147 lines
17 KiB
XML
147 lines
17 KiB
XML
<document id="19A34D127D28C9A13D6C60DBBC4AD058" ID-DOI="10.5281/zenodo.169943" ID-GBIF-Dataset="a80741a3-d6db-4474-a7e6-79293fb0e80f" ID-ISSN="11755326" ID-Zenodo-Dep="169943" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459748760006" checkinUser="plazi" docAuthor="Will, Kipling W." docDate="2005" docId="5D21241C5C7B486D3F184CD7FE22F9C9" docLanguage="en" docName="zt01049p017.pdf" docOrigin="Zootaxa 1049" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Stolonis scortensis Will, new species" docType="treatment" docVersion="8" lastPageNumber="11" masterDocId="A1185C645C7348673E104913FFD3FFC7" masterDocTitle="The Neotropical genera Oxycrepis Reiche and Stolonis Motschulsky: a taxonomic review, key to the described species and description of new Stolonis species from Ecuador (Coleoptera: Carabidae: Loxandrini)" masterLastPageNumber="17" masterPageNumber="1" pageNumber="9" updateTime="1698211146980" updateUser="plazi">
|
||
<mods:mods id="ACF24349A0C521BF9D582BCEE44EECE8" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="03D44B81180A0CED1E7D377C9C6B4712">
|
||
<mods:title id="29274CA9EB674BA5113BF8751AA54AFE">The Neotropical genera Oxycrepis Reiche and Stolonis Motschulsky: a taxonomic review, key to the described species and description of new Stolonis species from Ecuador (Coleoptera: Carabidae: Loxandrini)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="254F39BEC8A5A6E61495D9C4D4515C09" type="personal">
|
||
<mods:role id="17ECD6AC4DDC1100A81F8BD6F9020F5F">
|
||
<mods:roleTerm id="9B9220BFA74FB118268C414243F05007">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="D624B23FF53C7855DB1CD80B81B6D75F">Will, Kipling W.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="91951904AA2219D8316452665767033F">text</mods:typeOfResource>
|
||
<mods:relatedItem id="9A3C8013F51880D9B424AE2AAF3F395E" type="host">
|
||
<mods:titleInfo id="69133C9D57272A93AD2F1E5CAE8FB59C">
|
||
<mods:title id="D172C6E5B5C61BDABD93E68B3F8FD74E">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="032767F297F68485B2889BF30940BFBC">
|
||
<mods:date id="B3046D0BBE92850430D25C8B268D5A5B">2005</mods:date>
|
||
<mods:detail id="A5390259F3C9F4D08D9EA0830D49B586" type="volume">
|
||
<mods:number id="2620EE46C0F07A8B510AA6D4DCC8E8B4">1049</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="373F475FB504C0FA6BD321D9BB1DA36C" unit="page">
|
||
<mods:start id="57B7E6D4099E163A1EEEB52CFF59E67F">1</mods:start>
|
||
<mods:end id="D858C9E5BF2E4CCEF0F10A26A6325748">17</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="9D1ACCCBBCB5486AD29DBCC8F60312EC">journal article</mods:classification>
|
||
<mods:identifier id="06B229F81A633AA1C0F126F34614E5E8" type="DOI">10.5281/zenodo.169943</mods:identifier>
|
||
<mods:identifier id="8073F0B481B935D3C88E1C21947D526B" type="GBIF-Dataset">a80741a3-d6db-4474-a7e6-79293fb0e80f</mods:identifier>
|
||
<mods:identifier id="D7CD3672793666FC4B57FC19F532DE34" type="ISSN">11755326</mods:identifier>
|
||
<mods:identifier id="14081E6C848FF94DCE981F4AEBF8B70A" type="Zenodo-Dep">169943</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="5D21241C5C7B486D3F184CD7FE22F9C9" ID-DOI="http://doi.org/10.5281/zenodo.6266240" ID-GBIF-Taxon="119337164" ID-Zenodo-Dep="6266240" LSID="urn:lsid:plazi:treatment:5D21241C5C7B486D3F184CD7FE22F9C9" httpUri="http://treatment.plazi.org/id/5D21241C5C7B486D3F184CD7FE22F9C9" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
|
||
<subSubSection id="9D92C6815C7B486F3F184CD7FE02F9C6" pageId="8" pageNumber="9" type="nomenclature">
|
||
<paragraph id="D537950A5C7B486F3F184CD7FD11FA1A" blockId="8.[264,1324,1475,1897]" box="[264,706,1475,1502]" pageId="8" pageNumber="9">
|
||
<heading id="8E7F22665C7B486F3F184CD7FD11FA1A" bold="true" box="[264,706,1475,1502]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
|
||
<emphasis id="E7FC49185C7B486F3F184CD7FD11FA1A" bold="true" box="[264,706,1475,1502]" pageId="8" pageNumber="9">
|
||
<taxonomicName id="1288EE895C7B486F3F184CD7FDF6FA1A" ID-CoL="8LNLR" authority="Will" authorityName="Will" box="[264,549,1475,1502]" class="Insecta" family="Carabidae" genus="Stolonis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="scortensis" status="sp. nov.">
|
||
<emphasis id="E7FC49185C7B486F3F184CD7FE34FA19" bold="true" box="[264,487,1476,1502]" italics="true" pageId="8" pageNumber="9">Stolonis scortensis</emphasis>
|
||
Will
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="FCCFF4635C7B486F3C224CD0FD11FA1A" box="[562,706,1475,1501]" pageId="8" pageNumber="9" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="D537950A5C7B486F3F184CF4FE02F9C6" blockId="8.[264,1324,1475,1897]" box="[264,465,1511,1537]" pageId="8" pageNumber="9">
|
||
(
|
||
<figureCitation id="4DB3898F5C7B486F3F014CF4FEB0F9C6" box="[273,355,1511,1537]" captionStart="FIGURE 2" captionStartId="8.[264,367,1326,1350]" captionTargetBox="[337,1321,462,1293]" captionTargetId="figure@8.[267,1326,452,1303]" captionTargetPageId="8" captionText="FIGURE 2. Pronota of: A— Stolonis tapiai, B— Stolonis scortensis, C— Stolonis spinosus, D— Stolonis yasuni, E— Stolonis catenarius." httpUri="https://zenodo.org/record/169945/files/figure.png" pageId="8" pageNumber="9">Figs. 2</figureCitation>
|
||
B, 3E–H)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9D92C6815C7B486F3F184F1DFE09F936" pageId="8" pageNumber="9" type="materials_examined">
|
||
<paragraph id="D537950A5C7B486F3F184F1DFE09F936" blockId="8.[264,1324,1475,1897]" pageId="8" pageNumber="9">
|
||
<emphasis id="E7FC49185C7B486F3F184F1DFE6DF9EF" bold="true" box="[264,446,1550,1576]" pageId="8" pageNumber="9">
|
||
<typeStatus id="0A332BA85C7B486F3F184F1DFE90F9EF" box="[264,323,1550,1576]" pageId="8" pageNumber="9">Type</typeStatus>
|
||
Material
|
||
</emphasis>
|
||
.
|
||
<typeStatus id="0A332BA85C7B486F3FC54F1CFDA7F9EE" box="[469,628,1551,1577]" pageId="8" pageNumber="9" type="holotype">HOLOTYPE</typeStatus>
|
||
: ɗ (
|
||
<collectionCode id="B3990DCF5C7B486F3CBF4F1CFCD4F9EE" LSID="urn:lsid:biocol.org:col:1013" box="[687,775,1551,1577]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="8" pageNumber="9">EMEC</collectionCode>
|
||
), dissected genitalia in vial, portions of the abdomen and a single hind leg stored in ethanol separate from point mounted body and genitalia. Labelled: “
|
||
<collectingCountry id="AD9FD59A5C7B486F3C144F4CFD49F9BE" box="[516,666,1631,1657]" name="Ecuador" pageId="8" pageNumber="9">ECUADOR</collectingCountry>
|
||
: SUCUMBIOS: Reserva Faunistica Cuyabeno, Nuevo Mundo cabins along Rio Cuyabeno at jcn with Lago Agrio–Tipishca HWY
|
||
<date id="A136B3CA5C7B486F3A954F94FAFFF966" box="[1157,1324,1671,1697]" pageId="8" pageNumber="9" value="1994-04-19" valueMax="1994-04-29" valueMin="1994-04-19">19–29.iv.1994</date>
|
||
W. Maddison #WPM 94021”/ “
|
||
<collectionCode id="B3990DCF5C7B486F3C8F4FBCFD25F90E" LSID="urn:lsid:biocol.org:col:1013" box="[671,758,1711,1737]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="8" pageNumber="9">EMEC</collectionCode>
|
||
1004673”. Ethanol stored portion additionally labeled "kww38".
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9D92C6815C7B486F3F284FEDFE73F8AE" pageId="8" pageNumber="9" type="etymology">
|
||
<paragraph id="D537950A5C7B486F3F284FEDFE73F8AE" blockId="8.[264,1324,1475,1897]" pageId="8" pageNumber="9">
|
||
<emphasis id="E7FC49185C7B486F3F284FEDFE6EF8DF" bold="true" box="[312,445,1790,1816]" pageId="8" pageNumber="9">Etymology</emphasis>
|
||
. Specific epithet formed by the junction of two Latin words,
|
||
<emphasis id="E7FC49185C7B486F3A9A4FECFEF8F887" italics="true" pageId="8" pageNumber="9">scorteus, a, um</emphasis>
|
||
, "leather" and
|
||
<emphasis id="E7FC49185C7B486F3FF44E34FD86F887" box="[484,597,1831,1856]" italics="true" pageId="8" pageNumber="9">ensis, is,</emphasis>
|
||
m "sword" and refers to the leathery rugosities of the male median lobe.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="81F7C5825C7A486E3F184FA7FE53F8D6" httpUri="https://zenodo.org/record/169946/files/figure.png" pageId="9" pageNumber="10" targetBox="[284,1304,346,1671]" targetPageId="9">
|
||
<paragraph id="D537950A5C7A486E3F184FA7FE53F8D6" blockId="9.[264,1323,1716,1809]" pageId="9" pageNumber="10">
|
||
<emphasis id="E7FC49185C7A486E3F184FA7FE58F90B" bold="true" box="[264,395,1716,1740]" pageId="9" pageNumber="10">FIGURE 3.</emphasis>
|
||
Aedeagus dorsal, left lateral, right lateral and ventral views of: A–D—
|
||
<taxonomicName id="1288EE895C7A486E3A994FA6FAF7F90B" box="[1161,1316,1717,1740]" class="Insecta" family="Carabidae" genus="Stolonis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="tapiai">
|
||
<emphasis id="E7FC49185C7A486E3A994FA6FAF7F90B" box="[1161,1316,1717,1740]" italics="true" pageId="9" pageNumber="10">Stolonis tapiai</emphasis>
|
||
</taxonomicName>
|
||
, E–H—
|
||
<taxonomicName id="1288EE895C7A486E3F444FC4FDF9F929" box="[340,554,1751,1774]" class="Insecta" family="Carabidae" genus="Stolonis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="scortensis">
|
||
<emphasis id="E7FC49185C7A486E3F444FC4FDF9F929" box="[340,554,1751,1774]" italics="true" pageId="9" pageNumber="10">Stolonis scortensis</emphasis>
|
||
</taxonomicName>
|
||
, I–L—
|
||
<taxonomicName id="1288EE895C7A486E3C984FC4FC83F929" box="[648,848,1751,1774]" class="Insecta" family="Carabidae" genus="Stolonis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="spinosus">
|
||
<emphasis id="E7FC49185C7A486E3C984FC4FC83F929" box="[648,848,1751,1774]" italics="true" pageId="9" pageNumber="10">Stolonis spinosus</emphasis>
|
||
</taxonomicName>
|
||
, M–P—
|
||
<taxonomicName id="1288EE895C7A486E3DAC4FC4FBBEF929" box="[956,1133,1751,1774]" class="Insecta" family="Carabidae" genus="Stolonis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="yasuni">
|
||
<emphasis id="E7FC49185C7A486E3DAC4FC4FBBEF929" box="[956,1133,1751,1774]" italics="true" pageId="9" pageNumber="10">Stolonis yasuni</emphasis>
|
||
</taxonomicName>
|
||
, Q–T—
|
||
<taxonomicName id="1288EE895C7A486E3AC54FC4FEABF8D6" class="Insecta" family="Carabidae" genus="Stolonis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="catenarius">
