566 lines
65 KiB
XML
566 lines
65 KiB
XML
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<mods:title id="45FF56C1703CCADFCDE16E3E2BE5A48C">Taxonomic revision of the Neotropical diving beetles genus Laccodytes Régimbart, 1895 (Coleoptera: Dytiscidae)</mods:title>
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<mods:namePart id="93630D111C431F5E6F66D89B0011C721">Toledo, Mario</mods:namePart>
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<mods:namePart id="AE1998164991D0BD188C8722353EBCA1">Spangler, Paul J.</mods:namePart>
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<mods:namePart id="A613107E20978EC049858983CA9853E2">Balke, Michael</mods:namePart>
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<mods:title id="83D81320F27C67C5A9A02D3D846E492F">Zootaxa</mods:title>
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<mods:date id="61CA20863D673446194CF4DF956C276B">2010</mods:date>
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<mods:number id="7F1A5C91A2404BA5CD8780517DFBD54D">2347</mods:number>
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<treatment id="11578788FE5DFFC77DB5FAB1146BDD41" ID-DOI="http://doi.org/10.5281/zenodo.6198356" ID-GBIF-Taxon="124955818" ID-Zenodo-Dep="6198356" LSID="urn:lsid:plazi:treatment:11578788FE5DFFC77DB5FAB1146BDD41" httpUri="http://treatment.plazi.org/id/11578788FE5DFFC77DB5FAB1146BDD41" lastPageId="5" lastPageNumber="42" pageId="1" pageNumber="38">
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<subSubSection id="D1E46515FE5DFFC37DB5FAB1140CDD0E" box="[151,514,1346,1372]" pageId="1" pageNumber="38" type="nomenclature">
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<paragraph id="9941369EFE5DFFC37DB5FAB1140CDD0E" blockId="1.[151,514,1346,1372]" box="[151,514,1346,1372]" pageId="1" pageNumber="38">
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<heading id="C20981F2FE5DFFC37DB5FAB1140CDD0E" bold="true" box="[151,514,1346,1372]" fontSize="11" level="1" pageId="1" pageNumber="38" reason="1">
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<emphasis id="AB8AEA8CFE5DFFC37DB5FAB1140CDD0E" bold="true" box="[151,514,1346,1372]" pageId="1" pageNumber="38">
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Genus
|
||
<taxonomicName id="5EFE4D1DFE5DFFC37DCCFAB1140CDD0E" ID-CoL="59RJ" authority="Regimbart" authorityName="Regimbart" box="[238,514,1346,1372]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="38" phylum="Arthropoda" rank="genus">
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<emphasis id="AB8AEA8CFE5DFFC37DCCFAB1177ADD0E" bold="true" box="[238,372,1346,1372]" italics="true" pageId="1" pageNumber="38">Laccodytes</emphasis>
|
||
Régimbart
|
||
</taxonomicName>
|
||
</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="D1E46515FE5DFFC37DB5FA7F154ADDB1" pageId="1" pageNumber="38" type="reference_group">
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<paragraph id="9941369EFE5DFFC37DB5FA7F154ADDB1" blockId="1.[151,1436,1420,1507]" pageId="1" pageNumber="38">
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<treatmentCitationGroup id="B9EE11B0FE5DFFC37DB5FA7F154ADDB1" pageId="1" pageNumber="38">
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<taxonomicName id="5EFE4D1DFE5DFFC37DB5FA7F1419DDF1" ID-CoL="59RJ" authority="Regimbart (1895: 345)" authorityName="Regimbart" authorityPageNumber="345" authorityYear="1895" box="[151,535,1420,1443]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="38" phylum="Arthropoda" rank="genus">
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<emphasis id="AB8AEA8CFE5DFFC37DB5FA7F1701DDF1" box="[151,271,1420,1443]" italics="true" pageId="1" pageNumber="38">Laccodytes</emphasis>
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<treatmentCitation id="185F108FFE5DFFC37C3BFA7F1419DDF1" author="Regimbart" box="[281,535,1420,1443]" page="345" pageId="1" pageNumber="38" year="1895">
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<bibRefCitation id="FD6F4B6FFE5DFFC37C3BFA7F1419DDF1" author="Regimbart" box="[281,535,1420,1443]" pageId="1" pageNumber="58" refString="Regimbart, M. (1895) Dytiscides trouves dans les Tabacs par les soins de M. Antoine Grouvelle. Annales de la Societe Entomologique de France, 64, 321 - 348, 1 pl." type="journal article" year="1895">Régimbart (1895: 345)</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
|
||
;
|
||
<treatmentCitation id="185F108FFE5DFFC37F07FA7F150ADDF1" author="Peschet" box="[549,772,1420,1443]" page="146" pageId="1" pageNumber="38" year="1919">
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<bibRefCitation id="FD6F4B6FFE5DFFC37F07FA7F150ADDF1" author="Peschet" box="[549,772,1420,1443]" pageId="1" pageNumber="58" refString="Peschet, R. (1919) Description d'un Dytiscidae nouveaux de l'Amerique du Sud (Col.). Bulletin de la Societe Entomologique de France, 1919 (8), 145 - 147." type="journal article" year="1919">Peschet (1919: 146)</bibRefCitation>
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</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="185F108FFE5DFFC37E31FA7F1226DDF1" author="Zimmermann" box="[787,1064,1420,1443]" page="119" pageId="1" pageNumber="38" year="1919">
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<bibRefCitation id="FD6F4B6FFE5DFFC37E31FA7F15FFDDF1" author="Zimmermann" box="[787,1009,1420,1443]" pageId="1" pageNumber="58" refString="Zimmermann, A. (1919) Die Schwimmkafer des Deutschen Entomologischen Museum in Berlin-Dahlem. Archiv fur Naturgeschichte, 83 (12) (1917), 107 - 249." type="journal article" year="1919">Zimmermann (1919</bibRefCitation>
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||
: 119
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||
</treatmentCitation>
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||
; 1920: 29);
|
||
<treatmentCitation id="185F108FFE5DFFC37991FA7F1399DDF1" author="Guignot" box="[1203,1431,1420,1443]" page="138" pageId="1" pageNumber="38" year="1937">
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||
<bibRefCitation id="FD6F4B6FFE5DFFC37991FA7F1399DDF1" author="Guignot" box="[1203,1431,1420,1443]" pageId="1" pageNumber="58" refString="Guignot, F. (1937) Contribution a l'etude des Laccophilinae (Col. Dytiscidae). Bulletin de la Societe entomologique de France, 42 (9), 137 - 143." type="journal article" year="1937">Guignot (1937: 138)</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="185F108FFE5DFFC37DE5FA5F140EDD91" author="Balfour-Browne" box="[199,512,1452,1475]" page="104" pageId="1" pageNumber="38" year="1939">
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||
<bibRefCitation id="FD6F4B6FFE5DFFC37DE5FA5F140EDD91" author="Balfour-Browne" box="[199,512,1452,1475]" pageId="1" pageNumber="58" refString="Balfour-Browne, J. (1939) A contribution to the study of the Dytiscidae I. (Coleoptera, Adephaga). Annals and Magazine of natural History, Ser. 11, 3, 97 - 114." type="journal article" year="1939">Balfour-Browne (1939: 104)</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="185F108FFE5DFFC37F2FFA5F14DFDD91" author="Young" box="[525,721,1452,1475]" page="48" pageId="1" pageNumber="38" year="1954">
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||
