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<document id="C7C175F0992FA7D618B35FD313649983" ID-CLB-Dataset="5915" ID-DOI="10.11646/zootaxa.3764.2.4" ID-GBIF-Dataset="b39bc65b-aa5a-4c56-a04a-3b0a28a5f039" ID-ISSN="1175-5326" ID-Zenodo-Dep="5045612" ID-ZooBank="60207805-8A9F-4C48-9DD7-27649E2A4BE6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623111510211" checkinUser="felipe" docAuthor="Castro, Peter &amp; Naruse, Tohru" docDate="2014" docId="C5176E384D3D577100C0FEECFE8AE78E" docLanguage="en" docName="zt03764p180.pdf" docOrigin="Zootaxa 3764 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Latreillopsis okala Castro &amp; Naruse 2014, n. sp." docType="treatment" docVersion="7" lastPageNumber="175" masterDocId="392E16404D3C57770057FFABFFFFE50B" masterDocTitle="New species of Latreillopsis Henderson, 1888 (Brachyura: Homolidae) and Neopalicus Moosa &amp; Serène, 1981 (Brachyura: Palicidae) from the Hawaiian Islands" masterLastPageNumber="180" masterPageNumber="169" pageNumber="170" updateTime="1699002576160" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="BBB87DCCF24A2E5806DEB40C0FB625FC">New species of Latreillopsis Henderson, 1888 (Brachyura: Homolidae) and Neopalicus Moosa &amp; Serène, 1981 (Brachyura: Palicidae) from the Hawaiian Islands</mods:title>
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<mods:namePart id="034D530C3FAF14CA36BAEFC48B1ED4DD">Castro, Peter</mods:namePart>
<mods:affiliation id="D89A9107670C69DB2DA1B440C1553413">Biological Sciences Department, California State Polytechnic University, Pomona, California 91768 - 4032, USA.</mods:affiliation>
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<mods:namePart id="FA57CC6666E27F8B6332C89BC0275A72">Naruse, Tohru</mods:namePart>
<mods:affiliation id="A95A274206CE4C4C07A54AD87E3F0C18">Tropical Biosphere Research Center, Iriomote Station, University of the Ryukyus, 870 Uehara, Taketomi, Okinawa 907 - 1541, Japan. E-mail: naruse @ lab. u-ryukyu. ac. jp</mods:affiliation>
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<mods:date id="3FE236D1316D5EA4E8C02F6CD6E42A8A">2014</mods:date>
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<treatment id="C5176E384D3D577100C0FEECFE8AE78E" ID-DOI="http://doi.org/10.5281/zenodo.5045492" ID-GBIF-Taxon="183227355" ID-Zenodo-Dep="5045492" LSID="urn:lsid:plazi:treatment:C5176E384D3D577100C0FEECFE8AE78E" httpUri="http://treatment.plazi.org/id/C5176E384D3D577100C0FEECFE8AE78E" lastPageId="6" lastPageNumber="175" pageId="1" pageNumber="170">
<subSubSection id="05A48CA54D3D577600C0FEECFE3FE469" box="[151,448,327,354]" pageId="1" pageNumber="170" type="nomenclature">
<paragraph id="4D01DF2E4D3D577600C0FEECFE3FE469" blockId="1.[151,448,327,388]" box="[151,448,327,354]" pageId="1" pageNumber="170">
<heading id="164968424D3D577600C0FEECFE3FE469" bold="true" box="[151,448,327,354]" fontSize="11" level="1" pageId="1" pageNumber="170" reason="1">
<emphasis id="7FCA033C4D3D577600C0FEECFE3FE469" bold="true" box="[151,448,327,354]" pageId="1" pageNumber="170">
<taxonomicName id="8ABEA4AD4D3D577600C0FEECFE88E46A" ID-CoL="6P4HQ" authority="Castro &amp; Naruse, 2014" authorityName="Castro &amp; Naruse" authorityYear="2014" box="[151,375,327,353]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="170" phylum="Arthropoda" rank="species" species="okala" status="sp. nov.">
<emphasis id="7FCA033C4D3D577600C0FEECFE88E46A" bold="true" box="[151,375,327,353]" italics="true" pageId="1" pageNumber="170">Latreillopsis okala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="64F9BE474D3D57760129FEE3FE3FE469" box="[382,448,328,354]" pageId="1" pageNumber="170" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="05A48CA54D3D577600C0FEC0FEEEE48F" box="[151,273,363,388]" pageId="1" pageNumber="170" type="description">
<paragraph id="4D01DF2E4D3D577600C0FEC0FEEEE48F" blockId="1.[151,448,327,388]" box="[151,273,363,388]" pageId="1" pageNumber="170">
(
<figureCitation id="D585C3AB4D3D577600C8FEC0FEF7E48F" box="[159,264,363,388]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,249,1983,2005]" captionStartId-1="4.[151,249,1280,1302]" captionStartId-2="5.[151,249,1877,1899]" captionTargetBox-0="[164,1423,298,1961]" captionTargetBox-1="[151,1436,193,1259]" captionTargetBox-2="[180,1414,213,1841]" captionTargetId-0="figure-95@3.[164,1423,298,1961]" captionTargetId-1="figure-387@4.[151,1436,193,1259]" captionTargetId-2="figure-71@5.[172,1414,193,1856]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Latreillopsis okala n. sp. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, habitus, dorsal view; b, color in life (photograph by P. Fiene)." captionText-1="FIGURE 2. Latreillopsis okala n. sp. species. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, cephalothorax, ventral view; b, outer surface of propodus and dactylus, outer view." captionText-2="FIGURE 3. Latreillopsis okala n. sp. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, cephalothorax, dorsal view (wartlike granules are not drawn); b, left anterior part of ventral surface of cephalothorax (wart-like granules are not drawn); c, left G1, ventral view; d, left G2, ventral view. Scales: ad = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5045614" figureDoi-1="http://doi.org/10.5281/zenodo.5045616" figureDoi-2="http://doi.org/10.5281/zenodo.5045618" httpUri-0="https://zenodo.org/record/5045614/files/figure.png" httpUri-1="https://zenodo.org/record/5045616/files/figure.png" httpUri-2="https://zenodo.org/record/5045618/files/figure.png" pageId="1" pageNumber="170">Figs. 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="05A48CA54D3D577600C0FE1AFD38E705" pageId="1" pageNumber="170" type="reference_group">
<paragraph id="4D01DF2E4D3D577600C0FE1AFCB6E4CC" blockId="1.[151,876,433,526]" box="[151,841,433,455]" pageId="1" pageNumber="170">
<treatmentCitationGroup id="6DAEF8004D3D577600C0FE1AFCB6E4CC" box="[151,841,433,455]" pageId="1" pageNumber="170">
<taxonomicName id="8ABEA4AD4D3D577600F5FE1AFE33E4CC" authorityName="Kubo" authorityYear="1936" box="[162,460,433,455]" class="Malacostraca" family="Homolidae" genus="Homola" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="170" phylum="Arthropoda" rank="species" species="majora" subGenus="Parhomola">
<emphasis id="7FCA033C4D3D577600F5FE1AFF0BE4CC" box="[162,244,433,455]" italics="true" pageId="1" pageNumber="170">Homola</emphasis>
(
<emphasis id="7FCA033C4D3D57760155FE1AFE8BE4CC" box="[258,372,433,455]" italics="true" pageId="1" pageNumber="170">Parhomola</emphasis>
)
<emphasis id="7FCA033C4D3D577601D4FE19FE33E4CC" box="[387,460,434,455]" italics="true" pageId="1" pageNumber="170">majora</emphasis>
</taxonomicName>
?” —
<treatmentCitation id="CC1FF93F4D3D57760259FE1AFD08E4CC" author="Edmondson, C. H." box="[526,759,433,455]" page="202" pageId="1" pageNumber="170" year="1951">
<bibRefCitation id="292FA2DF4D3D57760259FE1AFD08E4CC" author="Edmondson, C. H." box="[526,759,433,455]" pageId="1" pageNumber="170" pagination="183 - 243" refId="ref6433" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of Bernice P. Bishop Museum, 20 (13), 183 - 243." type="journal article" year="1951">Edmondson 1951: 202</bibRefCitation>
</treatmentCitation>
, fig. 10.
</treatmentCitationGroup>
</paragraph>
<paragraph id="4D01DF2E4D3D577600C0FE7FFC93E4E1" blockId="1.[151,876,433,526]" box="[151,876,468,490]" pageId="1" pageNumber="170">
<emphasis id="7FCA033C4D3D577600C0FE7FFF5AE4E1" box="[151,165,468,490]" italics="true" pageId="1" pageNumber="170"></emphasis>
?
<emphasis id="7FCA033C4D3D577600E1FE7FFE40E4E1" box="[182,447,468,490]" italics="true" pageId="1" pageNumber="170">
<taxonomicName id="8ABEA4AD4D3D577600E1FE7FFE47E4E1" ID-CoL="3SLTX" authorityName="Guinot &amp; Richer de Forges" authorityYear="1995" box="[182,440,468,490]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="170" phylum="Arthropoda" rank="species" species="cornuta">Latreillopsis aff. cornuta</taxonomicName>
</emphasis>
<bibRefCitation id="292FA2DF4D3D577601B1FE7FFC97E4E1" author="Guinot, D. &amp; Richer de Forges, B." box="[486,872,468,490]" pageId="1" pageNumber="170" pagination="283 - 517" refId="ref6581" refString="Guinot, D. &amp; Richer de Forges, B. (1995) Crustacea Decapoda Brachyura: Revision de la famille des Homolidae de Haan, 1839. In: Crosnier, A. (Ed.), Resultats des campagnes MUSORSTOM, vol. 13. Memoires du Museum national d'Histoire naturelle (Paris), 163, pp. 283 - 517." type="journal article" year="1995">Guinot &amp; Richer de Forges 1995: 414</bibRefCitation>
.
</paragraph>
<paragraph id="4D01DF2E4D3D577600C0FE53FD38E705" blockId="1.[151,876,433,526]" box="[151,711,504,526]" pageId="1" pageNumber="170">
“ (?)
<taxonomicName id="8ABEA4AD4D3D5776009CFE53FE60E705" ID-CoL="3SLTX" authorityName="Guinot &amp; Richer de Forges" authorityYear="1995" box="[203,415,504,526]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="170" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="7FCA033C4D3D5776009CFE53FE60E705" box="[203,415,504,526]" italics="true" pageId="1" pageNumber="170">Latreillopsis cornuta</emphasis>
</taxonomicName>
” —
<bibRefCitation id="292FA2DF4D3D57760187FE53FD8FE705" author="Castro, P." box="[464,624,504,526]" pageId="1" pageNumber="170" pagination="1 - 154" refId="ref6255" refString="Castro, P. (2011) Catalog of the anomuran and brachyuran crabs (Crustacea: Decapoda: Anomura, Brachyura) of the Hawaiian Islands. Zootaxa, 2947, 1 - 154." type="journal article" year="2011">Castro 2011: 35</bibRefCitation>
[in list].
</paragraph>
</subSubSection>
<subSubSection id="05A48CA54D3D577600C0FD95FB69E771" pageId="1" pageNumber="170" type="materials_examined">
<paragraph id="4D01DF2E4D3D577600C0FD95FB69E771" blockId="1.[151,1436,574,2011]" pageId="1" pageNumber="170">
<emphasis id="7FCA033C4D3D577600C0FD95FE7DE75C" bold="true" box="[151,386,574,599]" inLineHeading="true" pageId="1" pageNumber="170" reason="1">Material examined.</emphasis>
<materialsCitation id="FDD6D5734D3D577601DBFD95FB69E771" ID-GBIF-Occurrence="3312226301" collectingDate="2112-10-26" collectionCode="USNM" collectorName="Mike Severns &amp; Shirley Speer" elevation="91" location="southwestern Maui" pageId="1" pageNumber="170" specimenCode="USNM 1220315" specimenCount="1" typeStatus="holotype">
<typeStatus id="9205618C4D3D577601DBFD95FE0BE75C" box="[396,500,574,599]" pageId="1" pageNumber="170" type="holotype">Holotype</typeStatus>
male 6.0 mm [5.4 mm not including rostrum] × 4.0 mm, off
<location id="486189F54D3D577604EAFD95FA67E75C" LSID="urn:lsid:plazi:treatment:C5176E384D3D577100C0FEECFE8AE78E:486189F54D3D577604EAFD95FA67E75C" box="[1213,1432,574,599]" name="southwestern Maui" pageId="1" pageNumber="170">southwestern Maui</location>
, dredged,
<elevation id="C693381D4D3D577600A8FDCAFEC7E772" box="[255,312,609,634]" metricMagnitude="1" metricUnit="m" metricValue="9.1" pageId="1" pageNumber="170" unit="m" value="91.0">91 m</elevation>
(
<elevation id="C693381D4D3D57760111FDCAFE71E771" box="[326,398,609,634]" metricMagnitude="1" metricUnit="m" metricValue="9.144" pageId="1" pageNumber="170" unit="ft" value="300.0">300 ft</elevation>
),
<collectorName id="E04BBAF84D3D577601F5FDCAFDC3E771" box="[418,572,609,634]" pageId="1" pageNumber="170">Mike Severns</collectorName>
&amp;
<collectorName id="E04BBAF84D3D57760237FDCAFD07E771" box="[608,760,609,634]" pageId="1" pageNumber="170">Shirley Speer</collectorName>
coll.,
<date id="3900F9EE4D3D57760369FDCAFC45E771" box="[830,954,609,634]" pageId="1" pageNumber="170" value="2112-10-26">
<collectingDate id="294400064D3D57760369FDCAFC45E771" box="[830,954,609,634]" pageId="1" pageNumber="170" value="2112-10-26">26.10.2112</collectingDate>
</date>
(
<specimenCode id="1D1877554D3D5776039EFDCAFB75E771" box="[969,1162,609,634]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="170" type="Museum">USNM 1220315</specimenCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="05A48CA54D3D577600C0FD2FFEEDE37C" pageId="1" pageNumber="170" type="description">
<paragraph id="4D01DF2E4D3D577600C0FD2FFE6AE69F" blockId="1.[151,1436,574,2011]" pageId="1" pageNumber="170">
<emphasis id="7FCA033C4D3D577600C0FD2FFE07E796" bold="true" box="[151,504,644,669]" inLineHeading="true" pageId="1" pageNumber="170" reason="1">Description of male holotype.</emphasis>
Carapace (
<figureCitation id="D585C3AB4D3D5776022EFD2FFD43E796" box="[633,700,644,669]" captionStart="FIGURE 1" captionStartId="3.[151,249,1983,2005]" captionTargetBox="[164,1423,298,1961]" captionTargetId="figure-95@3.[164,1423,298,1961]" captionTargetPageId="3" captionText="FIGURE 1. Latreillopsis okala n. sp. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, habitus, dorsal view; b, color in life (photograph by P. Fiene)." figureDoi="http://doi.org/10.5281/zenodo.5045614" httpUri="https://zenodo.org/record/5045614/files/figure.png" pageId="1" pageNumber="170">Fig. 1</figureCitation>
,