|
||
<emphasis id="E7FC49185C7A486E3AC54FC4FEABF8D6" italics="true" pageId="9" pageNumber="10">Stolonis catenarius</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="9D92C6815C79486D3F284808FCF8FE69" pageId="10" pageNumber="11" type="diagnosis">
|
||
<paragraph id="D537950A5C79486D3F284808FCF8FE69" blockId="10.[264,1324,283,1550]" pageId="10" pageNumber="11">
|
||
<emphasis id="E7FC49185C79486D3F284808FE63FEF2" bold="true" box="[312,432,283,309]" pageId="10" pageNumber="11">Diagnosis</emphasis>
|
||
. Differs from other
|
||
<taxonomicName id="1288EE895C79486D3C8F480FFD2FFEF2" box="[671,764,284,309]" class="Insecta" family="Carabidae" genus="Stolonis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E7FC49185C79486D3C8F480FFD2FFEF2" box="[671,764,284,309]" italics="true" pageId="10" pageNumber="11">Stolonis</emphasis>
|
||
</taxonomicName>
|
||
species by the combination of broad pronotum with narrowly expanded lateral margins, low rounded, but produced, angular process at the hind setae (
|
||
<figureCitation id="4DB3898F5C79486D3F87487FFE32FE41" box="[407,481,364,390]" captionStart="FIGURE 2" captionStartId="8.[264,367,1326,1350]" captionTargetBox="[337,1321,462,1293]" captionTargetId="figure@8.[267,1326,452,1303]" captionTargetPageId="8" captionText="FIGURE 2. Pronota of: A— Stolonis tapiai, B— Stolonis scortensis, C— Stolonis spinosus, D— Stolonis yasuni, E— Stolonis catenarius." httpUri="https://zenodo.org/record/169945/files/figure.png" pageId="10" pageNumber="11">Fig. 2</figureCitation>
|
||
B) and immaculate elytra. The laterally wrinkled median lobe of the aedeagus is diagnostic for males (
|
||
<figureCitation id="4DB3898F5C79486D3C834887FD31FE69" box="[659,738,404,430]" captionStart="FIGURE 3" captionStartId="9.[264,368,1716,1740]" captionTargetBox="[284,1304,346,1671]" captionTargetId="figure@9.[264,1323,329,1690]" captionTargetPageId="9" captionText="FIGURE 3. Aedeagus dorsal, left lateral, right lateral and ventral views of: A – D— Stolonis tapiai, E – H— Stolonis scortensis, I – L— Stolonis spinosus, M – P— Stolonis yasuni, Q – T— Stolonis catenarius." httpUri="https://zenodo.org/record/169946/files/figure.png" pageId="10" pageNumber="11">Figs. 3</figureCitation>
|
||
E–H).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9D92C6815C79486D3F2848A8FE22F9C9" pageId="10" pageNumber="11" type="description">
|
||
<paragraph id="D537950A5C79486D3F2848A8FE6EFD89" blockId="10.[264,1324,283,1550]" pageId="10" pageNumber="11">
|
||
<emphasis id="E7FC49185C79486D3F2848A8FE1AFE12" bold="true" box="[312,457,443,469]" pageId="10" pageNumber="11">Description</emphasis>
|
||
. Length
|
||
<quantity id="127038EF5C79486D3C4148AFFD74FE11" box="[593,679,444,470]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="10" pageNumber="11" unit="mm" value="5.9">5.9mm</quantity>
|
||
. Dorsal surface black, shiny, lightly iridescent; antennomeres 1–3 brunneous, 4–6, base of 7 and 11 black, apex of 7 and 8–10 white; mouth parts brunneous and legs flavotestaceous with coxae darker, concolorous with ventral surface.
|
||
</paragraph>
|
||
<paragraph id="D537950A5C79486D3F284B4FFDC5FD01" blockId="10.[264,1324,283,1550]" pageId="10" pageNumber="11">Form of head average build; eyes moderately prominent; frontal impressions not well marked, very short, punctiform, frons between eyes shiny, scarcely iridescent near base, microlines not evident.</paragraph>
|
||
<paragraph id="D537950A5C79486D3F284BC7FDBFFBB9" blockId="10.[264,1324,283,1550]" pageId="10" pageNumber="11">
|
||
Pronotum (
|
||
<figureCitation id="4DB3898F5C79486D3FAD4BC7FDD5FD29" box="[445,518,724,750]" captionStart="FIGURE 2" captionStartId="8.[264,367,1326,1350]" captionTargetBox="[337,1321,462,1293]" captionTargetId="figure@8.[267,1326,452,1303]" captionTargetPageId="8" captionText="FIGURE 2. Pronota of: A— Stolonis tapiai, B— Stolonis scortensis, C— Stolonis spinosus, D— Stolonis yasuni, E— Stolonis catenarius." httpUri="https://zenodo.org/record/169945/files/figure.png" pageId="10" pageNumber="11">Fig. 2</figureCitation>
|
||
B); clearly broader than long; narrowly constricted at base; anterior submarginal sulcus deep and complete; basal impressions short broad and not clearly defined apically; lateral margins broadly rounded and moderately explanate, of almost equal width throughout; low rounded, but produced, angular process at hind setae; base punctate between and in basal impressions. Elytra elongate oval, length
|
||
<quantity id="127038EF5C79486D3A654A67FB18FC49" box="[1141,1227,884,910]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="10" pageNumber="11" unit="mm" value="3.5">3.5mm</quantity>
|
||
, overall width
|
||
<quantity id="127038EF5C79486D3F434A8FFE7BFC71" box="[339,424,924,950]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="10" pageNumber="11" unit="mm" value="2.3">2.3mm</quantity>
|
||
, slightly convex, rounded and notably sinuate near apex; humeri sloped and rounded; striae densely, shallowly punctate; intervals broad, slightly convex, shallowly crenulate. Legs, moderately slender; meso and metatarsi with evident, but shallow, external sulcus; fifth tarsomeres ventrally glabrous. Ventral surface shiny, clearly iridescent; mesosternum with 7 small, shallow punctures; metasternum laterally with 7 shallow, very small punctures.
|
||
</paragraph>
|
||
<paragraph id="D537950A5C79486D3F284D9FFE22F9C9" blockId="10.[264,1324,283,1550]" pageId="10" pageNumber="11">
|
||
Base of abdominal sternum II with row of shallowly impressed punctures; metepisternum longer than wide (l/w= 1.5); base of sterna IV–VII with band of dull, coarse irregular microsculpture; sterna IV–VI with one pair paramedial setae; in male one pair paramedial setae on VII. Aedeagus (
|
||
<figureCitation id="4DB3898F5C79486D3D004C17FCB5FAD9" box="[784,870,1284,1310]" captionStart="FIGURE 3" captionStartId="9.[264,368,1716,1740]" captionTargetBox="[284,1304,346,1671]" captionTargetId="figure@9.[264,1323,329,1690]" captionTargetPageId="9" captionText="FIGURE 3. Aedeagus dorsal, left lateral, right lateral and ventral views of: A – D— Stolonis tapiai, E – H— Stolonis scortensis, I – L— Stolonis spinosus, M – P— Stolonis yasuni, Q – T— Stolonis catenarius." httpUri="https://zenodo.org/record/169946/files/figure.png" pageId="10" pageNumber="11">Figs. 3</figureCitation>
|
||
E–H) with median lobe stout, small narrowed and blunt tip, slightly reflexed ventrally, ostium dorsal and very large; ventral surface broadly membranous medially; median lobe laterally irregularly strigate; endophallus in repose with 4 spines near apex, 2 subapical spines, and two spines located medially, arranged as in figures 3E–H. Females unknown so reproductive tract unstudied. Pygidial gland reservoir form simple with no additional lobes, chemical compounds produced unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |