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<document id="DB04C64AB3B7FF3252BD6B7B05E363C7" ID-DOI="10.11646/zootaxa.5319.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="8182108" ID-ZooBank="45CD8E29-3320-42E2-A8A8-3A36BC0F41CE" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1690276500814" checkinUser="plazi" docAuthor="Hashizume, Takuto, Yamamoto, Shûhei &amp; Maruyama, Munetoshi" docDate="2023" docId="090C879DFFE78E17899AFCDBACE9FEC7" docLanguage="en" docName="zootaxa.5319.1.2.pdf" docOrigin="Zootaxa 5319 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5319.1.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudatheta taiwanensis Pace 2008" docType="treatment" docVersion="2" lastPageNumber="35" masterDocId="F535FFE5FFE08E1E890DFFDAA87BFFDA" masterDocTitle="Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan" masterLastPageNumber="47" masterPageNumber="27" pageNumber="34" updateTime="1690276987895" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="AD7A3891BA19BED09B6DA07F46BE8E7F">Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan</mods:title>
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<mods:namePart id="535DC68518D3D07EB7A3A8B75829DC56">Hashizume, Takuto</mods:namePart>
<mods:affiliation id="3216FE8D0E7FDE3E7B1451E8163802F3">Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka, 819 - 0395 Japan.</mods:affiliation>
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<mods:namePart id="01C72DFCDF194DB1699CA3B881BB3B80">Yamamoto, Shûhei</mods:namePart>
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<mods:affiliation id="15BBBED77543C631F13361A07E862727">The Hokkaido University Museum, Hokkaido University, Kita 10, Nishi 8, Kita-ku, Sapporo, 060 - 0810 Japan. s. yamamoto. 64 @ gmail. com; https: // orcid. org / 0000 - 0002 - 4162 - 8457</mods:affiliation>
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<mods:affiliation id="382E7F82101481C6F211E29948CF50B8">The Kyushu University Museum, Fukuoka, 812 - 8581 Japan.</mods:affiliation>
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<taxonomicName id="46A54D08FFE78E19899AFCDBAA37FCC1" ID-CoL="4NLJV" authority="Pace, 2008" authorityName="Pace" authorityYear="2008" box="[151,588,769,796]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="34" phylum="Arthropoda" rank="species" species="taiwanensis">
<emphasis id="B3D1EA99FFE78E19899AFCDBAA37FCC1" bold="true" box="[151,588,769,796]" pageId="7" pageNumber="34">
<emphasis id="B3D1EA99FFE78E19899AFCDBA9BAFCC6" bold="true" box="[151,449,769,796]" italics="true" pageId="7" pageNumber="34">Pseudatheta taiwanensis</emphasis>
<bibRefCitation id="E5344B7AFFE78E1988C5FCD8AA37FCC1" author="Pace, R." box="[456,588,769,796]" pageId="7" pageNumber="34" pagination="137 - 156" refId="ref16257" refString="Pace, R. (2008) Pronomaeini, Homalotini, Bolitocharini, Diestotini e Falagriini di Taiwan (Coleoptera, Staphylinidae). Bollettino del Museo Civico di Storia Naturale di Verona. Botanica Zoologia, 32, 137 - 156." type="journal article" year="2008">Pace, 2008</bibRefCitation>
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(
<figureCitation id="199E2A0EFFE78E198992FCFCA97AFC9A" box="[159,257,806,832]" captionStart="FIGURE 1" captionStartId="3.[151,250,1944,1968]" captionTargetBox="[178,1424,196,1901]" captionTargetId="figure-21@3.[164,1424,179,1919]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Pseudatheta spp. A. Pseudatheta crenulicauda (Bernhauer, 1907), comb. nov.; B. Pseudatheta taiwanensis Pace, 2008; C. Pseudatheta hilaris (Sharp, 1888), comb. nov., lectotype of Oxypoda hiralis; D. Pseudatheta thailandensis Pace, 1992; E. Pseudatheta cooteri Pace, 1998; F. Pseudatheta elegans Cameron, 1920. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8182112" httpUri="https://zenodo.org/record/8182112/files/figure.png" pageId="7" pageNumber="34">Figs. 1B</figureCitation>
,
<figureCitation id="199E2A0EFFE78E198800FCFCA966FC9A" box="[269,285,806,832]" captionStart="FIGURE 4" captionStartId="7.[151,250,627,651]" captionTargetBox="[164,1419,309,603]" captionTargetId="figure-90@7.[151,1436,309,603]" captionTargetPageId="7" captionText="FIGURE 4. Pseudatheta taiwanensis Pace, 2008. A. male elytra (white arrow indicates granules); B. male tergite VII (white arrow indicates granule); C. male sternite VI (white arrow indicates medial lobe)." figureDoi="http://doi.org/10.5281/zenodo.8182120" httpUri="https://zenodo.org/record/8182120/files/figure.png" pageId="7" pageNumber="34">4</figureCitation>
