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<document id="978E8B5502D7BB40BB2211EBB3736056" ID-DOI="10.11646/zootaxa.5319.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="8182108" ID-ZooBank="45CD8E29-3320-42E2-A8A8-3A36BC0F41CE" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1690276500814" checkinUser="plazi" docAuthor="Hashizume, Takuto, Yamamoto, Shûhei &amp; Maruyama, Munetoshi" docDate="2023" docId="090C879DFFEA8E15899AFB6CABF8FDC3" docLanguage="en" docName="zootaxa.5319.1.2.pdf" docOrigin="Zootaxa 5319 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5319.1.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudatheta thailandensis Pace 1992" docType="treatment" docVersion="2" lastPageNumber="38" masterDocId="F535FFE5FFE08E1E890DFFDAA87BFFDA" masterDocTitle="Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan" masterLastPageNumber="47" masterPageNumber="27" pageNumber="37" updateTime="1690276987895" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="45EC7AF7FDF216993BDFD36737FA46C1">Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan</mods:title>
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<mods:namePart id="5979A31A8CB4A3081B754C2F1C85D98E">Hashizume, Takuto</mods:namePart>
<mods:affiliation id="FD47958CFC413B6CD286B96EB32B4C89">Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka, 819 - 0395 Japan.</mods:affiliation>
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<mods:namePart id="91B9E34418776D1DFC4BFE77D77F4B99">Yamamoto, Shûhei</mods:namePart>
<mods:nameIdentifier id="A719E22347C38B119EF46C6F0A3448B4" type="ORCID">0000-0002-4162-8457</mods:nameIdentifier>
<mods:affiliation id="4A237FDD0D8E3188AE454A44202C4FEA">The Hokkaido University Museum, Hokkaido University, Kita 10, Nishi 8, Kita-ku, Sapporo, 060 - 0810 Japan. s. yamamoto. 64 @ gmail. com; https: // orcid. org / 0000 - 0002 - 4162 - 8457</mods:affiliation>
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<mods:namePart id="5FEAEF75C4BC15BC6BC82A8F6EDED6D0">Maruyama, Munetoshi</mods:namePart>
<mods:affiliation id="FAD6A7E48F26A557EF6B264FC83E72DF">The Kyushu University Museum, Fukuoka, 812 - 8581 Japan.</mods:affiliation>
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<treatment id="090C879DFFEA8E15899AFB6CABF8FDC3" ID-DOI="http://doi.org/10.5281/zenodo.8182138" ID-Zenodo-Dep="8182138" LSID="urn:lsid:plazi:treatment:090C879DFFEA8E15899AFB6CABF8FDC3" httpUri="http://treatment.plazi.org/id/090C879DFFEA8E15899AFB6CABF8FDC3" lastPageId="11" lastPageNumber="38" pageId="10" pageNumber="37">
<subSubSection id="C9BF6500FFEA8E14899AFB6CAA1BFB0A" box="[151,608,1206,1233]" pageId="10" pageNumber="37" type="nomenclature">
<paragraph id="811A368BFFEA8E14899AFB6CAA1BFB0A" blockId="10.[151,608,1206,1269]" box="[151,608,1206,1233]" pageId="10" pageNumber="37">
<heading id="DA5281E7FFEA8E14899AFB6CAA1BFB0A" bold="true" box="[151,608,1206,1233]" fontSize="11" level="1" pageId="10" pageNumber="37" reason="1">
<taxonomicName id="46A54D08FFEA8E14899AFB6CAA1BFB0A" ID-CoL="4NLJW" authority="Pace, 1992" authorityName="Pace" authorityYear="1992" box="[151,608,1206,1233]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="37" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis id="B3D1EA99FFEA8E14899AFB6CAA1BFB0A" bold="true" box="[151,608,1206,1233]" pageId="10" pageNumber="37">
<emphasis id="B3D1EA99FFEA8E14899AFB6CA9AFFB0B" bold="true" box="[151,468,1206,1233]" italics="true" pageId="10" pageNumber="37">Pseudatheta thailandensis</emphasis>
<bibRefCitation id="E5344B7AFFEA8E1488D1FB6DAA1BFB0A" author="Pace, R." box="[476,608,1206,1233]" pageId="10" pageNumber="37" pagination="227 - 268" refId="ref16061" refString="Pace, R. (1992) Aleocharinae della Thailandia (Coleoptera Staphylinidae). Bollettino del Museo Civico di Storia Naturale di Verona, 16, 227 - 268." type="journal article" year="1992">Pace, 1992</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C9BF6500FFEA8E14899AFB01A920FB2F" box="[151,347,1243,1269]" pageId="10" pageNumber="37" type="description">
<paragraph id="811A368BFFEA8E14899AFB01A920FB2F" blockId="10.[151,608,1206,1269]" box="[151,347,1243,1269]" pageId="10" pageNumber="37">
(
<figureCitation id="199E2A0EFFEA8E148992FB01A979FB2F" box="[159,258,1243,1269]" captionStart="FIGURE 1" captionStartId="3.[151,250,1944,1968]" captionTargetBox="[178,1424,196,1901]" captionTargetId="figure-21@3.[164,1424,179,1919]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Pseudatheta spp. A. Pseudatheta crenulicauda (Bernhauer, 1907), comb. nov.; B. Pseudatheta taiwanensis Pace, 2008; C. Pseudatheta hilaris (Sharp, 1888), comb. nov., lectotype of Oxypoda hiralis; D. Pseudatheta thailandensis Pace, 1992; E. Pseudatheta cooteri Pace, 1998; F. Pseudatheta elegans Cameron, 1920. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8182112" httpUri="https://zenodo.org/record/8182112/files/figure.png" pageId="10" pageNumber="37">Figs. 1D</figureCitation>
,
<figureCitation id="199E2A0EFFEA8E148802FB01A928FB2F" box="[271,339,1243,1269]" captionStart="FIGURE 6" captionStartId="9.[151,250,1687,1711]" captionTargetBox="[167,1423,323,1651]" captionTargetId="figure-90@9.[151,1436,309,1663]" captionTargetPageId="9" captionText="FIGURE 6. Pseudatheta spp. A: Ps. hilaris (Sharp, 1888), comb. nov., BD: Ps. thailandensis Pace, 1992. A, D. spermatheca; B. male tergite VIII in dorsal view; C. median lobe of aedeagus in lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8182130" httpUri="https://zenodo.org/record/8182130/files/figure.png" pageId="10" pageNumber="37">6BD</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C9BF6500FFEA8E14899AFAC0AB20FA8A" pageId="10" pageNumber="37" type="reference_group">
<paragraph id="811A368BFFEA8E14899AFAC0AB20FA8A" blockId="10.[151,1437,1306,1361]" pageId="10" pageNumber="37">
<treatmentCitationGroup id="A1B511A5FFEA8E14899AFAC0AB20FA8A" pageId="10" pageNumber="37">
<treatmentCitation id="0004109AFFEA8E14899AFAC0AA34FAE8" author="Pace, R." box="[151,591,1306,1330]" page="239" pageId="10" pageNumber="37" year="1992">
<taxonomicName id="46A54D08FFEA8E14899AFAC0AA34FAE8" ID-CoL="4NLJW" authority="Pace, 1992: 239" authorityName="Pace" authorityPageNumber="239" authorityYear="1992" box="[151,591,1306,1330]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="37" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis id="B3D1EA99FFEA8E14899AFAC0A9D8FAE8" box="[151,419,1306,1330]" italics="true" pageId="10" pageNumber="37">Pseudatheta thailandensis</emphasis>
<bibRefCitation id="E5344B7AFFEA8E1488A7FAC0AA34FAE8" author="Pace, R." box="[426,591,1306,1330]" pageId="10" pageNumber="37" pagination="227 - 268" refId="ref16061" refString="Pace, R. (1992) Aleocharinae della Thailandia (Coleoptera Staphylinidae). Bollettino del Museo Civico di Storia Naturale di Verona, 16, 227 - 268." type="journal article" year="1992">Pace, 1992: 239</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description;
<typeStatus id="5E1E8829FFEA8E148A39FAC0AB1BFAE8" box="[820,864,1306,1330]" pageId="10" pageNumber="37">type</typeStatus>
locality: “Chiang Mai Prov., Doi Suthep”);
<treatmentCitation id="0004109AFFEA8E148C28FAC0A89BFA8A" author="Pace, R." page="66" pageId="10" pageNumber="37" year="2004">
<bibRefCitation id="E5344B7AFFEA8E148C28FAC0A89BFA8A" author="Pace, R." pageId="10" pageNumber="37" pagination="63 - 76" refId="ref16139" refString="Pace, R. (2004) Specie nuove o poco note di Homalotini, Silusini, Bolitocharini, Diestotini e Autaliini della Cina e della Thailandia (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 111 (1), 63 - 76. https: // doi. org / 10.5962 / bhl. part. 80227" type="journal article" year="2004">Pace, 2004: 66</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="F9B2761BFFEA8E1489E3FAE2A955FA8A" box="[238,302,1336,1360]" name="China" pageId="10" pageNumber="37">China</collectingCountry>
,
<collectingRegion id="4361F869FFEA8E148835FAE2A9F5FA8A" box="[312,398,1336,1360]" country="China" name="Shaanxi" pageId="10" pageNumber="37">Shaanxi</collectingRegion>
);
<treatmentCitation id="0004109AFFEA8E1488ADFAE2AAA7FA8A" author="Schulke, M. &amp; Smetana, A." box="[416,732,1336,1361]" page="634" pageId="10" pageNumber="37" year="2015">
<bibRefCitation id="E5344B7AFFEA8E1488ADFAE2AAA7FA8A" author="Schulke, M. &amp; Smetana, A." box="[416,732,1336,1361]" pageId="10" pageNumber="37" refId="ref16621" refString="Schulke, M. &amp; Smetana, A. (2015) Staphylinidae. In: L ˆ bl, I. &amp; L ˆ bl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Staphylinoidea. Revised and updated edition. Brill, Leiden / Boston, pp. 304 - 1134." type="book" year="2015">Schülke &amp; Smetana, 2015: 634</bibRefCitation>
</treatmentCitation>
(catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C9BF6500FFEA8E14899AFAA3AB98F8C5" pageId="10" pageNumber="37" type="materials_examined">
<paragraph id="811A368BFFEA8E14899AFAA3A95BFA6D" blockId="10.[151,1437,1401,2040]" pageId="10" pageNumber="37">
<emphasis id="B3D1EA99FFEA8E14899AFAA3A9F9FA49" bold="true" box="[151,386,1401,1427]" pageId="10" pageNumber="37">Material examined.</emphasis>
<materialsCitation id="31CD3CD6FFEA8E148886FAA3A967FA6D" collectingDate="1983-10-20" collectionCode="EUM" collectorName="M. Sakai" country="Thailand" elevation="1560" location="Doi Inthanon" municipality="Chaing Mai" pageId="10" pageNumber="37" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<emphasis id="B3D1EA99FFEA8E148886FAA3AA59FA49" bold="true" box="[395,546,1401,1427]" pageId="10" pageNumber="37">
<collectingCountry id="F9B2761BFFEA8E148886FAA3AA59FA49" box="[395,546,1401,1427]" name="Thailand" pageId="10" pageNumber="37">THAILAND</collectingCountry>
</emphasis>
:
<specimenCount id="97A3FD02FFEA8E148B3FFAA3AAF7FA49" box="[562,652,1401,1427]" pageId="10" pageNumber="37" type="male">2 males</specimenCount>
,
<specimenCount id="97A3FD02FFEA8E148B94FAA3AA84FA49" box="[665,767,1401,1427]" pageId="10" pageNumber="37" type="female">1 female</specimenCount>
,
<location id="847A6050FFEA8E148A06FAA3ABDFFA49" LSID="urn:lsid:plazi:treatment:090C879DFFEA8E15899AFB6CABF8FDC3:847A6050FFEA8E148A06FAA3ABDFFA49" box="[779,932,1401,1427]" country="Thailand" municipality="Chaing Mai" name="Doi Inthanon" pageId="10" pageNumber="37">Doi Inthanon</location>
,
<collectingMunicipality id="617EACF1FFEA8E148ABCFAA3AC41FA49" box="[945,1082,1401,1427]" pageId="10" pageNumber="37">Chaing Mai</collectingMunicipality>
,
<quantity id="465D9B6EFFEA8E148D4AFAA3ACE4FA4E" box="[1095,1183,1401,1428]" metricMagnitude="3" metricUnit="m" metricValue="1.56" pageId="10" pageNumber="37" unit="m" value="1560.0">
<elevation id="0A88D1B8FFEA8E148D4AFAA3ACE4FA4E" box="[1095,1183,1401,1428]" metricMagnitude="3" metricUnit="m" metricValue="1.56" pageId="10" pageNumber="37" unit="m" value="1560.0">1560 m</elevation>
</quantity>
,
<date id="F51B104BFFEA8E148DA0FAA3AD52FA49" box="[1197,1321,1401,1428]" pageId="10" pageNumber="37" value="1983-10-20">
<collectingDate id="E55FE9A3FFEA8E148DA0FAA3AD52FA49" box="[1197,1321,1401,1428]" pageId="10" pageNumber="37" value="1983-10-20">20 X 1983</collectingDate>
</date>
,
<collectorName id="2C50535DFFEA8E148C3BFAA0ADE7FA49" box="[1334,1436,1401,1428]" pageId="10" pageNumber="37">M. Sakai</collectorName>
leg. (
<collectionCode id="E7B4AE4EFFEA8E1489C2FA47A968FA6D" box="[207,275,1437,1463]" pageId="10" pageNumber="37">EUM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="811A368BFFEA8E1489CAFA1BA940F9F9" blockId="10.[151,1437,1401,2040]" pageId="10" pageNumber="37">
<emphasis id="B3D1EA99FFEA8E1489CAFA1BA9A6FA01" bold="true" box="[199,477,1473,1499]" pageId="10" pageNumber="37">Additional description.</emphasis>
Measurements (n = 1): BL ≈ 2.07; FBL: 1.22; HL: 0.36; HW: 0.43;
<collectionCode id="E7B4AE4EFFEA8E148DF5FA18AD66FA06" box="[1272,1309,1474,1500]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="10" pageNumber="37" type="Museum">PL</collectionCode>
: 0.52;
<collectionCode id="E7B4AE4EFFEA8E148C66FA18ADE3FA06" box="[1387,1432,1474,1500]" country="Thailand" name="Paleontological Collections" pageId="10" pageNumber="37">PW</collectionCode>
: 0.60; EL: 0.43; EW: 0.72. Ratios (n = 1): HW/HL: 1.18,
<collectionCode id="E7B4AE4EFFEA8E148A1AFA3FAB38FA25" box="[791,835,1509,1535]" country="Thailand" name="Paleontological Collections" pageId="10" pageNumber="37">PW</collectionCode>
/
<collectionCode id="E7B4AE4EFFEA8E148A41FA3FAB15FA25" box="[844,878,1509,1535]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="10" pageNumber="37" type="Museum">PL</collectionCode>
: 1.44, EW/EL: 1.67,
<collectionCode id="E7B4AE4EFFEA8E148D51FA3FACF3FA25" box="[1116,1160,1509,1535]" country="Thailand" name="Paleontological Collections" pageId="10" pageNumber="37">PW</collectionCode>
/HW: 1.39, EL/
<collectionCode id="E7B4AE4EFFEA8E148C3AFA3FAD21FA25" box="[1335,1370,1509,1535]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="10" pageNumber="37" type="Museum">PL</collectionCode>
: 1.03, EW/
<collectionCode id="E7B4AE4EFFEA8E1489C1F9D3A883F9F9" box="[204,248,1545,1571]" country="Thailand" name="Paleontological Collections" pageId="10" pageNumber="37">PW</collectionCode>
: 1.20.
</paragraph>
<paragraph id="811A368BFFEA8E1489CAF9F7ACDEF9B1" blockId="10.[151,1437,1401,2040]" pageId="10" pageNumber="37">Relative length of antennomeres IXI (n = 1): 20: 24: 15: 10: 12: 10: 12: 12: 12: 11: 34. Ratio of length/width of antennomeres IXI (n = 1): 1.79: 2.36: 1.50: 0.68: 0.76: 0.65: 0.66: 0.63: 0.58: 0.55: 1.61.</paragraph>
<paragraph id="811A368BFFEA8E1489CAF9AFAB98F8C5" blockId="10.[151,1437,1401,2040]" pageId="10" pageNumber="37">
Male. Elytra with a pair of small tubercles at about posterior 1/3 on sutural margin. Sternite
<collectionCode id="E7B4AE4EFFEA8E148DF1F9ACAD61F94A" box="[1276,1306,1654,1680]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="10" pageNumber="37">VI</collectionCode>
with small medial lobe on posterior margin. Tergite VII with a tubercle on posteromedian area. Tergite VIII (
<figureCitation id="199E2A0EFFEA8E148DE3F943AD3EF969" box="[1262,1349,1689,1715]" captionStart="FIGURE 6" captionStartId="9.[151,250,1687,1711]" captionTargetBox="[167,1423,323,1651]" captionTargetId="figure-90@9.[151,1436,309,1663]" captionTargetPageId="9" captionText="FIGURE 6. Pseudatheta spp. A: Ps. hilaris (Sharp, 1888), comb. nov., BD: Ps. thailandensis Pace, 1992. A, D. spermatheca; B. male tergite VIII in dorsal view; C. median lobe of aedeagus in lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8182130" httpUri="https://zenodo.org/record/8182130/files/figure.png" pageId="10" pageNumber="37">Fig. 6B</figureCitation>
) with a tubercle on median area; four processes on each side of posterior margin, outer ones slightly curved, longer and sharper than the others. Aedeagus as in
<figureCitation id="199E2A0EFFEA8E148B40F93BAAD8F921" box="[589,675,1761,1787]" captionStart="FIGURE 6" captionStartId="9.[151,250,1687,1711]" captionTargetBox="[167,1423,323,1651]" captionTargetId="figure-90@9.[151,1436,309,1663]" captionTargetPageId="9" captionText="FIGURE 6. Pseudatheta spp. A: Ps. hilaris (Sharp, 1888), comb. nov., BD: Ps. thailandensis Pace, 1992. A, D. spermatheca; B. male tergite VIII in dorsal view; C. median lobe of aedeagus in lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8182130" httpUri="https://zenodo.org/record/8182130/files/figure.png" pageId="10" pageNumber="37">Fig. 6C</figureCitation>
; median lobe with large basal bulb; apical process relatively slender, narrowed apically in lateral view, apex pointed; flagellum moderately long.
</paragraph>
</subSubSection>
<subSubSection id="C9BF6500FFEA8E1489CAF8F3AAECF899" box="[199,663,1833,1859]" pageId="10" pageNumber="37" type="distribution">
<paragraph id="811A368BFFEA8E1489CAF8F3AAECF899" blockId="10.[151,1437,1401,2040]" box="[199,663,1833,1859]" pageId="10" pageNumber="37">
<emphasis id="B3D1EA99FFEA8E1489CAF8F3A91AF899" bold="true" box="[199,353,1833,1859]" pageId="10" pageNumber="37">Distribution.</emphasis>
<collectingCountry id="F9B2761BFFEA8E14886AF8F3A9B6F899" box="[359,461,1833,1859]" name="Thailand" pageId="10" pageNumber="37">Thailand</collectingCountry>
,
<collectingCountry id="F9B2761BFFEA8E1488D5F8F3AA60F899" box="[472,539,1833,1859]" name="China" pageId="10" pageNumber="37">China</collectingCountry>
(
<collectingRegion id="4361F869FFEA8E148B27F8F3AAF1F899" box="[554,650,1833,1859]" country="China" name="Shaanxi" pageId="10" pageNumber="37">Shaanxi</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C9BF6500FFEA8E1589CAF897ABF8FDC3" lastPageId="11" lastPageNumber="38" pageId="10" pageNumber="37" type="discussion">
<paragraph id="811A368BFFEA8E1589CAF897AD0CFEC7" blockId="10.[151,1437,1401,2040]" lastBlockId="11.[151,1436,151,537]" lastPageId="11" lastPageNumber="38" pageId="10" pageNumber="37">
<emphasis id="B3D1EA99FFEA8E1489CAF897A940F8BD" bold="true" box="[199,315,1869,1895]" pageId="10" pageNumber="37">Remarks.</emphasis>
This species was described only from a female and could not be distinguished from its allied species based on the spermatheca.According to the original description by
<bibRefCitation id="E5344B7AFFEA8E148A7BF8A8AB87F851" author="Pace, R." box="[886,1020,1905,1932]" pageId="10" pageNumber="37" pagination="227 - 268" refId="ref16061" refString="Pace, R. (1992) Aleocharinae della Thailandia (Coleoptera Staphylinidae). Bollettino del Museo Civico di Storia Naturale di Verona, 16, 227 - 268." type="journal article" year="1992">Pace (1992)</bibRefCitation>
, the median part of the spermatheca is short, but this description may have derived from a deformed specimen. In many aleocharine species, spermathecae are easily deformed immediately after mounting in mounting medium. Later,
<bibRefCitation id="E5344B7AFFEA8E148D14F860ACD8F809" author="Pace, R." box="[1049,1187,1977,2004]" pageId="10" pageNumber="37" pagination="63 - 76" refId="ref16139" refString="Pace, R. (2004) Specie nuove o poco note di Homalotini, Silusini, Bolitocharini, Diestotini e Autaliini della Cina e della Thailandia (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 111 (1), 63 - 76. https: // doi. org / 10.5962 / bhl. part. 80227" type="journal article" year="2004">Pace (2004)</bibRefCitation>
recorded this species from
<collectingCountry id="F9B2761BFFEA8E1489D8F807A960F82D" box="[213,283,2013,2039]" name="China" pageId="10" pageNumber="37">China</collectingCountry>
, but it was again a female specimen. In this study, we were able to examine specimens of both sexes of the genus
<taxonomicName id="46A54D08FFEB8E158804FF4DA9E8FF6B" authorityName="Cameron" authorityYear="1920" box="[265,403,151,177]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="B3D1EA99FFEB8E158804FF4DA9E8FF6B" box="[265,403,151,177]" italics="true" pageId="11" pageNumber="38">Pseudatheta</emphasis>
</taxonomicName>
, which were similar to
<taxonomicName id="46A54D08FFEB8E158B91FF4DAB20FF6B" authorityName="Pace" authorityYear="1992" box="[668,859,151,177]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="38" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis id="B3D1EA99FFEB8E158B91FF4DAB20FF6B" box="[668,859,151,177]" italics="true" pageId="11" pageNumber="38">Ps. thailandensis</emphasis>
</taxonomicName>
collected in an area close to the
<typeStatus id="5E1E8829FFEB8E158DC4FF4DAC82FF6B" box="[1225,1273,151,177]" pageId="11" pageNumber="38">type</typeStatus>
locality of
<taxonomicName id="46A54D08FFEB8E158C74FF4DA957FF0F" authorityName="Pace" authorityYear="1992" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="38" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis id="B3D1EA99FFEB8E158C74FF4DA957FF0F" italics="true" pageId="11" pageNumber="38">Ps. thailandensis</emphasis>
</taxonomicName>
. Since there are areas where more than one species of this genus is distributed, such as
<collectingCountry id="F9B2761BFFEB8E158DF8FF61AD43FF0F" box="[1269,1336,187,213]" name="Japan" pageId="11" pageNumber="38">Japan</collectingCountry>
, whether these specimens are
<taxonomicName id="46A54D08FFEB8E158875FF05AA4DFF23" authorityName="Pace" authorityYear="1992" box="[376,566,223,249]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="38" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis id="B3D1EA99FFEB8E158875FF05AA4DFF23" box="[376,566,223,249]" italics="true" pageId="11" pageNumber="38">Ps. thailandensis</emphasis>
</taxonomicName>
is unclear. Nonetheless, since this species is the only known
<taxonomicName id="46A54D08FFEB8E158DD9FF05AD12FF23" authorityName="Pace" authorityYear="1992" box="[1236,1385,223,249]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="38" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis id="B3D1EA99FFEB8E158DD9FF05AD12FF23" box="[1236,1385,223,249]" italics="true" pageId="11" pageNumber="38">thailandensis</emphasis>
</taxonomicName>
-like species from
<collectingCountry id="F9B2761BFFEB8E158821FED9A9E9FEC7" box="[300,402,259,285]" name="Thailand" pageId="11" pageNumber="38">Thailand</collectingCountry>
, we tentatively identified the specimens as
<taxonomicName id="46A54D08FFEB8E158A75FED9AC4CFEC7" authorityName="Pace" authorityYear="1992" box="[888,1079,259,285]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="38" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis id="B3D1EA99FFEB8E158A75FED9AC4CFEC7" box="[888,1079,259,285]" italics="true" pageId="11" pageNumber="38">Ps. thailandensis</emphasis>
</taxonomicName>
and described the aedeagus.
</paragraph>
<paragraph id="811A368BFFEB8E1589CAFEFDABF8FDC3" blockId="11.[151,1436,151,537]" pageId="11" pageNumber="38">
This species can be distinguished from its allies (
<taxonomicName id="46A54D08FFEB8E158BF1FEFDABC0FE9B" authorityName="Hashizume &amp; Yamamoto &amp; Maruyama" authorityYear="2023" baseAuthorityName="Bernhauer" baseAuthorityYear="1907" box="[764,955,295,321]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="38" phylum="Arthropoda" rank="species" species="crenulicauda">
<emphasis id="B3D1EA99FFEB8E158BF1FEFDABC0FE9B" box="[764,955,295,321]" italics="true" pageId="11" pageNumber="38">Ps. crenulicauda</emphasis>
</taxonomicName>
,
<taxonomicName id="46A54D08FFEB8E158AC7FEFDAC01FE9A" authorityName="Pace" authorityYear="2008" box="[970,1146,295,320]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="38" phylum="Arthropoda" rank="species" species="taiwanensis">
<emphasis id="B3D1EA99FFEB8E158AC7FEFDAC01FE9A" box="[970,1146,295,320]" italics="true" pageId="11" pageNumber="38">Ps. taiwanensis</emphasis>
</taxonomicName>
) by the apical process of the aedeagus, which tapers gradually toward the apex. However,
<taxonomicName id="46A54D08FFEB8E158A7CFE91AD74FEBF" authority="(Bernhauer, 1915)" baseAuthorityName="Bernhauer" baseAuthorityYear="1915" box="[881,1295,330,357]" class="Insecta" family="Staphylinidae" genus="Phymatura" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="38" phylum="Arthropoda" rank="species" species="bigranipennis">
<emphasis id="B3D1EA99FFEB8E158A7CFE91AC41FEBF" box="[881,1082,331,357]" italics="true" pageId="11" pageNumber="38">Ph. bigranipennis</emphasis>
(Bernhauer, 1915)
</taxonomicName>
, redescribed by
<bibRefCitation id="E5344B7AFFEB8E1589CDFEB5A91DFE53" author="Benick, G." box="[192,358,367,393]" pageId="11" pageNumber="38" pagination="335 - 340" refId="ref15097" refString="Benick, G. (1980) Weitere neue Atheten aus Sowjet-Russland (Col. Staphyl.). Nouvelle Revue d'Entomologie, 10 (4), 335 - 340." type="journal article" year="1980">Benick (1980)</bibRefCitation>
and
<bibRefCitation id="E5344B7AFFEB8E1588AAFEB5AAEBFE53" author="Kim, Y. - H. &amp; Ahn, K. - J." box="[423,656,367,393]" pageId="11" pageNumber="38" pagination="189 - 196" refId="ref15557" refString="Kim, Y. - H. &amp; Ahn, K. - J. (2016) Taxonomy of the genus Phymatura J. Sahlberg (Coleoptera: Staphylinidae: Aleocharinae) in the Korean Peninsula. Zootaxa, 4061 (2), 189 - 196. https: // doi. org / 10.11646 / zootaxa. 4061.2.9" type="journal article" year="2016">Kim &amp; Ahn (2016)</bibRefCitation>
, cannot be distinguished from
<taxonomicName id="46A54D08FFEB8E158D09FEB5ACB3FE53" authorityName="Pace" authorityYear="1992" box="[1028,1224,367,393]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="38" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis id="B3D1EA99FFEB8E158D09FEB5ACB3FE53" box="[1028,1224,367,393]" italics="true" pageId="11" pageNumber="38">Ps. thailandensis</emphasis>
</taxonomicName>
judging from the illustrations and description, and specimens described by the authors are considered identical. Since
<bibRefCitation id="E5344B7AFFEB8E158DF6FE49ADE7FE77" author="Benick, G." box="[1275,1436,403,429]" pageId="11" pageNumber="38" pagination="335 - 340" refId="ref15097" refString="Benick, G. (1980) Weitere neue Atheten aus Sowjet-Russland (Col. Staphyl.). Nouvelle Revue d'Entomologie, 10 (4), 335 - 340." type="journal article" year="1980">Benick (1980)</bibRefCitation>
and
<bibRefCitation id="E5344B7AFFEB8E1589C5FE6DA9D8FE0B" author="Kim, Y. - H. &amp; Ahn, K. - J." box="[200,419,439,465]" pageId="11" pageNumber="38" pagination="189 - 196" refId="ref15557" refString="Kim, Y. - H. &amp; Ahn, K. - J. (2016) Taxonomy of the genus Phymatura J. Sahlberg (Coleoptera: Staphylinidae: Aleocharinae) in the Korean Peninsula. Zootaxa, 4061 (2), 189 - 196. https: // doi. org / 10.11646 / zootaxa. 4061.2.9" type="journal article" year="2016">Kim &amp; Ahn (2016)</bibRefCitation>
did not state that they examined the
<typeStatus id="5E1E8829FFEB8E158A40FE6DAB06FE0B" box="[845,893,439,465]" pageId="11" pageNumber="38">type</typeStatus>
specimen, we cannot rule out the possibility of a misidentification. Thus, we refrain from taxonomic treatment of
<taxonomicName id="46A54D08FFEB8E158A8AFE01AC2AFE2F" baseAuthorityName="Bernhauer" baseAuthorityYear="1915" box="[903,1105,475,501]" class="Insecta" family="Staphylinidae" genus="Phymatura" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="38" phylum="Arthropoda" rank="species" species="bigranipennis">
<emphasis id="B3D1EA99FFEB8E158A8AFE01AC2AFE2F" box="[903,1105,475,501]" italics="true" pageId="11" pageNumber="38">Ph. bigranipennis</emphasis>
</taxonomicName>
at this time. The relationship between
<taxonomicName id="46A54D08FFEB8E1589F6FE25A9C1FDC3" authorityName="Pace" authorityYear="1992" box="[251,442,511,537]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="38" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis id="B3D1EA99FFEB8E1589F6FE25A9C1FDC3" box="[251,442,511,537]" italics="true" pageId="11" pageNumber="38">Ps. thailandensis</emphasis>
</taxonomicName>
and
<taxonomicName id="46A54D08FFEB8E1588FCFE25AA1FFDC3" box="[497,612,511,537]" class="Insecta" family="Staphylinidae" genus="Oxypoda" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="38" phylum="Arthropoda" rank="species" species="hilaris">
<emphasis id="B3D1EA99FFEB8E1588FCFE25AA1FFDC3" box="[497,612,511,537]" italics="true" pageId="11" pageNumber="38">Ps. hilaris</emphasis>
</taxonomicName>
could not be determined.
</paragraph>
</subSubSection>
</treatment>
</document>