343 lines
39 KiB
XML
343 lines
39 KiB
XML
<document id="0A0F3DEB3467612FCADE4C74FE440D73" ID-DOI="10.11646/zootaxa.5319.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="8182108" ID-ZooBank="45CD8E29-3320-42E2-A8A8-3A36BC0F41CE" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1690276500814" checkinUser="plazi" docAuthor="Hashizume, Takuto, Yamamoto, Shûhei & Maruyama, Munetoshi" docDate="2023" docId="090C879DFFEB8E12899AFDB0AB3FFE0B" docLanguage="en" docName="zootaxa.5319.1.2.pdf" docOrigin="Zootaxa 5319 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5319.1.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudatheta cooteri Pace 1998" docType="treatment" docVersion="2" lastPageNumber="38" masterDocId="F535FFE5FFE08E1E890DFFDAA87BFFDA" masterDocTitle="Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan" masterLastPageNumber="47" masterPageNumber="27" pageNumber="38" updateTime="1690276987895" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CF1AACDFC9071669EFA28CC8CBF3FE37">Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan</mods:title>
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<mods:namePart id="767611E9EB439138F2C046E959989B2D">Hashizume, Takuto</mods:namePart>
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<mods:affiliation id="734D3A5AD809A9466E93E6615EB3D52B">Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka, 819 - 0395 Japan.</mods:affiliation>
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<mods:namePart id="B55E52970549C8A35A47A6E0AD053EF3">Yamamoto, Shûhei</mods:namePart>
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<mods:affiliation id="246589DE1D2C63C6308F9DE9E6590DD9">The Hokkaido University Museum, Hokkaido University, Kita 10, Nishi 8, Kita-ku, Sapporo, 060 - 0810 Japan. s. yamamoto. 64 @ gmail. com; https: // orcid. org / 0000 - 0002 - 4162 - 8457</mods:affiliation>
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<mods:nameIdentifier id="8092659E43E1F4E9298EEDFEFD89A11F" type="email">s.yamamoto.64@gmail.com</mods:nameIdentifier>
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<mods:namePart id="905A46ECD35D55836053673695C0A1BA">Maruyama, Munetoshi</mods:namePart>
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<mods:affiliation id="BB3BCC1059B2AEA0F019702F4ADE357C">The Kyushu University Museum, Fukuoka, 812 - 8581 Japan.</mods:affiliation>
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<mods:title id="736142D1FEB9BA472EEB08EFC32D5E8D">Zootaxa</mods:title>
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<mods:date id="5C4652AE8DBDBF6920564B91C14CFB3C">2023</mods:date>
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<mods:number id="EB013A37E8927E925C33A83141F86E28">2023-07-24</mods:number>
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<taxonomicName id="46A54D08FFEB8E15899AFDB0AA69FD5E" ID-CoL="4NLJ9" authority="Pace, 1998" authorityName="Pace" authorityYear="1998" box="[151,530,618,645]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="38" phylum="Arthropoda" rank="species" species="cooteri">
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<emphasis id="B3D1EA99FFEB8E15899AFDB0AA69FD5E" bold="true" box="[151,530,618,645]" pageId="11" pageNumber="38">
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<emphasis id="B3D1EA99FFEB8E15899AFDB0A9FDFD5F" bold="true" box="[151,390,618,645]" italics="true" pageId="11" pageNumber="38">Pseudatheta cooteri</emphasis>
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<bibRefCitation id="E5344B7AFFEB8E158880FDB1AA69FD5E" author="Pace, R." box="[397,530,618,645]" pageId="11" pageNumber="38" pagination="139 - 220" refId="ref16092" refString="Pace, R. (1998) Aleocharinae della Cina: Parte I (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 105 (1), 139 - 220. https: // doi. org / 10.5962 / bhl. part. 80035" type="journal article" year="1998">Pace, 1998</bibRefCitation>
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<paragraph id="811A368BFFEB8E15899AFD55A921FD73" blockId="11.[151,860,618,717]" box="[151,346,655,681]" pageId="11" pageNumber="38">
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(
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<figureCitation id="199E2A0EFFEB8E158992FD55A884FD73" box="[159,255,655,681]" captionStart="FIGURE 1" captionStartId="3.[151,250,1944,1968]" captionTargetBox="[178,1424,196,1901]" captionTargetId="figure-21@3.[164,1424,179,1919]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Pseudatheta spp. A. Pseudatheta crenulicauda (Bernhauer, 1907), comb. nov.; B. Pseudatheta taiwanensis Pace, 2008; C. Pseudatheta hilaris (Sharp, 1888), comb. nov., lectotype of Oxypoda hiralis; D. Pseudatheta thailandensis Pace, 1992; E. Pseudatheta cooteri Pace, 1998; F. Pseudatheta elegans Cameron, 1920. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8182112" httpUri="https://zenodo.org/record/8182112/files/figure.png" pageId="11" pageNumber="38">Figs. 1E</figureCitation>
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,
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<figureCitation id="199E2A0EFFEB8E158801FD55A92AFD73" box="[268,337,655,681]" captionStart="FIGURE 7" captionStartId="12.[151,250,1890,1914]" captionTargetBox="[161,1422,497,1858]" captionTargetId="figure-190@12.[151,1436,489,1866]" captionTargetPageId="12" captionText="FIGURE 7. Pseudatheta spp. A–D: Ps. cooteri Pace, 1998, E–G: Ps. elegans Cameron, 1920. A, E. male tergite VIII in dorsal view; B, F. median lobe of aedeagus in lateral view; D, G. spermatheca; C. ventral process of median lobe of aedeagus in lateral view, a: Yaku-shima Is., b: Iriomote-jima Is. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8182134" httpUri="https://zenodo.org/record/8182134/files/figure.png" pageId="11" pageNumber="38">7A–D</figureCitation>
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)
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<paragraph id="811A368BFFEB8E15899AFD69AB27FD17" blockId="11.[151,860,618,717]" box="[151,860,691,717]" pageId="11" pageNumber="38">[Japanese name: Minami-hime-kinokotsuyakeshi-hanekakushi]</paragraph>
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<treatmentCitation id="0004109AFFEB8E15899AFD2DAA72FCD5" author="Pace, R." box="[151,521,759,783]" page="208" pageId="11" pageNumber="38" year="1998">
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<taxonomicName id="46A54D08FFEB8E15899AFD2DAA72FCD5" ID-CoL="4NLJ9" authority="Pace, 1998: 208" authorityName="Pace" authorityPageNumber="208" authorityYear="1998" box="[151,521,759,783]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="38" phylum="Arthropoda" rank="species" species="cooteri">
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<emphasis id="B3D1EA99FFEB8E15899AFD2DA919FCD5" box="[151,354,759,783]" italics="true" pageId="11" pageNumber="38">Pseudatheta cooteri</emphasis>
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<bibRefCitation id="E5344B7AFFEB8E158865FD22AA72FCD5" author="Pace, R." box="[360,521,759,783]" pageId="11" pageNumber="38" pagination="139 - 220" refId="ref16092" refString="Pace, R. (1998) Aleocharinae della Cina: Parte I (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 105 (1), 139 - 220. https: // doi. org / 10.5962 / bhl. part. 80035" type="journal article" year="1998">Pace, 1998: 208</bibRefCitation>
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(original description;
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locality: “
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<collectingCountry id="F9B2761BFFEB8E158A70FD2DABC5FCD5" box="[893,958,759,783]" name="China" pageId="11" pageNumber="38">China</collectingCountry>
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,
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<collectingRegion id="4361F869FFEB8E158ACAFD2DAC2AFCD5" box="[967,1105,759,783]" country="China" name="Jiangsu" pageId="11" pageNumber="38">Jiangsu Prov.</collectingRegion>
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, Nanjing Zijinshan”):
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<treatmentCitation id="0004109AFFEB8E158C3CFD2DA9EAFCF4" author="Schulke, M. & Smetana, A." page="634" pageId="11" pageNumber="38" year="2015">
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<bibRefCitation id="E5344B7AFFEB8E158C3CFD2DA9EAFCF4" author="Schulke, M. & Smetana, A." pageId="11" pageNumber="38" refId="ref16621" refString="Schulke, M. & Smetana, A. (2015) Staphylinidae. In: L ˆ bl, I. & L ˆ bl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Staphylinoidea. Revised and updated edition. Brill, Leiden / Boston, pp. 304 - 1134." type="book" year="2015">Schülke & Smetana, 2015: 634</bibRefCitation>
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(catalogue);
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<treatmentCitation id="0004109AFFEB8E158B1AFCCCAAC2FCF4" author="Pace, R." box="[535,697,790,814]" page="301" pageId="11" pageNumber="38" year="2016">
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<bibRefCitation id="E5344B7AFFEB8E158B1AFCCCAAC2FCF4" author="Pace, R." box="[535,697,790,814]" pageId="11" pageNumber="38" pagination="295 - 336" refId="ref16439" refString="Pace, R. (2016) Aleocharinae della Cina al " Naturkundemuseum " di Erfurt (Insecta: Coleoptera: Staphyinidae). Veroffentlichungen des Naturkundemuseums Erfurt - VERNATE, 35, 295 - 336." type="journal article" year="2016">Pace, 2016: 301</bibRefCitation>
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(
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).
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<collectingRegion id="4361F869FFEB8E15899AFC86A884FCAC" box="[151,255,860,886]" pageId="11" pageNumber="38">Material</collectingRegion>
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examined.
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<collectingCountry id="F9B2761BFFEB8E158881FC86A99DFCAC" box="[396,486,860,886]" name="Japan" pageId="11" pageNumber="38">JAPAN</collectingCountry>
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: [
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<collectingRegion id="4361F869FFEB8E1588F2FC86AAC0FCAC" box="[511,699,860,886]" pageId="11" pageNumber="38">Kagoshima-ken</collectingRegion>
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]:
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<collectorName id="2C50535DFFEB8E158BD8FC86ABF9FCAD" box="[725,898,860,887]" pageId="11" pageNumber="38">Yaku-shima Is.</collectorName>
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:
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,
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<collectingDate id="E55FE9A3FFEB8E158C3DFC86A896FC40" pageId="11" pageNumber="38" value="2006-09-25" valueMax="2006-10-23" valueMin="2006-09-25">25 IX–23 X 2006</collectingDate>
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,
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.
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<collectorName id="2C50535DFFEB8E158819FC5AA9FFFC40" box="[276,388,896,922]" pageId="11" pageNumber="38">Yamauchi</collectorName>
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<emphasis id="B3D1EA99FFEB8E158886FC5BA9C7FC40" box="[395,444,896,922]" italics="true" pageId="11" pageNumber="38">et al</emphasis>
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. leg, (
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,
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||
<date id="F51B104BFFEB8E158D36FC5AACBCFC40" box="[1083,1223,896,923]" pageId="11" pageNumber="38" value="2021-07-21">
|
||
<collectingDate id="E55FE9A3FFEB8E158D36FC5AACBCFC40" box="[1083,1223,896,923]" pageId="11" pageNumber="38" value="2021-07-21">21 VII 2021</collectingDate>
|
||
</date>
|
||
,
|
||
<collectionCode id="E7B4AE4EFFEB8E158DDCFC5BAC98FC41" box="[1233,1251,897,923]" name="Tavera, Department of Geology and Geophysics" pageId="11" pageNumber="38">T</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<collectorName id="2C50535DFFEB8E158DE3FC5AAD17FC40" box="[1262,1388,896,922]" pageId="11" pageNumber="38">Hashizume</collectorName>
|
||
leg. (
|
||
<collectionCode id="E7B4AE4EFFEB8E1589ADFC7EA89DFC64" box="[160,230,932,958]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="11" pageNumber="38" type="Herbarium">KUM</collectionCode>
|
||
);
|
||
<materialsCitation id="31CD3CD6FFEB8E1589F6FC7FA9F4FC39" collectingDate="1993-11-03" collectionCode="KUM, V" collectorName="Okinawa-jima Is. & M. Kimura" country="Japan" county="Takazato" location="Yona" municipality="Ogimi-son" pageId="11" pageNumber="38" specimenCount="4" specimenCount-male="4" stateProvince="Okinawa-ken">
|
||
[
|
||
<emphasis id="B3D1EA99FFEB8E158809FC7EA9DDFC64" bold="true" box="[260,422,932,958]" pageId="11" pageNumber="38">
|
||
<collectingRegion id="4361F869FFEB8E158809FC7EA9DDFC64" box="[260,422,932,958]" pageId="11" pageNumber="38">Okinawa-ken</collectingRegion>
|
||
</emphasis>
|
||
]:
|
||
<collectorName id="2C50535DFFEB8E1588B2FC7EAAFEFC65" box="[447,645,932,959]" pageId="11" pageNumber="38">Okinawa-jima Is.</collectorName>
|
||
:
|
||
<specimenCount id="97A3FD02FFEB8E158B9FFC7EAA97FC64" box="[658,748,932,958]" pageId="11" pageNumber="38" type="male">2 males</specimenCount>
|
||
,
|
||
<collectingCounty id="687B4E07FFEB8E158BF5FC7EAB1BFC64" box="[760,864,932,958]" pageId="11" pageNumber="38">Takazato</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="617EACF1FFEB8E158A66FC7FAB9DFC1A" box="[875,998,933,960]" pageId="11" pageNumber="38">Ôgimi-son</collectingMunicipality>
|
||
,
|
||
<date id="F51B104BFFEB8E158AFFFC7EAC13FC64" box="[1010,1128,932,959]" pageId="11" pageNumber="38" value="1993-11-03">
|
||
<collectingDate id="E55FE9A3FFEB8E158AFFFC7EAC13FC64" box="[1010,1128,932,959]" pageId="11" pageNumber="38" value="1993-11-03">3 XI 1993</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="2C50535DFFEB8E158D79FC7FAC89FC64" box="[1140,1266,932,959]" pageId="11" pageNumber="38">M. Kimura</collectorName>
|
||
leg. (
|
||
<collectionCode id="E7B4AE4EFFEB8E158C39FC7EAD01FC64" box="[1332,1402,932,958]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="11" pageNumber="38" type="Herbarium">KUM</collectionCode>
|
||
);
|
||
<specimenCount id="97A3FD02FFEB8E158C83FC7EA8A0FC38" pageId="11" pageNumber="38" type="male">2 males</specimenCount>
|
||
,
|
||
<location id="847A6050FFEB8E1589EBFC13A958FC39" LSID="urn:lsid:plazi:treatment:090C879DFFEB8E12899AFDB0AB3FFE0B:847A6050FFEB8E1589EBFC13A958FC39" box="[230,291,969,995]" country="Japan" county="Takazato" municipality="Ogimi-son" name="Yona" pageId="11" pageNumber="38" stateProvince="Okinawa-ken">Yona</location>
|
||
,
|
||
<date id="F51B104BFFEB8E158823FC12A9F4FC39" box="[302,399,968,995]" pageId="11" pageNumber="38" value="1974-05-25" valueMax="1974-05-27" valueMin="1974-05-25">
|
||
25–27
|
||
<collectionCode id="E7B4AE4EFFEB8E158876FC13A9F4FC39" box="[379,399,969,995]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="11" pageNumber="38" type="Museum">V</collectionCode>
|
||
</date>
|
||
</materialsCitation>
|
||
<materialsCitation id="31CD3CD6FFEB8E15889BFC12A9F9FBDC" collectingDate="1984-03-27" collectionCode="KUM" collectorName="M. Sato & Iriomote-jima Is. & S. Nomura" country="Japan" location="Kampiree" pageId="11" pageNumber="38" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Okinawa-ken">
|
||
1974,
|
||
<collectorName id="2C50535DFFEB8E1588D1FC13AA4EFC38" box="[476,565,968,995]" pageId="11" pageNumber="38">M. Sato</collectorName>
|
||
leg. (
|
||
<collectionCode id="E7B4AE4EFFEB8E158B78FC12AAC7FC38" box="[629,700,968,994]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="11" pageNumber="38" type="Herbarium">KUM</collectionCode>
|
||
);
|
||
<collectorName id="2C50535DFFEB8E158BDDFC12ABEBFC39" box="[720,912,968,995]" pageId="11" pageNumber="38">Iriomote-jima Is.</collectorName>
|
||
:
|
||
<specimenCount id="97A3FD02FFEB8E158A90FC12AB8DFC38" box="[925,1014,968,994]" pageId="11" pageNumber="38" type="male">2 males</specimenCount>
|
||
,
|
||
<specimenCount id="97A3FD02FFEB8E158D0FFC12AC0AFC38" box="[1026,1137,968,994]" pageId="11" pageNumber="38" type="female">2 females</specimenCount>
|
||
,
|
||
<location id="847A6050FFEB8E158D71FC12AC96FC38" LSID="urn:lsid:plazi:treatment:090C879DFFEB8E12899AFDB0AB3FFE0B:847A6050FFEB8E158D71FC12AC96FC38" box="[1148,1261,968,994]" country="Japan" name="Kampiree" pageId="11" pageNumber="38" stateProvince="Okinawa-ken">Kampiree</location>
|
||
,
|
||
<date id="F51B104BFFEB8E158DF4FC12AD01FC38" box="[1273,1402,968,995]" pageId="11" pageNumber="38" value="1984-03-27">
|
||
<collectingDate id="E55FE9A3FFEB8E158DF4FC12AD01FC38" box="[1273,1402,968,995]" pageId="11" pageNumber="38" value="1984-03-27">27 III 1984</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="2C50535DFFEB8E158C8BFC12A888FBDD" pageId="11" pageNumber="38">S. Nomura</collectorName>
|
||
leg. (
|
||
<collectionCode id="E7B4AE4EFFEB8E15883EFC36A902FBDC" box="[307,377,1004,1030]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="11" pageNumber="38" type="Herbarium">KUM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="811A368BFFEB8E1589CAFBCAAC74FBF0" blockId="11.[151,1437,860,2039]" box="[199,1039,1040,1067]" pageId="11" pageNumber="38">
|
||
<materialsCitation id="31CD3CD6FFEB8E1589CAFBCAAAB5FBF1" box="[199,718,1040,1067]" collectionCode="V" country="Taiwan" location="Taipei" pageId="11" pageNumber="38" specimenCount="3" specimenCount-male="3" stateProvince="Taipei">
|
||
<emphasis id="B3D1EA99FFEB8E1589CAFBCAA943FBF0" bold="true" box="[199,312,1040,1066]" pageId="11" pageNumber="38">
|
||
<collectingCountry id="F9B2761BFFEB8E1589CAFBCAA943FBF0" box="[199,312,1040,1066]" name="Taiwan" pageId="11" pageNumber="38">TAIWAN</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="97A3FD02FFEB8E15884AFBCAA9DBFBF0" box="[327,416,1040,1066]" pageId="11" pageNumber="38" type="male">3 males</specimenCount>
|
||
, Wulai,
|
||
<collectingRegion id="4361F869FFEB8E1588F1FBCAAA39FBF0" box="[508,578,1040,1066]" country="Taiwan" name="Taipei" pageId="11" pageNumber="38">Taipei</collectingRegion>
|
||
Hsien,
|
||
<date id="F51B104BFFEB8E158B9AFBCAAAB5FBF1" box="[663,718,1040,1067]" pageId="11" pageNumber="38" value="1972-05-17">
|
||
17
|
||
<collectionCode id="E7B4AE4EFFEB8E158BB7FBCBAAB5FBF1" box="[698,718,1041,1067]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="11" pageNumber="38" type="Museum">V</collectionCode>
|
||
</date>
|
||
</materialsCitation>
|
||
<materialsCitation id="31CD3CD6FFEB8E158BD8FBCAAC70FBF0" box="[725,1035,1040,1067]" collectionCode="EUM" collectorName="Sakai" country="Taiwan" pageId="11" pageNumber="38" specimenCount="1">
|
||
1972, M,
|
||
<collectorName id="2C50535DFFEB8E158A4FFBCAABFBFBF0" box="[834,896,1040,1066]" pageId="11" pageNumber="38">Sakai</collectorName>
|
||
leg. (
|
||
<collectionCode id="E7B4AE4EFFEB8E158AB2FBCAAC78FBF0" box="[959,1027,1040,1066]" pageId="11" pageNumber="38">EUM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C9BF6500FFEB8E1589CAFBEEAA83F8BC" pageId="11" pageNumber="38" type="description">
|
||
<paragraph id="811A368BFFEB8E1589CAFBEEAB53FBA9" blockId="11.[151,1437,860,2039]" pageId="11" pageNumber="38">
|
||
<emphasis id="B3D1EA99FFEB8E1589CAFBEEA90EFB94" bold="true" box="[199,373,1076,1102]" pageId="11" pageNumber="38">Redescription.</emphasis>
|
||
Measurements (n = 5): BL ≈ 1.65–1.84; FBL, 0.91–1.00; HL, 0.28–0.34; HW, 0.35–0.39; PL, 0.31–0.35; PW, 0.46–0.50; EL, 0.30–0.35; EW, 0.55–0.60.
|
||
</paragraph>
|
||
<paragraph id="811A368BFFEB8E1589CAFBA6ACDEFB60" blockId="11.[151,1437,860,2039]" pageId="11" pageNumber="38">Relative length of antennomeres I–XI (n = 1): 25: 27: 19: 10: 10: 10: 11: 11: 13: 13: 35. Ratio of length/width of antennomeres I–XI (n = 1): 1.94: 2.21: 1.80: 0.87: 0.66: 0.64: 0.59: 0.58: 0.63: 0.61: 1.62.</paragraph>
|
||
<paragraph id="811A368BFFEB8E1589CAFB1EA9E1FAD8" blockId="11.[151,1437,860,2039]" pageId="11" pageNumber="38">
|
||
Body (
|
||
<figureCitation id="199E2A0EFFEB8E15881AFB1EA914FB04" box="[279,367,1220,1246]" captionStart="FIGURE 1" captionStartId="3.[151,250,1944,1968]" captionTargetBox="[178,1424,196,1901]" captionTargetId="figure-21@3.[164,1424,179,1919]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Pseudatheta spp. A. Pseudatheta crenulicauda (Bernhauer, 1907), comb. nov.; B. Pseudatheta taiwanensis Pace, 2008; C. Pseudatheta hilaris (Sharp, 1888), comb. nov., lectotype of Oxypoda hiralis; D. Pseudatheta thailandensis Pace, 1992; E. Pseudatheta cooteri Pace, 1998; F. Pseudatheta elegans Cameron, 1920. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8182112" httpUri="https://zenodo.org/record/8182112/files/figure.png" pageId="11" pageNumber="38">Fig. 1E</figureCitation>
|
||
) reddish brown; head darker; elytra dark brown except for the anterior 1/4–1/3; abdominal segments V–VI darker.
|
||
</paragraph>
|
||
<paragraph id="811A368BFFEB8E1589CAFAD6AB80FAB4" blockId="11.[151,1437,860,2039]" pageId="11" pageNumber="38">Head slightly transverse, HW/HL: 1.12–1.36; surface densely covered with setae. Antenna with antennomeres I–III and XI longer than wide, antennomeres IV–X wider than long; antennomeres I and II with rounded distal end, antennomeres III–X strongly angulated near distal end, antennomere XI oval.</paragraph>
|
||
<paragraph id="811A368BFFEB8E1589CAFAA3AA5FF99C" blockId="11.[151,1437,860,2039]" pageId="11" pageNumber="38">Pronotum transverse, PW/PL: 1.43–1.50, PW/HW: 1.29–1.33; surface densely covered with setae, finely punctured, without microsculpture; posterior margin arcuate. Elytra wider than long, EW/EL: 1.71–1.84, EL/PL: 0.92–1.00, EW/PW: 1.13–1.24; surface densely covered with setae and finely punctured; posterior margin sinuate near posterolateral corners. Mesoventrite without longitudinal carina; mesoventral process extended to slightly beyond middle of mesocoxal cavities, with pointed apex; metaventral process shorter than mesoventral process, with rounded apex; isthmus absent.</paragraph>
|
||
<paragraph id="811A368BFFEB8E1589CAF98AAC71F9B1" blockId="11.[151,1437,860,2039]" box="[199,1034,1616,1643]" pageId="11" pageNumber="38">Abdomen slightly narrowed posteriad; surface densely covered with setae.</paragraph>
|
||
<paragraph id="811A368BFFEB8E1589CAF9AEA972F8C4" blockId="11.[151,1437,860,2039]" pageId="11" pageNumber="38">
|
||
Male. Elytra with a pair of small tubercles at about posterior 1/3 on sutural margin. Tergite VII with a tubercle on anteromedian area. Tergite VIII (
|
||
<figureCitation id="199E2A0EFFEB8E158B38F942AAEBF968" box="[565,656,1688,1714]" captionStart="FIGURE 7" captionStartId="12.[151,250,1890,1914]" captionTargetBox="[161,1422,497,1858]" captionTargetId="figure-190@12.[151,1436,489,1866]" captionTargetPageId="12" captionText="FIGURE 7. Pseudatheta spp. A–D: Ps. cooteri Pace, 1998, E–G: Ps. elegans Cameron, 1920. A, E. male tergite VIII in dorsal view; B, F. median lobe of aedeagus in lateral view; D, G. spermatheca; C. ventral process of median lobe of aedeagus in lateral view, a: Yaku-shima Is., b: Iriomote-jima Is. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8182134" httpUri="https://zenodo.org/record/8182134/files/figure.png" pageId="11" pageNumber="38">Fig. 7A</figureCitation>
|
||
) with a tubercle on median area; six processes on posterior margin. Aedeagus as in
|
||
<figureCitation id="199E2A0EFFEB8E15885EF966A9D6F90C" box="[339,429,1724,1750]" captionStart="FIGURE 7" captionStartId="12.[151,250,1890,1914]" captionTargetBox="[161,1422,497,1858]" captionTargetId="figure-190@12.[151,1436,489,1866]" captionTargetPageId="12" captionText="FIGURE 7. Pseudatheta spp. A–D: Ps. cooteri Pace, 1998, E–G: Ps. elegans Cameron, 1920. A, E. male tergite VIII in dorsal view; B, F. median lobe of aedeagus in lateral view; D, G. spermatheca; C. ventral process of median lobe of aedeagus in lateral view, a: Yaku-shima Is., b: Iriomote-jima Is. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8182134" httpUri="https://zenodo.org/record/8182134/files/figure.png" pageId="11" pageNumber="38">Fig. 7C</figureCitation>
|
||
; median lobe with large basal bulb; apical process elongated, simply curved ventrally, slightly widened at basal part; ventral process large, apex pointed, with a distinct angle on dorsa side; flagellum very long.
|
||
</paragraph>
|
||
<paragraph id="811A368BFFEB8E1589CAF8F2AA83F8BC" blockId="11.[151,1437,860,2039]" pageId="11" pageNumber="38">
|
||
Female. Spermatheca (
|
||
<figureCitation id="199E2A0EFFEB8E1588CBF8F2AA66F898" box="[454,541,1832,1858]" captionStart="FIGURE 7" captionStartId="12.[151,250,1890,1914]" captionTargetBox="[161,1422,497,1858]" captionTargetId="figure-190@12.[151,1436,489,1866]" captionTargetPageId="12" captionText="FIGURE 7. Pseudatheta spp. A–D: Ps. cooteri Pace, 1998, E–G: Ps. elegans Cameron, 1920. A, E. male tergite VIII in dorsal view; B, F. median lobe of aedeagus in lateral view; D, G. spermatheca; C. ventral process of median lobe of aedeagus in lateral view, a: Yaku-shima Is., b: Iriomote-jima Is. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8182134" httpUri="https://zenodo.org/record/8182134/files/figure.png" pageId="11" pageNumber="38">Fig. 7D</figureCitation>
|
||
) curved twice; distal portion large, nearly conical shape, slightly curved; median portion elongate, curved; proximal portion very small.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C9BF6500FFEB8E1589CAF8AAA913F874" pageId="11" pageNumber="38" type="distribution">
|
||
<paragraph id="811A368BFFEB8E1589CAF8AAA913F874" blockId="11.[151,1437,860,2039]" pageId="11" pageNumber="38">
|
||
<emphasis id="B3D1EA99FFEB8E1589CAF8AAA91AF850" bold="true" box="[199,353,1904,1930]" pageId="11" pageNumber="38">Distribution.</emphasis>
|
||
<collectingCountry id="F9B2761BFFEB8E158862F8ABA9D4F851" box="[367,431,1905,1931]" name="Japan" pageId="11" pageNumber="38">Japan</collectingCountry>
|
||
(Yaku-shima Is., Okinawa-jima Is., Iriomote-jima Is.),
|
||
<collectingCountry id="F9B2761BFFEB8E158D5DF8AAACD8F850" box="[1104,1187,1904,1930]" name="Taiwan" pageId="11" pageNumber="38">Taiwan</collectingCountry>
|
||
—new record;
|
||
<collectingCountry id="F9B2761BFFEB8E158C54F8AAADE7F850" box="[1369,1436,1904,1930]" name="China" pageId="11" pageNumber="38">China</collectingCountry>
|
||
(
|
||
<collectingRegion id="4361F869FFEB8E158992F84EA88CF874" box="[159,247,1940,1966]" country="China" name="Jiangsu" pageId="11" pageNumber="38">Jiangsu</collectingRegion>
|
||
,
|
||
<collectingRegion id="4361F869FFEB8E15880CF84EA927F874" box="[257,348,1940,1966]" country="China" name="Yunnan" pageId="11" pageNumber="38">Yunnan</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C9BF6500FFEB8E1289CAF862AB3FFE0B" lastPageId="12" lastPageNumber="39" pageId="11" pageNumber="38" type="discussion">
|
||
<paragraph id="811A368BFFEB8E1289CAF862AB3FFE0B" blockId="11.[151,1437,860,2039]" lastBlockId="12.[151,1437,151,465]" lastPageId="12" lastPageNumber="39" pageId="11" pageNumber="38">
|
||
<emphasis id="B3D1EA99FFEB8E1589CAF862A946F808" bold="true" box="[199,317,1976,2002]" pageId="11" pageNumber="38">Remarks.</emphasis>
|
||
We report the first discovery of males of
|
||
<taxonomicName id="46A54D08FFEB8E158A01F863ABFCF808" authorityName="Pace" authorityYear="1998" box="[780,903,1977,2002]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="38" phylum="Arthropoda" rank="species" species="cooteri">
|
||
<emphasis id="B3D1EA99FFEB8E158A01F863ABFCF808" box="[780,903,1977,2002]" italics="true" pageId="11" pageNumber="38">Ps. cooteri</emphasis>
|
||
</taxonomicName>
|
||
and describe them here. Variations in the shape of the ventral process of the median lobe of the aedeagus were observed (
|
||
<figureCitation id="199E2A0EFFEB8E158ADCF806AC51F82C" box="[977,1066,2012,2038]" captionStart="FIGURE 7" captionStartId="12.[151,250,1890,1914]" captionTargetBox="[161,1422,497,1858]" captionTargetId="figure-190@12.[151,1436,489,1866]" captionTargetPageId="12" captionText="FIGURE 7. Pseudatheta spp. A–D: Ps. cooteri Pace, 1998, E–G: Ps. elegans Cameron, 1920. A, E. male tergite VIII in dorsal view; B, F. median lobe of aedeagus in lateral view; D, G. spermatheca; C. ventral process of median lobe of aedeagus in lateral view, a: Yaku-shima Is., b: Iriomote-jima Is. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8182134" httpUri="https://zenodo.org/record/8182134/files/figure.png" pageId="11" pageNumber="38">Fig. 7B</figureCitation>
|
||
). The aedeagus and spermatheca of
|
||
<taxonomicName id="46A54D08FFEC8E1289B5FF4DA947FF6A" authorityName="Pace" authorityYear="2008" box="[184,316,151,176]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="39" phylum="Arthropoda" rank="species" species="spinosa">
|
||
<emphasis id="B3D1EA99FFEC8E1289B5FF4DA947FF6A" box="[184,316,151,176]" italics="true" pageId="12" pageNumber="39">Ps. spinosa</emphasis>
|
||
</taxonomicName>
|
||
, a species from
|
||
<collectingCountry id="F9B2761BFFEC8E1288F2FF4DAA28FF6B" box="[511,595,151,177]" name="Taiwan" pageId="12" pageNumber="39">Taiwan</collectingCountry>
|
||
described by
|
||
<bibRefCitation id="E5344B7AFFEC8E128BF5FF4DABFDFF6B" author="Pace, R." box="[760,902,151,177]" pageId="12" pageNumber="39" pagination="137 - 156" refId="ref16257" refString="Pace, R. (2008) Pronomaeini, Homalotini, Bolitocharini, Diestotini e Falagriini di Taiwan (Coleoptera, Staphylinidae). Bollettino del Museo Civico di Storia Naturale di Verona. Botanica Zoologia, 32, 137 - 156." type="journal article" year="2008">Pace (2008)</bibRefCitation>
|
||
, are similar to those of
|
||
<taxonomicName id="46A54D08FFEC8E128D92FF4DAD66FF6A" authorityName="Pace" authorityYear="1998" box="[1183,1309,151,176]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="39" phylum="Arthropoda" rank="species" species="cooteri">
|
||
<emphasis id="B3D1EA99FFEC8E128D92FF4DAD66FF6A" box="[1183,1309,151,176]" italics="true" pageId="12" pageNumber="39">Ps. cooteri</emphasis>
|
||
</taxonomicName>
|
||
. However, judging from the illustrations in that report,
|
||
<taxonomicName id="46A54D08FFEC8E128B85FF61AB78FF0E" authorityName="Pace" authorityYear="1998" box="[648,771,187,212]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="39" phylum="Arthropoda" rank="species" species="cooteri">
|
||
<emphasis id="B3D1EA99FFEC8E128B85FF61AB78FF0E" box="[648,771,187,212]" italics="true" pageId="12" pageNumber="39">Ps. cooteri</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="46A54D08FFEC8E128D38FF61ACCDFF0E" authorityName="Pace" authorityYear="2008" box="[1077,1206,187,212]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="39" phylum="Arthropoda" rank="species" species="spinosa">
|
||
<emphasis id="B3D1EA99FFEC8E128D38FF61ACCDFF0E" box="[1077,1206,187,212]" italics="true" pageId="12" pageNumber="39">Ps. spinosa</emphasis>
|
||
</taxonomicName>
|
||
by the longer apical process of the median lobe of its aedeagus and by the longer, weakly curved distal portion of the spermatheca. We were able to examine specimens of
|
||
<taxonomicName id="46A54D08FFEC8E128B64FED9AA8DFEC7" authorityName="Cameron" authorityYear="1920" box="[617,758,259,285]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="39" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B3D1EA99FFEC8E128B64FED9AA8DFEC7" box="[617,758,259,285]" italics="true" pageId="12" pageNumber="39">Pseudatheta</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F9B2761BFFEC8E128A32FED9ABE8FEC7" box="[831,915,259,285]" name="Taiwan" pageId="12" pageNumber="39">Taiwan</collectingCountry>
|
||
and found that the aedeagi were identical to those of
|
||
<taxonomicName id="46A54D08FFEC8E1289F1FEFDA903FE9A" authorityName="Pace" authorityYear="1998" box="[252,376,295,320]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="39" phylum="Arthropoda" rank="species" species="cooteri">
|
||
<emphasis id="B3D1EA99FFEC8E1289F1FEFDA903FE9A" box="[252,376,295,320]" italics="true" pageId="12" pageNumber="39">Ps. cooteri</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F9B2761BFFEC8E1288CDFEFDAA7EFE9B" box="[448,517,295,321]" name="Japan" pageId="12" pageNumber="39">Japan</collectingCountry>
|
||
. We consider the former specimens to be
|
||
<taxonomicName id="46A54D08FFEC8E128AE5FEFDAC1EFE9A" authorityName="Pace" authorityYear="1998" box="[1000,1125,295,320]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="39" phylum="Arthropoda" rank="species" species="cooteri">
|
||
<emphasis id="B3D1EA99FFEC8E128AE5FEFDAC1EFE9A" box="[1000,1125,295,320]" italics="true" pageId="12" pageNumber="39">Ps. cooteri</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F9B2761BFFEC8E128DA1FEFDAD78FE9B" box="[1196,1283,295,321]" name="Taiwan" pageId="12" pageNumber="39">Taiwan</collectingCountry>
|
||
, although we were unable to compare the spermathecae because females were not collected with the male specimens. While it is possible that the two species are distributed in
|
||
<collectingCountry id="F9B2761BFFEC8E128BE4FEB5AB3BFE53" box="[745,832,367,393]" name="Taiwan" pageId="12" pageNumber="39">Taiwan</collectingCountry>
|
||
, the slight differences may have instead been due to intraspecific variation, deformation of the specimens, or misinterpretations in sketching. Therefore, we cannot rule out the possibility that these two species are synonyms.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |