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<mods:title id="32C4FF4AEAEEF8BA8881B21090947B85">Revision of the Genera Pseudatheta Cameron and Phymatura Sahlberg (Coleoptera, Staphylinidae, Aleocharinae) from Japan</mods:title>
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<mods:namePart id="682E2864A45EFC6ACDD7B777B521E170">Hashizume, Takuto</mods:namePart>
<mods:affiliation id="12A66A232610C0F60469FDDAF3415800">Entomological Laboratory, Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka, 819 - 0395 Japan.</mods:affiliation>
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<mods:namePart id="D99A33C2E5FD52135C08E39E2CB031F8">Yamamoto, Shûhei</mods:namePart>
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<mods:affiliation id="8CB9953ECED3E8911659E33D4EA5104B">The Hokkaido University Museum, Hokkaido University, Kita 10, Nishi 8, Kita-ku, Sapporo, 060 - 0810 Japan. s. yamamoto. 64 @ gmail. com; https: // orcid. org / 0000 - 0002 - 4162 - 8457</mods:affiliation>
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<mods:namePart id="3CD3DE5893FA0EF04B806F105F0ADB88">Maruyama, Munetoshi</mods:namePart>
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<heading id="DA5281E7FFE28E1C899AFA40AB2BFA6F" bold="true" box="[151,848,1434,1461]" fontSize="11" level="1" pageId="2" pageNumber="29" reason="1">
<emphasis id="B3D1EA99FFE28E1C899AFA40AB2BFA6F" bold="true" box="[151,848,1434,1461]" pageId="2" pageNumber="29">
<taxonomicName id="46A54D08FFE28E1C899AFA40AAC7FA6E" authority="(Bernhauer, 1907)" authorityName="Hashizume &amp; Yamamoto &amp; Maruyama" authorityYear="2023" baseAuthorityName="Bernhauer" baseAuthorityYear="1907" box="[151,700,1434,1461]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="crenulicauda" status="comb. nov.">
<emphasis id="B3D1EA99FFE28E1C899AFA40A9A8FA6F" bold="true" box="[151,467,1434,1461]" italics="true" pageId="2" pageNumber="29">Pseudatheta crenulicauda</emphasis>
(
<bibRefCitation id="E5344B7AFFE28E1C88EFFA40AACFFA6E" author="Bernhauer, M." box="[482,692,1434,1460]" pageId="2" pageNumber="29" pagination="371 - 414" refId="ref15129" refString="Bernhauer, M. (1907) Zur Staphylinidenfauna von Japan. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 57, 371 - 414." type="journal article" year="1907">Bernhauer, 1907</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicNameLabel id="A8E257E2FFE28E1C8BC7FA41AB2BFA6F" box="[714,848,1435,1461]" pageId="2" pageNumber="29" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C9BF6500FFE28E1C899AFA61A908FA09" box="[151,371,1467,1491]" pageId="2" pageNumber="29" type="description">
<paragraph id="811A368BFFE28E1C899AFA61A908FA09" blockId="2.[151,848,1434,1528]" box="[151,371,1467,1491]" pageId="2" pageNumber="29">
(
<figureCitation id="199E2A0EFFE28E1C8992FA61A883FA09" box="[159,248,1467,1491]" captionStart="FIGURE 1" captionStartId="3.[151,250,1944,1968]" captionTargetBox="[178,1424,196,1901]" captionTargetId="figure-21@3.[164,1424,179,1919]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Pseudatheta spp. A. Pseudatheta crenulicauda (Bernhauer, 1907), comb. nov.; B. Pseudatheta taiwanensis Pace, 2008; C. Pseudatheta hilaris (Sharp, 1888), comb. nov., lectotype of Oxypoda hiralis; D. Pseudatheta thailandensis Pace, 1992; E. Pseudatheta cooteri Pace, 1998; F. Pseudatheta elegans Cameron, 1920. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8182112" httpUri="https://zenodo.org/record/8182112/files/figure.png" pageId="2" pageNumber="29">Figs. 1A</figureCitation>
, 
<figureCitation id="199E2A0EFFE28E1C8809FA61A958FA09" box="[260,291,1467,1491]" captionStart="FIGURE 2" captionStartId="4.[151,250,1825,1849]" captionTargetBox="[151,1436,194,1800]" captionTargetId="figure-35@4.[151,1436,181,1800]" captionTargetPageId="4" captionText="FIGURE 2. Habitus of Phymatura spp. and type specimens examined. A. Phymatura japonica Cameron, 1933, stat. rev.; B. Phymatura russa Assing, 2021; C. lectotype of Atheta (Datomicra) crenulicauda Bernhauer, 1907 (=Pseudatheta crenulicauda comb. nov.); D. labels of C; E. holotype of Phymatura japonica Cameron, 1933, stat. rev. Scale bars: 1.0 mm for A–B." figureDoi="http://doi.org/10.5281/zenodo.8182114" httpUri="https://zenodo.org/record/8182114/files/figure.png" pageId="2" pageNumber="29">2C</figureCitation>
, 
<figureCitation id="199E2A0EFFE28E1C8823FA61A917FA09" box="[302,364,1467,1491]" captionStart="FIGURE 5" captionStartId="8.[151,250,1626,1650]" captionTargetBox="[161,1430,250,1592]" captionTargetId="figure-73@8.[151,1436,237,1603]" captionTargetPageId="8" captionText="FIGURE 5. Pseudatheta spp. A–C: Pseudatheta crenulicauda (Bernhauer, 1907), comb. nov., D–F: Pseudatheta taiwanensis Pace, 2008. A, D. male tergite VIII in dorsal view; B, E. median lobe of aedeagus in lateral view; C, F. spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8182126" httpUri="https://zenodo.org/record/8182126/files/figure.png" pageId="2" pageNumber="29">5A–C</figureCitation>
)
</paragraph>
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<subSubSection id="C9BF6500FFE28E1C899AFA04AB79FA2D" box="[151,770,1501,1528]" pageId="2" pageNumber="29" type="vernacular_names">
<paragraph id="811A368BFFE28E1C899AFA04AB79FA2D" blockId="2.[151,848,1434,1528]" box="[151,770,1501,1528]" pageId="2" pageNumber="29">[Japanese name: Miiro-kinokotsuyakeshi-hanekakushi]</paragraph>
</subSubSection>
<subSubSection id="C9BF6500FFE28E1C899AF9C1ABA6F9E9" box="[151,989,1563,1587]" pageId="2" pageNumber="29" type="reference_group">
<paragraph id="811A368BFFE28E1C899AF9C1ABA6F9E9" blockId="2.[151,1437,1563,1863]" box="[151,989,1563,1587]" pageId="2" pageNumber="29">
<treatmentCitationGroup id="A1B511A5FFE28E1C899AF9C1ABA6F9E9" box="[151,989,1563,1587]" pageId="2" pageNumber="29">
<treatmentCitation id="0004109AFFE28E1C899AF9C1AA89F9E9" author="Bernhauer, M." box="[151,754,1563,1587]" page="398" pageId="2" pageNumber="29" year="1907">
<taxonomicName id="46A54D08FFE28E1C899AF9C1AA89F9E9" ID-CoL="JC4W" authority="Bernhauer, 1907: 398" authorityName="Bernhauer" authorityPageNumber="398" authorityYear="1907" box="[151,754,1563,1587]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="crenulicauda" subGenus="Datomicra">
<emphasis id="B3D1EA99FFE28E1C899AF9C1A8A2F9E9" box="[151,217,1563,1587]" italics="true" pageId="2" pageNumber="29">Atheta</emphasis>
(
<emphasis id="B3D1EA99FFE28E1C89FDF9C6A925F9E9" box="[240,350,1564,1587]" italics="true" pageId="2" pageNumber="29">Datomicra</emphasis>
) 
<emphasis id="B3D1EA99FFE28E1C8879F9C1A982F9E9" box="[372,505,1563,1587]" italics="true" pageId="2" pageNumber="29">crenulicauda</emphasis>
<bibRefCitation id="E5344B7AFFE28E1C8B0AF9C1AA89F9E9" author="Bernhauer, M." box="[519,754,1563,1587]" pageId="2" pageNumber="29" pagination="371 - 414" refId="ref15129" refString="Bernhauer, M. (1907) Zur Staphylinidenfauna von Japan. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 57, 371 - 414." type="journal article" year="1907">Bernhauer, 1907: 398</bibRefCitation>
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(original description;
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</paragraph>
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<paragraph id="811A368BFFE28E1C8AE6F9C6ACF6F988" blockId="2.[151,1437,1563,1863]" pageId="2" pageNumber="29">
<typeStatus id="5E1E8829FFE28E1C8AE6F9C6AC6CF9EE" box="[1003,1047,1564,1588]" pageId="2" pageNumber="29">type</typeStatus>
locality: “Bukenji” [Yokohama-shi, Kanagawa-ken, central Honshu, 
<collectingCountry id="F9B2761BFFE28E1C8B1EF9E0AA28F988" box="[531,595,1594,1618]" name="Japan" pageId="2" pageNumber="29">Japan</collectingCountry>
]); 
<treatmentCitation id="0004109AFFE28E1C8B67F9E0AB57F988" author="Sawada, K." box="[618,812,1594,1618]" page="217" pageId="2" pageNumber="29" year="1977">
<bibRefCitation id="E5344B7AFFE28E1C8B67F9E0AB57F988" author="Sawada, K." box="[618,812,1594,1618]" pageId="2" pageNumber="29" pagination="171 - 222" refId="ref16510" refString="Sawada, K. (1977) Studies on the genus Atheta Thomson and its allies (Coleoptera, Staphylinidae) III: Japanese species described by the previous authors. Contributions from the Biological Laboratory, Kyoto University, 25 (2), 171 - 222." type="journal article" year="1977">Sawada, 1977: 217</bibRefCitation>
</treatmentCitation>
(as a synonym of 
<taxonomicName id="46A54D08FFE28E1C8AEBF9E0AC05F988" baseAuthorityName="Sharp" baseAuthorityYear="1888" box="[998,1150,1594,1618]" class="Insecta" family="Staphylinidae" genus="Platyola" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="oligotinula">
<emphasis id="B3D1EA99FFE28E1C8AEBF9E0AC05F988" box="[998,1150,1594,1618]" italics="true" pageId="2" pageNumber="29">Ph. oligotinula</emphasis>
</taxonomicName>
).
</paragraph>
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<subSubSection id="C9BF6500FFE28E1C899AF982AA96F9AA" box="[151,749,1624,1649]" pageId="2" pageNumber="29" type="reference_group">
<paragraph id="811A368BFFE28E1C899AF982AA96F9AA" blockId="2.[151,1437,1563,1863]" box="[151,749,1624,1649]" pageId="2" pageNumber="29">
<treatmentCitationGroup id="A1B511A5FFE28E1C899AF982AA96F9AA" box="[151,749,1624,1649]" pageId="2" pageNumber="29">
<taxonomicName id="46A54D08FFE28E1C899AF982AA2AF9AB" ID-CoL="77H4C" authority="(Bernhauer, 1907)" baseAuthorityName="Bernhauer" baseAuthorityYear="1907" box="[151,593,1624,1649]" class="Insecta" family="Staphylinidae" genus="Phymatura" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="crenulicauda">
<emphasis id="B3D1EA99FFE28E1C899AF982A9E9F9AA" box="[151,402,1624,1648]" italics="true" pageId="2" pageNumber="29">Phymatura crenulicauda</emphasis>
(
<bibRefCitation id="E5344B7AFFE28E1C8890F982AA32F9AB" author="Bernhauer, M." box="[413,585,1624,1649]" pageId="2" pageNumber="29" pagination="371 - 414" refId="ref15129" refString="Bernhauer, M. (1907) Zur Staphylinidenfauna von Japan. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 57, 371 - 414." type="journal article" year="1907">Bernhauer, 1907</bibRefCitation>
)
</taxonomicName>
; 
<treatmentCitation id="0004109AFFE28E1C8B57F983AA96F9AA" author="Pace, R." box="[602,749,1624,1649]" page="54" pageId="2" pageNumber="29" year="1984">
<bibRefCitation id="E5344B7AFFE28E1C8B57F983AA96F9AA" author="Pace, R." box="[602,749,1624,1649]" pageId="2" pageNumber="29" pagination="51 - 57" refId="ref15893" refString="Pace, R. (1984) Note su alcune specie del genere Platyola Muls. &amp; Rey e generi affini del Giappone e del Gabon (Coleoptera Staphylinidae). Lavori della Societa veneziana di scienze naturali, 9, 51 - 57." type="journal article" year="1984">Pace, 1984: 54</bibRefCitation>
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</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C9BF6500FFE28E1C8BFFF982ABD6F955" pageId="2" pageNumber="29" type="discussion">
<paragraph id="811A368BFFE28E1C8BFFF982ABD6F955" blockId="2.[151,1437,1563,1863]" pageId="2" pageNumber="29">
(transferred to the genus 
<taxonomicName id="46A54D08FFE28E1C8AE7F982AC20F9AA" box="[1002,1115,1624,1648]" class="Insecta" family="Staphylinidae" genus="Phymatura" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B3D1EA99FFE28E1C8AE7F982AC20F9AA" box="[1002,1115,1624,1648]" italics="true" pageId="2" pageNumber="29">Phymatura</emphasis>
</taxonomicName>
from the genus 
<taxonomicName id="46A54D08FFE28E1C8DF7F982AD47F9AA" box="[1274,1340,1624,1648]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B3D1EA99FFE28E1C8DF7F982AD47F9AA" box="[1274,1340,1624,1648]" italics="true" pageId="2" pageNumber="29">Atheta</emphasis>
</taxonomicName>
); 
<treatmentCitation id="0004109AFFE28E1C8C5DF982A917F955" author="Shibata, Y. &amp; Maruyama, M. &amp; Hoshina, H. &amp; Kishimoto, T. &amp; Naomi, S. - I. &amp; Nomura, S. &amp; Puthz, V &amp; Shimada, T. &amp; Watanabe, Y. &amp; Yamamoto, S." page="115" pageId="2" pageNumber="29" year="2013">
<bibRefCitation id="E5344B7AFFE28E1C8C5DF982A917F955" author="Shibata, Y. &amp; Maruyama, M. &amp; Hoshina, H. &amp; Kishimoto, T. &amp; Naomi, S. - I. &amp; Nomura, S. &amp; Puthz, V &amp; Shimada, T. &amp; Watanabe, Y. &amp; Yamamoto, S." pageId="2" pageNumber="29" pagination="69 - 218" refId="ref16780" refString="Shibata, Y., Maruyama, M., Hoshina, H., Kishimoto, T., Naomi, S. - I., Nomura, S., Puthz, V, Shimada, T., Watanabe, Y. &amp; Yamamoto, S. (2013) Catalogue of Japanese Staphylinidae (Insecta: Coleoptera). Bulletin of the Kyushu University Museum, 11, 69 - 218." type="journal article" year="2013">
Shibata 
<emphasis id="B3D1EA99FFE28E1C89CAF9A2A88FF955" box="[199,244,1655,1679]" italics="true" pageId="2" pageNumber="29">et al</emphasis>
., 2013: 115
</bibRefCitation>
</treatmentCitation>
(catalogue); 
<treatmentCitation id="0004109AFFE28E1C88FCF9ADAB56F955" author="Schulke, M. &amp; Smetana, A." box="[497,813,1655,1679]" page="633" pageId="2" pageNumber="29" year="2015">
<bibRefCitation id="E5344B7AFFE28E1C88FCF9ADAB56F955" author="Schulke, M. &amp; Smetana, A." box="[497,813,1655,1679]" pageId="2" pageNumber="29" refId="ref16621" refString="Schulke, M. &amp; Smetana, A. (2015) Staphylinidae. In: L ˆ bl, I. &amp; L ˆ bl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Staphylinoidea. Revised and updated edition. Brill, Leiden / Boston, pp. 304 - 1134." type="book" year="2015">Schülke &amp; Smetana, 2015: 633</bibRefCitation>
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(catalogue).
</paragraph>
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<subSubSection id="C9BF6500FFE28E1C899AF94CAA20F917" pageId="2" pageNumber="29" type="reference_group">
<paragraph id="811A368BFFE28E1C899AF94CABC4F974" blockId="2.[151,1437,1563,1863]" box="[151,959,1686,1710]" pageId="2" pageNumber="29">
<treatmentCitationGroup id="A1B511A5FFE28E1C899AF94CABC4F974" box="[151,959,1686,1710]" pageId="2" pageNumber="29">
“ 
<treatmentCitation id="0004109AFFE28E1C89AFF94CAA18F974" author="Sawada, K." box="[162,611,1686,1710]" page="217" pageId="2" pageNumber="29" year="1977">
<taxonomicName id="46A54D08FFE28E1C89AFF94CAA18F974" authority="&quot;: Sawada, 1977: 217" authorityName="Sawada" authorityPageNumber="217" authorityYear="1977" box="[162,611,1686,1710]" class="Insecta" family="Staphylinidae" genus="Phymatura" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="oligotinula">
<emphasis id="B3D1EA99FFE28E1C89AFF94CA9F3F974" box="[162,392,1686,1710]" italics="true" pageId="2" pageNumber="29">Phymatura oligotinula</emphasis>
”: 
<bibRefCitation id="E5344B7AFFE28E1C88ACF94CAA18F974" author="Sawada, K." box="[417,611,1686,1710]" pageId="2" pageNumber="29" pagination="171 - 222" refId="ref16510" refString="Sawada, K. (1977) Studies on the genus Atheta Thomson and its allies (Coleoptera, Staphylinidae) III: Japanese species described by the previous authors. Contributions from the Biological Laboratory, Kyoto University, 25 (2), 171 - 222." type="journal article" year="1977">Sawada, 1977: 217</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(redescription; misidentification).
</treatmentCitationGroup>
</paragraph>
<paragraph id="811A368BFFE28E1C899AF96FAA20F917" blockId="2.[151,1437,1563,1863]" box="[151,603,1716,1741]" pageId="2" pageNumber="29">
<treatmentCitationGroup id="A1B511A5FFE28E1C899AF96FAA20F917" box="[151,603,1716,1741]" pageId="2" pageNumber="29">
“ 
<treatmentCitation id="0004109AFFE28E1C89AFF96EAA20F917" author="Pace, R." box="[162,603,1716,1741]" page="242" pageId="2" pageNumber="29" year="1992">
<taxonomicName id="46A54D08FFE28E1C89AFF96EAA20F917" ID-CoL="4NLJK" authority="Pace, 1992: 242" authorityName="Pace" authorityPageNumber="242" authorityYear="1992" box="[162,603,1716,1741]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="oligotinula">
<emphasis id="B3D1EA99FFE28E1C89AFF96EA9E3F916" box="[162,408,1716,1740]" italics="true" pageId="2" pageNumber="29">Pseudatheta oligotinula</emphasis>
”: 
<bibRefCitation id="E5344B7AFFE28E1C88BEF96FAA20F917" author="Pace, R." box="[435,603,1717,1741]" pageId="2" pageNumber="29" pagination="227 - 268" refId="ref16061" refString="Pace, R. (1992) Aleocharinae della Thailandia (Coleoptera Staphylinidae). Bollettino del Museo Civico di Storia Naturale di Verona, 16, 227 - 268." type="journal article" year="1992">Pace, 1992: 242</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C9BF6500FFE28E1C8B69F96EA961F931" pageId="2" pageNumber="29" type="discussion">
<paragraph id="811A368BFFE28E1C8B69F96EA961F931" blockId="2.[151,1437,1563,1863]" pageId="2" pageNumber="29">
(transferred to genus 
<taxonomicName id="46A54D08FFE28E1C8A49F96EABBAF916" authorityName="Cameron" authorityYear="1920" box="[836,961,1716,1740]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B3D1EA99FFE28E1C8A49F96EABBAF916" box="[836,961,1716,1740]" italics="true" pageId="2" pageNumber="29">Pseudatheta</emphasis>
</taxonomicName>
from 
<taxonomicName id="46A54D08FFE28E1C8D08F96EAC0DF916" box="[1029,1142,1716,1740]" class="Insecta" family="Staphylinidae" genus="Phymatura" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B3D1EA99FFE28E1C8D08F96EAC0DF916" box="[1029,1142,1716,1740]" italics="true" pageId="2" pageNumber="29">Phymatura</emphasis>
</taxonomicName>
; probably based on 
<bibRefCitation id="E5344B7AFFE28E1C8C40F96EA96DF931" author="Sawada, K." pageId="2" pageNumber="29" pagination="171 - 222" refId="ref16510" refString="Sawada, K. (1977) Studies on the genus Atheta Thomson and its allies (Coleoptera, Staphylinidae) III: Japanese species described by the previous authors. Contributions from the Biological Laboratory, Kyoto University, 25 (2), 171 - 222." type="journal article" year="1977">Sawada (1977))</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C9BF6500FFE28E1C899AF928AAACF8D0" box="[151,727,1778,1802]" pageId="2" pageNumber="29" type="reference_group">
<paragraph id="811A368BFFE28E1C899AF928AAACF8D0" blockId="2.[151,1437,1563,1863]" box="[151,727,1778,1802]" pageId="2" pageNumber="29">
<treatmentCitationGroup id="A1B511A5FFE28E1C899AF928AAACF8D0" box="[151,727,1778,1802]" pageId="2" pageNumber="29">
<treatmentCitation id="0004109AFFE28E1C899AF928A984F8D0" author="Pace, R." box="[151,511,1778,1802]" page="268" pageId="2" pageNumber="29" year="2010">
<taxonomicName id="46A54D08FFE28E1C899AF928A984F8D0" ID-CoL="4NLJQ" authority="Pace, 2010: 268" authorityName="Pace" authorityPageNumber="268" authorityYear="2010" box="[151,511,1778,1802]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B3D1EA99FFE28E1C899AF928A920F8D0" box="[151,347,1778,1802]" italics="true" pageId="2" pageNumber="29">Pseudatheta similis</emphasis>
<bibRefCitation id="E5344B7AFFE28E1C886DF928A984F8D0" author="Pace, R." box="[352,511,1778,1802]" pageId="2" pageNumber="29" pagination="259 - 280" refId="ref16300" refString="Pace, R. (2010) Biodiversita delle Aleocharinae della Cina: Placusini, Homalotini, Bolitocharini, Eusteniamorphini e Falagriini 1 (Coleoptera, Staphylinidae). Beitrage zur entomologie, 60 (2), 259 - 280. https: // doi. org / 10.21248 / contrib. entomol. 60.2.259 - 280" type="journal article" year="2010">Pace, 2010: 268</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description;
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C9BF6500FFE28E1C8BD1F928AB53F89D" pageId="2" pageNumber="29" type="distribution">
<paragraph id="811A368BFFE28E1C8BD1F928AB53F89D" blockId="2.[151,1437,1563,1863]" pageId="2" pageNumber="29">
<typeStatus id="5E1E8829FFE28E1C8BD1F928AB73F8D0" box="[732,776,1778,1802]" pageId="2" pageNumber="29">type</typeStatus>
locality: “ 
<collectingCountry id="F9B2761BFFE28E1C8A60F928ABD4F8D0" box="[877,943,1778,1802]" name="China" pageId="2" pageNumber="29">China</collectingCountry>
: 
<collectingRegion id="4361F869FFE28E1C8ABAF928AC70F8D0" box="[951,1035,1778,1802]" country="China" name="Sichuan" pageId="2" pageNumber="29">Sichuan</collectingRegion>
, Ganzi Prefecture, Daxue Shan, Gongga Shan, Hailougou glacier park, 
<geoCoordinate id="E491504CFFE28E1C8B08F8CBAA1DF8F3" box="[517,614,1809,1833]" degrees="102" direction="east" minutes="04" orientation="longitude" pageId="2" pageNumber="29" precision="925" value="102.066666">102°04’E</geoCoordinate>
<geoCoordinate id="E491504CFFE28E1C8B62F8CAAAB2F8F2" box="[623,713,1808,1832]" degrees="29" direction="north" minutes="36" orientation="latitude" pageId="2" pageNumber="29" precision="925" value="29.6">29°36’N</geoCoordinate>
, river valley, ca. 
<quantity id="465D9B6EFFE28E1C8A70F8CBABC8F8F2" box="[893,947,1808,1833]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="29" unit="km" value="1.0">1 km</quantity>
above camp I, 
<quantity id="465D9B6EFFE28E1C8D5AF8CBACD1F8F3" box="[1111,1194,1809,1833]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="29" unit="m" value="2100.0">2100 m</quantity>
”); 
<treatmentCitation id="0004109AFFE28E1C8DC5F8CAA956F89D" author="Schulke, M. &amp; Smetana, A." page="634" pageId="2" pageNumber="29" year="2015">
<bibRefCitation id="E5344B7AFFE28E1C8DC5F8CAA956F89D" author="Schulke, M. &amp; Smetana, A." pageId="2" pageNumber="29" refId="ref16621" refString="Schulke, M. &amp; Smetana, A. (2015) Staphylinidae. In: L ˆ bl, I. &amp; L ˆ bl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Staphylinoidea. Revised and updated edition. Brill, Leiden / Boston, pp. 304 - 1134." type="book" year="2015">Schülke &amp; Smetana, 2015: 634</bibRefCitation>
</treatmentCitation>
(catalogue); 
<treatmentCitation id="0004109AFFE28E1C88BEF8EAAA2EF89D" author="Pace, R." box="[435,597,1839,1863]" page="301" pageId="2" pageNumber="29" year="2016">
<bibRefCitation id="E5344B7AFFE28E1C88BEF8EAAA2EF89D" author="Pace, R." box="[435,597,1839,1863]" pageId="2" pageNumber="29" pagination="295 - 336" refId="ref16439" refString="Pace, R. (2016) Aleocharinae della Cina al &quot; Naturkundemuseum &quot; di Erfurt (Insecta: Coleoptera: Staphyinidae). Veroffentlichungen des Naturkundemuseums Erfurt - VERNATE, 35, 295 - 336." type="journal article" year="2016">Pace, 2016: 301</bibRefCitation>
</treatmentCitation>
(
<collectingRegion id="4361F869FFE28E1C8B6EF8F5AACCF89D" box="[611,695,1839,1863]" country="China" name="Yunnan" pageId="2" pageNumber="29">Yunnan</collectingRegion>
). 
<emphasis id="B3D1EA99FFE28E1C8BCAF8F5AB53F89D" bold="true" box="[711,808,1839,1863]" pageId="2" pageNumber="29">Syn. nov.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C9BF6500FFE28E18899AF8AAAA2AFD73" lastPageId="6" lastPageNumber="33" pageId="2" pageNumber="29" type="materials_examined">
<paragraph id="811A368BFFE28E1C899AF8AAAC67F82C" blockId="2.[151,1437,1904,2039]" pageId="2" pageNumber="29">
<emphasis id="B3D1EA99FFE28E1C899AF8AAAA46F850" bold="true" box="[151,573,1904,1930]" pageId="2" pageNumber="29">Material examined. Type material.</emphasis>
<taxonomicName id="46A54D08FFE28E1C8B4BF8AAABB9F850" authorityName="Bernhauer" authorityYear="1907" box="[582,962,1904,1930]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="crenulicauda" subGenus="Datomicra">
<emphasis id="B3D1EA99FFE28E1C8B4BF8AAAAF4F850" box="[582,655,1904,1930]" italics="true" pageId="2" pageNumber="29">Atheta</emphasis>
(
<emphasis id="B3D1EA99FFE28E1C8BAFF8ABAB60F850" box="[674,795,1905,1930]" italics="true" pageId="2" pageNumber="29">Datomicra</emphasis>
) 
<emphasis id="B3D1EA99FFE28E1C8A22F8AAABB9F850" box="[815,962,1904,1930]" italics="true" pageId="2" pageNumber="29">crenulicauda</emphasis>
</taxonomicName>
: 
<materialsCitation id="31CD3CD6FFE28E1C8ADEF8ABAC67F82C" collectionCode="FMNH" collectorName="M. Bernhauer" country="Japan" location="Japan" pageId="2" pageNumber="29" specimenCount="1" typeStatus="lectotype">
<typeStatus id="5E1E8829FFE28E1C8ADEF8ABAC3CF851" box="[979,1095,1905,1931]" pageId="2" pageNumber="29">Lectotype</typeStatus>
: here designated, male “Hans Sauter 4000 / Bukenji 10 XI 05 / Unter Schilfhaufen [handwritten] // Bukenji / 
<collectingCountry id="F9B2761BFFE28E1C8D48F84FACF3F875" box="[1093,1160,1941,1967]" name="Japan" pageId="2" pageNumber="29">Japan</collectingCountry>
. Sauter [handwritten] // crenulicauda / Brnh. Typ. [handwritten] // Chicago NHMus / 
<collectorName id="2C50535DFFE28E1C8A6EF863AC78F808" box="[867,1027,1976,2003]" pageId="2" pageNumber="29">M. Bernhauer</collectorName>
/ Collection” (abdominal segments VIII–X and aedeagus were dissected and mounted in Euparal by 
<collectionCode id="E7B4AE4EFFE28E1C8A62F806ABD8F82C" box="[879,931,2012,2038]" country="France" name="University of Montpellier" pageId="2" pageNumber="29">MM</collectionCode>
) (
<collectionCode id="E7B4AE4EFFE28E1C8AB4F806AC6BF82C" box="[953,1040,2012,2038]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="2" pageNumber="29" type="Museum">FMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<caption id="D5DA6603FFE38E1D899AF842AD15F822" ID-DOI="http://doi.org/10.5281/zenodo.8182112" ID-Zenodo-Dep="8182112" httpUri="https://zenodo.org/record/8182112/files/figure.png" pageId="3" pageNumber="30" startId="3.[151,250,1944,1968]" targetBox="[178,1424,196,1901]" targetPageId="3" targetType="figure">
<paragraph id="811A368BFFE38E1D899AF842AD15F822" blockId="3.[151,1436,1944,2041]" pageId="3" pageNumber="30">
<emphasis id="B3D1EA99FFE38E1D899AF842A962F86A" bold="true" box="[151,281,1944,1968]" pageId="3" pageNumber="30">FIGURE 1.</emphasis>
Habitus of 
<taxonomicName id="46A54D08FFE38E1D88ACF843AA0FF86B" box="[417,628,1945,1969]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="B3D1EA99FFE38E1D88ACF843AA65F86B" box="[417,542,1945,1969]" italics="true" pageId="3" pageNumber="30">Pseudatheta</emphasis>
spp. A
</taxonomicName>
. 
<taxonomicName id="46A54D08FFE38E1D8B8EF843AC24F86B" authority="(Bernhauer, 1907)" authorityName="Hashizume &amp; Yamamoto &amp; Maruyama" authorityYear="2023" baseAuthorityName="Bernhauer" baseAuthorityYear="1907" box="[643,1119,1945,1969]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="crenulicauda" status="comb. nov.">
<emphasis id="B3D1EA99FFE38E1D8B8EF843ABE9F86B" box="[643,914,1945,1969]" italics="true" pageId="3" pageNumber="30">Pseudatheta crenulicauda</emphasis>
(
<bibRefCitation id="E5344B7AFFE38E1D8AA9F843AC2CF86B" author="Bernhauer, M." box="[932,1111,1945,1969]" pageId="3" pageNumber="30" pagination="371 - 414" refId="ref15129" refString="Bernhauer, M. (1907) Zur Staphylinidenfauna von Japan. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 57, 371 - 414." type="journal article" year="1907">Bernhauer, 1907</bibRefCitation>
)
</taxonomicName>
, 
<emphasis id="B3D1EA99FFE38E1D8D62F843AC93F86B" bold="true" box="[1135,1256,1945,1969]" pageId="3" pageNumber="30">
<taxonomicNameLabel id="A8E257E2FFE38E1D8D62F843AC93F86B" box="[1135,1256,1945,1969]" pageId="3" pageNumber="30" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
; B. 
<taxonomicName id="46A54D08FFE38E1D8C12F843A9EBF80F" authority="Pace, 2008" authorityName="Pace" authorityYear="2008" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="taiwanensis">
<emphasis id="B3D1EA99FFE38E1D8C12F843A974F80E" italics="true" pageId="3" pageNumber="30">Pseudatheta taiwanensis</emphasis>
<bibRefCitation id="E5344B7AFFE38E1D8814F867A9EBF80F" author="Pace, R." box="[281,400,1981,2005]" pageId="3" pageNumber="30" pagination="137 - 156" refId="ref16257" refString="Pace, R. (2008) Pronomaeini, Homalotini, Bolitocharini, Diestotini e Falagriini di Taiwan (Coleoptera, Staphylinidae). Bollettino del Museo Civico di Storia Naturale di Verona. Botanica Zoologia, 32, 137 - 156." type="journal article" year="2008">Pace, 2008</bibRefCitation>
</taxonomicName>
; C. 
<taxonomicName id="46A54D08FFE38E1D88B2F867AB5EF80F" authority="(Sharp, 1888)" authorityName="Hashizume &amp; Yamamoto &amp; Maruyama" authorityYear="2023" baseAuthorityName="Sharp" baseAuthorityYear="1888" box="[447,805,1981,2005]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="hilaris" status="comb. nov.">
<emphasis id="B3D1EA99FFE38E1D88B2F867AAF2F80F" box="[447,649,1981,2005]" italics="true" pageId="3" pageNumber="30">Pseudatheta hilaris</emphasis>
(
<bibRefCitation id="E5344B7AFFE38E1D8B97F867AB65F80F" author="Sharp, D. S." box="[666,798,1981,2005]" pageId="3" pageNumber="30" pagination="277 - 295" refId="ref16732" refString="Sharp, D. S. (1888) The Staphylinidae of Japan. Annales and Magazine of Natural History, Series 6, 3, 277 - 295 + 369 - 387 + 451 - 464. https: // doi. org / 10.1080 / 00222938809487515" type="journal article" year="1888">Sharp, 1888</bibRefCitation>
)
</taxonomicName>
, 
<emphasis id="B3D1EA99FFE38E1D8A3EF867ABD0F80F" bold="true" box="[819,939,1981,2005]" pageId="3" pageNumber="30">
<taxonomicNameLabel id="A8E257E2FFE38E1D8A3EF867ABD0F80F" box="[819,939,1981,2005]" pageId="3" pageNumber="30" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, lectotype of 
<taxonomicName id="46A54D08FFE38E1D8D4EF867AC91F80F" box="[1091,1258,1981,2005]" class="Insecta" family="Staphylinidae" genus="Oxypoda" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="hiralis">
<emphasis id="B3D1EA99FFE38E1D8D4EF867AC91F80F" box="[1091,1258,1981,2005]" italics="true" pageId="3" pageNumber="30">Oxypoda hiralis</emphasis>
</taxonomicName>
; D. 
<taxonomicName id="46A54D08FFE38E1D8C12F867A9ECF823" authority="Pace, 1992" authorityName="Pace" authorityYear="1992" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis id="B3D1EA99FFE38E1D8C12F867A965F823" italics="true" pageId="3" pageNumber="30">Pseudatheta thailandensis</emphasis>
<bibRefCitation id="E5344B7AFFE38E1D8828F83BA9ECF823" author="Pace, R." box="[293,407,2017,2041]" pageId="3" pageNumber="30" pagination="227 - 268" refId="ref16061" refString="Pace, R. (1992) Aleocharinae della Thailandia (Coleoptera Staphylinidae). Bollettino del Museo Civico di Storia Naturale di Verona, 16, 227 - 268." type="journal article" year="1992">Pace, 1992</bibRefCitation>
</taxonomicName>
; E. 
<taxonomicName id="46A54D08FFE38E1D88B0F83BAB7AF823" authority="Pace, 1998" authorityName="Pace" authorityYear="1998" box="[445,769,2017,2041]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="cooteri">
<emphasis id="B3D1EA99FFE38E1D88B0F83BAAF2F822" box="[445,649,2017,2041]" italics="true" pageId="3" pageNumber="30">Pseudatheta cooteri</emphasis>
<bibRefCitation id="E5344B7AFFE38E1D8B82F83BAB7AF823" author="Pace, R." box="[655,769,2017,2041]" pageId="3" pageNumber="30" pagination="139 - 220" refId="ref16092" refString="Pace, R. (1998) Aleocharinae della Cina: Parte I (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 105 (1), 139 - 220. https: // doi. org / 10.5962 / bhl. part. 80035" type="journal article" year="1998">Pace, 1998</bibRefCitation>
</taxonomicName>
; F. 
<taxonomicName id="46A54D08FFE38E1D8A29F83BACE3F823" authority="Cameron, 1920" authorityName="Cameron" authorityYear="1920" box="[804,1176,2017,2041]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="B3D1EA99FFE38E1D8A29F83BAB8EF823" box="[804,1013,2017,2041]" italics="true" pageId="3" pageNumber="30">Pseudatheta elegans</emphasis>
<bibRefCitation id="E5344B7AFFE38E1D8AF6F83BACE3F823" author="Cameron, M." box="[1019,1176,2017,2041]" pageId="3" pageNumber="30" pagination="212 - 284" refId="ref15189" refString="Cameron, M. (1920) New species of Staphylinidae from Singapore. Part III. Transactions of the Entomological Society of London, 1920, 212 - 284. https: // doi. org / 10.1111 / j. 1365 - 2311.1920. tb 00214. x" type="journal article" year="1920">Cameron, 1920</bibRefCitation>
</taxonomicName>
. Scale bars: 1.0 mm.
</paragraph>
</caption>
<caption id="D5DA6603FFE48E1A899AF8FBAD2EF85B" ID-DOI="http://doi.org/10.5281/zenodo.8182114" ID-Zenodo-Dep="8182114" httpUri="https://zenodo.org/record/8182114/files/figure.png" pageId="4" pageNumber="31" startId="4.[151,250,1825,1849]" targetBox="[151,1436,194,1800]" targetPageId="4" targetType="figure">
<paragraph id="811A368BFFE48E1A899AF8FBAD2EF85B" blockId="4.[151,1436,1825,1921]" pageId="4" pageNumber="31">
<emphasis id="B3D1EA99FFE48E1A899AF8FBA96FF8E3" bold="true" box="[151,276,1825,1849]" pageId="4" pageNumber="31">FIGURE 2.</emphasis>
Habitus of 
<taxonomicName id="46A54D08FFE48E1A8883F8FBAA4AF8E2" box="[398,561,1825,1849]" class="Insecta" family="Staphylinidae" genus="Phymatura" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B3D1EA99FFE48E1A8883F8FBA984F8E3" box="[398,511,1825,1849]" italics="true" pageId="4" pageNumber="31">Phymatura</emphasis>
spp.
</taxonomicName>
and type specimens examined. A. 
<taxonomicName id="46A54D08FFE48E1A8A9BF8FBAD75F8E3" authority="Cameron, 1933" authorityName="Cameron" authorityYear="1933" box="[918,1294,1825,1849]" class="Insecta" family="Staphylinidae" genus="Phymatura" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="japonica" status="stat. rev.">
<emphasis id="B3D1EA99FFE48E1A8A9BF8FBAC1CF8E3" box="[918,1127,1825,1849]" italics="true" pageId="4" pageNumber="31">Phymatura japonica</emphasis>
<bibRefCitation id="E5344B7AFFE48E1A8D62F8FBAD75F8E3" author="Cameron, M." box="[1135,1294,1825,1849]" pageId="4" pageNumber="31" pagination="168 - 175" refId="ref15272" refString="Cameron, M. (1933) New species of Staphylinidae (Col.) from Japan. Entomologist's Monthly Magazine, 69, 168 - 175 + 208 - 219." type="journal article" year="1933">Cameron, 1933</bibRefCitation>
</taxonomicName>
, 
<emphasis id="B3D1EA99FFE48E1A8C17F8FBAD0DF8E3" bold="true" box="[1306,1398,1825,1849]" pageId="4" pageNumber="31">
<taxonomicNameLabel id="A8E257E2FFE48E1A8C17F8FBAD0DF8E3" box="[1306,1398,1825,1849]" pageId="4" pageNumber="31">stat. rev.</taxonomicNameLabel>
</emphasis>
; B. 
<taxonomicName id="46A54D08FFE48E1A899AF89FA9A8F887" authority="Assing, 2021" authorityName="Assing" authorityYear="2021" box="[151,467,1861,1885]" class="Insecta" family="Staphylinidae" genus="Phymatura" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="russa">
<emphasis id="B3D1EA99FFE48E1A899AF89FA93EF887" box="[151,325,1861,1885]" italics="true" pageId="4" pageNumber="31">Phymatura russa</emphasis>
<bibRefCitation id="E5344B7AFFE48E1A8846F89FA9A8F887" author="Assing, V." box="[331,467,1861,1885]" pageId="4" pageNumber="31" pagination="329 - 334" refId="ref15059" refString="Assing, V. (2021) Two new species of Phymatura from China (Coleoptera: Staphylinidae: Aleocharinae). Acta Musei Moraviae, Science biologocae, 106 (2), 329 - 334." type="journal article" year="2021">Assing, 2021</bibRefCitation>
</taxonomicName>
; C. lectotype of 
<taxonomicName id="46A54D08FFE48E1A8B75F89FAC03F887" authority="Bernhauer, 1907" authorityName="Bernhauer" authorityYear="1907" box="[632,1144,1861,1885]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="crenulicauda" subGenus="Datomicra">
<emphasis id="B3D1EA99FFE48E1A8B75F89FAAC1F887" box="[632,698,1861,1885]" italics="true" pageId="4" pageNumber="31">Atheta</emphasis>
(
<emphasis id="B3D1EA99FFE48E1A8BC4F89CAB4CF887" box="[713,823,1862,1885]" italics="true" pageId="4" pageNumber="31">Datomicra</emphasis>
) 
<emphasis id="B3D1EA99FFE48E1A8A48F89FABB1F887" box="[837,970,1861,1885]" italics="true" pageId="4" pageNumber="31">crenulicauda</emphasis>
<bibRefCitation id="E5344B7AFFE48E1A8ADDF89FAC03F887" author="Bernhauer, M." box="[976,1144,1861,1885]" pageId="4" pageNumber="31" pagination="371 - 414" refId="ref15129" refString="Bernhauer, M. (1907) Zur Staphylinidenfauna von Japan. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 57, 371 - 414." type="journal article" year="1907">Bernhauer, 1907</bibRefCitation>
</taxonomicName>
(= 
<taxonomicName id="46A54D08FFE48E1A8D99F89FADE7F887" authorityName="Hashizume &amp; Yamamoto &amp; Maruyama" authorityYear="2023" baseAuthorityName="Bernhauer" baseAuthorityYear="1907" box="[1172,1436,1861,1885]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="crenulicauda" status="comb. nov.">
<emphasis id="B3D1EA99FFE48E1A8D99F89FADE7F887" box="[1172,1436,1861,1885]" italics="true" pageId="4" pageNumber="31">Pseudatheta crenulicauda</emphasis>
</taxonomicName>
<emphasis id="B3D1EA99FFE48E1A899AF8B3A970F85B" bold="true" box="[151,267,1897,1921]" pageId="4" pageNumber="31">
<taxonomicNameLabel id="A8E257E2FFE48E1A899AF8B3A970F85B" box="[151,267,1897,1921]" pageId="4" pageNumber="31" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
); D. labels of C; E. holotype of 
<taxonomicName id="46A54D08FFE48E1A8B5FF8B3ABBDF85B" authority="Cameron, 1933" authorityName="Cameron" authorityYear="1933" box="[594,966,1897,1921]" class="Insecta" family="Staphylinidae" genus="Phymatura" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="japonica" status="stat. rev.">
<emphasis id="B3D1EA99FFE48E1A8B5FF8B3AB59F85B" box="[594,802,1897,1921]" italics="true" pageId="4" pageNumber="31">Phymatura japonica</emphasis>
<bibRefCitation id="E5344B7AFFE48E1A8A25F8B3ABBDF85B" author="Cameron, M." box="[808,966,1897,1921]" pageId="4" pageNumber="31" pagination="168 - 175" refId="ref15272" refString="Cameron, M. (1933) New species of Staphylinidae (Col.) from Japan. Entomologist's Monthly Magazine, 69, 168 - 175 + 208 - 219." type="journal article" year="1933">Cameron, 1933</bibRefCitation>
</taxonomicName>
, 
<emphasis id="B3D1EA99FFE48E1A8ADCF8B3AC57F85B" bold="true" box="[977,1068,1897,1921]" pageId="4" pageNumber="31">
<taxonomicNameLabel id="A8E257E2FFE48E1A8ADCF8B3AC57F85B" box="[977,1068,1897,1921]" pageId="4" pageNumber="31">stat. rev.</taxonomicNameLabel>
</emphasis>
Scale bars: 1.0 mm for A–B.
</paragraph>
</caption>
<caption id="D5DA6603FFE58E1B899AF84DA985F82D" ID-DOI="http://doi.org/10.5281/zenodo.8182118" ID-Zenodo-Dep="8182118" httpUri="https://zenodo.org/record/8182118/files/figure.png" pageId="5" pageNumber="32" startId="5.[151,250,1943,1967]" targetBox="[214,1374,181,1919]" targetPageId="5" targetType="figure">
<paragraph id="811A368BFFE58E1B899AF84DA985F82D" blockId="5.[151,1437,1943,2039]" pageId="5" pageNumber="32">
<emphasis id="B3D1EA99FFE58E1B899AF84DA968F875" bold="true" box="[151,275,1943,1967]" pageId="5" pageNumber="32">FIGURE 3.</emphasis>
Syntypes of 
<taxonomicName id="46A54D08FFE58E1B8897F84DAB69F875" authority="Cameron, 1920" authorityName="Cameron" authorityYear="1920" box="[410,786,1943,1967]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="B3D1EA99FFE58E1B8897F84DAA10F875" box="[410,619,1943,1967]" italics="true" pageId="5" pageNumber="32">Pseudatheta elegans</emphasis>
<bibRefCitation id="E5344B7AFFE58E1B8B7FF84DAB69F875" author="Cameron, M." box="[626,786,1943,1967]" pageId="5" pageNumber="32" pagination="212 - 284" refId="ref15189" refString="Cameron, M. (1920) New species of Staphylinidae from Singapore. Part III. Transactions of the Entomological Society of London, 1920, 212 - 284. https: // doi. org / 10.1111 / j. 1365 - 2311.1920. tb 00214. x" type="journal article" year="1920">Cameron, 1920</bibRefCitation>
</taxonomicName>
. A. habitus in dorsal view; B. median lobe of aedeagus of A; C. labels of A; D. habitus in dorsal view and dissected body parts; E. median lobe of aedeagus of D; F. labels of D. (photo by K. Matsumoto, used with permission).
</paragraph>
</caption>
<paragraph id="811A368BFFE68E1889CAFF4DAA93FF23" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="33">
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<typeStatus id="5E1E8829FFE68E1889CAFF4DA91FFF6B" box="[199,356,151,177]" pageId="6" pageNumber="33">Paralectotype</typeStatus>
: 
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, “crenulicauda / Bernh. Cotypus [handwritten] // Unzen. 2600’ / 
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. Sauter [handwritten] // Chicago NHMus / 
<collectorName id="2C50535DFFE68E188B12FF61AAC0FF0F" box="[543,699,187,213]" pageId="6" pageNumber="33">M. Bernhauer</collectorName>
/ Collection” (abdominal segments VIII–X and spermatheca were dissected and mounted in Euparal by 
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) (
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)
</materialsCitation>
.
</paragraph>
<paragraph id="811A368BFFE68E1889CAFED8AA2AFD73" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="33">
<emphasis id="B3D1EA99FFE68E1889CAFED8AA48FEC6" bold="true" box="[199,563,258,284]" pageId="6" pageNumber="33">
<collectingRegion id="4361F869FFE68E1889CAFED8A93CFEC6" box="[199,327,258,284]" pageId="6" pageNumber="33">Additional</collectingRegion>
material examined.
</emphasis>
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</emphasis>
: [
<emphasis id="B3D1EA99FFE68E188BA7FED8AB40FEC6" bold="true" box="[682,827,258,284]" pageId="6" pageNumber="33">
<collectingRegion id="4361F869FFE68E188BA7FED8AB40FEC6" box="[682,827,258,284]" pageId="6" pageNumber="33">Aomori-ken</collectingRegion>
</emphasis>
]: 
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, 
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, 
<location id="847A6050FFE68E188D28FED9ACD1FEC7" LSID="urn:lsid:plazi:treatment:090C879DFFE28E19899AFA40AC27FEC6:847A6050FFE68E188D28FED9ACD1FEC7" box="[1061,1194,259,285]" country="Japan" municipality="Yasumiya" name="Towada-shi" pageId="6" pageNumber="33" stateProvince="Aomori-ken">Towada-shi</location>
, 
<date id="F51B104BFFE68E188DB9FED8AD2DFEC7" box="[1204,1366,258,285]" pageId="6" pageNumber="33" value="1966-08-04" valueMax="1966-08-05" valueMin="1966-08-04">
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</date>
, M. &amp; M. 
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</materialsCitation>
. 
<materialsCitation id="31CD3CD6FFE68E1889D7FEF2A9D7FE9B" box="[218,428,295,323]" collectionCode="KUM" country="Japan" location="Ch" pageId="6" pageNumber="33" specimenCount="1" stateProvince="Aomori-ken">
<location id="847A6050FFE68E1889D7FEF2A885FE99" LSID="urn:lsid:plazi:treatment:090C879DFFE28E19899AFA40AC27FEC6:847A6050FFE68E1889D7FEF2A885FE99" box="[218,254,296,323]" country="Japan" name="Ch" pageId="6" pageNumber="33" stateProvince="Aomori-ken">Ch</location>
̊jô leg. (
<collectionCode id="E7B4AE4EFFE68E188853FEFDA9DFFE9B" box="[350,420,295,321]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="6" pageNumber="33" type="Herbarium">KUM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="31CD3CD6FFE68E1888BAFEFDADE3FE9B" box="[439,1432,294,321]" collectingDate="2009-08-20" collectionCode="KUM" collectorName="J. Aoki" country="Japan" location="Taiwa-cho" municipality="Yoshida" pageId="6" pageNumber="33" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Miyagi-ken">
[
<emphasis id="B3D1EA99FFE68E1888CDFEFCAA32FE9A" bold="true" box="[448,585,294,320]" pageId="6" pageNumber="33">
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</emphasis>
]: 
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, 
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, 
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, 
<location id="847A6050FFE68E188A94FEFDAC69FE9B" LSID="urn:lsid:plazi:treatment:090C879DFFE28E19899AFA40AC27FEC6:847A6050FFE68E188A94FEFDAC69FE9B" box="[921,1042,295,321]" country="Japan" municipality="Yoshida" name="Taiwa-cho" pageId="6" pageNumber="33" stateProvince="Miyagi-ken">Taiwa-chô</location>
, 
<date id="F51B104BFFE68E188D10FEFDACCBFE9B" box="[1053,1200,295,321]" pageId="6" pageNumber="33" value="2009-08-20">
<collectingDate id="E55FE9A3FFE68E188D10FEFDACCBFE9B" box="[1053,1200,295,321]" pageId="6" pageNumber="33" value="2009-08-20">20 VIII 2009</collectingDate>
</date>
, 
<collectorName id="2C50535DFFE68E188DB6FEFDAD71FE9B" box="[1211,1290,295,321]" pageId="6" pageNumber="33">J. Aoki</collectorName>
leg. (
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)
</materialsCitation>
; 
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[
<emphasis id="B3D1EA99FFE68E1889ACFE90A926FEBE" bold="true" box="[161,349,330,356]" pageId="6" pageNumber="33">
<collectingRegion id="4361F869FFE68E1889ACFE90A926FEBE" box="[161,349,330,356]" pageId="6" pageNumber="33">Fukushima-ken</collectingRegion>
</emphasis>
]: 
<specimenCount id="97A3FD02FFE68E188875FE91A9B3FEBF" box="[376,456,331,357]" pageId="6" pageNumber="33" type="male">1 male</specimenCount>
, 
<collectingMunicipality id="617EACF1FFE68E1888DBFE91AAB2FEBF" box="[470,713,331,357]" pageId="6" pageNumber="33">Shitokigawa-keikoku</collectingMunicipality>
, 
<location id="847A6050FFE68E188BDBFE91AB38FEBF" LSID="urn:lsid:plazi:treatment:090C879DFFE28E19899AFA40AC27FEC6:847A6050FFE68E188BDBFE91AB38FEBF" box="[726,835,331,357]" country="Japan" municipality="Shitokigawa-keikoku" name="Iwaki-shi" pageId="6" pageNumber="33" stateProvince="Fukushima-ken">Iwaki-shi</location>
, 
<date id="F51B104BFFE68E188A5CFE91ABF1FEBF" box="[849,906,331,357]" pageId="6" pageNumber="33" value="1992-05-24">
24 
<collectionCode id="E7B4AE4EFFE68E188A7BFE91ABF1FEBF" box="[886,906,331,357]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="33" type="Museum">V</collectionCode>
</date>
</materialsCitation>
1992, 
<collectorName id="2C50535DFFE68E188AD1FE91AC47FEBF" box="[988,1084,331,357]" pageId="6" pageNumber="33">K. Haga</collectorName>
leg. (
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); 
<materialsCitation id="31CD3CD6FFE68E188DD3FE91A993FE53" collectionCode="V" country="Japan" location="Watarase" pageId="6" pageNumber="33" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Tochigi-ken">
[
<emphasis id="B3D1EA99FFE68E188DEAFE90AD0FFEBE" bold="true" box="[1255,1396,330,356]" pageId="6" pageNumber="33">
<collectingRegion id="4361F869FFE68E188DEAFE90AD0FFEBE" box="[1255,1396,330,356]" pageId="6" pageNumber="33">Tochigi-ken</collectingRegion>
</emphasis>
]: 
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, 
<specimenCount id="97A3FD02FFE68E1889D1FEB5A93BFE53" box="[220,320,367,393]" pageId="6" pageNumber="33" type="female">1 female</specimenCount>
, 
<location id="847A6050FFE68E188846FEB5A9CFFE53" LSID="urn:lsid:plazi:treatment:090C879DFFE28E19899AFA40AC27FEC6:847A6050FFE68E188846FEB5A9CFFE53" box="[331,436,367,393]" country="Japan" name="Watarase" pageId="6" pageNumber="33" stateProvince="Tochigi-ken">Watarase</location>
, 
<date id="F51B104BFFE68E1888B2FEB5A993FE53" box="[447,488,367,393]" pageId="6" pageNumber="33" value="1979-05-06">
6 
<collectionCode id="E7B4AE4EFFE68E1888D9FEB5A993FE53" box="[468,488,367,393]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="33" type="Museum">V</collectionCode>
</date>
</materialsCitation>
1979, 
<collectorName id="2C50535DFFE68E188B38FEB5AAFDFE53" box="[565,646,367,393]" pageId="6" pageNumber="33">M. Tao</collectorName>
leg. (
<collectionCode id="E7B4AE4EFFE68E188BC8FEB5AB77FE53" box="[709,780,367,393]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="6" pageNumber="33" type="Herbarium">KUM</collectionCode>
); 
<materialsCitation id="31CD3CD6FFE68E188A2DFEB5A946FE77" collectingDate="1997-08-09" collectionCode="T" country="Japan" location="Naganohara-machi" municipality="Naganohara" pageId="6" pageNumber="33" specimenCount="1" specimenCount-male="1" stateProvince="Gunma-ken">
[
<emphasis id="B3D1EA99FFE68E188A24FEB4ABC3FE52" bold="true" box="[809,952,366,392]" pageId="6" pageNumber="33">
<collectingRegion id="4361F869FFE68E188A24FEB4ABC3FE52" box="[809,952,366,392]" pageId="6" pageNumber="33">Gunma-ken</collectingRegion>
</emphasis>
]: 
<specimenCount id="97A3FD02FFE68E188ADCFEB5AC64FE53" box="[977,1055,367,393]" pageId="6" pageNumber="33" type="male">1 male</specimenCount>
, 
<collectingMunicipality id="617EACF1FFE68E188D26FEB5ACCEFE53" box="[1067,1205,367,393]" pageId="6" pageNumber="33">Naganohara</collectingMunicipality>
, 
<location id="847A6050FFE68E188DCDFEB5ADE3FE53" LSID="urn:lsid:plazi:treatment:090C879DFFE28E19899AFA40AC27FEC6:847A6050FFE68E188DCDFEB5ADE3FE53" box="[1216,1432,367,393]" country="Japan" municipality="Naganohara" name="Naganohara-machi" pageId="6" pageNumber="33" stateProvince="Gunma-ken">Naganohara-machi</location>
, 
<date id="F51B104BFFE68E18899AFE49A95BFE77" box="[151,288,403,429]" pageId="6" pageNumber="33" value="1997-08-09">
<collectingDate id="E55FE9A3FFE68E18899AFE49A95BFE77" box="[151,288,403,429]" pageId="6" pageNumber="33" value="1997-08-09">9 VIII 1997</collectingDate>
</date>
, 
<collectionCode id="E7B4AE4EFFE68E188826FE49A946FE77" box="[299,317,403,429]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="33">T</collectionCode>
</materialsCitation>
. Kishimoto leg. (TKc); 
<materialsCitation id="31CD3CD6FFE68E188B40FE49A95BFE0B" collectingDate="2000-07-16" collectionCode="KUM" collectorName="K. Toyoda" country="Japan" location="Yoshimi-machi" pageId="6" pageNumber="33" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Saitama-ken">
[
<emphasis id="B3D1EA99FFE68E188B5BFE48AA96FE76" bold="true" box="[598,749,402,428]" pageId="6" pageNumber="33">
<collectingRegion id="4361F869FFE68E188B5BFE48AA96FE76" box="[598,749,402,428]" pageId="6" pageNumber="33">Saitama-ken</collectingRegion>
</emphasis>
]: 
<specimenCount id="97A3FD02FFE68E188A0BFE49AB2EFE77" box="[774,853,403,429]" pageId="6" pageNumber="33" type="male">1 male</specimenCount>
, 
<specimenCount id="97A3FD02FFE68E188A6FFE49ABBCFE77" box="[866,967,403,429]" pageId="6" pageNumber="33" type="female">1 female</specimenCount>
, 
<location id="847A6050FFE68E188ADFFE49ACFBFE77" LSID="urn:lsid:plazi:treatment:090C879DFFE28E19899AFA40AC27FEC6:847A6050FFE68E188ADFFE49ACFBFE77" box="[978,1152,403,429]" country="Japan" name="Yoshimi-machi" pageId="6" pageNumber="33" stateProvince="Saitama-ken">Yoshimi-machi</location>
, 
<date id="F51B104BFFE68E188D81FE49AD61FE77" box="[1164,1306,403,429]" pageId="6" pageNumber="33" value="2000-07-16">
<collectingDate id="E55FE9A3FFE68E188D81FE49AD61FE77" box="[1164,1306,403,429]" pageId="6" pageNumber="33" value="2000-07-16">16 VII 2000</collectingDate>
</date>
, 
<collectorName id="2C50535DFFE68E188C2BFE49ADE6FE77" box="[1318,1437,403,429]" pageId="6" pageNumber="33">K. Toyoda</collectorName>
leg. (
<collectionCode id="E7B4AE4EFFE68E1889DCFE6DA963FE0B" box="[209,280,439,465]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="6" pageNumber="33" type="Herbarium">KUM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="31CD3CD6FFE68E188820FE6DAB08FE0B" box="[301,883,439,465]" collectionCode="V" country="Japan" location="Chichibu-shi" municipality="Mitsumine" pageId="6" pageNumber="33" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Saitama-ken">
<specimenCount id="97A3FD02FFE68E188820FE6DA9F3FE0B" box="[301,392,439,465]" pageId="6" pageNumber="33" type="male">3 males</specimenCount>
, 
<specimenCount id="97A3FD02FFE68E188898FE6DAA7EFE0B" box="[405,517,439,465]" pageId="6" pageNumber="33" type="female">2 females</specimenCount>
, 
<collectingMunicipality id="617EACF1FFE68E188B1FFE6DAAF5FE0B" box="[530,654,439,465]" pageId="6" pageNumber="33">Mitsumine</collectingMunicipality>
, 
<location id="847A6050FFE68E188B96FE6DAB54FE0B" LSID="urn:lsid:plazi:treatment:090C879DFFE28E19899AFA40AC27FEC6:847A6050FFE68E188B96FE6DAB54FE0B" box="[667,815,439,465]" country="Japan" municipality="Mitsumine" name="Chichibu-shi" pageId="6" pageNumber="33" stateProvince="Saitama-ken">Chichibu-shi</location>
, 
<date id="F51B104BFFE68E188A36FE6DAB08FE0B" box="[827,883,439,465]" pageId="6" pageNumber="33" value="1997-05-28">
28 
<collectionCode id="E7B4AE4EFFE68E188A52FE6DAB08FE0B" box="[863,883,439,465]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="33" type="Museum">V</collectionCode>
</date>
</materialsCitation>
1997, 
<collectorName id="2C50535DFFE68E188ACEFE6DAC1EFE0B" box="[963,1125,439,465]" pageId="6" pageNumber="33">M. Maruyama</collectorName>
leg. (
<collectionCode id="E7B4AE4EFFE68E188DAAFE6DAC95FE0B" box="[1191,1262,439,465]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="6" pageNumber="33" type="Herbarium">KUM</collectionCode>
); 
<materialsCitation id="31CD3CD6FFE68E188C09FE6DA9E4FE2F" collectionCode="V" country="Japan" location="Noda-shi" pageId="6" pageNumber="33" specimenCount="1" specimenCount-male="1" stateProvince="Chiba-ken">
[
<emphasis id="B3D1EA99FFE68E188C00FE6CADF0FE0A" bold="true" box="[1293,1419,438,464]" pageId="6" pageNumber="33">
<collectingRegion id="4361F869FFE68E188C00FE6CADF0FE0A" box="[1293,1419,438,464]" pageId="6" pageNumber="33">Chiba-ken</collectingRegion>
</emphasis>
]: 
<specimenCount id="97A3FD02FFE68E18899AFE01A89DFE2F" box="[151,230,475,501]" pageId="6" pageNumber="33" type="male">1 male</specimenCount>
, 
<location id="847A6050FFE68E1889FFFE01A927FE2F" LSID="urn:lsid:plazi:treatment:090C879DFFE28E19899AFA40AC27FEC6:847A6050FFE68E1889FFFE01A927FE2F" box="[242,348,475,501]" country="Japan" name="Noda-shi" pageId="6" pageNumber="33" stateProvince="Chiba-ken">Noda-shi</location>
, 
<date id="F51B104BFFE68E188865FE01A9E4FE2F" box="[360,415,475,501]" pageId="6" pageNumber="33" value="1991-05-17">
17 
<collectionCode id="E7B4AE4EFFE68E188886FE01A9E4FE2F" box="[395,415,475,501]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="33" type="Museum">V</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="31CD3CD6FFE68E1888AAFE01A984FE2F" box="[423,511,475,501]" collectionCode="T" country="Japan" isTailEnumeration="true" location="Noda-shi" pageId="6" pageNumber="33" specimenCount="1" stateProvince="Chiba-ken">
1991, 
<collectionCode id="E7B4AE4EFFE68E1888E0FE01A984FE2F" box="[493,511,475,501]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="33">T</collectionCode>
</materialsCitation>
. Kishimoto leg. (TKc); 
<materialsCitation id="31CD3CD6FFE68E188A02FE01A9C2FDC3" collectingDate="2010-07-17" collectionCode="KUM" collectorName="J. Aoki" country="Japan" location="Naka-cho" municipality="Shikibidani" pageId="6" pageNumber="33" specimenCount="1" specimenCount-male="1" stateProvince="Tokushima-ken">
[
<emphasis id="B3D1EA99FFE68E188A15FE00ABA9FE2E" bold="true" box="[792,978,474,500]" pageId="6" pageNumber="33">
<collectingRegion id="4361F869FFE68E188A15FE00ABA9FE2E" box="[792,978,474,500]" pageId="6" pageNumber="33">Tokushima-ken</collectingRegion>
</emphasis>
]: 
<specimenCount id="97A3FD02FFE68E188AE6FE01AC41FE2F" box="[1003,1082,475,501]" pageId="6" pageNumber="33" type="male">1 male</specimenCount>
, 
<collectingMunicipality id="617EACF1FFE68E188D4BFE01ACB1FE2F" box="[1094,1226,475,501]" pageId="6" pageNumber="33">Shikibidani</collectingMunicipality>
, 
<location id="847A6050FFE68E188DDBFE01AD3DFE2F" LSID="urn:lsid:plazi:treatment:090C879DFFE28E19899AFA40AC27FEC6:847A6050FFE68E188DDBFE01AD3DFE2F" box="[1238,1350,475,501]" country="Japan" municipality="Shikibidani" name="Naka-cho" pageId="6" pageNumber="33" stateProvince="Tokushima-ken">Naka-chô</location>
, 
<date id="F51B104BFFE68E188C5FFE01A8A9FDC3" pageId="6" pageNumber="33" value="2010-07-17">
<collectingDate id="E55FE9A3FFE68E188C5FFE01A8A9FDC3" pageId="6" pageNumber="33" value="2010-07-17">17 VII 2010</collectingDate>
</date>
, 
<collectorName id="2C50535DFFE68E1889D0FE25A957FDC3" box="[221,300,511,537]" pageId="6" pageNumber="33">J. Aoki</collectorName>
leg. (
<collectionCode id="E7B4AE4EFFE68E188866FE25A9CAFDC3" box="[363,433,511,537]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="6" pageNumber="33" type="Herbarium">KUM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="31CD3CD6FFE68E1888C9FE25AA54FDE7" collectingDate="1962-04-29" collectingDateMax="1975-07-18" collectingDateMin="1962-04-29" collectionCode="EUM" collectorName="S. Hisamatsu &amp; A. Yonetsu" country="Japan" location="Komenono" pageId="6" pageNumber="33" specimenCount="1" specimenCount-male="1" stateProvince="Ehime-ken">
[
<emphasis id="B3D1EA99FFE68E1888C0FE24AA2BFDC2" bold="true" box="[461,592,510,536]" pageId="6" pageNumber="33">
<collectingRegion id="4361F869FFE68E1888C0FE24AA2BFDC2" box="[461,592,510,536]" pageId="6" pageNumber="33">Ehime-ken</collectingRegion>
</emphasis>
]: 
<specimenCount id="97A3FD02FFE68E188B65FE25AACEFDC3" box="[616,693,511,537]" pageId="6" pageNumber="33" type="male">1 male</specimenCount>
, Sara-ga-mine, 
<date id="F51B104BFFE68E188A68FE25AB9DFDC3" box="[869,998,511,537]" pageId="6" pageNumber="33" value="1962-04-29">
<collectingDate id="E55FE9A3FFE68E188A68FE25AB9DFDC3" box="[869,998,511,537]" pageId="6" pageNumber="33" value="1962-04-29">29 IV 1962</collectingDate>
</date>
, 
<collectorName id="2C50535DFFE68E188AFDFE25ACF8FDC3" box="[1008,1155,511,537]" pageId="6" pageNumber="33">S. Hisamatsu</collectorName>
leg. (
<collectionCode id="E7B4AE4EFFE68E188DCFFE25AD7DFDC3" box="[1218,1286,511,537]" pageId="6" pageNumber="33">EUM</collectionCode>
); 
<location id="847A6050FFE68E188C14FE25ADE3FDC3" LSID="urn:lsid:plazi:treatment:090C879DFFE28E19899AFA40AC27FEC6:847A6050FFE68E188C14FE25ADE3FDC3" box="[1305,1432,511,537]" country="Japan" name="Komenono" pageId="6" pageNumber="33" stateProvince="Ehime-ken">Komenono</location>
, 
<date id="F51B104BFFE68E18899AFDF9A95AFDE7" box="[151,289,546,573]" pageId="6" pageNumber="33" value="1975-07-18">
<collectingDate id="E55FE9A3FFE68E18899AFDF9A95AFDE7" box="[151,289,546,573]" pageId="6" pageNumber="33" value="1975-07-18">18 VII 1975</collectingDate>
</date>
, 
<collectorName id="2C50535DFFE68E188827FDF9A9DEFDE7" box="[298,421,547,573]" pageId="6" pageNumber="33">A. Yonetsu</collectorName>
leg. (
<collectionCode id="E7B4AE4EFFE68E1888EEFDF9AA5CFDE7" box="[483,551,547,573]" pageId="6" pageNumber="33">EUM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="31CD3CD6FFE68E188B37FDF9AB0DFDE7" box="[570,886,547,573]" collectionCode="VI" country="Japan" pageId="6" pageNumber="33" specimenCount="1" specimenCount-male="1" stateProvince="Ehime-ken">
<specimenCount id="97A3FD02FFE68E188B37FDF9AAFCFDE7" box="[570,647,547,573]" pageId="6" pageNumber="33" type="male">1 male</specimenCount>
, Sara-ga-mine, 
<date id="F51B104BFFE68E188A3BFDF9AB0DFDE7" box="[822,886,547,573]" pageId="6" pageNumber="33" value="1959-06-27">
27 
<collectionCode id="E7B4AE4EFFE68E188A55FDF9AB0DFDE7" box="[856,886,547,573]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="6" pageNumber="33">VI</collectionCode>
</date>
</materialsCitation>
1959, 
<collectorName id="2C50535DFFE68E188ACCFDF9AC62FDE7" box="[961,1049,547,573]" pageId="6" pageNumber="33">M. Satô</collectorName>
leg. (
<collectionCode id="E7B4AE4EFFE68E188D5BFDF9ACE1FDE7" box="[1110,1178,547,573]" pageId="6" pageNumber="33">EUM</collectionCode>
); 
<materialsCitation id="31CD3CD6FFE68E188DA0FDF9A9ACFDBB" collectionCode="V" country="Japan" location="Yusuhara-cho" pageId="6" pageNumber="33" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Kochi-ken">
[
<emphasis id="B3D1EA99FFE68E188DBAFDF8AD48FDE6" bold="true" box="[1207,1331,546,572]" pageId="6" pageNumber="33">
<collectingRegion id="4361F869FFE68E188DBAFDF8AD48FDE6" box="[1207,1331,546,572]" pageId="6" pageNumber="33">Kochi-ken</collectingRegion>
</emphasis>
]: 
<specimenCount id="97A3FD02FFE68E188C46FDF9ADE3FDE7" box="[1355,1432,547,573]" pageId="6" pageNumber="33" type="male">1 male</specimenCount>
, 
<specimenCount id="97A3FD02FFE68E18899AFD9DA887FDBB" box="[151,252,583,609]" pageId="6" pageNumber="33" type="female">1 female</specimenCount>
, 
<location id="847A6050FFE68E18880BFD9DA9D8FDBB" LSID="urn:lsid:plazi:treatment:090C879DFFE28E19899AFA40AC27FEC6:847A6050FFE68E18880BFD9DA9D8FDBB" box="[262,419,583,609]" country="Japan" name="Yusuhara-cho" pageId="6" pageNumber="33" stateProvince="Kochi-ken">Yusuhara-chô</location>
, 
<date id="F51B104BFFE68E1888A3FD9CA9ACFDBB" box="[430,471,582,609]" pageId="6" pageNumber="33" value="2010-05-05">
5 
<collectionCode id="E7B4AE4EFFE68E1888CEFD9DA9ACFDBB" box="[451,471,583,609]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="33" type="Museum">V</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="31CD3CD6FFE68E1888D3FD9DAA4DFDBB" box="[478,566,583,609]" collectionCode="T" country="Japan" isTailEnumeration="true" location="Yusuhara-cho" pageId="6" pageNumber="33" specimenCount="1" stateProvince="Kochi-ken">
2010, 
<collectionCode id="E7B4AE4EFFE68E188B29FD9DAA4DFDBB" box="[548,566,583,609]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="33">T</collectionCode>
</materialsCitation>
. &amp; 
<materialsCitation id="31CD3CD6FFE68E188B53FD9DAA0BFDBB" box="[606,624,583,609]" collectionCode="T" country="Japan" isTailEnumeration="true" location="Yusuhara-cho" pageId="6" pageNumber="33" specimenCount="1" stateProvince="Kochi-ken">
<collectionCode id="E7B4AE4EFFE68E188B53FD9DAA0BFDBB" box="[606,624,583,609]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="33">T</collectionCode>
</materialsCitation>
. 
<collectorName id="2C50535DFFE68E188B71FD9DAAB0FDBB" box="[636,715,583,609]" pageId="6" pageNumber="33">Miyata</collectorName>
leg. (
<collectionCode id="E7B4AE4EFFE68E188A06FD9DAB29FDBB" box="[779,850,583,609]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="6" pageNumber="33" type="Herbarium">KUM</collectionCode>
); 
<materialsCitation id="31CD3CD6FFE68E188A6BFD9DA8B4FD5F" collectionCode="V" country="Japan" location="Yusuhara-cho" municipality="Kamioriwatashi" pageId="6" pageNumber="33" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Kochi-ken">
<specimenCount id="97A3FD02FFE68E188A6BFD9DABC4FDBB" box="[870,959,583,609]" pageId="6" pageNumber="33" type="male">2 males</specimenCount>
, 
<specimenCount id="97A3FD02FFE68E188AC6FD9DAC54FDBB" box="[971,1071,583,609]" pageId="6" pageNumber="33" type="female">1 female</specimenCount>
, 
<collectingMunicipality id="617EACF1FFE68E188D36FD9DAC8AFDBB" box="[1083,1265,583,609]" pageId="6" pageNumber="33">Kamioriwatashi</collectingMunicipality>
, 
<location id="847A6050FFE68E188DF6FD9DADE3FDBB" LSID="urn:lsid:plazi:treatment:090C879DFFE28E19899AFA40AC27FEC6:847A6050FFE68E188DF6FD9DADE3FDBB" box="[1275,1432,583,609]" country="Japan" municipality="Kamioriwatashi" name="Yusuhara-cho" pageId="6" pageNumber="33" stateProvince="Kochi-ken">Yusuhara-chô</location>
, 
<date id="F51B104BFFE68E18899AFDB1A8B4FD5F" box="[151,207,619,645]" pageId="6" pageNumber="33" value="1997-05-24">
24 
<collectionCode id="E7B4AE4EFFE68E1889B6FDB1A8B4FD5F" box="[187,207,619,645]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="33" type="Museum">V</collectionCode>
</date>
</materialsCitation>
1997, 
<collectorName id="2C50535DFFE68E188813FDB1A9FFFD5F" box="[286,388,619,645]" pageId="6" pageNumber="33">M. Sakai</collectorName>
leg. (
<collectionCode id="E7B4AE4EFFE68E1888C8FDB1AA72FD5F" box="[453,521,619,645]" pageId="6" pageNumber="33">EUM</collectionCode>
); [
<emphasis id="B3D1EA99FFE68E188B2AFDB0AAB1FD5E" bold="true" box="[551,714,618,644]" pageId="6" pageNumber="33">
<collectingRegion id="4361F869FFE68E188B2AFDB0AAB1FD5E" box="[551,714,618,644]" pageId="6" pageNumber="33">Nagasaki-ken</collectingRegion>
</emphasis>
]: Tsushima Is.: 
<materialsCitation id="31CD3CD6FFE68E188A8CFDB1A8A3FD73" collectionCode="VI" country="Japan" location="Tsushima-shi" municipality="Kamiagatamachi Sasuna" pageId="6" pageNumber="33" specimenCount="1" specimenCount-male="1" stateProvince="Nagasaki-ken">
<specimenCount id="97A3FD02FFE68E188A8CFDB1ABABFD5F" box="[897,976,619,645]" pageId="6" pageNumber="33" type="male">1 male</specimenCount>
, 
<collectingMunicipality id="617EACF1FFE68E188AD0FDB1AC8FFD5F" box="[989,1268,619,645]" pageId="6" pageNumber="33">Kamiagatamachi Sasuna</collectingMunicipality>
, 
<location id="847A6050FFE68E188C0DFDB1ADE3FD5F" LSID="urn:lsid:plazi:treatment:090C879DFFE28E19899AFA40AC27FEC6:847A6050FFE68E188C0DFDB1ADE3FD5F" box="[1280,1432,619,645]" country="Japan" municipality="Kamiagatamachi Sasuna" name="Tsushima-shi" pageId="6" pageNumber="33" stateProvince="Nagasaki-ken">Tsushima-shi</location>
, 
<date id="F51B104BFFE68E18899AFD55A8A3FD73" box="[151,216,655,681]" pageId="6" pageNumber="33" value="2022-06-18">
18 
<collectionCode id="E7B4AE4EFFE68E1889B7FD55A8A3FD73" box="[186,216,655,681]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="6" pageNumber="33">VI</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="31CD3CD6FFE68E1889D3FD55A94DFD73" box="[222,310,655,681]" collectionCode="T" country="Japan" isTailEnumeration="true" location="Tsushima-shi" municipality="Kamiagatamachi Sasuna" pageId="6" pageNumber="33" specimenCount="1" stateProvince="Nagasaki-ken">
2022, 
<collectionCode id="E7B4AE4EFFE68E188829FD55A94DFD73" box="[292,310,655,681]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="33">T</collectionCode>
</materialsCitation>
. 
<collectorName id="2C50535DFFE68E18884CFD55A9C4FD73" box="[321,447,655,681]" pageId="6" pageNumber="33">Hashizume</collectorName>
leg. (
<collectionCode id="E7B4AE4EFFE68E1888F3FD55AA3EFD73" box="[510,581,655,681]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="6" pageNumber="33" type="Herbarium">KUM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C9BF6500FFE68E1889CAFD68ADE9F9D3" pageId="6" pageNumber="33" type="description">
<paragraph id="811A368BFFE68E1889CAFD68A9E8FCCF" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="33">
<emphasis id="B3D1EA99FFE68E1889CAFD68A90EFD16" bold="true" box="[199,373,690,716]" pageId="6" pageNumber="33">Redescription.</emphasis>
Measurements (n = 5): BL ≈ 1.86–2.39; FBL, 1.00–1.22; HL, 0.34–0.38; HW, 0.38–0.44; PL, 0.35–0.42; PW, 0.50–0.59; EL, 0.38–0.48; EW, 0.60–0.73. (
<typeStatus id="5E1E8829FFE68E188A35FD0DABD1FD2B" box="[824,938,727,753]" pageId="6" pageNumber="33">Lectotype</typeStatus>
of 
<taxonomicName id="46A54D08FFE68E188AC2FD0DAC9EFD2B" authority=": BL" authorityName="BL" box="[975,1253,727,753]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="33" phylum="Arthropoda" rank="species" species="crenulicauda">
<emphasis id="B3D1EA99FFE68E188AC2FD0DACC8FD2B" box="[975,1203,727,753]" italics="true" pageId="6" pageNumber="33">Atheta crenulicauda</emphasis>
: BL
</taxonomicName>
≈ 2.0; PL, 0.40; PW, 0.55; HTL, 0.41).
</paragraph>
<paragraph id="811A368BFFE68E1889CAFCC5ACDEFC87" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="33">Relative length of antennomeres I–XI (n = 1): 25: 24: 21: 11: 11: 11: 11: 10: 11: 12: 37. Ratio of length/width of antennomeres I–XI (n = 1): 1.84: 2.07: 2.16: 0.88: 0.71: 0.68: 0.64: 0.55: 0.57: 0.63: 1.83.</paragraph>
<paragraph id="811A368BFFE68E1889CAFCBDA94EFC7F" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="33">
Body (
<figureCitation id="199E2A0EFFE68E18881BFCBDA907FC5B" box="[278,380,871,897]" captionStart="FIGURE 1" captionStartId="3.[151,250,1944,1968]" captionTargetBox="[178,1424,196,1901]" captionTargetId="figure-21@3.[164,1424,179,1919]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Pseudatheta spp. A. Pseudatheta crenulicauda (Bernhauer, 1907), comb. nov.; B. Pseudatheta taiwanensis Pace, 2008; C. Pseudatheta hilaris (Sharp, 1888), comb. nov., lectotype of Oxypoda hiralis; D. Pseudatheta thailandensis Pace, 1992; E. Pseudatheta cooteri Pace, 1998; F. Pseudatheta elegans Cameron, 1920. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8182112" httpUri="https://zenodo.org/record/8182112/files/figure.png" pageId="6" pageNumber="33">Figs. 1A</figureCitation>
, 
<figureCitation id="199E2A0EFFE68E188886FCBDA9D7FC5B" box="[395,428,871,897]" captionStart="FIGURE 2" captionStartId="4.[151,250,1825,1849]" captionTargetBox="[151,1436,194,1800]" captionTargetId="figure-35@4.[151,1436,181,1800]" captionTargetPageId="4" captionText="FIGURE 2. Habitus of Phymatura spp. and type specimens examined. A. Phymatura japonica Cameron, 1933, stat. rev.; B. Phymatura russa Assing, 2021; C. lectotype of Atheta (Datomicra) crenulicauda Bernhauer, 1907 (=Pseudatheta crenulicauda comb. nov.); D. labels of C; E. holotype of Phymatura japonica Cameron, 1933, stat. rev. Scale bars: 1.0 mm for A–B." figureDoi="http://doi.org/10.5281/zenodo.8182114" httpUri="https://zenodo.org/record/8182114/files/figure.png" pageId="6" pageNumber="33">2C</figureCitation>
) reddish brown; head darker; posterolateral areas of elytra darker; abdominal segments V–VII darker.
</paragraph>
<paragraph id="811A368BFFE68E1889CAFC75AB80FBCB" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="33">Head almost as long as wide, HW/HL: 1.11–1.26; surface densely covered with setae.Antenna with antennomeres I–III and XI longer than wide, antennomeres IV–X wider than long; antennomeres I and II with rounded distal end, antennomeres III–X strongly angulated near distal end, antennomere XI oval.</paragraph>
<paragraph id="811A368BFFE68E1889CAFBC1ACAEFB33" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="33">Pronotum transverse, PW/PL: 1.36–1.50, PW/HW: 1.32–1.41; surface densely covered with setae, finely punctured, without microsculpture; posterior margin slightly bisinuate. Elytra wider than long, EW/EL: 1.60–1.70, EL/PL: 1.00–1.10, EW/PW: 1.20–1.24; surface densely covered with setae and finely punctured; posterior margin sinuate near posterolateral corners. Hind wings well developed; flabellum with one seta. Mesoventrite without longitudinal carina; mesoventral process extended to slightly beyond middle of mesocoxal cavities, with pointed apex; metaventral process shorter than mesoventral process, with rounded apex; isthmus present.</paragraph>
<paragraph id="811A368BFFE68E1889CAFB29AC71FAD7" blockId="6.[151,1437,151,2013]" box="[199,1034,1267,1293]" pageId="6" pageNumber="33">Abdomen slightly narrowed posteriad; surface densely covered with setae.</paragraph>
<paragraph id="811A368BFFE68E1889CAFACDAC12FA1B" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="33">
Male. Elytra with a pair of small tubercles at about posterior 1/3 on sutural margin. Sternite VI with small medial lobe on posterior margin. Tergite VII with a tubercle on posteromedian area. Tergite VIII (
<figureCitation id="199E2A0EFFE68E188DE0FAE1AD3EFA8F" box="[1261,1349,1338,1365]" captionStart="FIGURE 5" captionStartId="8.[151,250,1626,1650]" captionTargetBox="[161,1430,250,1592]" captionTargetId="figure-73@8.[151,1436,237,1603]" captionTargetPageId="8" captionText="FIGURE 5. Pseudatheta spp. A–C: Pseudatheta crenulicauda (Bernhauer, 1907), comb. nov., D–F: Pseudatheta taiwanensis Pace, 2008. A, D. male tergite VIII in dorsal view; B, E. median lobe of aedeagus in lateral view; C, F. spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8182126" httpUri="https://zenodo.org/record/8182126/files/figure.png" pageId="6" pageNumber="33">Fig. 5A</figureCitation>
) with a tubercle on median area; three to five processes on each side of posterior margin, outer ones slightly curved, longer and sharper than the others. Aedeagus as in 
<figureCitation id="199E2A0EFFE68E188B76FA59AAAAFA47" box="[635,721,1410,1437]" captionStart="FIGURE 5" captionStartId="8.[151,250,1626,1650]" captionTargetBox="[161,1430,250,1592]" captionTargetId="figure-73@8.[151,1436,237,1603]" captionTargetPageId="8" captionText="FIGURE 5. Pseudatheta spp. A–C: Pseudatheta crenulicauda (Bernhauer, 1907), comb. nov., D–F: Pseudatheta taiwanensis Pace, 2008. A, D. male tergite VIII in dorsal view; B, E. median lobe of aedeagus in lateral view; C, F. spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8182126" httpUri="https://zenodo.org/record/8182126/files/figure.png" pageId="6" pageNumber="33">Fig. 5B</figureCitation>
; median lobe with large basal bulb; apical process broad, dilated in lateral view, apical end thin and elongated, apex pointed; flagellum moderately long.
</paragraph>
<paragraph id="811A368BFFE68E1889CAFA11ADE9F9D3" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="33">
Female. Spermatheca (
<figureCitation id="199E2A0EFFE68E1888C1FA11AA5EFA3F" box="[460,549,1482,1509]" captionStart="FIGURE 5" captionStartId="8.[151,250,1626,1650]" captionTargetBox="[161,1430,250,1592]" captionTargetId="figure-73@8.[151,1436,237,1603]" captionTargetPageId="8" captionText="FIGURE 5. Pseudatheta spp. A–C: Pseudatheta crenulicauda (Bernhauer, 1907), comb. nov., D–F: Pseudatheta taiwanensis Pace, 2008. A, D. male tergite VIII in dorsal view; B, E. median lobe of aedeagus in lateral view; C, F. spermatheca. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.8182126" httpUri="https://zenodo.org/record/8182126/files/figure.png" pageId="6" pageNumber="33">Fig. 5C</figureCitation>
) curved twice, with a transverse band-like structure at base of distal portion; distal portion elongated, U-shaped; median portion elongated, slightly curved; proximal portion longer than wide.
</paragraph>
</subSubSection>
<subSubSection id="C9BF6500FFE68E1889CAF9C8ACC4F9F7" box="[199,1215,1554,1581]" pageId="6" pageNumber="33" type="distribution">
<paragraph id="811A368BFFE68E1889CAF9C8ACC4F9F7" blockId="6.[151,1437,151,2013]" box="[199,1215,1554,1581]" pageId="6" pageNumber="33">
<emphasis id="B3D1EA99FFE68E1889CAF9C8A91AF9F6" bold="true" box="[199,353,1554,1580]" pageId="6" pageNumber="33">Distribution.</emphasis>
<collectingCountry id="F9B2761BFFE68E18886AF9C9A9DCF9F7" box="[359,423,1555,1581]" name="Japan" pageId="6" pageNumber="33">Japan</collectingCountry>
(Honshu, Shikoku, Tsushima Is., Kyushu?); 
<collectingCountry id="F9B2761BFFE68E188A93F9C9AB9AF9F7" box="[926,993,1555,1581]" name="China" pageId="6" pageNumber="33">China</collectingCountry>
(
<collectingRegion id="4361F869FFE68E188AFDF9C9AC34F9F7" box="[1008,1103,1555,1581]" country="China" name="Sichuan" pageId="6" pageNumber="33">Sichuan</collectingRegion>
, 
<collectingRegion id="4361F869FFE68E188D54F9C9ACCFF9F7" box="[1113,1204,1555,1581]" country="China" name="Yunnan" pageId="6" pageNumber="33">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C9BF6500FFE68E1989CAF9EDAC27FEC6" lastPageId="7" lastPageNumber="34" pageId="6" pageNumber="33" type="discussion">
<paragraph id="811A368BFFE68E1989CAF9EDAC27FEC6" blockId="6.[151,1437,151,2013]" lastBlockId="7.[151,1436,151,285]" lastPageId="7" lastPageNumber="34" pageId="6" pageNumber="33">
<emphasis id="B3D1EA99FFE68E1889CAF9EDA940F98B" bold="true" box="[199,315,1591,1617]" pageId="6" pageNumber="33">Remarks.</emphasis>
<bibRefCitation id="E5344B7AFFE68E18884CF9EDA992F98B" author="Sawada, K." box="[321,489,1591,1617]" pageId="6" pageNumber="33" pagination="171 - 222" refId="ref16510" refString="Sawada, K. (1977) Studies on the genus Atheta Thomson and its allies (Coleoptera, Staphylinidae) III: Japanese species described by the previous authors. Contributions from the Biological Laboratory, Kyoto University, 25 (2), 171 - 222." type="journal article" year="1977">Sawada (1977)</bibRefCitation>
redescribed this species in detail as 
<taxonomicName id="46A54D08FFE68E188A76F9EDAC59F98B" authorityName="&quot;: Pace" authorityYear="1992" box="[891,1058,1591,1617]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="33" phylum="Arthropoda" rank="species" species="oligotinula">
<emphasis id="B3D1EA99FFE68E188A76F9EDAC59F98B" box="[891,1058,1591,1617]" italics="true" pageId="6" pageNumber="33">Ph. oligotinula</emphasis>
</taxonomicName>
. Based on its characteristics (e.g., small body size, absence of the longitudinal carina on the mesoventrite, and the shape of the median lobe of the aedeagus and the spermatheca), this species belongs to 
<taxonomicName id="46A54D08FFE68E188A02F9A5ABE2F943" authorityName="Cameron" authorityYear="1920" box="[783,921,1663,1689]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B3D1EA99FFE68E188A02F9A5ABE2F943" box="[783,921,1663,1689]" italics="true" pageId="6" pageNumber="33">Pseudatheta</emphasis>
</taxonomicName>
rather than to 
<taxonomicName id="46A54D08FFE68E188D4EF9A5ACC4F943" box="[1091,1215,1663,1689]" class="Insecta" family="Staphylinidae" genus="Phymatura" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B3D1EA99FFE68E188D4EF9A5ACC4F943" box="[1091,1215,1663,1689]" italics="true" pageId="6" pageNumber="33">Phymatura</emphasis>
</taxonomicName>
. The illustration of the median lobe of the aedeagus of 
<taxonomicName id="46A54D08FFE68E188B2FF979AAEFF967" authorityName="Pace" authorityYear="2010" box="[546,660,1699,1725]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="33" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B3D1EA99FFE68E188B2FF979AAEFF967" box="[546,660,1699,1725]" italics="true" pageId="6" pageNumber="33">Ps. similis</emphasis>
</taxonomicName>
, shown by 
<bibRefCitation id="E5344B7AFFE68E188A19F979ABE6F967" author="Pace, R." box="[788,925,1699,1725]" pageId="6" pageNumber="33" pagination="259 - 280" refId="ref16300" refString="Pace, R. (2010) Biodiversita delle Aleocharinae della Cina: Placusini, Homalotini, Bolitocharini, Eusteniamorphini e Falagriini 1 (Coleoptera, Staphylinidae). Beitrage zur entomologie, 60 (2), 259 - 280. https: // doi. org / 10.21248 / contrib. entomol. 60.2.259 - 280" type="journal article" year="2010">Pace (2010)</bibRefCitation>
, fully agrees with that of 
<taxonomicName id="46A54D08FFE68E188DB4F979AD0EF967" box="[1209,1397,1699,1725]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="33" phylum="Arthropoda" rank="species" species="crenulicauda">
<emphasis id="B3D1EA99FFE68E188DB4F979AD0EF967" box="[1209,1397,1699,1725]" italics="true" pageId="6" pageNumber="33">Ps. crenulicauda</emphasis>
</taxonomicName>
(as 
<taxonomicName id="46A54D08FFE68E18899AF91DA970F93B" authorityName="Pace" authorityYear="2010" box="[151,267,1735,1761]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="33" phylum="Arthropoda" rank="species" species="similis" status="syn. nov.">
<emphasis id="B3D1EA99FFE68E18899AF91DA970F93B" box="[151,267,1735,1761]" italics="true" pageId="6" pageNumber="33">Ps. similis</emphasis>
</taxonomicName>
<emphasis id="B3D1EA99FFE68E18881EF91DA900F93B" bold="true" box="[275,379,1735,1761]" pageId="6" pageNumber="33">
<taxonomicNameLabel id="A8E257E2FFE68E18881EF91DA900F93B" box="[275,379,1735,1761]" pageId="6" pageNumber="33">syn. nov.</taxonomicNameLabel>
</emphasis>
). 
<bibRefCitation id="E5344B7AFFE68E18889EF91DAA60F93B" author="Pace, R." box="[403,539,1735,1761]" pageId="6" pageNumber="33" pagination="295 - 336" refId="ref16439" refString="Pace, R. (2016) Aleocharinae della Cina al &quot; Naturkundemuseum &quot; di Erfurt (Insecta: Coleoptera: Staphyinidae). Veroffentlichungen des Naturkundemuseums Erfurt - VERNATE, 35, 295 - 336." type="journal article" year="2016">Pace (2016)</bibRefCitation>
recorded this species from Yunnan as 
<taxonomicName id="46A54D08FFE68E188AC2F91DAC38F93B" authorityName="Pace" authorityYear="2010" box="[975,1091,1735,1761]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="33" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B3D1EA99FFE68E188AC2F91DAC38F93B" box="[975,1091,1735,1761]" italics="true" pageId="6" pageNumber="33">Ps. similis</emphasis>
</taxonomicName>
. 
<bibRefCitation id="E5344B7AFFE68E188D5FF91DAD51F93B" author="Kim, Y. - H. &amp; Ahn, K. - J." box="[1106,1322,1735,1761]" pageId="6" pageNumber="33" refId="ref15503" refString="Kim, Y. - H. &amp; Ahn, K. - J. (2014) Insect Fauna of Korea. Vol. 12. No. 17. Arthropoda: Insecta; Coleoptera: Staphylinidae: Aleocharinae: Homalotini. National Institute of Biological Resources, Incheon, 113 pp." type="book" year="2014">Kim &amp; Ahn (2014)</bibRefCitation>
listed 
<taxonomicName id="46A54D08FFE68E188C7BF91DA951F8DF" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="33" phylum="Arthropoda" rank="species" species="crenulicauda">
<emphasis id="B3D1EA99FFE68E188C7BF91DA951F8DF" italics="true" pageId="6" pageNumber="33">Ph. crenulicauda</emphasis>
</taxonomicName>
as a Korean species, based on the record of “ 
<taxonomicName id="46A54D08FFE68E188A39F931ABBFF8DE" authorityName="Cameron" authorityYear="1933" box="[820,964,1771,1797]" class="Insecta" family="Staphylinidae" genus="Phymatura" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="33" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B3D1EA99FFE68E188A39F931ABBFF8DE" box="[820,964,1771,1797]" italics="true" pageId="6" pageNumber="33">Ph. japonica</emphasis>
</taxonomicName>
” from 
<collectingCountry id="F9B2761BFFE68E188D1AF931ACDEF8DF" box="[1047,1189,1771,1797]" name="North Korea" pageId="6" pageNumber="33">North Korea</collectingCountry>
in 
<bibRefCitation id="E5344B7AFFE68E188DC1F931AD10F8DF" author="Pasnik, G." box="[1228,1387,1771,1797]" pageId="6" pageNumber="33" pagination="185 - 234" refId="ref16476" refString="Pasnik, G. (2001) The North Korean Aleocharinae (Coleoptera, Staphylinidae): diversity and biogeography. Acta zoologica cracoviensia, 44 (3), 185 - 234." type="journal article" year="2001">Paśnik (2001)</bibRefCitation>
, but later found that the specimens examined in 
<bibRefCitation id="E5344B7AFFE68E188B70F8D5AB62F8F3" author="Pasnik, G." box="[637,793,1807,1833]" pageId="6" pageNumber="33" pagination="185 - 234" refId="ref16476" refString="Pasnik, G. (2001) The North Korean Aleocharinae (Coleoptera, Staphylinidae): diversity and biogeography. Acta zoologica cracoviensia, 44 (3), 185 - 234." type="journal article" year="2001">Paśnik (2001)</bibRefCitation>
were not 
<taxonomicName id="46A54D08FFE68E188A8AF8D5AD5EF8F3" authority="(Kim &amp; Ahn, 2016)" baseAuthorityName="Kim &amp; Ahn" baseAuthorityYear="2016" box="[903,1317,1807,1833]" class="Insecta" family="Staphylinidae" genus="Phymatura" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="33" phylum="Arthropoda" rank="species" species="crenuicauda">
<emphasis id="B3D1EA99FFE68E188A8AF8D5AC44F8F3" box="[903,1087,1807,1833]" italics="true" pageId="6" pageNumber="33">Ph. crenuicauda</emphasis>
(
<bibRefCitation id="E5344B7AFFE68E188D42F8D5AD66F8F3" author="Kim, Y. - H. &amp; Ahn, K. - J." box="[1103,1309,1807,1833]" pageId="6" pageNumber="33" pagination="189 - 196" refId="ref15557" refString="Kim, Y. - H. &amp; Ahn, K. - J. (2016) Taxonomy of the genus Phymatura J. Sahlberg (Coleoptera: Staphylinidae: Aleocharinae) in the Korean Peninsula. Zootaxa, 4061 (2), 189 - 196. https: // doi. org / 10.11646 / zootaxa. 4061.2.9" type="journal article" year="2016">Kim &amp; Ahn, 2016</bibRefCitation>
)
</taxonomicName>
. While the true species identity of these specimens is unknown, we tentatively consider this record as a record of 
<taxonomicName id="46A54D08FFE68E188C00F8E9ADE7F896" authorityName="Cameron" authorityYear="1933" box="[1293,1436,1843,1869]" class="Insecta" family="Staphylinidae" genus="Phymatura" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="33" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B3D1EA99FFE68E188C00F8E9ADE7F896" box="[1293,1436,1843,1869]" italics="true" pageId="6" pageNumber="33">Ph. japonica</emphasis>
</taxonomicName>
from 
<collectingCountry id="F9B2761BFFE68E1889D8F88DA91CF8AB" box="[213,359,1879,1905]" name="North Korea" pageId="6" pageNumber="33">North Korea</collectingCountry>
. This species can be distinguished from its congeners by the apically dilated apical process and the absence of a ventral process of the median lobe of the aedeagus. There is currently no practical key to distinguish the female of this species from the female of its allies [i.e., 
<taxonomicName id="46A54D08FFE68E188A3AF845ABD0F863" authorityName="Hashizume &amp; Yamamoto &amp; Maruyama" authorityYear="2023" baseAuthorityName="Sharp" baseAuthorityYear="1888" box="[823,939,1951,1977]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="33" phylum="Arthropoda" rank="species" species="hilaris">
<emphasis id="B3D1EA99FFE68E188A3AF845ABD0F863" box="[823,939,1951,1977]" italics="true" pageId="6" pageNumber="33">Ps. hilaris</emphasis>
</taxonomicName>
, 
<taxonomicName id="46A54D08FFE68E188AB7F845AC13F862" authorityName="Pace" authorityYear="2008" box="[954,1128,1951,1976]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="33" phylum="Arthropoda" rank="species" species="taiwanensis">
<emphasis id="B3D1EA99FFE68E188AB7F845AC13F862" box="[954,1128,1951,1976]" italics="true" pageId="6" pageNumber="33">Ps. taiwanensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="46A54D08FFE68E188D7AF845AD4CF863" authorityName="Pace" authorityYear="1992" box="[1143,1335,1951,1977]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="33" phylum="Arthropoda" rank="species" species="thailandensis">
<emphasis id="B3D1EA99FFE68E188D7AF845AD4CF863" box="[1143,1335,1951,1977]" italics="true" pageId="6" pageNumber="33">Ps. thailandensis</emphasis>
</taxonomicName>
, and 
<taxonomicName id="46A54D08FFE68E188C7BF845AA75F807" authority="(Bernhauer, 1915)" baseAuthorityName="Bernhauer" baseAuthorityYear="1915" class="Insecta" family="Staphylinidae" genus="Phymatura" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="33" phylum="Arthropoda" rank="species" species="bigranipennis">
<emphasis id="B3D1EA99FFE68E188C7BF845A948F807" italics="true" pageId="6" pageNumber="33">Ph. bigranipennis</emphasis>
(Bernhauer, 1915)
</taxonomicName>
]. Thus, despite our detailed morphological observations, these species could not be distinguished based on female specimens. The 
<typeStatus id="5E1E8829FFE78E198BB2FF4DAB2DFF6B" box="[703,854,151,177]" pageId="7" pageNumber="34">paralectotype</typeStatus>
of this species was collected from Unzen, Nagasaki, Kyushu, and is the only record of this species from Kyushu; however, because it was a female, it could not be positively identified. Further study is needed to allow morphologically based species identification in 
<taxonomicName id="46A54D08FFE78E198C1FFF05ADE7FF23" authorityName="Cameron" authorityYear="1920" box="[1298,1436,223,249]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B3D1EA99FFE78E198C1FFF05ADE7FF23" box="[1298,1436,223,249]" italics="true" pageId="7" pageNumber="34">Pseudatheta</emphasis>
</taxonomicName>
females, supported perhaps by DNA information (see also the remarks on 
<taxonomicName id="46A54D08FFE78E198ADBFED9AC37FEC7" authorityName="Hashizume &amp; Yamamoto &amp; Maruyama" authorityYear="2023" baseAuthorityName="Sharp" baseAuthorityYear="1888" box="[982,1100,259,285]" class="Insecta" family="Staphylinidae" genus="Pseudatheta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="34" phylum="Arthropoda" rank="species" species="hilaris">
<emphasis id="B3D1EA99FFE78E198ADBFED9AC37FEC7" box="[982,1100,259,285]" italics="true" pageId="7" pageNumber="34">Ph. hilaris</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>