,
<figureCitation id="199E2A0EFFE78E198824FCFFA911FC9A" box="[297,362,805,832]" captionStart="FIGURE 5" captionStartId="8.[151,250,1626,1650]" captionTargetBox="[161,1430,250,1592]" captionTargetId="figure-73@8.[151,1436,237,1603]" captionTargetPageId="8" captionText="FIGURE 5. Pseudatheta spp. A–C: Pseudatheta crenulicauda (Bernhauer, 1907), comb. nov., D–F: Pseudatheta taiwanensis Pace, 2008. A, D. male tergite VIII in dorsal view; B, E. median lobe of aedeagus in lateral view; C, F. spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8182126" httpUri="https://zenodo.org/record/8182126/files/figure.png" pageId="7" pageNumber="34">5D–F</figureCitation>
)
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<paragraph id="811A368BFFE78E19899AFC90AB21FCBE" blockId="7.[151,858,769,868]" box="[151,858,842,868]" pageId="7" pageNumber="34">[Japanese name: Taiwan-miiro-kinokotsuyakeshi-hanekakushi]</paragraph>
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<taxonomicName id="46A54D08FFE78E19899AFC5DAA46FC45" ID-CoL="4NLJV" authority="Pace, 2008: 146" authorityName="Pace" authorityPageNumber="146" authorityYear="2008" box="[151,573,903,928]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="34" phylum="Arthropoda" rank="species" species="taiwanensis">
<emphasis id="B3D1EA99FFE78E19899AFC5DA9E8FC45" box="[151,403,903,927]" italics="true" pageId="7" pageNumber="34">Pseudatheta taiwanensis</emphasis>
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<bibRefCitation id="E5344B7AFFE78E198897FC52AA46FC45" author="Pace, R." box="[410,573,903,928]" pageId="7" pageNumber="34" pagination="137 - 156" refId="ref16257" refString="Pace, R. (2008) Pronomaeini, Homalotini, Bolitocharini, Diestotini e Falagriini di Taiwan (Coleoptera, Staphylinidae). Bollettino del Museo Civico di Storia Naturale di Verona. Botanica Zoologia, 32, 137 - 156." type="journal article" year="2008">Pace, 2008: 146</bibRefCitation>
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(original description;
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locality: “
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,
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Hs., for. abv. Tona For. Sta.
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<collectingCountry id="F9B2761BFFE78E198894FC3FA988FC25" box="[409,499,997,1023]" name="Japan" pageId="7" pageNumber="34">JAPAN</collectingCountry>
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: [
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,
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,
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.
<collectorName id="2C50535DFFE78E198DE6FC3CAD12FBDA" box="[1259,1385,998,1024]" pageId="7" pageNumber="34">Hashizume</collectorName>
leg. (
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,
<collectorName id="2C50535DFFE78E19886CFBD1A995FBFC" box="[353,494,1035,1062]" pageId="7" pageNumber="34">Ôsawaguchi</collectorName>
, Nopporo-shinrin-kôen,
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,
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16–19
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2001,
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leg. (
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);
<materialsCitation id="31CD3CD6FFE78E198C3EFBD0ABE0FB92" collectingDate="2021-08-08" collectionCode="T" collectorName="Kannonzawa-rindo &amp; Minami-ku" country="Japan" location="Sapporo-shi" pageId="7" pageNumber="34" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Hokkaido">
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,
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,
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,
<collectorName id="2C50535DFFE78E1988DBFBF4AA2CFB92" box="[470,599,1070,1096]" pageId="7" pageNumber="34">Minami-ku</collectorName>
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,
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,
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.
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leg. (
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, same data, but
<date id="F51B104BFFE78E198A4DFBAFAB13FB4A" box="[832,872,1141,1168]" pageId="7" pageNumber="34" value="1998-05-05">
5
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1998 (
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, Hyô-no-sen,
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, I. H. B. C. (
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23
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2020,
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,
<date id="F51B104BFFE78E198A1EFB64AB40FB02" box="[787,827,1214,1240]" pageId="7" pageNumber="34" value="1971-05-07">
7
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1971,
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leg. (
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<collectingDate id="E55FE9A3FFE78E198C68FB67A8A9FB26" pageId="7" pageNumber="34" value="1968-10-15">15 X 1968</collectingDate>
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, M.-
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<collectorName id="2C50535DFFE78E19882AFB39A930FB24" box="[295,331,1251,1278]" pageId="7" pageNumber="34">Ch</collectorName>
̊jô leg. (
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<date id="F51B104BFFE78E19899AFADCA8B4FAFA" box="[151,207,1286,1312]" pageId="7" pageNumber="34" value="2021-05-14">
14
<collectionCode id="E7B4AE4EFFE78E1989B6FADCA8B4FAFA" box="[187,207,1286,1312]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="34" type="Museum">V</collectionCode>
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2021,
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leg. (
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31
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–10
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2011,
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leg. (
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).
</paragraph>
</subSubSection>
<subSubSection id="C9BF6500FFE78E1689CAFA97ADE9F890" lastPageId="8" lastPageNumber="35" pageId="7" pageNumber="34" type="description">
<paragraph id="811A368BFFE78E1989CAFA97AB56FA56" blockId="7.[151,1437,997,2032]" pageId="7" pageNumber="34">
<emphasis id="B3D1EA99FFE78E1989CAFA97A90EFABD" bold="true" box="[199,373,1357,1383]" pageId="7" pageNumber="34">Redescription.</emphasis>
Measurements (n = 5): BL ≈ 1.90–2.13; FBL: 1.09–1.25; HL: 0.34–0.38; HW: 0.39–0.44; PL: 0.38–0.40; PW: 0.53–0.59; EL: 0.36–0.43; EW: 0.62–0.72.
</paragraph>
<paragraph id="811A368BFFE78E1989CAFA4CACDEFA0E" blockId="7.[151,1437,997,2032]" pageId="7" pageNumber="34">Relative length of antennomeres I–XI (n = 1): 20: 22: 18: 10: 12: 13: 13: 11: 12: 12: 33. Ratio of length/width of antennomeres I–XI (n = 1): 1.90: 2.27: 1.86: 0.80: 0.73: 0.78: 0.77: 0.63: 0.73: 0.69: 1.79.</paragraph>
<paragraph id="811A368BFFE78E1989CAFA04A89EF9C6" blockId="7.[151,1437,997,2032]" pageId="7" pageNumber="34">
Body (
<figureCitation id="199E2A0EFFE78E198819FA04A910FA22" box="[276,363,1502,1528]" captionStart="FIGURE 1" captionStartId="3.[151,250,1944,1968]" captionTargetBox="[178,1424,196,1901]" captionTargetId="figure-21@3.[164,1424,179,1919]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Pseudatheta spp. A. Pseudatheta crenulicauda (Bernhauer, 1907), comb. nov.; B. Pseudatheta taiwanensis Pace, 2008; C. Pseudatheta hilaris (Sharp, 1888), comb. nov., lectotype of Oxypoda hiralis; D. Pseudatheta thailandensis Pace, 1992; E. Pseudatheta cooteri Pace, 1998; F. Pseudatheta elegans Cameron, 1920. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8182112" httpUri="https://zenodo.org/record/8182112/files/figure.png" pageId="7" pageNumber="34">Fig. 1B</figureCitation>
) reddish brown; head darker; posterolateral areas of elytra darker; abdominal segments V–VII darker.
</paragraph>
<paragraph id="811A368BFFE78E1989CAF9FCAB80F952" blockId="7.[151,1437,997,2032]" pageId="7" pageNumber="34">Head almost as long as wide, HW/HL:1.07–1.23; surface densely covered with setae.Antenna with antennomeres I–III and XI longer than wide, antennomeres IV–X wider than long; antennomeres I and II with rounded distal end, antennomeres III–X strongly angulated near distal end, antennomere XI oval.</paragraph>
<paragraph id="811A368BFFE78E1989CAF948ACAEF8BA" blockId="7.[151,1437,997,2032]" pageId="7" pageNumber="34">Pronotum transverse, PW/PL: 1.35–1.45, PW/HW: 1.31–1.41; surface densely covered with setae, finely punctured, without microsculpture; posterior margin slightly bisinuate. Elytra wider than long, EW/EL: 1.60–1.79, EL/PL: 0.97–1.06, EW/PW: 1.14–1.22; surface densely covered with setae and finely punctured; posterior margin sinuate near posterolateral corners. Hind wings well developed; flabellum with one seta. Mesoventrite without longitudinal carina; mesoventral process extended to slightly beyond middle of mesocoxal cavities, with rounded apex; metaventral process shorter than mesoventral process, with rounded apex; isthmus present.</paragraph>
<paragraph id="811A368BFFE78E1989CAF8B0AC71F85E" blockId="7.[151,1437,997,2032]" box="[199,1034,1898,1924]" pageId="7" pageNumber="34">Abdomen slightly narrowed posteriad; surface densely covered with setae.</paragraph>
<paragraph id="811A368BFFE78E1689CAF854AD05FF0F" blockId="7.[151,1437,997,2032]" lastBlockId="8.[151,1436,150,213]" lastPageId="8" lastPageNumber="35" pageId="7" pageNumber="34">
Male. Elytra with a pair of small tubercles at about posterior 1/3 on sutural margin. Sternite VI with small medial lobe on posterior margin. Tergite VII with a tubercle on posteromedian area. Tergite VIII (
<figureCitation id="199E2A0EFFE78E198DE0F868AD3EF816" box="[1261,1349,1969,1996]" captionStart="FIGURE 5" captionStartId="8.[151,250,1626,1650]" captionTargetBox="[161,1430,250,1592]" captionTargetId="figure-73@8.[151,1436,237,1603]" captionTargetPageId="8" captionText="FIGURE 5. Pseudatheta spp. A–C: Pseudatheta crenulicauda (Bernhauer, 1907), comb. nov., D–F: Pseudatheta taiwanensis Pace, 2008. A, D. male tergite VIII in dorsal view; B, E. median lobe of aedeagus in lateral view; C, F. spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8182126" httpUri="https://zenodo.org/record/8182126/files/figure.png" pageId="7" pageNumber="34">Fig. 5D</figureCitation>
) with a tubercle on median area; three to five processes on each side of posterior margin, outer ones slightly curved, longer and sharper than the others.Aedeagus as in
<figureCitation id="199E2A0EFFE88E168B75FF4DAAB6FF6B" box="[632,717,150,177]" captionStart="FIGURE 5" captionStartId="8.[151,250,1626,1650]" captionTargetBox="[161,1430,250,1592]" captionTargetId="figure-73@8.[151,1436,237,1603]" captionTargetPageId="8" captionText="FIGURE 5. Pseudatheta spp. A–C: Pseudatheta crenulicauda (Bernhauer, 1907), comb. nov., D–F: Pseudatheta taiwanensis Pace, 2008. A, D. male tergite VIII in dorsal view; B, E. median lobe of aedeagus in lateral view; C, F. spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8182126" httpUri="https://zenodo.org/record/8182126/files/figure.png" pageId="8" pageNumber="35">Fig. 5E</figureCitation>
; median lobe with large basal bulb; apical process broad, parallel to near apex, not dilated in lateral view, apical end thin and elongated, apex pointed; flagellum moderately long.
</paragraph>
<caption id="D5DA6603FFE88E16899AF980A897F960" ID-DOI="http://doi.org/10.5281/zenodo.8182126" ID-Zenodo-Dep="8182126" httpUri="https://zenodo.org/record/8182126/files/figure.png" pageId="8" pageNumber="35" startId="8.[151,250,1626,1650]" targetBox="[161,1430,250,1592]" targetPageId="8" targetType="figure">
<paragraph id="811A368BFFE88E16899AF980A897F960" blockId="8.[151,1437,1626,1722]" pageId="8" pageNumber="35">
<emphasis id="B3D1EA99FFE88E16899AF980A96FF9A8" bold="true" box="[151,276,1626,1650]" pageId="8" pageNumber="35">FIGURE 5.</emphasis>
<taxonomicName id="46A54D08FFE88E168816F980A999F9A8" box="[283,482,1626,1650]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="35" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="B3D1EA99FFE88E168816F980A9E3F9A8" box="[283,408,1626,1650]" italics="true" pageId="8" pageNumber="35">Pseudatheta</emphasis>
spp. A
</taxonomicName>
–C:
<taxonomicName id="46A54D08FFE88E168B01F980ABA3F9A8" authority="(Bernhauer, 1907)" authorityName="Hashizume &amp; Yamamoto &amp; Maruyama" authorityYear="2023" baseAuthorityName="Bernhauer" baseAuthorityYear="1907" box="[524,984,1626,1650]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="35" phylum="Arthropoda" rank="species" species="crenulicauda" status="comb. nov.">
<emphasis id="B3D1EA99FFE88E168B01F980AB6EF9A8" box="[524,789,1626,1650]" italics="true" pageId="8" pageNumber="35">Pseudatheta crenulicauda</emphasis>
(
<bibRefCitation id="E5344B7AFFE88E168A2EF980ABABF9A8" author="Bernhauer, M." box="[803,976,1626,1650]" pageId="8" pageNumber="35" pagination="371 - 414" refId="ref15129" refString="Bernhauer, M. (1907) Zur Staphylinidenfauna von Japan. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 57, 371 - 414." type="journal article" year="1907">Bernhauer, 1907</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="B3D1EA99FFE88E168AEEF980AC2CF9A8" bold="true" box="[995,1111,1626,1650]" pageId="8" pageNumber="35">
<taxonomicNameLabel id="A8E257E2FFE88E168AEEF980AC2CF9A8" box="[995,1111,1626,1650]" pageId="8" pageNumber="35" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, D–F:
<taxonomicName id="46A54D08FFE88E168DADF980A972F94C" authority="Pace, 2008" authorityName="Pace" authorityYear="2008" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="35" phylum="Arthropoda" rank="species" species="taiwanensis">
<emphasis id="B3D1EA99FFE88E168DADF980ADE6F9A8" box="[1184,1437,1626,1650]" italics="true" pageId="8" pageNumber="35">Pseudatheta taiwanensis</emphasis>
<bibRefCitation id="E5344B7AFFE88E16899AF9A5A972F94C" author="Pace, R." box="[151,265,1662,1686]" pageId="8" pageNumber="35" pagination="137 - 156" refId="ref16257" refString="Pace, R. (2008) Pronomaeini, Homalotini, Bolitocharini, Diestotini e Falagriini di Taiwan (Coleoptera, Staphylinidae). Bollettino del Museo Civico di Storia Naturale di Verona. Botanica Zoologia, 32, 137 - 156." type="journal article" year="2008">Pace, 2008</bibRefCitation>
</taxonomicName>
. A, D. male tergite VIII in dorsal view; B, E. median lobe of aedeagus in lateral view; C, F. spermatheca. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph id="811A368BFFE88E1689CAF8D6ADE9F890" blockId="8.[151,1437,1804,2011]" pageId="8" pageNumber="35">
Female. Spermatheca (
<figureCitation id="199E2A0EFFE88E1688C0F8D6AA59F8FD" box="[461,546,1804,1831]" captionStart="FIGURE 5" captionStartId="8.[151,250,1626,1650]" captionTargetBox="[161,1430,250,1592]" captionTargetId="figure-73@8.[151,1436,237,1603]" captionTargetPageId="8" captionText="FIGURE 5. Pseudatheta spp. A–C: Pseudatheta crenulicauda (Bernhauer, 1907), comb. nov., D–F: Pseudatheta taiwanensis Pace, 2008. A, D. male tergite VIII in dorsal view; B, E. median lobe of aedeagus in lateral view; C, F. spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8182126" httpUri="https://zenodo.org/record/8182126/files/figure.png" pageId="8" pageNumber="35">Fig. 5F</figureCitation>
) curved twice, with a transverse band-like structure at base of distal portion; distal portion elongated, U-shaped; median portion elongated, slightly curved; proximal portion longer than wide.
</paragraph>
</subSubSection>
<subSubSection id="C9BF6500FFE88E1689CAF88EAB80F8B4" box="[199,1019,1876,1903]" pageId="8" pageNumber="35" type="distribution">
<paragraph id="811A368BFFE88E1689CAF88EAB80F8B4" blockId="8.[151,1437,1804,2011]" box="[199,1019,1876,1903]" pageId="8" pageNumber="35">
<emphasis id="B3D1EA99FFE88E1689CAF88EA91AF8B4" bold="true" box="[199,353,1876,1902]" pageId="8" pageNumber="35">Distribution.</emphasis>
<collectingCountry id="F9B2761BFFE88E16886AF88FA9DCF8B5" box="[359,423,1877,1903]" name="Japan" pageId="8" pageNumber="35">Japan</collectingCountry>
(
<collectingRegion id="4361F869FFE88E1688BBF88EAA52F8B4" box="[438,553,1876,1902]" country="Japan" name="Hokkaido" pageId="8" pageNumber="35">Hokkaido</collectingRegion>
, Honshu, Kyushu)—new record;
<collectingCountry id="F9B2761BFFE88E168AACF88EAB8CF8B4" box="[929,1015,1876,1902]" name="Taiwan" pageId="8" pageNumber="35">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C9BF6500FFE88E1789CAF8A3ACE9FEC7" lastPageId="9" lastPageNumber="36" pageId="8" pageNumber="35" type="discussion">
<paragraph id="811A368BFFE88E1789CAF8A3ACE9FEC7" blockId="8.[151,1437,1804,2011]" lastBlockId="9.[151,1437,151,285]" lastPageId="9" lastPageNumber="36" pageId="8" pageNumber="35">
<emphasis id="B3D1EA99FFE88E1689CAF8A3A940F849" bold="true" box="[199,315,1913,1939]" pageId="8" pageNumber="35">Remarks.</emphasis>
This species can be distinguished from
<taxonomicName id="46A54D08FFE88E168A00F8A3ABB6F848" authorityName="Hashizume &amp; Yamamoto &amp; Maruyama" authorityYear="2023" baseAuthorityName="Bernhauer" baseAuthorityYear="1907" box="[781,973,1912,1938]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="35" phylum="Arthropoda" rank="species" species="crenulicauda">
<emphasis id="B3D1EA99FFE88E168A00F8A3ABB6F848" box="[781,973,1912,1938]" italics="true" pageId="8" pageNumber="35">Ps. crenulicauda</emphasis>
</taxonomicName>
by the non-dilated, parallel-sided apical process of its aedeagus. Most of the aedeagi of specimens from
<collectingCountry id="F9B2761BFFE88E168A79F847ABCFF86D" box="[884,948,1949,1975]" name="Japan" pageId="8" pageNumber="35">Japan</collectingCountry>
have an abruptly narrowed and protruding apical part on their apical process, which is not seen in Pace’s illustrations of the specimen from
<collectingCountry id="F9B2761BFFE88E168DFDF81AAD39F800" box="[1264,1346,1984,2010]" name="Taiwan" pageId="8" pageNumber="35">Taiwan</collectingCountry>
(2008). Similar aedeagi are seen in a few Japanese specimens, but it is likely that they were damaged. While we cannot rule out the possibility that the original description of
<taxonomicName id="46A54D08FFE98E178BCBFF61AB0FFF0E" authorityName="Pace" authorityYear="2008" box="[710,884,187,212]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="36" phylum="Arthropoda" rank="species" species="taiwanensis">
<emphasis id="B3D1EA99FFE98E178BCBFF61AB0FFF0E" box="[710,884,187,212]" italics="true" pageId="9" pageNumber="36">Ps. taiwanensis</emphasis>
</taxonomicName>
was based on a specimen with a damaged apical process of aedeagus, we nonetheless identified the Japanese specimen as
<taxonomicName id="46A54D08FFE98E178ADDFF05AC04FF22" authorityName="Pace" authorityYear="2008" box="[976,1151,223,248]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="36" phylum="Arthropoda" rank="species" species="taiwanensis">
<emphasis id="B3D1EA99FFE98E178ADDFF05AC04FF22" box="[976,1151,223,248]" italics="true" pageId="9" pageNumber="36">Ps. taiwanensis</emphasis>
</taxonomicName>
. The female morphology is not informative for distinguishing this species from
<taxonomicName id="46A54D08FFE98E178BFAFED9ABCFFEC7" authorityName="Hashizume &amp; Yamamoto &amp; Maruyama" authorityYear="2023" baseAuthorityName="Bernhauer" baseAuthorityYear="1907" box="[759,948,259,285]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="36" phylum="Arthropoda" rank="species" species="crenulicauda">
<emphasis id="B3D1EA99FFE98E178BFAFED9ABCFFEC7" box="[759,948,259,285]" italics="true" pageId="9" pageNumber="36">Ps. crenulicauda</emphasis>
</taxonomicName>
and other relatives.
</paragraph>
</subSubSection>
</treatment>
</document>
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