<bibRefCitation id="FD6F4B6FFE5DFFC37F2FFA5F14DFDD91" author="Young" box="[525,721,1452,1475]" pageId="1" pageNumber="58" refString="Young, F. N. (1954) The water beetles of Florida. University of Florida, Studies in Biological Sciences, Series 5, 1 - 238." type="book chapter" year="1954">Young (1954: 48)</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<bibRefCitation id="FD6F4B6FFE5DFFC37FFCFA5F15E2DD91" author="Steiner" box="[734,1004,1452,1475]" pageId="1" pageNumber="58" refString="Steiner, W. E. (1981) A new genus and a new species of Laccophilinae water beetle from Ecuador (Coleoptera: Dytiscidae). The Pan-Pacific Entomologist, 57, 251 - 259." type="journal article" year="1981">Steiner (1981: 257, 258)</bibRefCitation>
|
||
;
|
||
<treatmentCitation id="185F108FFE5DFFC37EDBFA5F12F3DD91" author="Brancucci" box="[1017,1277,1452,1475]" page="253" pageId="1" pageNumber="38" year="1983">
|
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<bibRefCitation id="FD6F4B6FFE5DFFC37EDBFA5F12F3DD91" author="Brancucci" box="[1017,1277,1452,1475]" pageId="1" pageNumber="58" refString="Brancucci, M. (1983 a) A new genus of the subfamily Laccophilinae (Coleoptera, Dytiscidae). Aquatic Insects, 5 (4), 251 - 254." type="journal article" year="1983" yearSuffix="a">Brancucci (1983a: 253)</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="185F108FFE5DFFC37828FA5F1731DDB1" author="Nilsson" page="299" pageId="1" pageNumber="38" year="1989">
|
||
<bibRefCitation id="FD6F4B6FFE5DFFC37828FA5F1707DDB1" author="Nilsson" pageId="1" pageNumber="58" refString="Nilsson, A. N., Roughley, R. E. & Brancucci, M. (1989) A review of the genus- and family-group names of the family Dytiscidae Leach (Coleoptera). Entomologica Scandinavica, 20, 287 - 316." type="journal article" year="1989">
|
||
Nilsson
|
||
<emphasis id="AB8AEA8CFE5DFFC37846FA5F1392DD91" box="[1380,1436,1452,1475]" italics="true" pageId="1" pageNumber="38">et al.</emphasis>
|
||
(1989
|
||
</bibRefCitation>
|
||
: 299
|
||
</treatmentCitation>
|
||
);
|
||
<bibRefCitation id="FD6F4B6FFE5DFFC37C70FA3F1452DDB1" author="Pederzani" box="[338,604,1484,1507]" pageId="1" pageNumber="58" refString="Pederzani, F. (1995) Keys to the identification of the genera and subgenera of adult Dytiscidae (sensu lato) of the world (Coleoptera: Dytiscidae). Atti dell'Accademia Roveretana degli Agiati 244 (1994), ser. VII, IV B, 5 - 83." type="book chapter" year="1995">Pederzani (1995: 43, 73)</bibRefCitation>
|
||
;
|
||
<treatmentCitation id="185F108FFE5DFFC37F4AFA3F154EDDB1" author="Nilsson" box="[616,832,1484,1507]" page="239" pageId="1" pageNumber="38" year="2001">
|
||
<bibRefCitation id="FD6F4B6FFE5DFFC37F4AFA3F154EDDB1" author="Nilsson" box="[616,832,1484,1507]" pageId="1" pageNumber="58" refString="Nilsson, A. N. (2001) World catalogue of insects. Vol. 3. Dytiscidae Coleoptera. Stenstrup, Apollo Books, 395 pp." type="book" year="2001">Nilsson (2001: 239)</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D1E46515FE5DFFC37DB5F9E11781DE01" pageId="1" pageNumber="38" type="materials_examined">
|
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<paragraph id="9941369EFE5DFFC37DB5F9E11781DE01" blockId="1.[151,1436,1554,2006]" pageId="1" pageNumber="38">
|
||
<emphasis id="AB8AEA8CFE5DFFC37DB5F9E11736DE7E" bold="true" box="[151,312,1554,1580]" pageId="1" pageNumber="38">
|
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<typeStatus id="4645883CFE5DFFC37DB5F9E116D8DE7E" box="[151,214,1554,1580]" pageId="1" pageNumber="38">Type</typeStatus>
|
||
species
|
||
</emphasis>
|
||
.
|
||
<taxonomicName id="5EFE4D1DFE5DFFC37C6AF9E01541DE7E" authority="Regimbart, 1895" authorityName="Regimbart" authorityYear="1895" box="[328,847,1554,1580]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="38" phylum="Arthropoda" rank="species" species="phalacroides">
|
||
<emphasis id="AB8AEA8CFE5DFFC37C6AF9E01479DE7E" box="[328,631,1555,1580]" italics="true" pageId="1" pageNumber="38">Laccodytes phalacroides</emphasis>
|
||
<bibRefCitation id="FD6F4B6FFE5DFFC37FA2F9E11541DE7E" author="Regimbart" box="[640,847,1554,1580]" pageId="1" pageNumber="58" refString="Regimbart, M. (1895) Dytiscides trouves dans les Tabacs par les soins de M. Antoine Grouvelle. Annales de la Societe Entomologique de France, 64, 321 - 348, 1 pl." type="journal article" year="1895">Régimbart, 1895</bibRefCitation>
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</taxonomicName>
|
||
, by subsequent designation (
|
||
<bibRefCitation id="FD6F4B6FFE5DFFC37992F9E11351DE7E" author="Guignot" box="[1200,1375,1554,1580]" pageId="1" pageNumber="58" refString="Guignot, F. (1937) Contribution a l'etude des Laccophilinae (Col. Dytiscidae). Bulletin de la Societe entomologique de France, 42 (9), 137 - 143." type="journal article" year="1937">Guignot 1937</bibRefCitation>
|
||
), see
|
||
<bibRefCitation id="FD6F4B6FFE5DFFC37DB5F9CA1785DE01" box="[151,395,1593,1619]" pageId="1" pageNumber="58" refString="Nilsson, A. N., Roughley, R. E. & Brancucci, M. (1989) A review of the genus- and family-group names of the family Dytiscidae Leach (Coleoptera). Entomologica Scandinavica, 20, 287 - 316." type="journal article">
|
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Nilsson
|
||
<emphasis id="AB8AEA8CFE5DFFC37DDAF9CA173CDE00" box="[248,306,1593,1618]" italics="true" pageId="1" pageNumber="38">et al.</emphasis>
|
||
(1989)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D1E46515FE5DFFC07DE4F9AC157FD9B8" lastPageId="2" lastPageNumber="39" pageId="1" pageNumber="38" type="diagnosis">
|
||
<paragraph id="9941369EFE5DFFC37DE4F9AC1493DEF2" blockId="1.[151,1436,1554,2006]" pageId="1" pageNumber="38">
|
||
<emphasis id="AB8AEA8CFE5DFFC37DE4F9AC1731DE2B" bold="true" box="[198,319,1631,1657]" pageId="1" pageNumber="38">Diagnosis</emphasis>
|
||
. Small beetles, TL 1.6–2.4 mm. Habitus (
|
||
<figureCitation id="01C52A1BFE5DFFC37E69F99315B9DE28" box="[843,951,1632,1658]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,270,2000,2024]" captionTargetBox="[168,1422,904,1961]" captionTargetId="figure@2.[151,1436,896,1975]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Habitus of: 1) Laccodytes rondonia sp. n.; 2) L. bassignanii sp. n." httpUri="https://zenodo.org/record/193407/files/figure.png" pageId="1" pageNumber="38">Figs 1–2</figureCitation>
|
||
) oval, dorsoventrally flattened. Usually without angle between pronotum and elytra.
|
||
</paragraph>
|
||
<paragraph id="9941369EFE5DFFC37DE4F95E1295DF84" blockId="1.[151,1436,1554,2006]" pageId="1" pageNumber="38">
|
||
Pronotum with base medially produced caudad, or almost straight.
|
||
<collectingCountry id="E1E9760EFE5DFFC37EF4F95E121FDE95" box="[982,1041,1709,1735]" name="India" pageId="1" pageNumber="38">Hind</collectingCountry>
|
||
angle either rounded or projected backwards to form a needle-shaped, acute tip. Surface with simple MR, without coarser punctation. Prosternal process broadly carinate, with a needle or thorn-like apical projection. Prosternum and head separated by a shallow step, therefore gula and medial line of prosternum apparently on the same level. Metacoxal process with hind lobes straight or slightly rounded, often separated by a more or less deep V-shaped notch. Elytra apically oval or truncate; narrowly bordered laterally. Epipleuron in both sexes broad, up to caudal margin of sternite 6 or 7, not enlarged apically. MR of dorsal and ventral side usually of polygonal meshes, their orientation often slightly to distinctly horizontal, especially so on elytra and metacoxae.
|
||
</paragraph>
|
||
<paragraph id="9941369EFE5EFFC07DE4FF6B1378D8AD" blockId="2.[151,1436,152,838]" pageId="2" pageNumber="39">
|
||
Fore and middle legs long and slender, tarsomeres 1–3 shorter than 4–5 together.
