147 lines
16 KiB
XML
147 lines
16 KiB
XML
<document id="0119234A9B30D50A8AAABB9052393C54" ID-DOI="10.5281/zenodo.193406" ID-GBIF-Dataset="5e8043a5-e685-4a21-b696-0f7a706ab223" ID-ISSN="1175-5326" ID-Zenodo-Dep="193406" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460138703176" checkinUser="plazi" docAuthor="Toledo, Mario, Spangler, Paul J. & Balke, Michael" docDate="2010" docId="11578788FE4CFFD37DB5FC6E1577D88A" docLanguage="en" docName="zt02347p058.pdf" docOrigin="Zootaxa 2347" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Laccodytes obscuratus Toledo, Spangler & Balke, 2010, sp.n." docType="treatment" docVersion="9" lastPageNumber="54" masterDocId="ED6EFFF0FE5CFFC27D22FFF3160ED852" masterDocTitle="Taxonomic revision of the Neotropical diving beetles genus Laccodytes Régimbart, 1895 (Coleoptera: Dytiscidae)" masterLastPageNumber="58" masterPageNumber="37" pageNumber="53" updateTime="1698235333712" updateUser="plazi">
|
||
<mods:mods id="2BAC94736B6FBDBF2A250922EBE37B77" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="99469E8E4D32E628DFF72697BAFEC31C">
|
||
<mods:title id="BDF80483BA87ED16FFA8BE5C6FED4890">Taxonomic revision of the Neotropical diving beetles genus Laccodytes Régimbart, 1895 (Coleoptera: Dytiscidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="9F893E4C5C23D6A1DDD475C7166677A7" type="personal">
|
||
<mods:role id="22562A8DB6CC91B1E96707A064E39CB9">
|
||
<mods:roleTerm id="BAA3124D785662DD3D1519A27970424F">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="0135DDA2EFABCCCDFB8486ED9BEDFAAF">Toledo, Mario</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="EB52A0587E7EB07ADDD665A2D51053E1" type="personal">
|
||
<mods:role id="8F86B5FA3E9389F9D9A8E21208A142D0">
|
||
<mods:roleTerm id="71B67CB3A7CDE0A0A89E2630C8D268A3">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="4A745EF6BF673BC04914E84B47A43388">Spangler, Paul J.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="1E9143E73C002F3F449CE821A2B3026C" type="personal">
|
||
<mods:role id="4567AEE5156D5DC0A64595E344F62168">
|
||
<mods:roleTerm id="AD56CEC72A6BA0380FF25AB94127AAEC">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="47637D9001AFDD194B297ED556FB17D6">Balke, Michael</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="040DB9C02EED7B61071C7530CC5A8F13">text</mods:typeOfResource>
|
||
<mods:relatedItem id="34AF72868FCAC84B32EE69756331D36B" type="host">
|
||
<mods:titleInfo id="760D986867DE186D885E48BC7BA1049E">
|
||
<mods:title id="81E244355E36E77068602219E19A46F0">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="668B6E5281F5BBC12055A81DDC22CAC5">
|
||
<mods:date id="AD3120B7C7071DDE0BE0D0CCD3E09BFE">2010</mods:date>
|
||
<mods:detail id="E7F64625DF158522CC910DB04DAC1713" type="volume">
|
||
<mods:number id="FB1F3975EE13B62CFDE55A000529E048">2347</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="3530FFFE5D20B14F61476906AAC7BF59" unit="page">
|
||
<mods:start id="F379AA81F99EF99C7B1F9344279DD082">37</mods:start>
|
||
<mods:end id="2193B00BA1ACAF7CFF19BCFA50F711CD">58</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="5BAD36BBFA6CB3E737245504A4A271F1">journal article</mods:classification>
|
||
<mods:identifier id="DCD326710FBACBAEC5ECB750FE302C4A" type="DOI">10.5281/zenodo.193406</mods:identifier>
|
||
<mods:identifier id="AB6B91B6484D846CC2B2B24234D26B9C" type="GBIF-Dataset">5e8043a5-e685-4a21-b696-0f7a706ab223</mods:identifier>
|
||
<mods:identifier id="A7A97977CC576F0231DAAAD83882E3B5" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="38A43062581F1DCA4A8BDF6194965022" type="Zenodo-Dep">193406</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="11578788FE4CFFD37DB5FC6E1577D88A" ID-DOI="http://doi.org/10.5281/zenodo.6198386" ID-GBIF-Taxon="124955832" ID-Zenodo-Dep="6198386" LSID="urn:lsid:plazi:treatment:11578788FE4CFFD37DB5FC6E1577D88A" httpUri="http://treatment.plazi.org/id/11578788FE4CFFD37DB5FC6E1577D88A" lastPageId="17" lastPageNumber="54" pageId="16" pageNumber="53">
|
||
<subSubSection id="D1E46515FE4CFFD27DB5FC6E1757DB88" pageId="16" pageNumber="53" type="nomenclature">
|
||
<paragraph id="9941369EFE4CFFD27DB5FC6E17E4DBE5" blockId="16.[151,490,925,986]" box="[151,490,925,951]" pageId="16" pageNumber="53">
|
||
<heading id="C20981F2FE4CFFD27DB5FC6E17E4DBE5" bold="true" box="[151,490,925,951]" fontSize="11" level="1" pageId="16" pageNumber="53" reason="1">
|
||
<emphasis id="AB8AEA8CFE4CFFD27DB5FC6E17E4DBE5" bold="true" box="[151,490,925,951]" pageId="16" pageNumber="53">
|
||
<taxonomicName id="5EFE4D1DFE4CFFD27DB5FC6E17A6DBE5" ID-CoL="3RLZJ" box="[151,424,925,951]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="53" phylum="Arthropoda" rank="species" species="obscuratus" status="sp. nov.">
|
||
<emphasis id="AB8AEA8CFE4CFFD27DB5FC6E17A6DBE5" bold="true" box="[151,424,925,951]" italics="true" pageId="16" pageNumber="53">Laccodytes obscuratus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B0B957F7FE4CFFD27C8DFC6E17E4DBE5" box="[431,490,925,951]" pageId="16" pageNumber="53" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="9941369EFE4CFFD27DB5FC331757DB88" blockId="16.[151,490,925,986]" box="[151,345,960,986]" pageId="16" pageNumber="53">Figs (35, 51, 62)</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D1E46515FE4CFFD27DB5FBFE1549DCC9" pageId="16" pageNumber="53" type="materials_examined">
|
||
<paragraph id="9941369EFE4CFFD27DB5FBFE154ADC75" blockId="16.[151,1437,1037,2030]" box="[151,836,1037,1063]" pageId="16" pageNumber="53">
|
||
<emphasis id="AB8AEA8CFE4CFFD27DB5FBFE1733DC75" bold="true" box="[151,317,1037,1063]" pageId="16" pageNumber="53">
|
||
<typeStatus id="4645883CFE4CFFD27DB5FBFE16DADC75" box="[151,212,1037,1063]" pageId="16" pageNumber="53">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="E1E9760EFE4CFFD27C66FBFE17CFDC75" box="[324,449,1037,1063]" name="Venezuela" pageId="16" pageNumber="53">Venezuela</collectingCountry>
|
||
: Amazonas, Cerro de la Neblina.
|
||
</paragraph>
|
||
<paragraph id="9941369EFE4CFFD27DE4FBC01549DCC9" blockId="16.[151,1437,1037,2030]" pageId="16" pageNumber="53">
|
||
<emphasis id="AB8AEA8CFE4CFFD27DE4FBC01770DC1F" bold="true" box="[198,382,1075,1101]" pageId="16" pageNumber="53">
|
||
<typeStatus id="4645883CFE4CFFD27DE4FBC0170DDC1F" box="[198,259,1075,1101]" pageId="16" pageNumber="53">Type</typeStatus>
|
||
material.
|
||
</emphasis>
|
||
<typeStatus id="4645883CFE4CFFD27CA5FBC717FADC1C" box="[391,500,1076,1102]" pageId="16" pageNumber="53" type="holotype">Holotype</typeStatus>
|
||
3 (
|
||
<collectionCode id="FFEFAE5BFE4CFFD27F07FBC7148FDC1C" LSID="urn:lsid:biocol.org:col:34871" box="[549,641,1076,1102]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="53">USNM</collectionCode>
|
||
):
|
||
<collectingCountry id="E1E9760EFE4CFFD27FB4FBC7151FDC1C" box="[662,785,1076,1102]" name="Venezuela" pageId="16" pageNumber="53">Venezuela</collectingCountry>
|
||
, TF Amazonas Cerro de la Neblina, basecamp
|
||
<quantity id="5E069B7BFE4CFFD2786BFBC71396DC1C" box="[1353,1432,1076,1102]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="16" pageNumber="53" unit="m" value="140.0">140 m</quantity>
|
||
,
|
||
<geoCoordinate id="FCCA5059FE4CFFD27DB5FBA916E2DC26" box="[151,236,1114,1140]" direction="north" orientation="latitude" pageId="16" pageNumber="53" precision="925" value="0.8333333">0°50'N</geoCoordinate>
|
||
<geoCoordinate id="FCCA5059FE4CFFD27DD6FBA91751DC26" box="[244,351,1114,1140]" direction="west" orientation="longitude" pageId="16" pageNumber="53" precision="925" value="-66.166664">66°10'W</geoCoordinate>
|
||
,
|
||
<date id="ED40105EFE4CFFD27C4FFBA917EADC26" box="[365,484,1114,1140]" pageId="16" pageNumber="53" value="1985-01-27">27.I.1985</date>
|
||
, P.J. Spangler, P.M. Spangler, R. Faitoute & W.E. Steiner (
|
||
<collectionCode id="FFEFAE5BFE4CFFD27994FBA9131ADC26" LSID="urn:lsid:biocol.org:col:34871" box="[1206,1300,1114,1140]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="53">USNM</collectionCode>
|
||
).
|
||
<typeStatus id="4645883CFE4CFFD2780BFBA91396DC26" box="[1321,1432,1114,1140]" pageId="16" pageNumber="53" type="paratype">Paratype</typeStatus>
|
||
: Same data as
|
||
<typeStatus id="4645883CFE4CFFD27C15FB721790DCC9" box="[311,414,1153,1179]" pageId="16" pageNumber="53" type="holotype">holotype</typeStatus>
|
||
except for
|
||
<date id="ED40105EFE4CFFD27F00FB721492DCC9" box="[546,668,1153,1179]" pageId="16" pageNumber="53" value="1985-02-27">27.II.1985</date>
|
||
(1 3
|
||
<collectionCode id="FFEFAE5BFE4CFFD27FC3FB721535DCC9" LSID="urn:lsid:biocol.org:col:34871" box="[737,827,1153,1179]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="16" pageNumber="53">USNM</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D1E46515FE4CFFD27DE4FB5417A9DF95" pageId="16" pageNumber="53" type="diagnosis">
|
||
<paragraph id="9941369EFE4CFFD27DE4FB54127BDCBA" blockId="16.[151,1437,1037,2030]" pageId="16" pageNumber="53">
|
||
<emphasis id="AB8AEA8CFE4CFFD27DE4FB541748DC93" bold="true" box="[198,326,1191,1217]" pageId="16" pageNumber="53">Diagnosis.</emphasis>
|
||
Habitus (
|
||
<figureCitation id="01C52A1BFE4CFFD27CE3FB5B1413DC90" box="[449,541,1192,1218]" captionStart="FIGURES 28 – 39" captionStartId="8.[151,272,1875,1899]" captionTargetBox="[154,1429,467,1844]" captionTargetId="figure@8.[151,1436,467,1848]" captionTargetPageId="8" captionText="FIGURES 28 – 39. Subschematic body shape and colour patterns: 28) Laccodytes apalodes; 29) L. rondonia; 30) L. americanus; 31) L. phalacroides; 32) L. takutuanus; 33 a – 33 b) L. neblinae (variation of elytral patterns); 34) L. bassignanii; 35) L. obscuratus; 36) L. olibroides; 37) L. androginus; 38) Laccodytes sp. 1.; 39) Laccodytes sp. 2." httpUri="https://zenodo.org/record/193411/files/figure.png" pageId="16" pageNumber="53">Fig. 35</figureCitation>
|
||
). Body length 1.6 mm. Body slightly drop-shaped, tip of elytron slightly truncate.
|
||
<collectingCountry id="E1E9760EFE4CFFD27C26FB3D1731DCBA" box="[260,319,1230,1256]" name="India" pageId="16" pageNumber="53">Hind</collectingCountry>
|
||
angle of pronotum rounded. No angle between pronotum and elytron.
|
||
</paragraph>
|
||
<paragraph id="9941369EFE4CFFD27DE4FB0615B5DDF8" blockId="16.[151,1437,1037,2030]" pageId="16" pageNumber="53">
|
||
Color. Head dark brown. Pronotum dark yellow; posterior 1/3 of prontoum darker, brownish. Elytron brown with obscure paler pattern of dark yellowish patches in the following positions (refer to
|
||
<figureCitation id="01C52A1BFE4CFFD27838FAEF137FDD64" box="[1306,1393,1308,1334]" captionStart="FIGURES 28 – 39" captionStartId="8.[151,272,1875,1899]" captionTargetBox="[154,1429,467,1844]" captionTargetId="figure@8.[151,1436,467,1848]" captionTargetPageId="8" captionText="FIGURES 28 – 39. Subschematic body shape and colour patterns: 28) Laccodytes apalodes; 29) L. rondonia; 30) L. americanus; 31) L. phalacroides; 32) L. takutuanus; 33 a – 33 b) L. neblinae (variation of elytral patterns); 34) L. bassignanii; 35) L. obscuratus; 36) L. olibroides; 37) L. androginus; 38) Laccodytes sp. 1.; 39) Laccodytes sp. 2." httpUri="https://zenodo.org/record/193411/files/figure.png" pageId="16" pageNumber="53">Fig. 35</figureCitation>
|
||
for explanation): a subbasal band and a larger apical patch. These patches are not contrasting and sometimes extremely obscure. The subbasal band reaches the margin of elytron, but hardly the anterior angle. Appendages yellowish. Venter, including epipleuron, dark brownish.
|
||
</paragraph>
|
||
<paragraph id="9941369EFE4CFFD27DE4FA451555DE4C" blockId="16.[151,1437,1037,2030]" pageId="16" pageNumber="53">Scultpure. The entire dorsal and ventral surface of the beetle with a faint MR. Fine and sparse dots are visible on metaventrite. No longitudinal lines or longish meshes visible. Few, transverse, shallow grooves are hardly visible at about the anterior third of metacoxal plates.</paragraph>
|
||
<paragraph id="9941369EFE4CFFD27DE4F9D914AFDE8D" blockId="16.[151,1437,1037,2030]" pageId="16" pageNumber="53">
|
||
Structures. Pronotum with narrow lateral bead; posterior angle rounded. Prosternum and prosternal process with small ridge; prosternal process broadly carinate and with a rather long, needle shaped, acute tip. Epipleuron broad up to level of sternite 6 or 7. Fore and middle legs very long and slender. Metatarsomeres 1– 4 with apico-lateral angle slightly lobed.
|
||
<collectingCountry id="E1E9760EFE4CFFD27F5BF96D14BADEEA" box="[633,692,1694,1720]" name="India" pageId="16" pageNumber="53">Hind</collectingCountry>
|
||
lobes of metacoxal process almost straight, with a very narrow V-shaped notch in the middle between them.
|
||
</paragraph>
|
||
<paragraph id="9941369EFE4CFFD27DE4F91F14EEDFF2" blockId="16.[151,1437,1037,2030]" pageId="16" pageNumber="53">
|
||
Male. Pro- and mesotarsi not dilated.
|
||
<collectingCountry id="E1E9760EFE4CFFD27F82F91F14D5DF54" box="[672,731,1772,1798]" name="India" pageId="16" pageNumber="53">Hind</collectingCountry>
|
||
margin of sternite 7 rounded, or very slightly tectiform. Aedeagus (
|
||
<figureCitation id="01C52A1BFE4CFFD27C3EF8E1177BDF7E" box="[284,373,1810,1836]" captionStart="FIGURES 48 – 53" captionStartId="11.[151,270,1543,1567]" captionTargetBox="[189,1376,223,1499]" captionTargetId="figure@11.[184,1403,197,1518]" captionTargetPageId="11" captionText="FIGURES 48 – 53. Male genitals in Laccodytes species: 48) L. takutuanus: a) median lobe of aedeagus in dorsal view, b) in lateral view, c-d) right and left parameres; 49) L. neblinae: a) median lobe of aedeagus in dorsal view, b) in lateral view, c-d) right and left parameres; 50) L. bassignanii: a) median lobe of aedeagus in dorsal view, b-c) in dorsal-lateral and lateral view, d-e) right and left parameres; 51) L. obscuratus: a) median lobe of aedeagus in dorsal view, b) in lateral view, c-d) right and left parameres; 52) L. olibroides: a) median lobe of aedeagus in lateral view, b-c) right an left parameres; 53) L. androginus: a) median lobe of aedeagus in dorsal view, b) in lateral view, c-d) right and left parameres." httpUri="https://zenodo.org/record/193414/files/figure.png" pageId="16" pageNumber="53">Fig. 51</figureCitation>
|
||
): median lobe, in lateral view, very long, thin and straight in the distal 2/3, ending with a rounded tip, gently bent down; the basal last third is broad. In dorsal view elongate and flat, slightly broadened on distal half, with tip rounded and gently bent right. Parameres different in shape but almost of the same size, both elongated, with a long apical seta.
|
||
</paragraph>
|
||
<paragraph id="9941369EFE4CFFD27DE4F85E17A9DF95" blockId="16.[151,1437,1037,2030]" box="[198,423,1965,1991]" pageId="16" pageNumber="53">Female. Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D1E46515FE4CFFD27DE4F82014C3DFBC" box="[198,717,2003,2030]" pageId="16" pageNumber="53" type="distribution">
|
||
<paragraph id="9941369EFE4CFFD27DE4F82014C3DFBC" blockId="16.[151,1437,1037,2030]" box="[198,717,2003,2030]" pageId="16" pageNumber="53">
|
||
<emphasis id="AB8AEA8CFE4CFFD27DE4F8201751DFBF" bold="true" box="[198,351,2003,2029]" pageId="16" pageNumber="53">Distribution</emphasis>
|
||
(
|
||
<figureCitation id="01C52A1BFE4CFFD27C4DF82717C9DFBC" box="[367,455,2004,2030]" captionStart="FIGURE 62" captionStartId="18.[151,255,1941,1965]" captionText="FIGURE 62. Known distribution of Laccodytes species." pageId="16" pageNumber="53">Fig. 62</figureCitation>
|
||
). Southern
|
||
<collectingCountry id="E1E9760EFE4CFFD27F6EF82714C6DFBC" box="[588,712,2004,2030]" name="Venezuela" pageId="16" pageNumber="53">Venezuela</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D1E46515FE4DFFD37DE4FF641577D88A" pageId="17" pageNumber="54" type="biology_ecology">
|
||
<paragraph id="9941369EFE4DFFD37DE4FF641577D88A" blockId="17.[198,1126,151,216]" pageId="17" pageNumber="54">
|
||
<emphasis id="AB8AEA8CFE4DFFD37DE4FF641724D8E3" bold="true" box="[198,298,151,177]" pageId="17" pageNumber="54">Biology.</emphasis>
|
||
The two specimens were seined from rocks in the rapids of Rio Baria.
|
||
<emphasis id="AB8AEA8CFE4DFFD37DE4FF4D17A3D88A" bold="true" box="[198,429,190,216]" pageId="17" pageNumber="54">Derivatio nominis.</emphasis>
|
||
From the dark coloration of the beetle.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |