treatments-xml/data/71/0E/87/710E8799FFFFFFDFFF3DF91CFDDAFCE7.xml
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<mods:title id="7C5865E58A73DBD17F752F96D353598B">Seven new species of the genus Lathrobium Gravenhorst (Coleoptera: Staphylinidae: Paederinae) from Central Honshu, Japan, with new distributional records</mods:title>
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<emphasis id="CBD3EA9DFFFFFFDCFF3DF91CFE69F979" bold="true" box="[151,488,1676,1703]" pageId="16" pageNumber="517">
<taxonomicName id="3EA74D0CFFFFFFDCFF3DF91CFE02F979" authority="Senda, 2024" authorityName="Senda" authorityYear="2024" box="[151,387,1676,1703]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="517" phylum="Arthropoda" rank="species" species="volkeri" status="sp. nov.">
<emphasis id="CBD3EA9DFFFFFFDCFF3DF91CFE02F979" bold="true" box="[151,387,1676,1703]" italics="true" pageId="16" pageNumber="517">Lathrobium volkeri</emphasis>
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<taxonomicNameLabel id="D0E057E6FFFFFFDCFE20F91DFE69F979" box="[394,488,1677,1703]" pageId="16" pageNumber="517" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="F918368FFFFFFFDCFF3DF920FEDEF915" blockId="16.[151,488,1676,1739]" box="[151,351,1712,1739]" pageId="16" pageNumber="517">
(
<figureCitation id="619C2A0AFFFFFFDCFF35F920FE83F915" box="[159,258,1712,1739]" captionStart="FIGURE 1" captionStartId="2.[151,250,1819,1843]" captionTargetBox="[162,1426,188,1793]" captionTargetId="figure-32@2.[151,1436,179,1793]" captionTargetPageId="2" captionText="FIGURE 1. Holotypes of Lathrobium spp., dorsal habitus: L. oopikkarianum sp. nov. (A); L. elongatiphallus sp. nov. (B); L. tomohidei sp. nov. (C); L. pegaseion sp. nov. (D); L. minoccidentale sp. nov. (E); L. kawamurai sp. nov. (F); L. volkeri sp. nov. (G). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10562431" httpUri="https://zenodo.org/record/10562431/files/figure.png" pageId="16" pageNumber="517">Figs. 1G</figureCitation>
,
<figureCitation id="619C2A0AFFFFFFDCFEA5F921FEACF915" box="[271,301,1713,1739]" captionStart="FIGURE 10" captionStartId="18.[151,250,1844,1868]" captionTargetBox="[164,1436,181,1801]" captionTargetId="figure-32@18.[151,1436,181,1819]" captionTargetPageId="18" captionText="FIGURE 10. Lathrobium volkeri sp. nov., male (AF) and female (GH): apical part of abdominal sternites (A, H); aedeagus, ventral view (B); ditto, lateral view (C); ditto, dorsal view (D); inverted endophallus of aedeagus, ventral view (E); ditto, lateral view (F); apical part of abdominal terga (G). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10562456" httpUri="https://zenodo.org/record/10562456/files/figure.png" pageId="16" pageNumber="517">10</figureCitation>
,
<figureCitation id="619C2A0AFFFFFFDCFE93F921FED7F915" box="[313,342,1713,1739]" captionStart="FIGURE 12" captionStartId="21.[151,250,1811,1835]" captionTargetBox="[243,1345,597,1787]" captionTargetId="figure-159@21.[243,1345,597,1787]" captionTargetPageId="21" captionText="FIGURE 12. Geographical distribution of Lathrobium kawamurai sp. nov., L. volkeri sp. nov., L. satoi Watanabe and their related congeners in central Honshu, Japan." figureDoi="http://doi.org/10.5281/zenodo.10562462" httpUri="https://zenodo.org/record/10562462/files/figure.png" pageId="16" pageNumber="517">12</figureCitation>
)
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material.
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:
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<collectingCountry id="81B0761FFFFFFFDCFE65F8ABFDACF88B" box="[463,557,1851,1877]" name="Japan" pageId="16" pageNumber="517">JAPAN</collectingCountry>
: Honshu:
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<specimenCount id="EFA1FD06FFFFFFDCFD05F8ACFD47F888" box="[687,710,1852,1878]" count="1" pageId="16" pageNumber="517" type="male"></specimenCount>
, “
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|
<collectingRegion id="3B63F86DFFFFFFDCFC39F8ABFB92F88B" box="[915,1043,1851,1877]" country="Japan" name="Aichi" pageId="16" pageNumber="517">Aichi Pref.</collectingRegion>
|
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<collectingDate id="9D5DE9A7FFFFFFDCFB84F8ABFB25F88B" box="[1070,1188,1851,1877]" pageId="16" pageNumber="517" value="1970-10-24">24.X.1970</collectingDate>
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|
<collectorName id="54525359FFFFFFDCFB6AF8ACFADAF88B" box="[1216,1371,1851,1878]" pageId="16" pageNumber="517">K. Yamagishi</collectorName>
leg.” [handwritten on white label], “
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|
<taxonomicName id="3EA74D0CFFFFFFDCFD09F8CFFCFCF8A7" authority="Senda, 2024" authorityName="Senda" authorityYear="2024" box="[675,893,1887,1913]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="517" phylum="Arthropoda" rank="species" species="volkeri" status="sp. nov.">
<emphasis id="CBD3EA9DFFFFFFDCFD09F8CFFCFCF8A7" box="[675,893,1887,1913]" italics="true" pageId="16" pageNumber="517">Lathrobium volkeri</emphasis>
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| Des. Y.
<bibRefCitation id="9D364B7EFFFFFFDCFBFAF8CFFB5EF8A7" author="Senda, Y." box="[1104,1247,1887,1913]" pageId="16" pageNumber="517" pagination="142 - 147" refId="ref17288" refString="Senda, Y. (2022) New species, new synonym and new records of the genus Lathrobium Gravenhorst of Eastern Honshu, Japan (Coleoptera: Staphylinidae: Paederinae). Japanese Journal of Systematic Entomology, 28, 142 - 147." type="journal article" year="2022">Senda, 2022</bibRefCitation>
” [printed on red label with black border]. (EUMJ).
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:
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<specimenCount id="EFA1FD06FFFFFFDCFD34F813FD58F843" box="[670,729,1923,1949]" count="3" pageId="16" pageNumber="517" type="male">3♂♂</specimenCount>
,
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,
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,
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,
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,
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,
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, alt.
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,
<collectorName id="54525359FFFFFFDCFD95F838FD0DF81A" box="[575,652,1960,1988]" pageId="16" pageNumber="517">T. Ôno</collectorName>
leg., NIAES-STAPH-JPN560, 561 (pcYS, NARO)
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.
</paragraph>
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<subSubSection id="B1BD6504FFFEFFDDFF6DFF06FCD7FE2B" pageId="17" pageNumber="518" type="diagnosis">
<paragraph id="F918368FFFFEFFDDFF6DFF06FCD7FE2B" blockId="17.[151,1437,150,2013]" pageId="17" pageNumber="518">
<emphasis id="CBD3EA9DFFFEFFDDFF6DFF06FEBBFF6E" bold="true" box="[199,314,150,176]" pageId="17" pageNumber="518">Diagnosis</emphasis>
. The new species is superficially similar to
<taxonomicName id="3EA74D0CFFFEFFDDFCBCFF07FB44FF6F" authority="Sharp, 1889" authorityName="Sharp" authorityYear="1889" box="[790,1221,151,177]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="518" phylum="Arthropoda" rank="species" species="brachypterum">
<emphasis id="CBD3EA9DFFFEFFDDFCBCFF07FBBAFF6F" box="[790,1083,151,177]" italics="true" pageId="17" pageNumber="518">Lathrobium brachypterum</emphasis>
Sharp, 1889
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but distinguishable from it by the following characteristics: 1) male abdominal sternite VIII weakly emarginated at middle of posterior margin; 2) aedeagal ventral process comparatively wide in apical half; 3) dorsal plate of aedeagus with small dorsal projection at apical 1/3 and small hook at apex (while in
<taxonomicName id="3EA74D0CFFFEFFDDFCA9FE93FC3DFEC3" authorityName="Sharp" authorityYear="1889" box="[771,956,259,285]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="518" phylum="Arthropoda" rank="species" species="brachypterum">
<emphasis id="CBD3EA9DFFFEFFDDFCA9FE93FC3DFEC3" box="[771,956,259,285]" italics="true" pageId="17" pageNumber="518">L. brachypterum</emphasis>
</taxonomicName>
: male abdominal sternite VIII subtruncated at middle of posterior margin; ventral process comparatively slender in apical half; dorsal plate of aedeagus with small dorsal projection at apical 1/4 and acute apex). The new species is also resembles
<taxonomicName id="3EA74D0CFFFEFFDDFBC8FEDBFF53FE57" authority="Watanabe, 2003" authorityName="Watanabe" authorityYear="2003" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="518" phylum="Arthropoda" rank="species" species="satoi">
<emphasis id="CBD3EA9DFFFEFFDDFBC8FEDBFAA3FEBA" box="[1122,1314,331,357]" italics="true" pageId="17" pageNumber="518">Lathrobium satoi</emphasis>
<bibRefCitation id="9D364B7EFFFEFFDDFA82FEDBFF53FE57" author="Watanabe, Y." pageId="17" pageNumber="518" pagination="151 - 155" refId="ref17663" refString="Watanabe, Y. (2003) Lathrobium satoi (Coleoptera, Staphylinidae), a new staphylinid beetle from Tokai district in Central Honshu, Japan. Satonius, Special Bulletin of the Japanese Society of Coleopterology, 2, 151 - 155." type="journal article" year="2003">Watanabe, 2003</bibRefCitation>
</taxonomicName>
, but is different from the latter by the following characteristics: 1) aedeagal ventral process weakly curved to right in apical half; 2) dorsal plate of aedeagus comparatively long, with small dorsal projection at apical 1/3 and small hook at apex (while in
<taxonomicName id="3EA74D0CFFFEFFDDFE74FE27FDB3FE0E" box="[478,562,439,464]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="518" phylum="Arthropoda" rank="species" species="satoi">
<emphasis id="CBD3EA9DFFFEFFDDFE74FE27FDB3FE0E" box="[478,562,439,464]" italics="true" pageId="17" pageNumber="518">L. satoi</emphasis>
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: aedeagal ventral process distinctly curved to right in apical half; dorsal plate of aedeagus comparatively short, with small hook at the apex).
</paragraph>
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<emphasis id="CBD3EA9DFFFEFFDDFF6DFE6EFED8FDC6" bold="true" box="[199,345,510,536]" pageId="17" pageNumber="518">Description.</emphasis>
Colouration: Body dark reddish brown to black; antennae, mandibles, labrum, basisternum, meso- and metavantrite reddish brown; labial palpi, maxillae, legs and abdominal segment IX (apical part) and X brownish yellow; elytra usually dark reddish brown in sutural and apical area.
</paragraph>
<paragraph id="F918368FFFFEFFDDFF6DFDFBFD46FC7B" blockId="17.[151,1437,150,2013]" pageId="17" pageNumber="518">
<emphasis id="CBD3EA9DFFFEFFDDFF6DFDFBFE80FD5B" box="[199,257,619,645]" italics="true" pageId="17" pageNumber="518">Male</emphasis>
. Body elongate, subparallel-sided, slightly flattened dorsally, shining. Head subhexagon in dorsal view, lateral sides gently arcuate, widest at basal 1/3 of posterior part of eyes, HL/HW ≈ 1.0; surface with coarse setiferous punctures and coriaceous microsculpture; punctures of surface becoming denser and less coarse in temporal area. Eyes small, about 0.3 times as long as temple, flat.Antennae moderate in length, AL/FBL ≈ 0.6; antennomere I widest at about apical half, dilated to widest point, thence feebly contracted toward apex; antennomere II constricted at the base; IIIX almost moniliform; antennomere XI fusiform, pointed at apex; length/width ratio of each antennomere I to XI as follows: 2.5, 1.7, 1.4, 1.3, 1.4, 1.4, 1.4, 1.4, 1.4, 1.4, 2.1; relative lengths of antennomeres from base to apex: 23.5: 12.5: 11.5: 11.0: 10.3: 10.3: 10.3: 10.3: 10.0: 10.0: 13.8, and relative widths: 14.6: 11.5: 12.3: 13.1: 11.5: 11.5: 11. 5: 11. 5: 11.2: 11.2: 10.0.
</paragraph>
<paragraph id="F918368FFFFEFFDDFF6DFC3FFD3EFB1B" blockId="17.[151,1437,150,2013]" pageId="17" pageNumber="518">Pronotum oblong, widest at anterior 1/5, PL/PW ≈ 1.2, PL/HL ≈ 1.4, PW/HW ≈ 1.2, lateral sides feebly arcuate, contracted from widest point to posterior corner; disc sparsely covered with coarse and setiferous shallow punctures except median smooth area; anterior margin very feebly emarginate at middle, broadly rounded at corners; posterior margin almost straight, rounded at corners. Scutellar shield impunctate, covered with indistinct microsculpture, sparsely covered with fine brownish pubescence. Elytra widest just before posterior corner, expanded weak arcuately from base to posterior corner, EL/EW ≈ 0.7, EL/PL ≈ 0.6, EW/PW ≈ 1.1; disc covered with shallow punctures, sparsely covered with fine brownish pubescence. Hind wings absent. Legs rather slender, covered with golden pubescence; protarsomeres IIV strongly dilated.</paragraph>
<paragraph id="F918368FFFFEFFDDFF6DFB5FFC49FA3B" blockId="17.[151,1437,150,2013]" pageId="17" pageNumber="518">
Abdomen covered with minute punctures, densely covered with dark brownish pubescence; sternite VI (
<figureCitation id="619C2A0AFFFEFFDDFAC5FB5FFF48FAD3" captionStart="FIGURE 10" captionStartId="18.[151,250,1844,1868]" captionTargetBox="[164,1436,181,1801]" captionTargetId="figure-32@18.[151,1436,181,1819]" captionTargetPageId="18" captionText="FIGURE 10. Lathrobium volkeri sp. nov., male (AF) and female (GH): apical part of abdominal sternites (A, H); aedeagus, ventral view (B); ditto, lateral view (C); ditto, dorsal view (D); inverted endophallus of aedeagus, ventral view (E); ditto, lateral view (F); apical part of abdominal terga (G). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10562456" httpUri="https://zenodo.org/record/10562456/files/figure.png" pageId="17" pageNumber="518">Fig. 10A</figureCitation>
) without median depression; sternite VII (
<figureCitation id="619C2A0AFFFEFFDDFD0DFB63FC8FFAD3" box="[679,782,1267,1293]" captionStart="FIGURE 10" captionStartId="18.[151,250,1844,1868]" captionTargetBox="[164,1436,181,1801]" captionTargetId="figure-32@18.[151,1436,181,1819]" captionTargetPageId="18" captionText="FIGURE 10. Lathrobium volkeri sp. nov., male (AF) and female (GH): apical part of abdominal sternites (A, H); aedeagus, ventral view (B); ditto, lateral view (C); ditto, dorsal view (D); inverted endophallus of aedeagus, ventral view (E); ditto, lateral view (F); apical part of abdominal terga (G). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10562456" httpUri="https://zenodo.org/record/10562456/files/figure.png" pageId="17" pageNumber="518">Fig. 10A</figureCitation>
) without median depression, but covered with short black setae in the posteromedial area, not emarginate at posterior margin; tergum VIII feebly arcuate at posterior margin; sternite VIII (
<figureCitation id="619C2A0AFFFEFFDDFE9BFAABFE16FA8B" box="[305,407,1339,1365]" captionStart="FIGURE 10" captionStartId="18.[151,250,1844,1868]" captionTargetBox="[164,1436,181,1801]" captionTargetId="figure-32@18.[151,1436,181,1819]" captionTargetPageId="18" captionText="FIGURE 10. Lathrobium volkeri sp. nov., male (AF) and female (GH): apical part of abdominal sternites (A, H); aedeagus, ventral view (B); ditto, lateral view (C); ditto, dorsal view (D); inverted endophallus of aedeagus, ventral view (E); ditto, lateral view (F); apical part of abdominal terga (G). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10562456" httpUri="https://zenodo.org/record/10562456/files/figure.png" pageId="17" pageNumber="518">Fig. 10A</figureCitation>
) with triangular median depression, which is covered with short black setae, with posterior margin weakly emarginated at middle; tergum IX setigerous except on basal 1/4, with posterolateral projections as long as 1/4 of tergum IX and slightly curved dorsally from apical 1/3 to pointed apex; sternite IX elongate, widest at basal 1/4, narrowed apically, setigerous in 2/3, with posterior margin feebly arcuate; tergum X trapezoidal, widest at basal corners, with few setae and pores, with posterior margin arcuate.
</paragraph>
<paragraph id="F918368FFFFEFFDDFF6DFA7FFE15F8F7" blockId="17.[151,1437,150,2013]" pageId="17" pageNumber="518">
Aedeagus (
<figureCitation id="619C2A0AFFFEFFDDFEE6FA7FFE58F9D7" box="[332,473,1519,1545]" captionStart="FIGURE 10" captionStartId="18.[151,250,1844,1868]" captionTargetBox="[164,1436,181,1801]" captionTargetId="figure-32@18.[151,1436,181,1819]" captionTargetPageId="18" captionText="FIGURE 10. Lathrobium volkeri sp. nov., male (AF) and female (GH): apical part of abdominal sternites (A, H); aedeagus, ventral view (B); ditto, lateral view (C); ditto, dorsal view (D); inverted endophallus of aedeagus, ventral view (E); ditto, lateral view (F); apical part of abdominal terga (G). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10562456" httpUri="https://zenodo.org/record/10562456/files/figure.png" pageId="17" pageNumber="518">Fig. 10BD</figureCitation>
) robust,
<quantity id="3E5F9B6AFFFEFFDDFDE0FA7FFD34F9D7" box="[586,693,1519,1545]" metricMagnitude="-3" metricUnit="m" metricValue="1.83" pageId="17" pageNumber="518" unit="mm" value="1.83">1.83 mm</quantity>
long, well sclerotised except membranous dorsal side; dorsal plate elongate and slender, symmetric, widest at apical 1/3, weakly curved ventrally at apical 1/3, gently dilated emarginately to widest point, thence emarginately contracted toward acute apex in dorsal view, with small dorsal projection at apical 1/3 and small hook at the apex; ventral process slightly asymmetrical, strongly curved ventrally at apical half, weakly curved to right in apical half, almost parallel-sided in basal 1/3, thence strongly contracted toward acute apex in ventral view, with fine longitudinal carina along midline in apical 1/3. Endophallus (
<figureCitation id="619C2A0AFFFEFFDDFAE1F933FF4AF93F" captionStart="FIGURE 10" captionStartId="18.[151,250,1844,1868]" captionTargetBox="[164,1436,181,1801]" captionTargetId="figure-32@18.[151,1436,181,1819]" captionTargetPageId="18" captionText="FIGURE 10. Lathrobium volkeri sp. nov., male (AF) and female (GH): apical part of abdominal sternites (A, H); aedeagus, ventral view (B); ditto, lateral view (C); ditto, dorsal view (D); inverted endophallus of aedeagus, ventral view (E); ditto, lateral view (F); apical part of abdominal terga (G). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10562456" httpUri="https://zenodo.org/record/10562456/files/figure.png" pageId="17" pageNumber="518">Fig. 10 EF</figureCitation>
): slightly longer than dorsal plate, simple, symmetric, without any membranous projection in everted condition, with ventrolateral teeth patch as
<figureCitation id="619C2A0AFFFEFFDDFE50F97BFDFBF8DB" box="[506,634,1771,1797]" captionStart="FIGURE 10" captionStartId="18.[151,250,1844,1868]" captionTargetBox="[164,1436,181,1801]" captionTargetId="figure-32@18.[151,1436,181,1819]" captionTargetPageId="18" captionText="FIGURE 10. Lathrobium volkeri sp. nov., male (AF) and female (GH): apical part of abdominal sternites (A, H); aedeagus, ventral view (B); ditto, lateral view (C); ditto, dorsal view (D); inverted endophallus of aedeagus, ventral view (E); ditto, lateral view (F); apical part of abdominal terga (G). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10562456" httpUri="https://zenodo.org/record/10562456/files/figure.png" pageId="17" pageNumber="518">Fig. 10EF</figureCitation>
; sclerotised copulatory piece lacking, but with numerous minute sclerites on dorsal side of apex.
</paragraph>
<paragraph id="F918368FFFFEFFDDFF6DF8A3FE55F803" blockId="17.[151,1437,150,2013]" pageId="17" pageNumber="518">
<emphasis id="CBD3EA9DFFFEFFDDFF6DF8A3FE9AF893" box="[199,283,1843,1869]" italics="true" pageId="17" pageNumber="518">Female</emphasis>
. Similar to male in general appearance. Length/width ratio of each antennomere I to XI as follows: 2.9, 1.2, 1.4, 1.3, 1.3, 1.3, 1.3, 1.2, 1.3, 1.3, 2.4; relative lengths of antennomeres from base to apex: 26.7: 10.0: 11.7: 10.8: 10.8: 10.8: 10.8: 10.0: 10.0: 10.0: 15.8, and relative widths: 13.8: 12.5: 12.5: 12.5: 12.5: 12.5: 12.5: 12.5: 11.3: 11.3: 10.0. HL/HW ≈ 1.0; AL/FBL ≈ 0.6; PL/PW ≈ 1.2; PL/HL ≈ 1.3; PW/HW ≈ 1.1; EL/EW ≈ 0.8; EL/PL ≈ 0.7; EW/PW ≈ 1.1.
</paragraph>
<caption id="ADD86607FFFDFFDEFF3DF8A4FCB9F84D" ID-DOI="http://doi.org/10.5281/zenodo.10562456" ID-Zenodo-Dep="10562456" httpUri="https://zenodo.org/record/10562456/files/figure.png" pageId="18" pageNumber="519" startId="18.[151,250,1844,1868]" targetBox="[164,1436,181,1801]" targetPageId="18" targetType="figure">
<paragraph id="F918368FFFFDFFDEFF3DF8A4FCB9F84D" blockId="18.[151,1436,1844,1940]" pageId="18" pageNumber="519">
<emphasis id="CBD3EA9DFFFDFFDEFF3DF8A4FEA1F892" bold="true" box="[151,288,1844,1868]" pageId="18" pageNumber="519">FIGURE 10.</emphasis>
<taxonomicName id="3EA74D0CFFFDFFDEFE8CF8A4FE6BF892" authority="Senda, 2024" authorityName="Senda" authorityYear="2024" box="[294,490,1844,1868]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="519" phylum="Arthropoda" rank="species" species="volkeri" status="sp. nov.">
<emphasis id="CBD3EA9DFFFDFFDEFE8CF8A4FE6BF892" box="[294,490,1844,1868]" italics="true" pageId="18" pageNumber="519">Lathrobium volkeri</emphasis>
</taxonomicName>
<emphasis id="CBD3EA9DFFFDFFDEFE5AF8A4FDC0F892" bold="true" box="[496,577,1844,1868]" pageId="18" pageNumber="519">
<taxonomicNameLabel id="D0E057E6FFFDFFDEFE5AF8A4FDC0F892" box="[496,577,1844,1868]" pageId="18" pageNumber="519" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male (AF) and female (GH): apical part of abdominal sternites (A, H); aedeagus, ventral view (B); ditto, lateral view (C); ditto, dorsal view (D); inverted endophallus of aedeagus, ventral view (E); ditto, lateral view (F); apical part of abdominal terga (G). Scale bars = 0.5 mm.
</paragraph>
</caption>
<paragraph id="F918368FFFFCFFDFFF6DFF07FDA2FE9F" blockId="19.[151,1437,151,825]" pageId="19" pageNumber="520">
Tergum VIII (
<figureCitation id="619C2A0AFFFCFFDFFECCFF07FE4FFF6F" box="[358,462,151,177]" captionStart="FIGURE 10" captionStartId="18.[151,250,1844,1868]" captionTargetBox="[164,1436,181,1801]" captionTargetId="figure-32@18.[151,1436,181,1819]" captionTargetPageId="18" captionText="FIGURE 10. Lathrobium volkeri sp. nov., male (AF) and female (GH): apical part of abdominal sternites (A, H); aedeagus, ventral view (B); ditto, lateral view (C); ditto, dorsal view (D); inverted endophallus of aedeagus, ventral view (E); ditto, lateral view (F); apical part of abdominal terga (G). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10562456" httpUri="https://zenodo.org/record/10562456/files/figure.png" pageId="19" pageNumber="520">Fig. 10G</figureCitation>
) weakly project at middle of posterior margin; sternite VIII (
<figureCitation id="619C2A0AFFFCFFDFFB2DFF07FB6FFF6F" box="[1159,1262,151,177]" captionStart="FIGURE 10" captionStartId="18.[151,250,1844,1868]" captionTargetBox="[164,1436,181,1801]" captionTargetId="figure-32@18.[151,1436,181,1819]" captionTargetPageId="18" captionText="FIGURE 10. Lathrobium volkeri sp. nov., male (AF) and female (GH): apical part of abdominal sternites (A, H); aedeagus, ventral view (B); ditto, lateral view (C); ditto, dorsal view (D); inverted endophallus of aedeagus, ventral view (E); ditto, lateral view (F); apical part of abdominal terga (G). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10562456" httpUri="https://zenodo.org/record/10562456/files/figure.png" pageId="19" pageNumber="520">Fig. 10H</figureCitation>
) subtruncate at middle of arcuate posterior margin; tergum IX (
<figureCitation id="619C2A0AFFFCFFDFFD0EFF2BFC88FF0B" box="[676,777,187,213]" captionStart="FIGURE 10" captionStartId="18.[151,250,1844,1868]" captionTargetBox="[164,1436,181,1801]" captionTargetId="figure-32@18.[151,1436,181,1819]" captionTargetPageId="18" captionText="FIGURE 10. Lathrobium volkeri sp. nov., male (AF) and female (GH): apical part of abdominal sternites (A, H); aedeagus, ventral view (B); ditto, lateral view (C); ditto, dorsal view (D); inverted endophallus of aedeagus, ventral view (E); ditto, lateral view (F); apical part of abdominal terga (G). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10562456" httpUri="https://zenodo.org/record/10562456/files/figure.png" pageId="19" pageNumber="520">Fig. 10G</figureCitation>
) similar to male, with thicker posterolateral projections and large subtriangular flap-like ventral plate on the base; tergum X (
<figureCitation id="619C2A0AFFFCFFDFFCCFFF4FFC4BFF27" box="[869,970,223,249]" captionStart="FIGURE 10" captionStartId="18.[151,250,1844,1868]" captionTargetBox="[164,1436,181,1801]" captionTargetId="figure-32@18.[151,1436,181,1819]" captionTargetPageId="18" captionText="FIGURE 10. Lathrobium volkeri sp. nov., male (AF) and female (GH): apical part of abdominal sternites (A, H); aedeagus, ventral view (B); ditto, lateral view (C); ditto, dorsal view (D); inverted endophallus of aedeagus, ventral view (E); ditto, lateral view (F); apical part of abdominal terga (G). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10562456" httpUri="https://zenodo.org/record/10562456/files/figure.png" pageId="19" pageNumber="520">Fig. 10G</figureCitation>
) similar to male. Gonocoxites (
<figureCitation id="619C2A0AFFFCFFDFFA8DFF4FFA0EFF27" box="[1319,1423,223,249]" captionStart="FIGURE 10" captionStartId="18.[151,250,1844,1868]" captionTargetBox="[164,1436,181,1801]" captionTargetId="figure-32@18.[151,1436,181,1819]" captionTargetPageId="18" captionText="FIGURE 10. Lathrobium volkeri sp. nov., male (AF) and female (GH): apical part of abdominal sternites (A, H); aedeagus, ventral view (B); ditto, lateral view (C); ditto, dorsal view (D); inverted endophallus of aedeagus, ventral view (E); ditto, lateral view (F); apical part of abdominal terga (G). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10562456" httpUri="https://zenodo.org/record/10562456/files/figure.png" pageId="19" pageNumber="520">Fig. 10H</figureCitation>
): lateral gonocoxal plate (distal and proximal gonocoxites fused) elongate, setigerous, with a few setae in apical area, with asymmetrically rounded apex.
</paragraph>
<paragraph id="F918368FFFFCFFDFFF6DFEDBFE7EFE0F" blockId="19.[151,1437,151,825]" pageId="19" pageNumber="520">
<emphasis id="CBD3EA9DFFFCFFDFFF6DFEDBFEF0FEBB" bold="true" box="[199,369,331,357]" italics="true" pageId="19" pageNumber="520">Measurements</emphasis>
. Male (n = 3)—BL:
<quantity id="3E5F9B6AFFFCFFDFFDFCFEDBFD78FEBB" box="[598,761,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="7.755" metricValueMax="7.83" metricValueMin="7.68" pageId="19" pageNumber="520" unit="mm" value="7.755" valueMax="7.83" valueMin="7.68">7.687.83 mm</quantity>
(7.77±
<quantity id="3E5F9B6AFFFCFFDFFCEDFEDBFC2EFEBB" box="[839,943,331,357]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="19" pageNumber="520" unit="mm" value="0.08">0.08 mm</quantity>
); FBL:
<quantity id="3E5F9B6AFFFCFFDFFBA9FEDBFB27FEBB" box="[1027,1190,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="3.2199999999999998" metricValueMax="3.32" metricValueMin="3.12" pageId="19" pageNumber="520" unit="mm" value="3.2199999999999998" valueMax="3.32" valueMin="3.12">3.123.32 mm</quantity>
(3.23±
<quantity id="3E5F9B6AFFFCFFDFFB59FEDBFADDFEBB" box="[1267,1372,331,357]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="520" unit="mm" value="0.1">0.10 mm</quantity>
); HL:
<quantity id="3E5F9B6AFFFCFFDFFF3DFEFFFEBCFE57" box="[151,317,367,393]" metricMagnitude="-4" metricUnit="m" metricValue="9.7" metricValueMax="10.1" metricValueMin="9.3" pageId="19" pageNumber="520" unit="mm" value="0.97" valueMax="1.01" valueMin="0.93">0.931.01 mm</quantity>
(0.97±
<quantity id="3E5F9B6AFFFCFFDFFE24FEFFFE7BFE57" box="[398,506,367,393]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="19" pageNumber="520" unit="mm" value="0.04">0.04 mm</quantity>
); HW:
<quantity id="3E5F9B6AFFFCFFDFFDE4FEFFFD75FE57" box="[590,756,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="1.015" metricValueMax="1.06" metricValueMin="0.97" pageId="19" pageNumber="520" unit="mm" value="1.015" valueMax="1.06" valueMin="0.97">0.971.06 mm</quantity>
(1.00±
<quantity id="3E5F9B6AFFFCFFDFFCEFFEFEFC30FE57" box="[837,945,366,393]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="19" pageNumber="520" unit="mm" value="0.05">0.05 mm</quantity>
); AL:
<quantity id="3E5F9B6AFFFCFFDFFC51FEFFFB20FE57" box="[1019,1185,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="1.845" metricValueMax="1.88" metricValueMin="1.81" pageId="19" pageNumber="520" unit="mm" value="1.845" valueMax="1.88" valueMin="1.81">1.811.88 mm</quantity>
(1.86±
<quantity id="3E5F9B6AFFFCFFDFFB5BFEFFFADFFE57" box="[1265,1374,367,393]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="19" pageNumber="520" unit="mm" value="0.04">0.04 mm</quantity>
); PL:
<quantity id="3E5F9B6AFFFCFFDFFF3DFE03FEBDFE73" box="[151,316,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="1.375" metricValueMax="1.41" metricValueMin="1.34" pageId="19" pageNumber="520" unit="mm" value="1.375" valueMax="1.41" valueMin="1.34">1.341.41 mm</quantity>
(1.37±
<quantity id="3E5F9B6AFFFCFFDFFE26FE03FE78FE73" box="[396,505,403,429]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="19" pageNumber="520" unit="mm" value="0.04">0.04 mm</quantity>
); PW:
<quantity id="3E5F9B6AFFFCFFDFFDE3FE03FD6CFE73" box="[585,749,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.19" metricValueMin="1.11" pageId="19" pageNumber="520" unit="mm" value="1.15" valueMax="1.19" valueMin="1.11">1.111.19 mm</quantity>
(1.15±
<quantity id="3E5F9B6AFFFCFFDFFC97FE02FC28FE73" box="[829,937,402,429]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="19" pageNumber="520" unit="mm" value="0.04">0.04 mm</quantity>
); EL:
<quantity id="3E5F9B6AFFFCFFDFFC58FE03FB16FE73" box="[1010,1175,403,429]" metricMagnitude="-4" metricUnit="m" metricValue="8.899999999999999" metricValueMax="9.399999999999999" metricValueMin="8.4" pageId="19" pageNumber="520" unit="mm" value="0.8899999999999999" valueMax="0.94" valueMin="0.84">0.840.94 mm</quantity>
(0.89±
<quantity id="3E5F9B6AFFFCFFDFFB43FE02FAD5FE73" box="[1257,1364,402,429]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="19" pageNumber="520" unit="mm" value="0.05">0.05 mm</quantity>
); EW:
<quantity id="3E5F9B6AFFFCFFDFFF3DFE27FEBBFE0F" box="[151,314,439,465]" metricMagnitude="-3" metricUnit="m" metricValue="1.21" metricValueMax="1.23" metricValueMin="1.19" pageId="19" pageNumber="520" unit="mm" value="1.21" valueMax="1.23" valueMin="1.19">1.191.23 mm</quantity>
(1.22±
<quantity id="3E5F9B6AFFFCFFDFFE22FE27FE73FE0F" box="[392,498,439,465]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="520" unit="mm" value="0.02">0.02 mm</quantity>
).
</paragraph>
<paragraph id="F918368FFFFCFFDFFF6DFE4BFED4FDBF" blockId="19.[151,1437,151,825]" pageId="19" pageNumber="520">
Female (n = 5)—BL:
<quantity id="3E5F9B6AFFFCFFDFFE6EFE4BFDE8FE2B" box="[452,617,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="7.654999999999999" metricValueMax="7.84" metricValueMin="7.47" pageId="19" pageNumber="520" unit="mm" value="7.654999999999999" valueMax="7.84" valueMin="7.47">7.477.84 mm</quantity>
(7.71±
<quantity id="3E5F9B6AFFFCFFDFFD10FE4AFCA7FE2B" box="[698,806,474,501]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="520" unit="mm" value="0.15">0.15 mm</quantity>
); FBL:
<quantity id="3E5F9B6AFFFCFFDFFC2AFE4BFBA5FE2B" box="[896,1060,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="3.195" metricValueMax="3.28" metricValueMin="3.11" pageId="19" pageNumber="520" unit="mm" value="3.195" valueMax="3.28" valueMin="3.11">3.113.28 mm</quantity>
(3.20±
<quantity id="3E5F9B6AFFFCFFDFFBDFFE4BFB61FE2B" box="[1141,1248,475,501]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="19" pageNumber="520" unit="mm" value="0.07">0.07 mm</quantity>
); HL:
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(0.96±
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); HW:
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(0.96±
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); AL:
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(1.81±
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); PL:
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(1.28±
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); PW:
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(1.06±
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); EL:
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(0.95±
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); EW:
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(1.15±
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).
</paragraph>
</subSubSection>
<subSubSection id="B1BD6504FFFCFFDFFF6DFDFAFB15FD5B" box="[199,1172,618,645]" pageId="19" pageNumber="520" type="distribution">
<paragraph id="F918368FFFFCFFDFFF6DFDFAFB15FD5B" blockId="19.[151,1437,151,825]" box="[199,1172,618,645]" pageId="19" pageNumber="520">
<emphasis id="CBD3EA9DFFFCFFDFFF6DFDFAFED8FD5A" bold="true" box="[199,345,618,644]" pageId="19" pageNumber="520">Distribution</emphasis>
.
<collectingCountry id="81B0761FFFFCFFDFFECDFDFBFE2AFD5B" box="[359,427,619,645]" name="Japan" pageId="19" pageNumber="520">Japan</collectingCountry>
: Central Honshu (northwestern area of
<collectingRegion id="3B63F86DFFFCFFDFFCC8FDFBFB9CFD5B" box="[866,1053,619,645]" country="Japan" name="Aichi" pageId="19" pageNumber="520">Aichi Prefecture</collectingRegion>
) (
<figureCitation id="619C2A0AFFFCFFDFFB99FDFBFB06FD5B" box="[1075,1159,619,645]" captionStart="FIGURE 12" captionStartId="21.[151,250,1811,1835]" captionTargetBox="[243,1345,597,1787]" captionTargetId="figure-159@21.[243,1345,597,1787]" captionTargetPageId="21" captionText="FIGURE 12. Geographical distribution of Lathrobium kawamurai sp. nov., L. volkeri sp. nov., L. satoi Watanabe and their related congeners in central Honshu, Japan." figureDoi="http://doi.org/10.5281/zenodo.10562462" httpUri="https://zenodo.org/record/10562462/files/figure.png" pageId="19" pageNumber="520">Fig. 12</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B1BD6504FFFCFFDFFF6DFD1FFD8DFD2F" pageId="19" pageNumber="520" type="etymology">
<paragraph id="F918368FFFFCFFDFFF6DFD1FFD8DFD2F" blockId="19.[151,1437,151,825]" pageId="19" pageNumber="520">
<emphasis id="CBD3EA9DFFFCFFDFFF6DFD1FFEC9FD77" bold="true" box="[199,328,655,681]" pageId="19" pageNumber="520">Etymology</emphasis>
. The specific name is dedicated to late Dr Volker Assing (Hannover), whose lifelong contributions to the taxonomy of Palaearctic and Oriental staphylinid beetles have been invaluable. Furthermore, this paper was started in collaboration with him.
</paragraph>
</subSubSection>
<subSubSection id="B1BD6504FFFCFFDFFF6DFD6AFDDAFCE7" pageId="19" pageNumber="520" type="discussion">
<paragraph id="F918368FFFFCFFDFFF6DFD6AFDDAFCE7" blockId="19.[151,1437,151,825]" pageId="19" pageNumber="520">
<emphasis id="CBD3EA9DFFFCFFDFFF6DFD6AFEC9FCCA" bold="true" box="[199,328,762,788]" pageId="19" pageNumber="520">Comments</emphasis>
. This new species is unequivocally classified in the
<taxonomicName id="3EA74D0CFFFCFFDFFC24FD6BFB35FCCB" authorityName="Sharp" authorityYear="1889" box="[910,1204,763,789]" class="Insecta" family="Staphylinidae" genus="Lathrobium" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="520" phylum="Arthropoda" rank="species" species="brachypterum">
<emphasis id="CBD3EA9DFFFCFFDFFC24FD6BFB35FCCB" box="[910,1204,763,789]" italics="true" pageId="19" pageNumber="520">Lathrobium brachypterum</emphasis>
</taxonomicName>
group (sensu
<bibRefCitation id="9D364B7EFFFCFFDFFAFBFD6BFF5EFCE7" author="Senda, Y." pageId="19" pageNumber="520" pagination="51 - 56" refId="ref17371" refString="Senda, Y. (2023 b) Lathrobium motonari, a new soil-inhabiting species of the genus Lathrobium (Coleoptera: Staphylinidae: Paederinae) from western Honshu, Japan. Journal of Insect Biodiversity, 43 (2), 51 - 56. https: // doi. org / 10.12976 / jib / 2023.43.2.2" type="journal article" year="2023">Senda, 2023b</bibRefCitation>
) based on its characteristic states.
</paragraph>
</subSubSection>
</treatment>
</document>