|
||
<collectingCountry id="E1E9760EFE5EFFC079A0FF6B12B3D8E0" box="[1154,1213,152,178]" name="India" pageId="2" pageNumber="39">Hind</collectingCountry>
|
||
legs with tarsi less broadened than in
|
||
<taxonomicName id="5EFE4D1DFE5EFFC07C5BFF4C1403D88A" box="[377,525,191,216]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AB8AEA8CFE5EFFC07C5BFF4C1403D88A" box="[377,525,191,216]" italics="true" pageId="2" pageNumber="39">Laccophilus</emphasis>
|
||
</taxonomicName>
|
||
but slightly more so than in
|
||
<taxonomicName id="5EFE4D1DFE5EFFC07E4FFF4C1202D88A" box="[877,1036,191,216]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AB8AEA8CFE5EFFC07E4FFF4C1202D88A" box="[877,1036,191,216]" italics="true" pageId="2" pageNumber="39">Neptosternus</emphasis>
|
||
</taxonomicName>
|
||
, with a antero-lateral expansion, more or less developed, on tarsomere 1–4. Metatibial spurs short, tip acuminate (truncate in
|
||
<taxonomicName id="5EFE4D1DFE5EFFC079F2FF16136BD8AC" box="[1232,1381,229,254]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="olibroides">
|
||
<emphasis id="AB8AEA8CFE5EFFC079F2FF16136BD8AC" box="[1232,1381,229,254]" italics="true" pageId="2" pageNumber="39">L. olibroides</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="9941369EFE5EFFC07DE4FEFF1281D9CE" blockId="2.[151,1436,152,838]" pageId="2" pageNumber="39">
|
||
Male with pro- and mesotarsus hardly or not dilated at all, bearing three pairs of stalked suction palettes ventrally: one pair on each of the 1st–3rd tarsomere (
|
||
<figureCitation id="01C52A1BFE5EFFC07E21FEC61542D91D" box="[771,844,309,335]" captionStart="FIGURES 3 – 16" captionStartId="3.[151,270,1860,1884]" captionTargetBox="[248,1343,197,1819]" captionTargetId="figure@3.[248,1348,197,1827]" captionTargetPageId="3" captionText="FIGURES 3 – 16. SEMs of some Laccodytes species structures: 3) segments 1 - 3 in male protarsus of L. americanus; 4) prosternal process of L. apalodes; 5) idem, L. americanus; 6) idem, L. neblinae; 7) metaventrite of L. apalodes, the arrow indicates the groove in which the tip of prosternal process fits; 8) idem, L. neblinae; 9) metacoxal process of L. apalodes; 10) idem, L. americanus; 11) idem, L. neblinae; 12) hind tarsus of L. apalodes (male); 13) idem, L. americanus (male); 14) idem, L. neblinae (female); 15) ovipositor of L. apalodes; 16) idem, L. americanus." httpUri="https://zenodo.org/record/193408/files/figure.png" pageId="2" pageNumber="39">Fig. 3</figureCitation>
|
||
). Foreclaws in both sexes subequal in length and shape. Sternite 7 caudad rounded or tectiform; or slightly (
|
||
<taxonomicName id="5EFE4D1DFE5EFFC07E6EFEAF15FAD927" box="[844,1012,348,373]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="americanus">
|
||
<emphasis id="AB8AEA8CFE5EFFC07E6EFEAF15FAD927" box="[844,1012,348,373]" italics="true" pageId="2" pageNumber="39">L. americanus</emphasis>
|
||
</taxonomicName>
|
||
), or deeply concave (
|
||
<taxonomicName id="5EFE4D1DFE5EFFC079D7FEAF1384D927" box="[1269,1418,348,373]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="olibroides">
|
||
<emphasis id="AB8AEA8CFE5EFFC079D7FEAF1384D927" box="[1269,1418,348,373]" italics="true" pageId="2" pageNumber="39">L. olibroides</emphasis>
|
||
</taxonomicName>
|
||
), or V-shaped (
|
||
<taxonomicName id="5EFE4D1DFE5EFFC07C17FE7017D6D9CE" box="[309,472,387,412]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="androginus">
|
||
<emphasis id="AB8AEA8CFE5EFFC07C17FE7017D6D9CE" box="[309,472,387,412]" italics="true" pageId="2" pageNumber="39">L. androginus</emphasis>
|
||
</taxonomicName>
|
||
) like in females. Aedeagus asymmetrical; parameres short.
|
||
</paragraph>
|
||
<paragraph id="9941369EFE5EFFC07DE4FE5A157FD9B8" blockId="2.[151,1436,152,838]" pageId="2" pageNumber="39">
|
||
Female with sternite 7 caudally with a central U- or V-shaped incision, more or less developed. Ovipositor serrate, but the “saw” consists of but a few teeth (
|
||
<figureCitation id="01C52A1BFE5EFFC07FF9FE23156DD9B8" box="[731,867,464,490]" captionStart="FIGURES 3 – 16" captionStartId="3.[151,270,1860,1884]" captionTargetBox="[248,1343,197,1819]" captionTargetId="figure@3.[248,1348,197,1827]" captionTargetPageId="3" captionText="FIGURES 3 – 16. SEMs of some Laccodytes species structures: 3) segments 1 - 3 in male protarsus of L. americanus; 4) prosternal process of L. apalodes; 5) idem, L. americanus; 6) idem, L. neblinae; 7) metaventrite of L. apalodes, the arrow indicates the groove in which the tip of prosternal process fits; 8) idem, L. neblinae; 9) metacoxal process of L. apalodes; 10) idem, L. americanus; 11) idem, L. neblinae; 12) hind tarsus of L. apalodes (male); 13) idem, L. americanus (male); 14) idem, L. neblinae (female); 15) ovipositor of L. apalodes; 16) idem, L. americanus." httpUri="https://zenodo.org/record/193408/files/figure.png" pageId="2" pageNumber="39">Figs 15–16</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D1E46515FE5EFFC77DE4FE0515AAD921" lastPageId="5" lastPageNumber="42" pageId="2" pageNumber="39" type="distribution">
|
||
<paragraph id="9941369EFE5EFFC07DE4FE051425DB14" blockId="2.[151,1436,152,838]" pageId="2" pageNumber="39">
|
||
<emphasis id="AB8AEA8CFE5EFFC07DE4FE05140DDA42" bold="true" box="[198,515,502,528]" pageId="2" pageNumber="39">Distribution and ecology.</emphasis>
|
||
<taxonomicName id="5EFE4D1DFE5EFFC07F2EFE04149EDA42" box="[524,656,503,528]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AB8AEA8CFE5EFFC07F2EFE04149EDA42" box="[524,656,503,528]" italics="true" pageId="2" pageNumber="39">Laccodytes</emphasis>
|
||
</taxonomicName>
|
||
is only known from tropical South
|
||
<collectingCountry id="E1E9760EFE5EFFC0791CFE0412A9DA43" box="[1086,1191,503,529]" name="United States of America" pageId="2" pageNumber="39">America</collectingCountry>
|
||
. Most of the species have been collected in and around the lands southern of the Orinoco basin, between the
|
||
<collectingCountry id="E1E9760EFE5EFFC079EAFDEE1326DA65" box="[1224,1320,541,567]" name="Guyana" pageId="2" pageNumber="39">Guyana</collectingCountry>
|
||
, northern
|
||
<collectingCountry id="E1E9760EFE5EFFC07DB5FDB716ECDA0C" box="[151,226,580,606]" name="Brazil" pageId="2" pageNumber="39">Brazil</collectingCountry>
|
||
and southern
|
||
<collectingCountry id="E1E9760EFE5EFFC07CB0FDB71401DA0C" box="[402,527,580,606]" name="Venezuela" pageId="2" pageNumber="39">Venezuela</collectingCountry>
|
||
(
|
||
<figureCitation id="01C52A1BFE5EFFC07F03FDB71470DA0C" box="[545,638,580,606]" captionStart="FIGURES 54 – 61" captionStartId="12.[151,273,1107,1131]" captionTargetBox="[153,1431,196,1081]" captionTargetId="figure@12.[151,1436,194,1083]" captionTargetPageId="12" captionText="FIGURES 54 – 61. Last abdominal segment (left male; right female) in Laccodytes species: 54) L. apalodes; 55) L. rondonia; 56) L. americanus; 57) L. phalacroides; 58) L. takutuanus; 59) L. neblinae; 60) L. bassignanii; 61) Laccodytes sp. 1 female." httpUri="https://zenodo.org/record/193415/files/figure.png" pageId="2" pageNumber="39">Fig. 61</figureCitation>
|
||
). Two species are known from southeastern
|
||
<collectingCountry id="E1E9760EFE5EFFC07986FDB712E1DA0C" box="[1188,1263,580,606]" name="Brazil" pageId="2" pageNumber="39">Brazil</collectingCountry>
|
||
and one from central-western
|
||
<collectingCountry id="E1E9760EFE5EFFC07C74FD9917ACDAD6" box="[342,418,618,644]" name="Brazil" pageId="2" pageNumber="39">Brazil</collectingCountry>
|
||
. At present most of the known species come from the range of the
|
||
<collectingCountry id="E1E9760EFE5EFFC079E7FD99132FDAD6" box="[1221,1313,618,644]" name="Guyana" pageId="2" pageNumber="39">Guyana</collectingCountry>
|
||
Shield, in
|
||
<collectingCountry id="E1E9760EFE5EFFC07DB5FD62171FDAF9" box="[151,273,657,683]" name="Venezuela" pageId="2" pageNumber="39">Venezuela</collectingCountry>
|
||
and
|
||
<collectingCountry id="E1E9760EFE5EFFC07C6EFD6217A2DAF9" box="[332,428,657,683]" name="Guyana" pageId="2" pageNumber="39">Guyana</collectingCountry>
|
||
. The mountain ranges of south-eastern
|
||
<collectingCountry id="E1E9760EFE5EFFC07EA6FD6215C3DAF9" box="[900,973,657,683]" name="Brazil" pageId="2" pageNumber="39">Brazil</collectingCountry>
|
||
(related to the Planalto Central) might represent another hot spot for this genus. The available ecological data suggest that
|
||
<taxonomicName id="5EFE4D1DFE5EFFC07991FD4B1330DA83" box="[1203,1342,696,721]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AB8AEA8CFE5EFFC07991FD4B1330DA83" box="[1203,1342,696,721]" italics="true" pageId="2" pageNumber="39">Laccodytes</emphasis>
|
||
</taxonomicName>
|
||
inhabit streams, often encountered among layers of decaying leafs and debris. We found specimens e.g. in moss on a boulder in strong current, in a water depth of about
|
||
<quantity id="5E069B7BFE5EFFC07E33FCF61552DB4D" box="[785,860,773,799]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="39" unit="cm" value="50.0">50 cm</quantity>
|
||
(
|
||
<collectingCountry id="E1E9760EFE5EFFC07E4EFCF615E1DB4D" box="[876,1007,773,799]" name="Venezuela" pageId="2" pageNumber="39">Venezuela</collectingCountry>
|
||
: Tobogan de la Selva). Sometimes, several species occur syntopically.
|
||
</paragraph>
|
||
<caption id="CD816616FE5EFFC07DB5F82315F3DFBA" box="[151,1021,2000,2024]" httpUri="https://zenodo.org/record/193407/files/figure.png" pageId="2" pageNumber="39" targetBox="[168,1422,904,1961]" targetPageId="2">
|
||
<paragraph id="9941369EFE5EFFC07DB5F82315F3DFBA" blockId="2.[151,1021,2000,2024]" box="[151,1021,2000,2024]" pageId="2" pageNumber="39">
|
||
<emphasis id="AB8AEA8CFE5EFFC07DB5F8231735DFBA" bold="true" box="[151,315,2000,2024]" pageId="2" pageNumber="39">FIGURES 1–2</emphasis>
|
||
. Habitus of:
|
||
<emphasis id="AB8AEA8CFE5EFFC07CE5F82317DADFBA" bold="true" box="[455,468,2000,2024]" pageId="2" pageNumber="39">1</emphasis>
|
||
)
|
||
<taxonomicName id="5EFE4D1DFE5EFFC07CC6F82214CADFBA" box="[484,708,2001,2024]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="rondonia" status="sp. nov.">
|
||
<emphasis id="AB8AEA8CFE5EFFC07CC6F82214CADFBA" box="[484,708,2001,2024]" italics="true" pageId="2" pageNumber="39">Laccodytes rondonia</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="AB8AEA8CFE5EFFC07FE8F82314F1DFBA" bold="true" box="[714,767,2000,2024]" pageId="2" pageNumber="39">
|
||
<taxonomicNameLabel id="B0B957F7FE5EFFC07FE8F82314F1DFBA" box="[714,767,2000,2024]" pageId="2" pageNumber="39" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
;
|
||
<emphasis id="AB8AEA8CFE5EFFC07E2FF8231514DFBA" bold="true" box="[781,794,2000,2024]" pageId="2" pageNumber="39">2</emphasis>
|
||
)
|
||
<taxonomicName id="5EFE4D1DFE5EFFC07E08F82215CFDFBA" box="[810,961,2001,2024]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="bassignanii" status="sp. nov.">
|
||
<emphasis id="AB8AEA8CFE5EFFC07E08F82215CFDFBA" box="[810,961,2001,2024]" italics="true" pageId="2" pageNumber="39">L. bassignanii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B0B957F7FE5EFFC07EEAF82315F3DFBA" box="[968,1021,2000,2024]" pageId="2" pageNumber="39" rank="species">sp.n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="CD816616FE5FFFC17DB5F8B71235DFB5" httpUri="https://zenodo.org/record/193408/files/figure.png" pageId="3" pageNumber="40" targetBox="[248,1343,197,1819]" targetPageId="3">
|
||
<paragraph id="9941369EFE5FFFC17DB5F8B71235DFB5" blockId="3.[151,1436,1860,2024]" pageId="3" pageNumber="40">
|
||
<emphasis id="AB8AEA8CFE5FFFC17DB5F8B71745DF0E" bold="true" box="[151,331,1860,1884]" pageId="3" pageNumber="40">FIGURES 3–16</emphasis>
|
||
. SEMs of some
|
||
<taxonomicName id="5EFE4D1DFE5FFFC17CDDF8B61479DF0E" box="[511,631,1861,1884]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AB8AEA8CFE5FFFC17CDDF8B61479DF0E" box="[511,631,1861,1884]" italics="true" pageId="3" pageNumber="40">Laccodytes</emphasis>
|
||
</taxonomicName>
|
||
species structures:
|
||
<emphasis id="AB8AEA8CFE5FFFC17E68F8B71559DF0E" bold="true" box="[842,855,1860,1884]" pageId="3" pageNumber="40">3</emphasis>
|
||
) segments 1-3 in male protarsus of
|
||
<taxonomicName id="5EFE4D1DFE5FFFC179FFF8B61379DF0E" box="[1245,1399,1861,1884]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="americanus">
|
||
<emphasis id="AB8AEA8CFE5FFFC179FFF8B61379DF0E" box="[1245,1399,1861,1884]" italics="true" pageId="3" pageNumber="40">L. americanus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="AB8AEA8CFE5FFFC178A7F8B7139CDF0E" bold="true" box="[1413,1426,1860,1884]" pageId="3" pageNumber="40">4</emphasis>
|
||
) prosternal process of
|
||
<taxonomicName id="5EFE4D1DFE5FFFC17C58F89A17FBDFD2" box="[378,501,1897,1920]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="apalodes">
|
||
<emphasis id="AB8AEA8CFE5FFFC17C58F89A17FBDFD2" box="[378,501,1897,1920]" italics="true" pageId="3" pageNumber="40">L. apalodes</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="AB8AEA8CFE5FFFC17F21F89B141EDFD2" bold="true" box="[515,528,1896,1920]" pageId="3" pageNumber="40">5</emphasis>
|
||
) idem,
|
||
<taxonomicName id="5EFE4D1DFE5FFFC17F40F89A14F7DFD2" box="[610,761,1897,1920]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="americanus">
|
||
<emphasis id="AB8AEA8CFE5FFFC17F40F89A14F7DFD2" box="[610,761,1897,1920]" italics="true" pageId="3" pageNumber="40">L. americanus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="AB8AEA8CFE5FFFC17E25F89B151ADFD2" bold="true" box="[775,788,1896,1920]" pageId="3" pageNumber="40">6</emphasis>
|
||
) idem,
|
||
<taxonomicName id="5EFE4D1DFE5FFFC17E47F89A15D3DFD2" box="[869,989,1897,1920]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="neblinae">
|
||
<emphasis id="AB8AEA8CFE5FFFC17E47F89A15D3DFD2" box="[869,989,1897,1920]" italics="true" pageId="3" pageNumber="40">L. neblinae</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="AB8AEA8CFE5FFFC17EC8F89B15F9DFD2" bold="true" box="[1002,1015,1896,1920]" pageId="3" pageNumber="40">7</emphasis>
|
||
) metaventrite of
|
||
<taxonomicName id="5EFE4D1DFE5FFFC1798CF89A1324DFD2" box="[1198,1322,1897,1920]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="apalodes">
|
||
<emphasis id="AB8AEA8CFE5FFFC1798CF89A1324DFD2" box="[1198,1322,1897,1920]" italics="true" pageId="3" pageNumber="40">L. apalodes</emphasis>
|
||
</taxonomicName>
|
||
, the arrow indicates the groove in which the tip of prosternal process fits;
|
||
<emphasis id="AB8AEA8CFE5FFFC17E10F8781531DFF1" bold="true" box="[818,831,1931,1955]" pageId="3" pageNumber="40">8</emphasis>
|
||
) idem,
|
||
<taxonomicName id="5EFE4D1DFE5FFFC17EB2F8781206DFF0" box="[912,1032,1931,1954]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="neblinae">
|
||
<emphasis id="AB8AEA8CFE5FFFC17EB2F8781206DFF0" box="[912,1032,1931,1954]" italics="true" pageId="3" pageNumber="40">L. neblinae</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="AB8AEA8CFE5FFFC17937F878122CDFF1" bold="true" box="[1045,1058,1931,1955]" pageId="3" pageNumber="40">9</emphasis>
|
||
) metacoxal process of
|
||
<taxonomicName id="5EFE4D1DFE5FFFC1783AF878139BDFF0" box="[1304,1429,1931,1954]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="apalodes">
|
||
<emphasis id="AB8AEA8CFE5FFFC1783AF878139BDFF0" box="[1304,1429,1931,1954]" italics="true" pageId="3" pageNumber="40">L. apalodes</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="AB8AEA8CFE5FFFC17DB5F85E16BCDF97" bold="true" box="[151,178,1965,1989]" pageId="3" pageNumber="40">10</emphasis>
|
||
) idem,
|
||
<taxonomicName id="5EFE4D1DFE5FFFC17C27F85D1790DF97" box="[261,414,1966,1989]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="americanus">
|
||
<emphasis id="AB8AEA8CFE5FFFC17C27F85D1790DF97" box="[261,414,1966,1989]" italics="true" pageId="3" pageNumber="40">L. americanus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="AB8AEA8CFE5FFFC17C8FF85E17CBDF97" bold="true" box="[429,453,1965,1989]" pageId="3" pageNumber="40">11</emphasis>
|
||
) idem,
|
||
<taxonomicName id="5EFE4D1DFE5FFFC17F38F85D149DDF97" box="[538,659,1966,1989]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="neblinae">
|
||
<emphasis id="AB8AEA8CFE5FFFC17F38F85D149DDF97" box="[538,659,1966,1989]" italics="true" pageId="3" pageNumber="40">L. neblinae</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="AB8AEA8CFE5FFFC17F80F85E14B3DF97" bold="true" box="[674,701,1965,1989]" pageId="3" pageNumber="40">12</emphasis>
|
||
) hind tarsus of
|
||
<taxonomicName id="5EFE4D1DFE5FFFC17E45F85D15EBDF97" box="[871,997,1966,1989]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="apalodes">
|
||
<emphasis id="AB8AEA8CFE5FFFC17E45F85D15EBDF97" box="[871,997,1966,1989]" italics="true" pageId="3" pageNumber="40">L. apalodes</emphasis>
|
||
</taxonomicName>
|
||
(male);
|
||
<emphasis id="AB8AEA8CFE5FFFC17962F85E1255DF97" bold="true" box="[1088,1115,1965,1989]" pageId="3" pageNumber="40">13</emphasis>
|
||
) idem,
|
||
<taxonomicName id="5EFE4D1DFE5FFFC1798CF85D1346DF97" box="[1198,1352,1966,1989]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="americanus">
|
||
<emphasis id="AB8AEA8CFE5FFFC1798CF85D1346DF97" box="[1198,1352,1966,1989]" italics="true" pageId="3" pageNumber="40">L. americanus</emphasis>
|
||
</taxonomicName>
|
||
(male);
|
||
<emphasis id="AB8AEA8CFE5FFFC17DB5F82316BCDFBA" bold="true" box="[151,178,2000,2024]" pageId="3" pageNumber="40">14</emphasis>
|
||
) idem,
|
||
<taxonomicName id="5EFE4D1DFE5FFFC17C26F8221772DFBA" box="[260,380,2001,2024]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="neblinae">
|
||
<emphasis id="AB8AEA8CFE5FFFC17C26F8221772DFBA" box="[260,380,2001,2024]" italics="true" pageId="3" pageNumber="40">L. neblinae</emphasis>
|
||
</taxonomicName>
|
||
(female);
|
||
<emphasis id="AB8AEA8CFE5FFFC17CC9F8231408DFBA" bold="true" box="[491,518,2000,2024]" pageId="3" pageNumber="40">15</emphasis>
|
||
) ovipositor of
|
||
<taxonomicName id="5EFE4D1DFE5FFFC17F86F822152FDFBA" box="[676,801,2001,2024]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="apalodes">
|
||
<emphasis id="AB8AEA8CFE5FFFC17F86F822152FDFBA" box="[676,801,2001,2024]" italics="true" pageId="3" pageNumber="40">L. apalodes</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="AB8AEA8CFE5FFFC17E0DF8231544DFBA" bold="true" box="[815,842,2000,2024]" pageId="3" pageNumber="40">16</emphasis>
|
||
) idem,
|
||
<taxonomicName id="5EFE4D1DFE5FFFC17EBEF822123BDFBA" box="[924,1077,2001,2024]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="americanus">
|
||
<emphasis id="AB8AEA8CFE5FFFC17EBEF822123BDFBA" box="[924,1077,2001,2024]" italics="true" pageId="3" pageNumber="40">L. americanus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="CD816616FE58FFC67DB5F882135FDFA3" httpUri="https://zenodo.org/record/193409/files/figure.png" pageId="4" pageNumber="41" targetBox="[163,1426,187,1879]" targetPageId="4">
|
||
<paragraph id="9941369EFE58FFC67DB5F882135FDFA3" blockId="4.[151,1437,1905,2034]" pageId="4" pageNumber="41">
|
||
<emphasis id="AB8AEA8CFE58FFC67DB5F8821758DFDB" bold="true" box="[151,342,1905,1929]" pageId="4" pageNumber="41">FIGURES 17–25</emphasis>
|
||
. SEMs of MR in some
|
||
<taxonomicName id="5EFE4D1DFE58FFC67F76F88214C2DFDA" box="[596,716,1905,1928]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AB8AEA8CFE58FFC67F76F88214C2DFDA" box="[596,716,1905,1928]" italics="true" pageId="4" pageNumber="41">Laccodytes</emphasis>
|
||
</taxonomicName>
|
||
species:
|
||
<emphasis id="AB8AEA8CFE58FFC67E0CF8821546DFDB" bold="true" box="[814,840,1905,1929]" pageId="4" pageNumber="41">17</emphasis>
|
||
) detail of elytron of
|
||
<taxonomicName id="5EFE4D1DFE58FFC67905F88212B1DFDA" box="[1063,1215,1905,1928]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="americanus">
|
||
<emphasis id="AB8AEA8CFE58FFC67905F88212B1DFDA" box="[1063,1215,1905,1928]" italics="true" pageId="4" pageNumber="41">L. americanus</emphasis>
|
||
</taxonomicName>
|
||
(magn. 2100 x);
|
||
<emphasis id="AB8AEA8CFE58FFC6785AF882139DDFDB" bold="true" box="[1400,1427,1905,1929]" pageId="4" pageNumber="41">18</emphasis>
|
||
) idem, (magn. 2600 x);
|
||
<emphasis id="AB8AEA8CFE58FFC67CB9F86717B9DFFE" bold="true" box="[411,439,1940,1964]" pageId="4" pageNumber="41">19</emphasis>
|
||
) metacoxa (MTX) and metaventrite (MTS) of
|
||
<taxonomicName id="5EFE4D1DFE58FFC67EEEF8661262DFFE" box="[972,1132,1941,1964]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="americanus">
|
||
<emphasis id="AB8AEA8CFE58FFC67EEEF8661262DFFE" box="[972,1132,1941,1964]" italics="true" pageId="4" pageNumber="41">L. americanus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="AB8AEA8CFE58FFC6795FF8671297DFFE" bold="true" box="[1149,1177,1940,1964]" pageId="4" pageNumber="41">20</emphasis>
|
||
) idem, detail at higher magnification;
|
||
<emphasis id="AB8AEA8CFE58FFC67C18F844175BDF9D" bold="true" box="[314,341,1975,1999]" pageId="4" pageNumber="41">21</emphasis>
|
||
) idem, detail of metaventrite (MTX) at higher magnification;
|
||
<emphasis id="AB8AEA8CFE58FFC67ED3F8441202DF9D" bold="true" box="[1009,1036,1975,1999]" pageId="4" pageNumber="41">22</emphasis>
|
||
) detail of elytron of
|
||
<taxonomicName id="5EFE4D1DFE58FFC679CEF84B1367DF9D" box="[1260,1385,1976,1999]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="apalodes">
|
||
<emphasis id="AB8AEA8CFE58FFC679CEF84B1367DF9D" box="[1260,1385,1976,1999]" italics="true" pageId="4" pageNumber="41">L. apalodes</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="AB8AEA8CFE58FFC6785AF844139DDF9D" bold="true" box="[1400,1427,1975,1999]" pageId="4" pageNumber="41">23</emphasis>
|
||
) detail of metacoxa of
|
||
<taxonomicName id="5EFE4D1DFE58FFC67C5DF82917F2DFA3" box="[383,508,2010,2033]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="apalodes">
|
||
<emphasis id="AB8AEA8CFE58FFC67C5DF82917F2DFA3" box="[383,508,2010,2033]" italics="true" pageId="4" pageNumber="41">L. apalodes</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="AB8AEA8CFE58FFC67F28F829142BDFA0" bold="true" box="[522,549,2010,2034]" pageId="4" pageNumber="41">24</emphasis>
|
||
) idem, detail at higher magnification;
|
||
<emphasis id="AB8AEA8CFE58FFC67E9DF82915D4DFA0" bold="true" box="[959,986,2010,2034]" pageId="4" pageNumber="41">25</emphasis>
|
||
) detail of metacoxa of
|
||
<taxonomicName id="5EFE4D1DFE58FFC679F0F8291347DFA3" box="[1234,1353,2010,2033]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="neblinae">
|
||
<emphasis id="AB8AEA8CFE58FFC679F0F8291347DFA3" box="[1234,1353,2010,2033]" italics="true" pageId="4" pageNumber="41">L. neblinae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9941369EFE59FFC77DE4FF6415AAD921" blockId="5.[151,1437,151,1299]" pageId="5" pageNumber="42">
|
||
<emphasis id="AB8AEA8CFE59FFC77DE4FF6417BFD8E3" bold="true" box="[198,433,151,177]" pageId="5" pageNumber="42">Taxonomic history</emphasis>
|
||
. After
|
||
<taxonomicName id="5EFE4D1DFE59FFC77F2EFF6B1532D8E0" authority="Leach, 1815" authorityName="Leach" authorityYear="1815" box="[524,828,152,178]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AB8AEA8CFE59FFC77F2EFF6B1495D8E3" box="[524,667,152,177]" italics="true" pageId="5" pageNumber="42">Laccophilus</emphasis>
|
||
<bibRefCitation id="FD6F4B6FFE59FFC77F85FF6B1532D8E0" author="Leach" box="[679,828,152,178]" pageId="5" pageNumber="58" refString="Leach, W. E. (1815) Entomology, pp. 57 - 172. In: Brewster D. Edinburgh Encyclopaedia. Vol. 9. Edinburgh: Balfour." type="book chapter" year="1815">Leach, 1815</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="5EFE4D1DFE59FFC77E5FFF6B12B9D8E0" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[893,1207,152,178]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AB8AEA8CFE59FFC77E5FFF6B1216D8E3" box="[893,1048,152,177]" italics="true" pageId="5" pageNumber="42">Neptosternus</emphasis>
|
||
<bibRefCitation id="FD6F4B6FFE59FFC77901FF6B12B9D8E0" author="Sharp" box="[1059,1207,152,178]" pageId="5" pageNumber="58" refString="Sharp, D. (1882) On aquatic carnivorous Coleoptera or Dytiscidae. The Scientific Transactions of the Royal Dublin Society (2) 2, 179 - 1003 + pls. 7 - 18." type="journal article" year="1882">Sharp, 1882</bibRefCitation>
|
||
</taxonomicName>
|
||
, this was the third genus of the
|
||
<taxonomicName id="5EFE4D1DFE59FFC77C10FF4D17D5D88A" box="[306,475,190,216]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="subFamily" subFamily="Laccophilinae">Laccophilinae</taxonomicName>
|
||
to be described (
|
||
<bibRefCitation id="FD6F4B6FFE59FFC77F8AFF4D1566D88A" author="Pederzani" box="[680,872,190,216]" pageId="5" pageNumber="58" refString="Pederzani, F. (1995) Keys to the identification of the genera and subgenera of adult Dytiscidae (sensu lato) of the world (Coleoptera: Dytiscidae). Atti dell'Accademia Roveretana degli Agiati 244 (1994), ser. VII, IV B, 5 - 83." type="book chapter" year="1995">Pederzani 1995</bibRefCitation>
|
||
). Since the original description, only two more species were added (
|
||
<bibRefCitation id="FD6F4B6FFE59FFC77CA4FF161428D8AD" author="Peschet" box="[390,550,229,255]" pageId="5" pageNumber="58" refString="Peschet, R. (1919) Description d'un Dytiscidae nouveaux de l'Amerique du Sud (Col.). Bulletin de la Societe Entomologique de France, 1919 (8), 145 - 147." type="journal article" year="1919">Peschet 1919</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="FD6F4B6FFE59FFC77F10FF1614D9D8AD" author="Guignot" box="[562,727,229,255]" pageId="5" pageNumber="58" refString="Guignot, F. (1955) Description de nouveaux Dytiscidae principalement de l'Amerique du Sud. Bulletin de l'Institut royal des Sciences naturelles de Belgique, 31 (27), 1 - 12." type="journal article" year="1955">Guignot 1955</bibRefCitation>
|
||
), and while the genus was occasionally mentioned in keys or catalogues, little has been published on
|
||
<taxonomicName id="5EFE4D1DFE59FFC77F48FEFF14E0D977" box="[618,750,268,293]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AB8AEA8CFE59FFC77F48FEFF14E0D977" box="[618,750,268,293]" italics="true" pageId="5" pageNumber="42">Laccodytes</emphasis>
|
||
</taxonomicName>
|
||
otherwise and confusion occurred related to the definition of this genus (
|
||
<bibRefCitation id="FD6F4B6FFE59FFC77C64FEC117EDD91E" author="Young" box="[326,483,306,332]" pageId="5" pageNumber="58" refString="Young, F. N. (1954) The water beetles of Florida. University of Florida, Studies in Biological Sciences, Series 5, 1 - 238." type="book chapter" year="1954">Young 1954</bibRefCitation>
|
||
). However, in a paper describing the genus
|
||
<taxonomicName id="5EFE4D1DFE59FFC77920FEC1128DD919" box="[1026,1155,306,331]" class="Insecta" family="Dytiscidae" genus="Napodytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AB8AEA8CFE59FFC77920FEC1128DD919" box="[1026,1155,306,331]" italics="true" pageId="5" pageNumber="42">Napodytes</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="E1E9760EFE59FFC779EDFEC11337D91E" box="[1231,1337,306,332]" name="Ecuador" pageId="5" pageNumber="42">Equador</collectingCountry>
|
||
,
|
||
<bibRefCitation id="FD6F4B6FFE59FFC77864FEC116EBD921" author="Steiner" pageId="5" pageNumber="58" refString="Steiner, W. E. (1981) A new genus and a new species of Laccophilinae water beetle from Ecuador (Coleoptera: Dytiscidae). The Pan-Pacific Entomologist, 57, 251 - 259." type="journal article" year="1981">Steiner (1981)</bibRefCitation>
|
||
suggested that
|
||
<taxonomicName id="5EFE4D1DFE59FFC77CB8FEAA1410D920" box="[410,542,345,370]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AB8AEA8CFE59FFC77CB8FEAA1410D920" box="[410,542,345,370]" italics="true" pageId="5" pageNumber="42">Laccodytes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="5EFE4D1DFE59FFC77F75FEAA14DDD920" box="[599,723,345,370]" class="Insecta" family="Dytiscidae" genus="Napodytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AB8AEA8CFE59FFC77F75FEAA14DDD920" box="[599,723,345,370]" italics="true" pageId="5" pageNumber="42">Napodytes</emphasis>
|
||
</taxonomicName>
|
||
are sister-groups.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D1E46515FE59FFC77DE4FE8C146BDD41" pageId="5" pageNumber="42" type="description">
|
||
<paragraph id="9941369EFE59FFC77DE4FE8C16C1DAD0" blockId="5.[151,1437,151,1299]" pageId="5" pageNumber="42">
|
||
<emphasis id="AB8AEA8CFE59FFC77DE4FE8C143DD9CB" bold="true" box="[198,563,383,409]" pageId="5" pageNumber="42">Phylogeny and Classification</emphasis>
|
||
. The small size, emarginate caudal margin of female sternite 7, the short segments 1–3 of pro and mesotarsus and the peculiar shape of the prosternal process are potential apomorphies for
|
||
<taxonomicName id="5EFE4D1DFE59FFC77C7CFE3E17ECD9B4" box="[350,482,461,486]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AB8AEA8CFE59FFC77C7CFE3E17ECD9B4" box="[350,482,461,486]" italics="true" pageId="5" pageNumber="42">Laccodytes</emphasis>
|
||
</taxonomicName>
|
||
. However, all of these characters also occur in
|
||
<taxonomicName id="5EFE4D1DFE59FFC7792AFE3E133BD9B5" authority="Steiner (1981)" authorityName="Steiner" authorityYear="1981" box="[1032,1333,461,487]" class="Insecta" family="Dytiscidae" genus="Napodytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AB8AEA8CFE59FFC7792AFE3E128AD9B4" box="[1032,1156,461,486]" italics="true" pageId="5" pageNumber="42">Napodytes</emphasis>
|
||
<bibRefCitation id="FD6F4B6FFE59FFC779A9FE3E133BD9B5" author="Steiner" box="[1163,1333,461,487]" pageId="5" pageNumber="58" refString="Steiner, W. E. (1981) A new genus and a new species of Laccophilinae water beetle from Ecuador (Coleoptera: Dytiscidae). The Pan-Pacific Entomologist, 57, 251 - 259." type="journal article" year="1981">Steiner (1981)</bibRefCitation>
|
||
</taxonomicName>
|
||
. For that reason, Steiner suggested that
|
||
<taxonomicName id="5EFE4D1DFE59FFC77CDBFE071473DA5F" box="[505,637,500,525]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AB8AEA8CFE59FFC77CDBFE071473DA5F" box="[505,637,500,525]" italics="true" pageId="5" pageNumber="42">Laccodytes</emphasis>
|
||
</taxonomicName>
|
||
+
|
||
<taxonomicName id="5EFE4D1DFE59FFC77FBEFE071516DA5F" box="[668,792,500,525]" class="Insecta" family="Dytiscidae" genus="Napodytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AB8AEA8CFE59FFC77FBEFE071516DA5F" box="[668,792,500,525]" italics="true" pageId="5" pageNumber="42">Napodytes</emphasis>
|
||
</taxonomicName>
|
||
are sister taxa.
|
||
<bibRefCitation id="FD6F4B6FFE59FFC77EECFE071278DA5C" author="Steiner" box="[974,1142,500,526]" pageId="5" pageNumber="58" refString="Steiner, W. E. (1981) A new genus and a new species of Laccophilinae water beetle from Ecuador (Coleoptera: Dytiscidae). The Pan-Pacific Entomologist, 57, 251 - 259." type="journal article" year="1981">Steiner (1981)</bibRefCitation>
|
||
characterized
|
||
<taxonomicName id="5EFE4D1DFE59FFC77802FE071395DA5F" box="[1312,1435,500,525]" class="Insecta" family="Dytiscidae" genus="Napodytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AB8AEA8CFE59FFC77802FE071395DA5F" box="[1312,1435,500,525]" italics="true" pageId="5" pageNumber="42">Napodytes</emphasis>
|
||
</taxonomicName>
|
||
based on two apomorphies, which are in fact autapomorphies for the only species of the genus,
|
||
<taxonomicName id="5EFE4D1DFE59FFC77860FDE91735DA09" authority="Steiner, 1981" authorityName="Steiner" authorityYear="1981" class="Insecta" family="Dytiscidae" genus="Napodytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="boki">
|
||
<emphasis id="AB8AEA8CFE59FFC77860FDE91392DA61" box="[1346,1436,538,563]" italics="true" pageId="5" pageNumber="42">N. boki</emphasis>
|
||
<bibRefCitation id="FD6F4B6FFE59FFC77DB5FDB21735DA09" author="Steiner" box="[151,315,577,603]" pageId="5" pageNumber="58" refString="Steiner, W. E. (1981) A new genus and a new species of Laccophilinae water beetle from Ecuador (Coleoptera: Dytiscidae). The Pan-Pacific Entomologist, 57, 251 - 259." type="journal article" year="1981">Steiner, 1981</bibRefCitation>
|
||
</taxonomicName>
|
||
: (1) some male antennomeres enlarged, and (2) hind tibia in both sexes with only one apical spur.
|
||
</paragraph>
|
||
<paragraph id="9941369EFE59FFC77DE7FD7D16F3DBE5" blockId="5.[151,1437,151,1299]" pageId="5" pageNumber="42">
|
||
On the other hand,
|
||
<taxonomicName id="5EFE4D1DFE59FFC77C8AFD7C1422DAFA" box="[424,556,655,680]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AB8AEA8CFE59FFC77C8AFD7C1422DAFA" box="[424,556,655,680]" italics="true" pageId="5" pageNumber="42">Laccodytes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="5EFE4D1DFE59FFC77F44FD7C14ECDAFA" box="[614,738,655,680]" class="Insecta" family="Dytiscidae" genus="Napodytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AB8AEA8CFE59FFC77F44FD7C14ECDAFA" box="[614,738,655,680]" italics="true" pageId="5" pageNumber="42">Napodytes</emphasis>
|
||
</taxonomicName>
|
||
appear externally similar to
|
||
<taxonomicName id="5EFE4D1DFE59FFC77916FD7C12C1DAFA" box="[1076,1231,655,680]" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AB8AEA8CFE59FFC77916FD7C12C1DAFA" box="[1076,1231,655,680]" italics="true" pageId="5" pageNumber="42">Neptosternus</emphasis>
|
||
</taxonomicName>
|
||
, with which they share: border of lateral sides of elytra visible in dorsal view, metatarsi poorly broadened, and long and slender fore and middle legs. This may suggest a closer relationship between these three genera. In a molecular phylogenetic analysis of the
|
||
<taxonomicName id="5EFE4D1DFE59FFC77CC9FCF11462DB4E" box="[491,620,770,796]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="family">Dytiscidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="5EFE4D1DFE59FFC77F5BFCF114F0DB49" box="[633,766,770,795]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AB8AEA8CFE59FFC77F5BFCF114F0DB49" box="[633,766,770,795]" italics="true" pageId="5" pageNumber="42">Laccodytes</emphasis>
|
||
</taxonomicName>
|
||
grouped with the Australian
|
||
<taxonomicName id="5EFE4D1DFE59FFC77978FCF112F7DB49" box="[1114,1273,770,795]" class="Insecta" family="Dytiscidae" genus="Australphilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AB8AEA8CFE59FFC77978FCF112F7DB49" box="[1114,1273,770,795]" italics="true" pageId="5" pageNumber="42">Australphilus</emphasis>
|
||
</taxonomicName>
|
||
, and both are in a clade with the other strictly running water
|
||
<taxonomicName id="5EFE4D1DFE59FFC77FF5FCDA157ADB11" box="[727,884,809,835]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
|
||
,
|
||
<taxonomicName id="5EFE4D1DFE59FFC77EA3FCDA122FDB10" box="[897,1057,809,834]" class="Insecta" family="Dytiscidae" genus="Philaccolilus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AB8AEA8CFE59FFC77EA3FCDA122FDB10" box="[897,1057,809,834]" italics="true" pageId="5" pageNumber="42">Philaccolilus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="5EFE4D1DFE59FFC77913FCDA12CEDB10" box="[1073,1216,809,834]" class="Insecta" family="Dytiscidae" genus="Philaccolus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AB8AEA8CFE59FFC77913FCDA12CEDB10" box="[1073,1216,809,834]" italics="true" pageId="5" pageNumber="42">Philaccolus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="5EFE4D1DFE59FFC779DEFCDA1787DB38" authority="Ribera et al. 2008" authorityName="Ribera et al." authorityYear="2008" class="Insecta" family="Dytiscidae" genus="Neptosternus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AB8AEA8CFE59FFC779DEFCDA1392DB10" box="[1276,1436,809,834]" italics="true" pageId="5" pageNumber="42">Neptosternus</emphasis>
|
||
(
|
||
<bibRefCitation id="FD6F4B6FFE59FFC77DBDFCA3178EDB38" author="Ribera" box="[159,384,848,874]" pageId="5" pageNumber="58" refString="Ribera, I., Vogler, A. P. & Balke, M. (2008) Phylogeny and diversification of diving beetles (Coleoptera: Dytiscidae). Cladistics, 24 (2008), 563 - 590." type="journal article" year="2008">
|
||
Ribera
|
||
<emphasis id="AB8AEA8CFE59FFC77DDBFCA31739DB3B" box="[249,311,848,873]" italics="true" pageId="5" pageNumber="42">et al.</emphasis>
|
||
2008
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. (Note: in that paper,
|
||
<taxonomicName id="5EFE4D1DFE59FFC77FB0FCA31517DB3B" box="[658,793,848,873]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AB8AEA8CFE59FFC77FB0FCA31517DB3B" box="[658,793,848,873]" italics="true" pageId="5" pageNumber="42">Laccodytes</emphasis>
|
||
</taxonomicName>
|
||
sp.2 is in fact
|
||
<taxonomicName id="5EFE4D1DFE59FFC77EEFFCA312E9DB3B" box="[973,1255,848,873]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="bassignanii">
|
||
<emphasis id="AB8AEA8CFE59FFC77EEFFCA312E9DB3B" box="[973,1255,848,873]" italics="true" pageId="5" pageNumber="42">Laccodytes bassignanii</emphasis>
|
||
</taxonomicName>
|
||
, and what was named
|
||
<taxonomicName id="5EFE4D1DFE59FFC77DCDFC841779DBC2" box="[239,375,887,912]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AB8AEA8CFE59FFC77DCDFC841779DBC2" box="[239,375,887,912]" italics="true" pageId="5" pageNumber="42">Laccodytes</emphasis>
|
||
</taxonomicName>
|
||
sp. 1 belongs to a new genus of small
|
||
<taxonomicName id="5EFE4D1DFE59FFC77E71FC8515E1DBC2" box="[851,1007,886,912]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
|
||
, which are however stagnant water species).
|
||
</paragraph>
|
||
<paragraph id="9941369EFE59FFC77DE4FC37146BDD41" blockId="5.[151,1437,151,1299]" pageId="5" pageNumber="42">
|
||
<taxonomicName id="5EFE4D1DFE59FFC77DE4FC371744DB8F" box="[198,330,964,989]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="AB8AEA8CFE59FFC77DE4FC371744DB8F" box="[198,330,964,989]" italics="true" pageId="5" pageNumber="42">Laccodytes</emphasis>
|
||
</taxonomicName>
|
||
can easily be grouped in two species-groups. The
|
||
<taxonomicName id="5EFE4D1DFE59FFC77E81FC371223DB8F" box="[931,1069,964,989]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="apalodes">
|
||
<emphasis id="AB8AEA8CFE59FFC77E81FC371223DB8F" box="[931,1069,964,989]" italics="true" pageId="5" pageNumber="42">L. apalodes</emphasis>
|
||
</taxonomicName>
|
||
-group with
|
||
<taxonomicName id="5EFE4D1DFE59FFC779E2FC371344DB8F" box="[1216,1354,964,989]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="apalodes">
|
||
<emphasis id="AB8AEA8CFE59FFC779E2FC371344DB8F" box="[1216,1354,964,989]" italics="true" pageId="5" pageNumber="42">L. apalodes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="5EFE4D1DFE59FFC778A6FC37170CDC51" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="rondonia">
|
||
<emphasis id="AB8AEA8CFE59FFC778A6FC37170CDC51" italics="true" pageId="5" pageNumber="42">L. rondonia</emphasis>
|
||
</taxonomicName>
|
||
is characterized by a long needle-like prosternal process extended caudally past the mesocoxae for a third of its length (
|
||
<figureCitation id="01C52A1BFE59FFC77C5AFBE217EADC79" box="[376,484,1041,1067]" captionStart="FIGURES 3 – 16" captionStartId="3.[151,270,1860,1884]" captionTargetBox="[248,1343,197,1819]" captionTargetId="figure@3.[248,1348,197,1827]" captionTargetPageId="3" captionText="FIGURES 3 – 16. SEMs of some Laccodytes species structures: 3) segments 1 - 3 in male protarsus of L. americanus; 4) prosternal process of L. apalodes; 5) idem, L. americanus; 6) idem, L. neblinae; 7) metaventrite of L. apalodes, the arrow indicates the groove in which the tip of prosternal process fits; 8) idem, L. neblinae; 9) metacoxal process of L. apalodes; 10) idem, L. americanus; 11) idem, L. neblinae; 12) hind tarsus of L. apalodes (male); 13) idem, L. americanus (male); 14) idem, L. neblinae (female); 15) ovipositor of L. apalodes; 16) idem, L. americanus." httpUri="https://zenodo.org/record/193408/files/figure.png" pageId="5" pageNumber="42">Figs 4, 7</figureCitation>
|
||
), the acute posterior pronotal angles (
|
||
<figureCitation id="01C52A1BFE59FFC77E88FBE2120ADC79" box="[938,1028,1041,1067]" captionStart="FIGURES 26 – 27" captionStartId="7.[151,270,879,903]" captionTargetBox="[151,1408,206,851]" captionTargetId="figure@7.[151,1436,197,854]" captionTargetPageId="7" captionText="FIGURES 26 – 27. Hind angle of pronotum of: 26) Laccodytes rondonia; 27) L. bassignanii." httpUri="https://zenodo.org/record/193410/files/figure.png" pageId="5" pageNumber="42">Fig. 26</figureCitation>
|
||
), a membranous expansion on the median lobe of the aedeagus (
|
||
<figureCitation id="01C52A1BFE59FFC77CD5FBCB1470DC00" box="[503,638,1080,1106]" captionStart="FIGURES 44 – 47" captionStartId="10.[151,270,1823,1847]" captionTargetBox="[180,1399,210,1765]" captionTargetId="figure@10.[171,1415,197,1798]" captionTargetPageId="10" captionText="FIGURES 44 – 47. Male genitals in Laccodytes species: 44) L. apalodes: a) median lobe of aedeagus in lateral view, b – c) right and left parameres; 45) L. rondonia: idem; 46) L. americanus: idem; 47) L. phalacroides: a) apical half of median lobe of aedeagus in dorsal view, b) median lobe in lateral view, c – d) right and left parameres;" httpUri="https://zenodo.org/record/193413/files/figure.png" pageId="5" pageNumber="42">Figs 44, 45</figureCitation>
|
||
) and the slightly dilated male pro- and mesotarsomeres 1–3. The
|
||
<taxonomicName id="5EFE4D1DFE59FFC778A6FBCB1721DC2A" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="phalacroides">
|
||
<emphasis id="AB8AEA8CFE59FFC778A6FBCB1721DC2A" italics="true" pageId="5" pageNumber="42">L. phalacroides</emphasis>
|
||
</taxonomicName>
|
||
-group includes all other species. Its members have a shorter tip of prosternal process not going beyond mesocoxae or very hardly so (
|
||
<figureCitation id="01C52A1BFE59FFC77F70FB7614D8DCCD" box="[594,726,1157,1183]" captionStart="FIGURES 3 – 16" captionStartId="3.[151,270,1860,1884]" captionTargetBox="[248,1343,197,1819]" captionTargetId="figure@3.[248,1348,197,1827]" captionTargetPageId="3" captionText="FIGURES 3 – 16. SEMs of some Laccodytes species structures: 3) segments 1 - 3 in male protarsus of L. americanus; 4) prosternal process of L. apalodes; 5) idem, L. americanus; 6) idem, L. neblinae; 7) metaventrite of L. apalodes, the arrow indicates the groove in which the tip of prosternal process fits; 8) idem, L. neblinae; 9) metacoxal process of L. apalodes; 10) idem, L. americanus; 11) idem, L. neblinae; 12) hind tarsus of L. apalodes (male); 13) idem, L. americanus (male); 14) idem, L. neblinae (female); 15) ovipositor of L. apalodes; 16) idem, L. americanus." httpUri="https://zenodo.org/record/193408/files/figure.png" pageId="5" pageNumber="42">Figs 5, 6, 8</figureCitation>
|
||
,
|
||
<figureCitation id="01C52A1BFE59FFC77FC1FB761554DCCD" box="[739,858,1157,1183]" captionStart="FIGURES 40 – 43" captionStartId="9.[151,276,898,922]" captionTargetBox="[156,1431,199,863]" captionTargetId="figure@9.[151,1436,194,874]" captionTargetPageId="9" captionText="FIGURES 40 – 43. Line drawings of prosternal process: 40) Laccodytes takutuanus; 41) L. phalacroides; 42) L. rondonia; 43) L. olibroides." httpUri="https://zenodo.org/record/193412/files/figure.png" pageId="5" pageNumber="42">40, 41, 43</figureCitation>
|
||
), rounded posterior angles of pronotum (
|
||
<figureCitation id="01C52A1BFE59FFC77815FB761380DCCD" box="[1335,1422,1157,1183]" captionStart="FIGURES 26 – 27" captionStartId="7.[151,270,879,903]" captionTargetBox="[151,1408,206,851]" captionTargetId="figure@7.[151,1436,197,854]" captionTargetPageId="7" captionText="FIGURES 26 – 27. Hind angle of pronotum of: 26) Laccodytes rondonia; 27) L. bassignanii." httpUri="https://zenodo.org/record/193410/files/figure.png" pageId="5" pageNumber="42">Fig. 27</figureCitation>
|
||
), no membranous expansions on the median lobe of the aedeagus and male pro- and mesotarsi not dilated. These character states, except for the latter, might in all cases represent the plesiomorphic state as they are generally present in other
|
||
<taxonomicName id="5EFE4D1DFE59FFC77CEBFB0A146FDD41" box="[457,609,1273,1299]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="tribe" tribe="Laccophilini">Laccophilini</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |