785]" gridcol="3" gridrow="7" pageId="5" pageNumber="46">0.34 0.36±0.04 (0.34–0.34) (0.31–0.39)</td>
<td id="F72D5A60FFE00055FC7CFD3165B3F94A" box="[980,1195,740,785]" gridcol="4" gridrow="7" pageId="5" pageNumber="46">0.37±0.02 0.33±0.04 (0.33–0.40) (0.29–0.37)</td>
<td id="F72D5A60FFE00055FB16FD316480F94A" box="[1214,1432,740,785]" gridcol="5" gridrow="7" pageId="5" pageNumber="46">0.34±0.01 0.32±0.01 (0.32–0.37) (0.29–0.35)</td>
</tr>
<tr id="B4FC331CFFE00055FF34FCFB6480F907" box="[156,1432,814,860]" gridrow="8" pageId="5" pageNumber="46">
<th id="F72D5A60FFE00055FF34FCFB61E2F907" box="[156,250,814,860]" gridcol="0" gridrow="8" pageId="5" pageNumber="46">EL:SVL</th>
<td id="F72D5A60FFE00055FEB4FCFB60F3F907" box="[284,491,814,860]" gridcol="1" gridrow="8" pageId="5" pageNumber="46">0.15±0.01 0.14 (0.14–0.16)</td>
<td id="F72D5A60FFE00055FDACFCFB63C0F907" box="[516,728,814,860]" gridcol="2" gridrow="8" pageId="5" pageNumber="46">0.13±0.01 0.12±0.01 (0.11–0.15) (0.10–0.13)</td>
<td id="F72D5A60FFE00055FD43FCFB62D8F907" box="[747,960,814,860]" gridcol="3" gridrow="8" pageId="5" pageNumber="46">0.11 0.12±0.01 (0.10–0.11) (0.10–0.13)</td>
<td id="F72D5A60FFE00055FC7CFCFB65B3F907" box="[980,1195,814,860]" gridcol="4" gridrow="8" pageId="5" pageNumber="46">0.12±0.00 0.11±0.01 (0.11–0.13) (0.10–0.12)</td>
<td id="F72D5A60FFE00055FB16FCFB6480F907" box="[1214,1432,814,860]" gridcol="5" gridrow="8" pageId="5" pageNumber="46">0.11±0.00 0.11±0.00 (0.10–0.12) (0.09–0.12)</td>
</tr>
<tr id="B4FC331CFFE00055FF34FCAD6480F9FD" box="[156,1432,888,934]" gridrow="9" pageId="5" pageNumber="46">
<th id="F72D5A60FFE00055FF34FCAD61E2F9FD" box="[156,250,888,934]" gridcol="0" gridrow="9" pageId="5" pageNumber="46">IOD:IN</th>
<td id="F72D5A60FFE00055FEB4FCAD60F3F9FD" box="[284,491,888,934]" gridcol="1" gridrow="9" pageId="5" pageNumber="46">0.88±0.03 0.88 (0.86–0.90)</td>
<td id="F72D5A60FFE00055FDACFCAD63C0F9FD" box="[516,728,888,934]" gridcol="2" gridrow="9" pageId="5" pageNumber="46">0.92±0.07 0.91±0.11 (0.81–1.00) (0.71–1.00)</td>
<td id="F72D5A60FFE00055FD43FCAD62D8F9FD" box="[747,960,888,934]" gridcol="3" gridrow="9" pageId="5" pageNumber="46">0.95 0.83±0.03 (0.95–0.95) (0.80–0.87)</td>
<td id="F72D5A60FFE00055FC7CFCAD65B3F9FD" box="[980,1195,888,934]" gridcol="4" gridrow="9" pageId="5" pageNumber="46">0.75±0.13 0.63±0.07 (0.61–0.96) (0.55–0.69)</td>
<td id="F72D5A60FFE00055FB16FCAD6480F9FD" box="[1214,1432,888,934]" gridcol="5" gridrow="9" pageId="5" pageNumber="46">0.77±0.10 0.64±0.08 (0.55–0.90) (0.53–0.78)</td>
</tr>
<tr id="B4FC331CFFE00055FF34FC176480F9AB" box="[156,1432,962,1008]" gridrow="10" pageId="5" pageNumber="46">
<th id="F72D5A60FFE00055FF34FC1761E2F9AB" box="[156,250,962,1008]" gridcol="0" gridrow="10" pageId="5" pageNumber="46">MBE:HL</th>
<td id="F72D5A60FFE00055FEB4FC1760F3F9AB" box="[284,491,962,1008]" gridcol="1" gridrow="10" pageId="5" pageNumber="46">0.18±0.01 0.19 (0.17–0.19)</td>
<td id="F72D5A60FFE00055FDACFC1763C0F9AB" box="[516,728,962,1008]" gridcol="2" gridrow="10" pageId="5" pageNumber="46">0.29±0.03 0.28±0.03 (0.27–0.32) (0.24–0.32)</td>
<td id="F72D5A60FFE00055FD43FC1762D8F9AB" box="[747,960,962,1008]" gridcol="3" gridrow="10" pageId="5" pageNumber="46">0.30 0.31±0.01 (0.30–0.31) (0.30–0.32)</td>
<td id="F72D5A60FFE00055FC7CFC1765B3F9AB" box="[980,1195,962,1008]" gridcol="4" gridrow="10" pageId="5" pageNumber="46">0.31±0.03 0.33±0.04 (0.27–0.35) (0.29–0.38)</td>
<td id="F72D5A60FFE00055FB16FC176480F9AB" box="[1214,1432,962,1008]" gridcol="5" gridrow="10" pageId="5" pageNumber="46">0.33±0.03 0.34±0.05 (0.27–0.38) (0.28–0.41)</td>
</tr>
<tr id="B4FC331CFFE00055FF34FBD96480FE61" box="[156,1432,1036,1082]" gridrow="11" pageId="5" pageNumber="46">
<th id="F72D5A60FFE00055FF34FBD961E2FE61" box="[156,250,1036,1082]" gridcol="0" gridrow="11" pageId="5" pageNumber="46">FAL:HAL</th>
<td id="F72D5A60FFE00055FEB4FBD960F3FE61" box="[284,491,1036,1082]" gridcol="1" gridrow="11" pageId="5" pageNumber="46">0.71±0.03 0.75 (0.69–0.73)</td>
<td id="F72D5A60FFE00055FDACFBD963C0FE61" box="[516,728,1036,1082]" gridcol="2" gridrow="11" pageId="5" pageNumber="46">0.85±0.08 0.87±0.09 (0.80–0.96) (0.79–0.95)</td>
<td id="F72D5A60FFE00055FD43FBD962D8FE61" box="[747,960,1036,1082]" gridcol="3" gridrow="11" pageId="5" pageNumber="46">0.86 0.83±0.02 (0.85–0.86) (0.80–0.84)</td>
<td id="F72D5A60FFE00055FC7CFBD965B3FE61" box="[980,1195,1036,1082]" gridcol="4" gridrow="11" pageId="5" pageNumber="46">1.00±0.01 0.97±0.02 (0.96–1.02) (0.95–1.00)</td>
<td id="F72D5A60FFE00055FB16FBD96480FE61" box="[1214,1432,1036,1082]" gridcol="5" gridrow="11" pageId="5" pageNumber="46">1.01±0.07 1.02±0.09 (0.92–1.21) (0.93–1.27)</td>
</tr>
<tr id="B4FC331CFFE00055FF34FB836480FEDF" box="[156,1432,1110,1156]" gridrow="12" pageId="5" pageNumber="46">
<th id="F72D5A60FFE00055FF34FB8361E2FEDF" box="[156,250,1110,1156]" gridcol="0" gridrow="12" pageId="5" pageNumber="46">FOL:TL</th>
<td id="F72D5A60FFE00055FEB4FB8360F3FEDF" box="[284,491,1110,1156]" gridcol="1" gridrow="12" pageId="5" pageNumber="46">1.04±0.01 1.04 (1.03–1.05)</td>
<td id="F72D5A60FFE00055FDACFB8363C0FEDF" box="[516,728,1110,1156]" gridcol="2" gridrow="12" pageId="5" pageNumber="46">1.01±0.02 0.97±0.01 (0.97–1.03) (0.92–1.03)</td>
<td id="F72D5A60FFE00055FD43FB8362D8FEDF" box="[747,960,1110,1156]" gridcol="3" gridrow="12" pageId="5" pageNumber="46">0.99 1.06±0.08 (0.99–0.99) (0.99–1.15)</td>
<td id="F72D5A60FFE00055FC7CFB8365B3FEDF" box="[980,1195,1110,1156]" gridcol="4" gridrow="12" pageId="5" pageNumber="46">1.01±0.08 1.00±0.03 (0.85–1.08) (0.97–1.03)</td>
<td id="F72D5A60FFE00055FB16FB836480FEDF" box="[1214,1432,1110,1156]" gridcol="5" gridrow="12" pageId="5" pageNumber="46">0.97±0.05 0.94±0.06 (0.90–1.07) (0.87–1.00)</td>
</tr>
<tr id="B4FC331CFFE00055FF34FB746480FF52" box="[156,1432,1185,1289]" gridrow="13" pageId="5" pageNumber="46">
<th id="F72D5A60FFE00055FF34FB7461E2FF52" box="[156,250,1185,1289]" gridcol="0" gridrow="13" pageId="5" pageNumber="46">Vo c a l marking on throat</th>
<td id="F72D5A60FFE00055FEB4FB7460F3FF52" box="[284,491,1185,1289]" gridcol="1" gridrow="13" pageId="5" pageNumber="46">Butterfly _____ shaped dark mak- ing</td>
<td id="F72D5A60FFE00055FDACFB7463C0FF52" box="[516,728,1185,1289]" gridcol="2" gridrow="13" pageId="5" pageNumber="46">Laterally _____ dark and medially pale</td>
<td id="F72D5A60FFE00055FD43FB7462D8FF52" box="[747,960,1185,1289]" gridcol="3" gridrow="13" pageId="5" pageNumber="46">Laterally _____ dark and medially pale</td>
<td id="F72D5A60FFE00055FC7CFB7465B3FF52" box="[980,1195,1185,1289]" gridcol="4" gridrow="13" pageId="5" pageNumber="46">Crescent _____ shaped dark mak- ing</td>
<td id="F72D5A60FFE00055FB16FB746480FF52" box="[1214,1432,1185,1289]" gridcol="5" gridrow="13" pageId="5" pageNumber="46">“W” ______ shaped dark marking</td>
</tr>
<tr id="B4FC331CFFE00055FF34FAF36480FF29" box="[156,1432,1318,1394]" gridrow="14" pageId="5" pageNumber="46">
<th id="F72D5A60FFE00055FF34FAF361E2FF29" box="[156,250,1318,1394]" gridcol="0" gridrow="14" pageId="5" pageNumber="46">Relative length of fingers</th>
<td id="F72D5A60FFE00055FEB4FAF360F3FF29" box="[284,491,1318,1394]" gridcol="1" gridrow="14" pageId="5" pageNumber="46">2&lt;4&lt;1&lt;3 2&lt;4&lt;1&lt;3</td>
<td id="F72D5A60FFE00055FDACFAF363C0FF29" box="[516,728,1318,1394]" gridcol="2" gridrow="14" pageId="5" pageNumber="46">2&lt;1&lt;4&lt;3 2&lt;1&lt;4&lt;3</td>
<td id="F72D5A60FFE00055FD43FAF362D8FF29" box="[747,960,1318,1394]" gridcol="3" gridrow="14" pageId="5" pageNumber="46">2=4&lt;1&lt;3 2=4&lt;1&lt;3</td>
<td id="F72D5A60FFE00055FC7CFAF365B3FF29" box="[980,1195,1318,1394]" gridcol="4" gridrow="14" pageId="5" pageNumber="46">1=2&lt;4&lt;3 1=2&lt;4&lt;3</td>
<td id="F72D5A60FFE00055FB16FAF36480FF29" box="[1214,1432,1318,1394]" gridcol="5" gridrow="14" pageId="5" pageNumber="46">2=4&lt;1&lt;3 2=4&lt;1&lt;3</td>
</tr>
<tr id="B4FC331CFFE00055FF34FA5B6480FC75" box="[156,1432,1422,1582]" gridrow="15" pageId="5" pageNumber="46">
<th id="F72D5A60FFE00055FF34FA5B61E2FC75" box="[156,250,1422,1582]" gridcol="0" gridrow="15" pageId="5" pageNumber="46">Middorsal line</th>
<td id="F72D5A60FFE00055FEB4FA5B60F3FC75" box="[284,491,1422,1582]" gridcol="1" gridrow="15" pageId="5" pageNumber="46">Distinct Distinct &amp; narrow &amp; narrow</td>
<td id="F72D5A60FFE00055FDACFA5B63C0FC75" box="[516,728,1422,1582]" gridcol="2" gridrow="15" pageId="5" pageNumber="46">Distinct Distinct &amp; narrow &amp; narrow</td>
<td id="F72D5A60FFE00055FD43FA5B62D8FC75" box="[747,960,1422,1582]" gridcol="3" gridrow="15" pageId="5" pageNumber="46">Distinct Distinct with same with same width width throughout throughout from snout from snout to vent to vent</td>
<td id="F72D5A60FFE00055FC7CFA5B65B3FC75" box="[980,1195,1422,1582]" gridcol="4" gridrow="15" pageId="5" pageNumber="46">Absent Absent</td>
<td id="F72D5A60FFE00055FB16FA5B6480FC75" box="[1214,1432,1422,1582]" gridcol="5" gridrow="15" pageId="5" pageNumber="46">Va ri ab l e Variable</td>
</tr>
</table>
</paragraph>
<paragraph id="0A73315EFFE0FFAFFF34F99E6492FC04" blockId="5.[156,1427,1611,1766]" box="[156,1418,1611,1631]" pageId="5" pageNumber="46">Skin fringe Weak and Weak and Ve ry i nd is - Ve ry i nd is - Absent Absent Vestige or Vestige or Distinct Distinct</paragraph>
<paragraph id="0A73315EFFE0FFAFFF34F9BD6584FC20" blockId="5.[156,1427,1611,1766]" box="[156,1180,1639,1659]" pageId="5" pageNumber="46">on outer side indistinct indistinct tinct tinct reduced reduced</paragraph>
<paragraph id="0A73315EFFE0FFAFFF34F95061F3FCC2" blockId="5.[156,1427,1611,1766]" box="[156,235,1667,1689]" pageId="5" pageNumber="46">of 5th toe</paragraph>
<paragraph id="0A73315EFFE0FFAFFF34F963648BFC91" blockId="5.[156,1427,1611,1766]" box="[156,1427,1718,1738]" pageId="5" pageNumber="46">
Body size 29.1–30.0 
<quantity id="CD349CBBFFE0FFAFFE3FF96360FFFC91" box="[407,487,1718,1738]" metricMagnitude="-2" metricUnit="m" metricValue="3.34" pageId="5" pageNumber="46" unit="mm" value="33.4">33.4 mm</quantity>
21.4–0.1 22.5–31.2 25.6–27.8 23.1–29.4 27.1–30.0 32.8–37.2 37.8–44.1 35.7–52.3
</paragraph>
<paragraph id="0A73315EFFE0FFAFFF34F9076093FD89" blockId="5.[156,1427,1611,1766]" lastBlockId="5.[151,1436,1870,2002]" pageId="5" pageNumber="46">
(SVL) mm mm mm mm mm mm mm mm mm Frequency ranges from 4–7.5 kHz with a dominant frequency around 4.1–5.1 kHz. 
<typeStatus id="D5778FFCFFE0FFAFFF6FF8A661E7FDD0" box="[199,255,1907,1931]" pageId="5" pageNumber="46">Type</typeStatus>
3 single note sound very similar to 
<typeStatus id="D5778FFCFFE0FFAFFD3EF8A663DEFDD0" box="[662,710,1907,1931]" pageId="5" pageNumber="46">type</typeStatus>
2. Duration of 5–11 ms (9.0±2.35, N=5) and inter-note duration of 5–6 ms (5.5±0.70, N=2). The frequency of 
<typeStatus id="D5778FFCFFE0FFAFFD0FF84263CFFDF4" box="[679,727,1943,1967]" pageId="5" pageNumber="46">type</typeStatus>
3 notes ranges from 1 to 6.5 KHz with a dominant frequency between 2 and 4 kHz.
</paragraph>
</subSubSection>
<caption id="5EB361D6FFE3FFACFF3FFCE065BBF917" box="[151,1187,821,844]" httpUri="https://zenodo.org/record/202488/files/figure.png" pageId="6" pageNumber="47" targetBox="[351,1238,200,788]" targetPageId="6">
<paragraph id="0A73315EFFE3FFACFF3FFCE065BBF917" blockId="6.[151,1187,821,844]" box="[151,1187,821,844]" pageId="6" pageNumber="47">
<emphasis id="38B8ED4CFFE3FFACFF3FFCE0600AF910" bold="true" box="[151,274,821,844]" pageId="6" pageNumber="47">FIGURE 4.</emphasis>
Sonogram (top) and oscillogram (bottom) of the advertisement call of 
<taxonomicName id="CDCC4ADDFFE3FFACFC4AFCE26587F917" box="[994,1183,823,844]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="asmati">Fejervarya asmati</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="42D662D5FFE3FFACFF6FFC47653AF995" pageId="6" pageNumber="47" type="etymology">
<paragraph id="0A73315EFFE3FFACFF6FFC47653AF995" blockId="6.[151,1436,914,2022]" pageId="6" pageNumber="47">
<emphasis id="38B8ED4CFFE3FFACFF6FFC476056F9F1" bold="true" box="[199,334,914,938]" pageId="6" pageNumber="47">Etymology.</emphasis>
—I take great pleasure in naming the new species as a patronym for my teacher and mentor Ghazi S. M. Asmat, in recognition of his endless contributions to my personal research.
</paragraph>
</subSubSection>
<subSubSection id="42D662D5FFE3FFACFF6FFC0C6451FCB2" pageId="6" pageNumber="47" type="distribution">
<paragraph id="0A73315EFFE3FFACFF6FFC0C6312FE61" blockId="6.[151,1436,914,2022]" pageId="6" pageNumber="47">
<emphasis id="38B8ED4CFFE3FFACFF6FFC0C6078F9A9" bold="true" box="[199,352,985,1010]" pageId="6" pageNumber="47">Distribution.</emphasis>
— 
<taxonomicName id="CDCC4ADDFFE3FFACFED4FC0E6352F9A9" box="[380,586,987,1010]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="asmati">
<emphasis id="38B8ED4CFFE3FFACFED4FC0E6352F9A9" box="[380,586,987,1010]" italics="true" pageId="6" pageNumber="47">Fejervarya asmati</emphasis>
</taxonomicName>
is known only from the 
<typeStatus id="D5778FFCFFE3FFACFCF7FC0E6297F9A8" box="[863,911,987,1011]" pageId="6" pageNumber="47">type</typeStatus>
locality (Fig. 1). A probable record exists also from Barisal district (
<geoCoordinate id="6FF85799FFE3FFACFE23FC2B6319FE4D" box="[395,513,1022,1046]" direction="north" orientation="latitude" pageId="6" pageNumber="47" precision="925" value="22.966667">22° 58´N</geoCoordinate>
, 
<geoCoordinate id="6FF85799FFE3FFACFDA5FC2B6366FE4D" box="[525,638,1022,1046]" direction="east" orientation="longitude" pageId="6" pageNumber="47" precision="925" value="90.21667">90° 13´E</geoCoordinate>
); here, similar advertisement calls were heard on 
<date id="7E72179EFFE3FFACFB1CFC2A6450FE4D" box="[1204,1352,1022,1047]" pageId="6" pageNumber="47" value="2009-05-03">May 3, 2009</date>
, but no voucher specimens are available.
</paragraph>
<paragraph id="0A73315EFFE3FFACFF6FFB93641FFF62" blockId="6.[151,1436,914,2022]" pageId="6" pageNumber="47">
<emphasis id="38B8ED4CFFE3FFACFF6FFB936090FE05" bold="true" box="[199,392,1093,1118]" pageId="6" pageNumber="47">Natural history.</emphasis>
—The new species were found in temporary pools along a small stream. Specimens have been observed breeding in a canal adjacent to 
<collectingCountry id="72DB71CEFFE3FFACFD09FBB86205FEDE" box="[673,797,1133,1157]" name="Bangladesh" pageId="6" pageNumber="47">Chittagong</collectingCountry>
University (
<date id="7E72179EFFE3FFACFC04FBB86546FEDD" box="[940,1118,1130,1158]" pageId="6" pageNumber="47">15th to 20th May</date>
, 2008). Gravid females preferred choruses rather than individual or few calling males. Courtship was initiated when a gravid female approached the calling male and made a physical contact with it. The male stopped calling and immediately jumped on the back of the female and clung to it by holding it below the armpits with the forearms and formed an axillary amplexus (
<figureCitation id="92F72DDBFFE3FFACFED8FB2860AFFF4E" box="[368,439,1277,1301]" captionStart="FIGURE 2" captionStartId="3.[151,250,1973,1996]" captionTargetBox="[156,1430,179,1949]" captionTargetId="figure@3.[156,1431,179,1949]" captionTargetPageId="3" captionText="FIGURE 2. Photographs in life of Fejervarya asmati sp. nov. A: Calling male. B: Mating of adult individuals. C: Ventral view of male." httpUri="https://zenodo.org/record/202486/files/figure.png" pageId="6" pageNumber="47">Fig. 2</figureCitation>
). The amplexed male in turn kicked the intruder with its hind limbs. The amplexed pair moved to a small, shallow water pool on the ground within the locality where the spawning occurred.
</paragraph>
<paragraph id="0A73315EFFE3FFACFF6FFA93659EFFFE" blockId="6.[151,1436,914,2022]" pageId="6" pageNumber="47">
In general, according to own observations, the courtship pattern is same for all observed species of 
<taxonomicName id="CDCC4ADDFFE3FFACFAB7FA936484FF06" box="[1311,1436,1350,1373]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="47" phylum="Chordata" rank="genus">
<emphasis id="38B8ED4CFFE3FFACFAB7FA936484FF06" box="[1311,1436,1350,1373]" italics="true" pageId="6" pageNumber="47">Fejervarya</emphasis>
</taxonomicName>
(
<taxonomicName id="CDCC4ADDFFE3FFACFF08FABF602AFFDA" box="[160,306,1385,1409]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="nepalensis">
<emphasis id="38B8ED4CFFE3FFACFF08FABF602AFFDA" box="[160,306,1385,1409]" italics="true" pageId="6" pageNumber="47">F. nepalensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="CDCC4ADDFFE3FFACFEE9FABF60B0FFDA" box="[321,424,1386,1409]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="pierrie">
<emphasis id="38B8ED4CFFE3FFACFEE9FABF60B0FFDA" box="[321,424,1386,1409]" italics="true" pageId="6" pageNumber="47">F. pierrie</emphasis>
</taxonomicName>
, 
<taxonomicName id="CDCC4ADDFFE3FFACFE1FFABF634EFFDA" box="[439,598,1385,1409]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="syhadrensis">
<emphasis id="38B8ED4CFFE3FFACFE1FFABF634EFFDA" box="[439,598,1385,1409]" italics="true" pageId="6" pageNumber="47">F. syhadrensis</emphasis>
</taxonomicName>
, and 
<taxonomicName id="CDCC4ADDFFE3FFACFD3CFABF6206FFDA" box="[660,798,1386,1409]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="teraiensis">
<emphasis id="38B8ED4CFFE3FFACFD3CFABF6206FFDA" box="[660,798,1386,1409]" italics="true" pageId="6" pageNumber="47">F. teraiensis</emphasis>
</taxonomicName>
) occurring at this locality. Further notes on conservation status and threats will be documented with other amphibians of 
<collectingCountry id="72DB71CEFFE3FFACFCCAFA5862FCFFFE" box="[866,996,1421,1445]" name="Bangladesh" pageId="6" pageNumber="47">Bangladesh</collectingCountry>
on elsewhere.
</paragraph>
<paragraph id="0A73315EFFE3FFACFF6FFA656451FCB2" blockId="6.[151,1436,914,2022]" pageId="6" pageNumber="47">
<emphasis id="38B8ED4CFFE3FFACFF6FFA6560D8FF92" bold="true" box="[199,448,1456,1481]" pageId="6" pageNumber="47">Sympatric species.—</emphasis>
As mentioned above, in the area of the 
<typeStatus id="D5778FFCFFE3FFACFCD5FA6762B5FF91" box="[893,941,1458,1482]" pageId="6" pageNumber="47">type</typeStatus>
locality of the new species, observations of the other four 
<taxonomicName id="CDCC4ADDFFE3FFACFE94FA0360A1FFB6" box="[316,441,1494,1517]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="47" phylum="Chordata" rank="genus">
<emphasis id="38B8ED4CFFE3FFACFE94FA0360A1FFB6" box="[316,441,1494,1517]" italics="true" pageId="6" pageNumber="47">Fejervarya</emphasis>
</taxonomicName>
species occurring in 
<collectingCountry id="72DB71CEFFE3FFACFD06FA006229FFB6" box="[686,817,1493,1517]" name="Bangladesh" pageId="6" pageNumber="47">Bangladesh</collectingCountry>
are also available (
<figureCitation id="92F72DDBFFE3FFACFBB9FA00654FFFB6" box="[1041,1111,1493,1517]" captionStart="FIGURE 5" captionStartId="7.[151,250,1653,1676]" captionTargetBox="[153,1436,630,1629]" captionTargetId="figure@7.[151,1436,630,1629]" captionTargetPageId="7" captionText="FIGURE 5. Four species of Fejervarya occurring in Bangladesh. A: F. nepalensis. B: F. pi e rr i e. C: F. syhadrensis. D: F. teraiensis." httpUri="https://zenodo.org/record/202489/files/figure.png" pageId="6" pageNumber="47">Fig. 5</figureCitation>
). As summarized in 
<tableCitation id="474E04E5FFE3FFACFAEFFA006484FFB6" box="[1351,1436,1493,1517]" captionStart="TABLE 1" captionStartId="5.[151,239,151,175]" captionTargetBox="[156,1432,244,1582]" captionTargetPageId="5" captionText="TABLE 1. Summary of qualitative diagnostic characters in Fejervarya asmati sp. nov. and specimens of other congeners. Morphological ratios given as mean ± standard deviation over range." httpUri="http://table.plazi.org/id/5EB361D6FFE0FFAFFF3FFF4262A8FA89" pageId="6" pageNumber="47" tableUuid="5EB361D6FFE0FFAFFF3FFF4262A8FA89">Table 1</tableCitation>
and in the following section, the five 
<taxonomicName id="CDCC4ADDFFE3FFACFDEBFA2F63A7FC4A" box="[579,703,1530,1553]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="47" phylum="Chordata" rank="genus">
<emphasis id="38B8ED4CFFE3FFACFDEBFA2F63A7FC4A" box="[579,703,1530,1553]" italics="true" pageId="6" pageNumber="47">Fejervarya</emphasis>
</taxonomicName>
species from this area can be distinguished by a combination of various morphological characters. Furthermore, we also have data on the advertisement calls of three of these species from the same area. Detailed bioacoustic analyses will be published elsewhere, but by general call structure it is easy to distinguish the calls of 
<taxonomicName id="CDCC4ADDFFE3FFACFDB1F9B363B5FC26" box="[537,685,1637,1661]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="nepalensis">
<emphasis id="38B8ED4CFFE3FFACFDB1F9B363B5FC26" box="[537,685,1637,1661]" italics="true" pageId="6" pageNumber="47">F. nepalensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="CDCC4ADDFFE3FFACFD16F9B36230FC26" box="[702,808,1638,1661]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="pierrie">
<emphasis id="38B8ED4CFFE3FFACFD16F9B36230FC26" box="[702,808,1638,1661]" italics="true" pageId="6" pageNumber="47">F. pierrie</emphasis>
</taxonomicName>
, 
<taxonomicName id="CDCC4ADDFFE3FFACFC90F9B362C0FC26" box="[824,984,1637,1661]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="syhadrensis">
<emphasis id="38B8ED4CFFE3FFACFC90F9B362C0FC26" box="[824,984,1637,1661]" italics="true" pageId="6" pageNumber="47">F. syhadrensis</emphasis>
</taxonomicName>
, and 
<taxonomicName id="CDCC4ADDFFE3FFACFBB3F9B365BDFC26" box="[1051,1189,1638,1661]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="teraiensis">
<emphasis id="38B8ED4CFFE3FFACFBB3F9B365BDFC26" box="[1051,1189,1638,1661]" italics="true" pageId="6" pageNumber="47">F. teraiensis</emphasis>
</taxonomicName>
from each other, and from 
<emphasis id="38B8ED4CFFE3FFACFF7DF95F6020FCFA" box="[213,312,1674,1697]" italics="true" pageId="6" pageNumber="47">F. a s m a t i</emphasis>
, confirming the species status of these taxa. All 
<taxonomicName id="CDCC4ADDFFE3FFACFCCCF95F62F9FCFA" box="[868,993,1674,1697]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="47" phylum="Chordata" rank="genus">
<emphasis id="38B8ED4CFFE3FFACFCCCF95F62F9FCFA" box="[868,993,1674,1697]" italics="true" pageId="6" pageNumber="47">Fejervarya</emphasis>
</taxonomicName>
species breed in rainy season (April to September), but it is rare to find them to share the same place in the same time. However, they share their breeding place with species of other genera, for example, 
<taxonomicName id="CDCC4ADDFFE3FFACFD1DF907629CFCB2" box="[693,900,1746,1769]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="asmati">
<emphasis id="38B8ED4CFFE3FFACFD1DF907629CFCB2" box="[693,900,1746,1769]" italics="true" pageId="6" pageNumber="47">Fejervarya asmati</emphasis>
</taxonomicName>
is usually found with 
<taxonomicName id="CDCC4ADDFFE3FFACFBD5F904645BFCB2" box="[1149,1347,1745,1769]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="ornata">
<emphasis id="38B8ED4CFFE3FFACFBD5F904645BFCB2" box="[1149,1347,1745,1769]" italics="true" pageId="6" pageNumber="47">Microhyla ornata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="42D662D5FFE3FFADFF6FF9216285F864" lastPageId="7" lastPageNumber="48" pageId="6" pageNumber="47" type="description">
<paragraph id="0A73315EFFE3FFADFF6FF9216001FB47" blockId="6.[151,1436,914,2022]" lastBlockId="7.[151,1436,151,576]" lastPageId="7" lastPageNumber="48" pageId="6" pageNumber="47">
<emphasis id="38B8ED4CFFE3FFACFF6FF9216074FD56" bold="true" box="[199,364,1780,1805]" pageId="6" pageNumber="47">Comparisons.</emphasis>
— 
<taxonomicName id="CDCC4ADDFFE3FFACFE20F923634FFD56" box="[392,599,1782,1805]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="asmati">
<emphasis id="38B8ED4CFFE3FFACFE20F923634FFD56" box="[392,599,1782,1805]" italics="true" pageId="6" pageNumber="47">Fejervarya asmati</emphasis>
</taxonomicName>
is distinguished from other congeners occurring in 
<collectingCountry id="72DB71CEFFE3FFACFB30F9206402FD56" box="[1176,1306,1781,1805]" name="Bangladesh" pageId="6" pageNumber="47">Bangladesh</collectingCountry>
as follows: butterfly shaped vocal marking on throat in male (vs. laterally dark and medially pale in both 
<taxonomicName id="CDCC4ADDFFE3FFACFB15F8CF6457FD6A" box="[1213,1359,1817,1841]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="nepalensis">
<emphasis id="38B8ED4CFFE3FFACFB15F8CF6457FD6A" box="[1213,1359,1817,1841]" italics="true" pageId="6" pageNumber="47">F. nepalensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="CDCC4ADDFFE3FFACFA2FF8CF61FBFD0E" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="pierrie">
<emphasis id="38B8ED4CFFE3FFACFA2FF8CF61FBFD0E" italics="true" pageId="6" pageNumber="47">F. pierrie</emphasis>
</taxonomicName>
, crescent shaped in 
<taxonomicName id="CDCC4ADDFFE3FFACFE6EF8EB637DFD0E" box="[454,613,1853,1877]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="syhadrensis">
<emphasis id="38B8ED4CFFE3FFACFE6EF8EB637DFD0E" box="[454,613,1853,1877]" italics="true" pageId="6" pageNumber="47">F. syhadrensis</emphasis>
</taxonomicName>
, “W” shaped in 
<taxonomicName id="CDCC4ADDFFE3FFACFC89F8EB62B1FD0E" box="[801,937,1854,1877]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="teraiensis">
<emphasis id="38B8ED4CFFE3FFACFC89F8EB62B1FD0E" box="[801,937,1854,1877]" italics="true" pageId="6" pageNumber="47">F. teraiensis</emphasis>
</taxonomicName>
); forearm length 72% hand length (vs. forearm length greater than 80– 95% hand length in both 
<taxonomicName id="CDCC4ADDFFE3FFACFCADF8B76281FD22" box="[773,921,1889,1913]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="nepalensis">
<emphasis id="38B8ED4CFFE3FFACFCADF8B76281FD22" box="[773,921,1889,1913]" italics="true" pageId="6" pageNumber="47">F. nepalensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="CDCC4ADDFFE3FFACFC7CF8B76526FD22" box="[980,1086,1890,1913]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="pierrie">
<emphasis id="38B8ED4CFFE3FFACFC7CF8B76526FD22" box="[980,1086,1890,1913]" italics="true" pageId="6" pageNumber="47">F. pierrie</emphasis>
</taxonomicName>
, forearm length equal to hand length in both 
<taxonomicName id="CDCC4ADDFFE3FFACFE93F85360C1FDC6" box="[315,473,1925,1949]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="syhadrensis">
<emphasis id="38B8ED4CFFE3FFACFE93F85360C1FDC6" box="[315,473,1925,1949]" italics="true" pageId="6" pageNumber="47">F. syhadrensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="CDCC4ADDFFE3FFACFDB9F8536382FDC6" box="[529,666,1926,1949]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="teraiensis">
<emphasis id="38B8ED4CFFE3FFACFDB9F8536382FDC6" box="[529,666,1926,1949]" italics="true" pageId="6" pageNumber="47">F. teraiensis</emphasis>
</taxonomicName>
); relative length of fingers, shortest to longest: 2&lt;4 &lt;1 &lt;3 (vs. relative length of fingers: 2= 4 &lt;1 &lt;
<quantity id="CD349CBBFFE3FFACFE57F87C6331FD9A" box="[511,553,1961,1985]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="6" pageNumber="47" unit="in" value="3.0">3 in</quantity>
both 
<taxonomicName id="CDCC4ADDFFE3FFACFDC1F87F63C8FD9A" box="[617,720,1962,1985]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="pierrie">
<emphasis id="38B8ED4CFFE3FFACFDC1F87F63C8FD9A" box="[617,720,1962,1985]" italics="true" pageId="6" pageNumber="47">F. pierrie</emphasis>
</taxonomicName>
and 
<taxonomicName id="CDCC4ADDFFE3FFACFCAFF87F6288FD9A" box="[775,912,1962,1985]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="teraiensis">
<emphasis id="38B8ED4CFFE3FFACFCAFF87F6288FD9A" box="[775,912,1962,1985]" italics="true" pageId="6" pageNumber="47">F. teraiensis</emphasis>
</taxonomicName>
; 2 &lt;1 &lt;4 &lt;
<quantity id="CD349CBBFFE3FFACFB8AF87C6555FD9A" box="[1058,1101,1961,1985]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="6" pageNumber="47" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="CDCC4ADDFFE3FFACFBFCF87F65FDFD9A" box="[1108,1253,1961,1985]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="nepalensis">
<emphasis id="38B8ED4CFFE3FFACFBFCF87F65FDFD9A" box="[1108,1253,1961,1985]" italics="true" pageId="6" pageNumber="47">F. nepalensis</emphasis>
</taxonomicName>
; 1= 2 &lt;4 &lt;
<quantity id="CD349CBBFFE3FFACFAD9F87C6484FD9A" box="[1393,1436,1961,1985]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="6" pageNumber="47" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="CDCC4ADDFFE3FFACFF3FF81B6020FDBE" box="[151,312,1997,2021]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="6" pageNumber="47" phylum="Chordata" rank="species" species="syhadrensis">
<emphasis id="38B8ED4CFFE3FFACFF3FF81B6020FDBE" box="[151,312,1997,2021]" italics="true" pageId="6" pageNumber="47">F. syhadrensis</emphasis>
</taxonomicName>
;); nostril much closer to snout tip than eye, distance from front of eyes to nostril over 1.5 times grater than nostril-snout length (nostril not much closer to snout in other congeners); internarial distance 1.5 times greater than nostril to snout tip distance (vs. below 1.5 times in both 
<taxonomicName id="CDCC4ADDFFE2FFADFC12FF686557FA8F" box="[954,1103,188,212]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="48" phylum="Chordata" rank="species" species="nepalensis">
<emphasis id="38B8ED4CFFE2FFADFC12FF686557FA8F" box="[954,1103,188,212]" italics="true" pageId="7" pageNumber="48">F. nepalensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="CDCC4ADDFFE2FFADFB22FF686433FA8F" box="[1162,1323,188,212]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="48" phylum="Chordata" rank="species" species="syhadrensis">
<emphasis id="38B8ED4CFFE2FFADFB22FF686433FA8F" box="[1162,1323,188,212]" italics="true" pageId="7" pageNumber="48">F. syhadrensis</emphasis>
</taxonomicName>
; approximately equal in 
<taxonomicName id="CDCC4ADDFFE2FFADFEE5FF3460ADFAA3" box="[333,437,225,248]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="48" phylum="Chordata" rank="species" species="pierrie">
<emphasis id="38B8ED4CFFE2FFADFEE5FF3460ADFAA3" box="[333,437,225,248]" italics="true" pageId="7" pageNumber="48">F. pierrie</emphasis>
</taxonomicName>
and 
<taxonomicName id="CDCC4ADDFFE2FFADFE45FF34636FFAA3" box="[493,631,225,248]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="48" phylum="Chordata" rank="species" species="teraiensis">
<emphasis id="38B8ED4CFFE2FFADFE45FF34636FFAA3" box="[493,631,225,248]" italics="true" pageId="7" pageNumber="48">F. teraiensis</emphasis>
</taxonomicName>
); MBE 18% of head length (vs. MBE 26– 41% of head length in other congeners).
</paragraph>
<paragraph id="0A73315EFFE2FFADFF6FFEFD6285F864" blockId="7.[151,1436,151,576]" pageId="7" pageNumber="48">
Moreover, 
<taxonomicName id="CDCC4ADDFFE2FFADFEECFEFC60B5FB1B" box="[324,429,297,320]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="48" phylum="Chordata" rank="species" species="asmati">
<emphasis id="38B8ED4CFFE2FFADFEECFEFC60B5FB1B" box="[324,429,297,320]" italics="true" pageId="7" pageNumber="48">F. asmati</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName id="CDCC4ADDFFE2FFADFD07FEFC6221FB1B" box="[687,825,297,320]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="48" phylum="Chordata" rank="species" species="teraiensis">
<emphasis id="38B8ED4CFFE2FFADFD07FEFC6221FB1B" box="[687,825,297,320]" italics="true" pageId="7" pageNumber="48">F. teraiensis</emphasis>
</taxonomicName>
by smaller size (SVL 
<quantity id="CD349CBBFFE2FFADFB96FEF265FBFB64" box="[1086,1251,295,319]" metricMagnitude="-2" metricUnit="m" metricValue="3.125" metricValueMax="3.34" metricValueMin="2.91" pageId="7" pageNumber="48" unit="mm" value="31.25" valueMax="33.4" valueMin="29.1">29.1–33.4 mm</quantity>
in 
<taxonomicName id="CDCC4ADDFFE2FFADFAA3FEFC646BFB1B" box="[1291,1395,297,320]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="48" phylum="Chordata" rank="species" species="asmati" status="sp. nov.">
<emphasis id="38B8ED4CFFE2FFADFAA3FEFC646BFB1B" box="[1291,1395,297,320]" italics="true" pageId="7" pageNumber="48">F. asmati</emphasis>
</taxonomicName>
<taxonomicNameLabel id="238B5037FFE2FFADFAD4FEFD61DEFB3F" pageId="7" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
; SVL 
<quantity id="CD349CBBFFE2FFADFEB9FE9960ADFB3F" box="[273,437,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" metricValueMax="5.2299999999999995" metricValueMin="3.57" pageId="7" pageNumber="48" unit="mm" value="44.0" valueMax="52.3" valueMin="35.7">35.7–52.3 mm</quantity>
in 
<taxonomicName id="CDCC4ADDFFE2FFADFE71FE98637AFB3F" box="[473,610,333,356]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="48" phylum="Chordata" rank="species" species="teraiensis">
<emphasis id="38B8ED4CFFE2FFADFE71FE98637AFB3F" box="[473,610,333,356]" italics="true" pageId="7" pageNumber="48">F. teraiensis</emphasis>
</taxonomicName>
); head is longer than wide (vs. head is longer than wide or equal in other congeners); snout length 37– 40% of head length (vs. snout length 40–51% of length in 
<taxonomicName id="CDCC4ADDFFE2FFADFBD6FEA465FEFBD3" box="[1150,1254,369,392]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="48" phylum="Chordata" rank="species" species="pierrie">
<emphasis id="38B8ED4CFFE2FFADFBD6FEA465FEFBD3" box="[1150,1254,369,392]" italics="true" pageId="7" pageNumber="48">F. pierrie</emphasis>
</taxonomicName>
, 
<taxonomicName id="CDCC4ADDFFE2FFADFB5DFEA4648DFBD3" box="[1269,1429,368,392]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="48" phylum="Chordata" rank="species" species="syhadrensis">
<emphasis id="38B8ED4CFFE2FFADFB5DFEA4648DFBD3" box="[1269,1429,368,392]" italics="true" pageId="7" pageNumber="48">F. syhadrensis</emphasis>
</taxonomicName>
, and 
<taxonomicName id="CDCC4ADDFFE2FFADFF6EFE406056FBF7" box="[198,334,405,428]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="48" phylum="Chordata" rank="species" species="teraiensis">
<emphasis id="38B8ED4CFFE2FFADFF6EFE406056FBF7" box="[198,334,405,428]" italics="true" pageId="7" pageNumber="48">F. teraiensis</emphasis>
</taxonomicName>
); middorsal line is narrow and distinct (vs. distinct with same width throughout from snout to vent in 
<taxonomicName id="CDCC4ADDFFE2FFADFF1FFE6C603AFB8B" box="[183,290,441,464]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="48" phylum="Chordata" rank="species" species="pierrie">
<emphasis id="38B8ED4CFFE2FFADFF1FFE6C603AFB8B" box="[183,290,441,464]" italics="true" pageId="7" pageNumber="48">F. pierrie</emphasis>
</taxonomicName>
, absent in 
<taxonomicName id="CDCC4ADDFFE2FFADFE0FFE6C6351FB8B" box="[423,585,440,464]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="48" phylum="Chordata" rank="species" species="syhadrensis">
<emphasis id="38B8ED4CFFE2FFADFE0FFE6C6351FB8B" box="[423,585,440,464]" italics="true" pageId="7" pageNumber="48">F. syhadrensis</emphasis>
</taxonomicName>
, variable in 
<taxonomicName id="CDCC4ADDFFE2FFADFD48FE6C6274FB8B" box="[736,876,441,464]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="48" phylum="Chordata" rank="species" species="teraiensis">
<emphasis id="38B8ED4CFFE2FFADFD48FE6C6274FB8B" box="[736,876,441,464]" italics="true" pageId="7" pageNumber="48">F. teraiensis</emphasis>
</taxonomicName>
); two red spot found on the middorsal line and another two red spot on each of the forelimbs usually found in male (red spot arrangements are not same in other congeners); weak and indistinct skin fringe on outer side of 5th toe (vs. distinct in 
<taxonomicName id="CDCC4ADDFFE2FFADFB82FDD165ABF840" box="[1066,1203,516,539]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="48" phylum="Chordata" rank="species" species="teraiensis">
<emphasis id="38B8ED4CFFE2FFADFB82FDD165ABF840" box="[1066,1203,516,539]" italics="true" pageId="7" pageNumber="48">F. teraiensis</emphasis>
</taxonomicName>
, absent in 
<taxonomicName id="CDCC4ADDFFE2FFADFA85FDD1648CF840" box="[1325,1428,516,539]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="48" phylum="Chordata" rank="species" species="pierrie">
<emphasis id="38B8ED4CFFE2FFADFA85FDD1648CF840" box="[1325,1428,516,539]" italics="true" pageId="7" pageNumber="48">F. pierrie</emphasis>
</taxonomicName>
, vestige or reduced in 
<taxonomicName id="CDCC4ADDFFE2FFADFE20FDFD633EF864" box="[392,550,551,575]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="48" phylum="Chordata" rank="species" species="syhadrensis">
<emphasis id="38B8ED4CFFE2FFADFE20FDFD633EF864" box="[392,550,551,575]" italics="true" pageId="7" pageNumber="48">F. syhadrensis</emphasis>
</taxonomicName>
, very indistinct in 
<taxonomicName id="CDCC4ADDFFE2FFADFD51FDFD6294F864" box="[761,908,551,575]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="7" pageNumber="48" phylum="Chordata" rank="species" species="nepalensis">
<emphasis id="38B8ED4CFFE2FFADFD51FDFD6210F864" box="[761,776,552,575]" italics="true" pageId="7" pageNumber="48">F</emphasis>
. 
<emphasis id="38B8ED4CFFE2FFADFCBEFDF26294F864" box="[790,908,551,575]" italics="true" pageId="7" pageNumber="48">nepalensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>