<figureCitation id="D585C3AB4D3D5776029CFD2FFD19E796" box="[715,742,644,669]" captionStart="FIGURE 3" captionStartId="5.[151,249,1877,1899]" captionTargetBox="[180,1414,213,1841]" captionTargetId="figure-71@5.[172,1414,193,1856]" captionTargetPageId="5" captionText="FIGURE 3. Latreillopsis okala n. sp. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, cephalothorax, dorsal view (wartlike granules are not drawn); b, left anterior part of ventral surface of cephalothorax (wart-like granules are not drawn); c, left G1, ventral view; d, left G2, ventral view. Scales: ad = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5045618" httpUri="https://zenodo.org/record/5045618/files/figure.png" pageId="1" pageNumber="170">3a</figureCitation>
) longitudinally rectangular, longer than wide (cl/cw = 1.5, cl [not including rostrum]/cw = 1.3); small size; dorsal regions strongly defined, granular. Mesogastric region elevated, surmounted by large median boss, 2 protogastric bosses. Cardiac region with high, wide boss. Anterior branchial region slightly inflated, with blunt tubercle. Subhepatic region (
<figureCitation id="D585C3AB4D3D577603B3FD45FBC8E60C" box="[996,1079,750,775]" captionStart="FIGURE 3" captionStartId="5.[151,249,1877,1899]" captionTargetBox="[180,1414,213,1841]" captionTargetId="figure-71@5.[172,1414,193,1856]" captionTargetPageId="5" captionText="FIGURE 3. Latreillopsis okala n. sp. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, cephalothorax, dorsal view (wartlike granules are not drawn); b, left anterior part of ventral surface of cephalothorax (wart-like granules are not drawn); c, left G1, ventral view; d, left G2, ventral view. Scales: ad = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5045618" httpUri="https://zenodo.org/record/5045618/files/figure.png" pageId="1" pageNumber="170">Fig. 3b</figureCitation>
) slightly inflated, with 3 teeth, anterior tooth strongest, anteriorly directed; second, third teeth small, laterally directed. Rostrum proportionally short, straight. Pseudorostral spines long, curved, slightly longer than rostrum, outer margins with several rounded tubercles plus short, tooth-like tubercle near base of spine. Posterolateral border with 2 small, truncate teeth, better developed on left side.
</paragraph>
<paragraph id="4D01DF2E4D3D57760090FC35FD91E6F6" blockId="1.[151,1436,574,2011]" pageId="1" pageNumber="170">Antennular, antennal articles bordered by short, rounded granules. Ocular peduncles (basophthalmite) long, thin, bordered by short, rounded granules; podophthalmite broad, bordered by few rounded granules. Cornea of eyes proportionally large, well developed.</paragraph>
<paragraph id="4D01DF2E4D3D57760090FBACFBD5E14F" blockId="1.[151,1436,574,2011]" pageId="1" pageNumber="170">
Third maxilliped (
<figureCitation id="D585C3AB4D3D577601CFFBACFE14E12B" box="[408,491,1031,1056]" captionStart="FIGURE 3" captionStartId="5.[151,249,1877,1899]" captionTargetBox="[180,1414,213,1841]" captionTargetId="figure-71@5.[172,1414,193,1856]" captionTargetPageId="5" captionText="FIGURE 3. Latreillopsis okala n. sp. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, cephalothorax, dorsal view (wartlike granules are not drawn); b, left anterior part of ventral surface of cephalothorax (wart-like granules are not drawn); c, left G1, ventral view; d, left G2, ventral view. Scales: ad = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5045618" httpUri="https://zenodo.org/record/5045618/files/figure.png" pageId="1" pageNumber="170">Fig. 3b</figureCitation>
) merus with large, rounded anterior tubercle, sharp but not pointed anteroexternal angle; ischium narrow, about as long as merus, smooth except few short granules.
</paragraph>
<paragraph id="4D01DF2E4D3D57760090FBE5FC3FE1A6" blockId="1.[151,1436,574,2011]" pageId="1" pageNumber="170">
Chelipeds (P1) (
<figureCitation id="D585C3AB4D3D577601D0FBE5FE30E16C" box="[391,463,1102,1127]" captionStart="FIGURE 2" captionStartId="4.[151,249,1280,1302]" captionTargetBox="[151,1436,193,1259]" captionTargetId="figure-387@4.[151,1436,193,1259]" captionTargetPageId="4" captionText="FIGURE 2. Latreillopsis okala n. sp. species. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, cephalothorax, ventral view; b, outer surface of propodus and dactylus, outer view." figureDoi="http://doi.org/10.5281/zenodo.5045616" httpUri="https://zenodo.org/record/5045616/files/figure.png" pageId="1" pageNumber="170">Fig. 2</figureCitation>
) long, slender, conspicuous rounded granules, spatulate setae along margins of all articles; large triangular tooth on proximal margin of left dactylus; large dark spot proximal to pollex; propodus enlarged distally; merus slightly larger than other articles, spines absent.
</paragraph>
<paragraph id="4D01DF2E4D3D57760090FB1CFACFE056" blockId="1.[151,1436,574,2011]" pageId="1" pageNumber="170">
Ambulatory legs (P2, P3 missing in
<typeStatus id="9205618C4D3D57760208FB1CFD3CE1DB" box="[607,707,1207,1232]" pageId="1" pageNumber="170" type="holotype">holotype</typeStatus>
) (
<figureCitation id="D585C3AB4D3D5776028DFB1CFCE4E1DB" box="[730,795,1207,1232]" captionStart="FIGURE 1" captionStartId="3.[151,249,1983,2005]" captionTargetBox="[164,1423,298,1961]" captionTargetId="figure-95@3.[164,1423,298,1961]" captionTargetPageId="3" captionText="FIGURE 1. Latreillopsis okala n. sp. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, habitus, dorsal view; b, color in life (photograph by P. Fiene)." figureDoi="http://doi.org/10.5281/zenodo.5045614" httpUri="https://zenodo.org/record/5045614/files/figure.png" pageId="1" pageNumber="170">Fig. 1</figureCitation>
) P4 merus longer, wider than other articles, conspicuous rounded granules along margins of all articles; dorsally-oriented distal tooth on merus; dactyli with row of sharp spines on ventral margins. P5 reaching base of pseudorostral spine when folded; with conspicuous rounded granules along margins of merus, carpus; propodus slightly curved, with short granules only on dorsal margin; 2 long, acute, at tandem, proximal teeth; 2 small, parallel, acute teeth distal to large teeth; dactylus smooth.
</paragraph>
<paragraph id="4D01DF2E4D3D57760090FACCFA81E0E1" blockId="1.[151,1436,574,2011]" pageId="1" pageNumber="170">
Male abdomen (
<figureCitation id="D585C3AB4D3D5776012AFACCFE32E08B" box="[381,461,1383,1408]" captionStart="FIGURE 2" captionStartId="4.[151,249,1280,1302]" captionTargetBox="[151,1436,193,1259]" captionTargetId="figure-387@4.[151,1436,193,1259]" captionTargetPageId="4" captionText="FIGURE 2. Latreillopsis okala n. sp. species. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, cephalothorax, ventral view; b, outer surface of propodus and dactylus, outer view." figureDoi="http://doi.org/10.5281/zenodo.5045616" httpUri="https://zenodo.org/record/5045616/files/figure.png" pageId="1" pageNumber="170">Fig. 2a</figureCitation>
) wide, all 6 somites freely articulating plus telson; homoloid press-button system, with prominence on abdominal sternite 4 locking with socket on abdominal somite 6. G1 (
<figureCitation id="D585C3AB4D3D57760405FA20FB5EE0AF" box="[1106,1185,1419,1444]" captionStart="FIGURE 3" captionStartId="5.[151,249,1877,1899]" captionTargetBox="[180,1414,213,1841]" captionTargetId="figure-71@5.[172,1414,193,1856]" captionTargetPageId="5" captionText="FIGURE 3. Latreillopsis okala n. sp. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, cephalothorax, dorsal view (wartlike granules are not drawn); b, left anterior part of ventral surface of cephalothorax (wart-like granules are not drawn); c, left G1, ventral view; d, left G2, ventral view. Scales: ad = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5045618" httpUri="https://zenodo.org/record/5045618/files/figure.png" pageId="1" pageNumber="170">Fig. 3c</figureCitation>
) triangular, with short basal part, tapering distally; lateral margin slightly rimmed in ventral view, mesial part folded onto ventral surface over proximal three-fourths of triangular part. G2 (
<figureCitation id="D585C3AB4D3D57760287FA7AFCDDE0E1" box="[720,802,1489,1514]" captionStart="FIGURE 3" captionStartId="5.[151,249,1877,1899]" captionTargetBox="[180,1414,213,1841]" captionTargetId="figure-71@5.[172,1414,193,1856]" captionTargetPageId="5" captionText="FIGURE 3. Latreillopsis okala n. sp. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, cephalothorax, dorsal view (wartlike granules are not drawn); b, left anterior part of ventral surface of cephalothorax (wart-like granules are not drawn); c, left G1, ventral view; d, left G2, ventral view. Scales: ad = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5045618" httpUri="https://zenodo.org/record/5045618/files/figure.png" pageId="1" pageNumber="170">Fig. 3d</figureCitation>
) with truncated distal part, U-shaped in
<emphasis id="7FCA033C4D3D577604B2FA7AFA81E0E1" bold="true" box="[1253,1406,1489,1514]" pageId="1" pageNumber="170">cross-section ̇</emphasis>
</paragraph>
<paragraph id="4D01DF2E4D3D57760090FA5FFC4CE306" blockId="1.[151,1436,574,2011]" box="[199,947,1524,1549]" pageId="1" pageNumber="170">
Complete abdomen (somite 6, telson in
<bibRefCitation id="292FA2DF4D3D577602D1FA5FFCB0E306" author="Edmondson, C. H." box="[646,847,1524,1549]" pageId="1" pageNumber="170" pagination="183 - 243" refId="ref6433" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of Bernice P. Bishop Museum, 20 (13), 183 - 243." type="journal article" year="1951">Edmondson 1951</bibRefCitation>
: fig. 10).
</paragraph>
<paragraph id="4D01DF2E4D3D57760090F9BCFE6FE33B" blockId="1.[151,1436,574,2011]" box="[199,400,1559,1584]" pageId="1" pageNumber="170">Female unknown.</paragraph>
<paragraph id="4D01DF2E4D3D57760090F990FEEDE37C" blockId="1.[151,1436,574,2011]" pageId="1" pageNumber="170">
<emphasis id="7FCA033C4D3D57760090F990FEEFE35F" bold="true" box="[199,272,1595,1620]" inLineHeading="true" pageId="1" pageNumber="170" reason="1">Color.</emphasis>
A photograph of the live
<typeStatus id="9205618C4D3D5776021FF990FD55E35F" box="[584,682,1595,1620]" pageId="1" pageNumber="170" type="holotype">holotype</typeStatus>
(
<figureCitation id="D585C3AB4D3D577602EAF990FCEEE35F" box="[701,785,1595,1620]" captionStart="FIGURE 1" captionStartId="3.[151,249,1983,2005]" captionTargetBox="[164,1423,298,1961]" captionTargetId="figure-95@3.[164,1423,298,1961]" captionTargetPageId="3" captionText="FIGURE 1. Latreillopsis okala n. sp. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, habitus, dorsal view; b, color in life (photograph by P. Fiene)." figureDoi="http://doi.org/10.5281/zenodo.5045614" httpUri="https://zenodo.org/record/5045614/files/figure.png" pageId="1" pageNumber="170">Fig. 1b</figureCitation>
) shows bright orange carapace and eyes, orange-brown pereopods.
</paragraph>
</subSubSection>
<subSubSection id="05A48CA54D3D57760090F92FFB5FE3CA" pageId="1" pageNumber="170" type="etymology">
<paragraph id="4D01DF2E4D3D57760090F92FFB5FE3CA" blockId="1.[151,1436,574,2011]" pageId="1" pageNumber="170">
<emphasis id="7FCA033C4D3D57760090F92FFEB1E396" bold="true" box="[199,334,1668,1693]" inLineHeading="true" pageId="1" pageNumber="170" reason="1">Etymology.</emphasis>
From
<emphasis id="7FCA033C4D3D577601F0F92AFE52E395" box="[423,429,1665,1694]" italics="true" pageId="1" pageNumber="170">̍</emphasis>
<taxonomicName id="8ABEA4AD4D3D577601FAF92FFD94E396" authority=", Hawaiian" authorityName="Hawaiian" box="[429,619,1668,1693]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="170" phylum="Arthropoda" rank="species" species="okala">
<emphasis id="7FCA033C4D3D577601FAF92FFE14E397" box="[429,491,1668,1692]" italics="true" pageId="1" pageNumber="170">ōkala</emphasis>
, Hawaiian
</taxonomicName>
for “rough” or “coarse,” in reference to the granular carapace and pereopods diagnostic for the new species. The name is treated as a Latin noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="05A48CA54D3D57740090F967FB54E5FC" lastPageId="3" lastPageNumber="172" pageId="1" pageNumber="170" type="discussion">
<paragraph id="4D01DF2E4D3D57750090F967FE1AE411" blockId="1.[151,1436,574,2011]" lastBlockId="2.[151,1437,151,2007]" lastPageId="2" lastPageNumber="171" pageId="1" pageNumber="170">
<emphasis id="7FCA033C4D3D57760090F967FEC4E3EE" bold="true" box="[199,315,1740,1765]" inLineHeading="true" pageId="1" pageNumber="170" reason="1">Remarks.</emphasis>
<bibRefCitation id="292FA2DF4D3D57760115F967FDACE3EE" author="Edmondson, C. H." box="[322,595,1740,1765]" pageId="1" pageNumber="170" pagination="183 - 243" refId="ref6433" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of Bernice P. Bishop Museum, 20 (13), 183 - 243." type="journal article" year="1951">Edmondson (1951: 203)</bibRefCitation>
examined two small specimens (female cl
<quantity id="8A4672CB4D3D5776046DF967FB82E3E8" box="[1082,1149,1740,1765]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="170" unit="mm" value="8.0">8 mm</quantity>
, second female specimen “of about the same size”) dredged from
<quantity id="8A4672CB4D3D57760225F944FD48E203" box="[626,695,1775,1800]" metricMagnitude="1" metricUnit="m" metricValue="3.6576" pageId="1" pageNumber="170" unit="ft" value="120.0">120 ft</quantity>
(
<quantity id="8A4672CB4D3D5776029DF944FCE1E203" box="[714,798,1775,1800]" metricMagnitude="1" metricUnit="m" metricValue="3.66" pageId="1" pageNumber="170" unit="m" value="36.6">36.6 m</quantity>
) off Oahu, Hawaii, and identified them as “probable juveniles of
<taxonomicName id="8ABEA4AD4D3D57760173F8B9FCFFE220" authority="Kubo, 1936" authorityName="Kubo" authorityYear="1936" box="[292,768,1810,1835]" class="Malacostraca" family="Homolidae" genus="Homola" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="170" phylum="Arthropoda" rank="species" species="majora" subGenus="Parhomola">
<emphasis id="7FCA033C4D3D57760173F8B9FE81E221" box="[292,382,1810,1834]" italics="true" pageId="1" pageNumber="170">Homola</emphasis>
(
<emphasis id="7FCA033C4D3D577601C7F8B9FDF2E221" box="[400,525,1810,1834]" italics="true" pageId="1" pageNumber="170">Parhomola</emphasis>
)
<emphasis id="7FCA033C4D3D57760248F8B8FD8FE221" box="[543,624,1811,1834]" italics="true" pageId="1" pageNumber="170">majora</emphasis>
Kubo, 1936
</taxonomicName>
” (
<typeStatus id="9205618C4D3D5776034CF8B9FCB2E220" box="[795,845,1810,1835]" pageId="1" pageNumber="170">type</typeStatus>
locality:
<collectingCountry id="35A99FBE4D3D577603EEF8B9FC01E220" box="[953,1022,1810,1835]" name="Japan" pageId="1" pageNumber="170">Japan</collectingCountry>
). Serène &amp; Lohavanijaya (1973: 31) believed that Edmondsons specimens, which they did not examine, were not juveniles and that they “belonged probably to
<taxonomicName id="8ABEA4AD4D3D57760177F8F2FE52E27A" authorityName="Henderson" authorityYear="1888" box="[288,429,1881,1905]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="170" phylum="Arthropoda" rank="genus">
<emphasis id="7FCA033C4D3D57760177F8F2FE52E27A" box="[288,429,1881,1905]" italics="true" pageId="1" pageNumber="170">Latreillopsis</emphasis>
</taxonomicName>
but to
<taxonomicName id="8ABEA4AD4D3D577601AAF8F2FD9EE27A" authorityName="Sakai" authorityYear="1936" box="[509,609,1881,1905]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="170" phylum="Arthropoda" rank="species" species="laciniata">
<emphasis id="7FCA033C4D3D577601AAF8F2FD9EE27A" box="[509,609,1881,1905]" italics="true" pageId="1" pageNumber="170">laciniata</emphasis>
</taxonomicName>
more than to
<taxonomicName id="8ABEA4AD4D3D577602A8F8F2FC95E27A" authorityName="Henderson" authorityYear="1888" box="[767,874,1881,1905]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="170" phylum="Arthropoda" rank="species" species="bispinosa">
<emphasis id="7FCA033C4D3D577602A8F8F2FC95E27A" box="[767,874,1881,1905]" italics="true" pageId="1" pageNumber="170">bispinosa</emphasis>
</taxonomicName>
.”
<bibRefCitation id="292FA2DF4D3D577603D2F8F3FABAE27A" author="Guinot, D. &amp; Richer de Forges, B." box="[901,1349,1880,1905]" pageId="1" pageNumber="170" pagination="283 - 517" refId="ref6581" refString="Guinot, D. &amp; Richer de Forges, B. (1995) Crustacea Decapoda Brachyura: Revision de la famille des Homolidae de Haan, 1839. In: Crosnier, A. (Ed.), Resultats des campagnes MUSORSTOM, vol. 13. Memoires du Museum national d'Histoire naturelle (Paris), 163, pp. 283 - 517." type="journal article" year="1995">Guinot &amp; Richer de Forges (1995: 414)</bibRefCitation>
did not examine Edmondsons specimens either and questionably identified them as “?
<taxonomicName id="8ABEA4AD4D3D57760435F8D7FA73E29F" authorityName="Guinot &amp; Richer de Forges" authorityYear="1995" box="[1122,1420,1916,1941]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="170" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="7FCA033C4D3D57760435F8D7FB10E29F" box="[1122,1263,1916,1940]" italics="true" pageId="1" pageNumber="170">Latreillopsis</emphasis>
aff.
<emphasis id="7FCA033C4D3D57760566F8D6FA73E29F" box="[1329,1420,1917,1940]" italics="true" pageId="1" pageNumber="170">cornuta</emphasis>
</taxonomicName>
<emphasis id="7FCA033C4D3D577605DBF8D6FA6FE29F" box="[1420,1424,1917,1940]" italics="true" pageId="1" pageNumber="170">.</emphasis>
<taxonomicName id="8ABEA4AD4D3D577600C0F834FCF0E2B3" authority="Guinot &amp; Richer de Forges, 1995" authorityName="Guinot &amp; Richer de Forges" authorityYear="1995" box="[151,783,1951,1976]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="170" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="7FCA033C4D3D577600C0F834FE7CE2BC" box="[151,387,1951,1975]" italics="true" pageId="1" pageNumber="170">Latreillopsis cornuta</emphasis>
<bibRefCitation id="292FA2DF4D3D577601DBF834FCF0E2B3" author="Guinot, D. &amp; Richer de Forges, B." box="[396,783,1951,1976]" pageId="1" pageNumber="170" pagination="283 - 517" refId="ref6581" refString="Guinot, D. &amp; Richer de Forges, B. (1995) Crustacea Decapoda Brachyura: Revision de la famille des Homolidae de Haan, 1839. In: Crosnier, A. (Ed.), Resultats des campagnes MUSORSTOM, vol. 13. Memoires du Museum national d'Histoire naturelle (Paris), 163, pp. 283 - 517." type="journal article" year="1995">Guinot &amp; Richer de Forges, 1995</bibRefCitation>
</taxonomicName>
, was described from an ovigerous female (13 ×
<quantity id="8A4672CB4D3D5776056CF834FA6BE2B3" box="[1339,1428,1951,1976]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="1" pageNumber="170" unit="mm" value="9.2">9.2 mm</quantity>
) from the Macclesfield Bank, South
<collectingCountry id="35A99FBE4D3D57760267F869FD8CE2D0" box="[560,627,1986,2011]" name="China" pageId="1" pageNumber="170">China</collectingCountry>
Sea.
<bibRefCitation id="292FA2DF4D3D577602E3F869FCA3E2D0" author="Gordon, I." box="[692,860,1986,2011]" pageId="1" pageNumber="170" pagination="201 - 253" refId="ref6465" refString="Gordon, I. (1950) Crustacea Dromiacea. Part I: Systematic account of the Dromiacea collected by the &quot; John Murray &quot; Expedition. Part II. The morphology of the spermatheca in certain Dromiacea. Scientific Reports of the John Murray Expedition 1933 - 34, 9 (3), 201 - 253." type="journal article" year="1950">Gordon (1950)</bibRefCitation>
also examined this specimen (as
<taxonomicName id="8ABEA4AD4D3D57760489F868FA9DE2D1" authorityName="Sakai" authorityYear="1936" box="[1246,1378,1986,2010]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="170" phylum="Arthropoda" rank="species" species="laciniata">
<emphasis id="7FCA033C4D3D57760489F868FA9DE2D1" box="[1246,1378,1986,2010]" italics="true" pageId="1" pageNumber="170">L. laciniata</emphasis>
</taxonomicName>
) and figured the posterior half of the thoracic sternum (
<bibRefCitation id="292FA2DF4D3E57750289FF3CFC83E5BB" author="Gordon, I." box="[734,892,151,176]" pageId="2" pageNumber="171" pagination="201 - 253" refId="ref6465" refString="Gordon, I. (1950) Crustacea Dromiacea. Part I: Systematic account of the Dromiacea collected by the &quot; John Murray &quot; Expedition. Part II. The morphology of the spermatheca in certain Dromiacea. Scientific Reports of the John Murray Expedition 1933 - 34, 9 (3), 201 - 253." type="journal article" year="1950">Gordon 1950</bibRefCitation>
: fig. 26A).
<bibRefCitation id="292FA2DF4D3E57750459FF3CFB26E5BB" author="Castro, P." box="[1038,1241,151,176]" pageId="2" pageNumber="171" pagination="1 - 154" refId="ref6255" refString="Castro, P. (2011) Catalog of the anomuran and brachyuran crabs (Crustacea: Decapoda: Anomura, Brachyura) of the Hawaiian Islands. Zootaxa, 2947, 1 - 154." type="journal article" year="2011">Castro (2011: 35)</bibRefCitation>
could not locate Edmondsons specimens originally deposited in the Bishop Museum, Honolulu, Hawaii (catalog number 5530;
<bibRefCitation id="292FA2DF4D3E577500C0FF75FE67E5FC" author="Edmondson, C. H." box="[151,408,222,247]" pageId="2" pageNumber="171" pagination="183 - 243" refId="ref6433" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of Bernice P. Bishop Museum, 20 (13), 183 - 243." type="journal article" year="1951">Edmondson 1951: 202</bibRefCitation>
) and followed Guinot &amp; Richer de Forges (1995) revision by listing Edmondsons record as “(?)
<emphasis id="7FCA033C4D3E577500BFFEAAFE25E412" box="[232,474,257,281]" italics="true" pageId="2" pageNumber="171">
<taxonomicName id="8ABEA4AD4D3E577500BFFEAAFE29E412" authorityName="Guinot &amp; Richer de Forges" authorityYear="1995" box="[232,470,257,281]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="cornuta">Latreillopsis cornuta</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="4D01DF2E4D3E57750090FE8FFAC7E646" blockId="2.[151,1437,151,2007]" pageId="2" pageNumber="171">
The similarities between Edmondsons specimens, based on his description and rather stylized drawings (
<bibRefCitation id="292FA2DF4D3E577500C9FEECFE5EE46B" author="Edmondson, C. H." box="[158,417,327,352]" pageId="2" pageNumber="171" pagination="183 - 243" refId="ref6433" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of Bernice P. Bishop Museum, 20 (13), 183 - 243." type="journal article" year="1951">Edmondson 1951: 202</bibRefCitation>
, fig. 10), and
<taxonomicName id="8ABEA4AD4D3E5775026BFEE2FD67E46B" authority="Castro &amp; Naruse, 2014" authorityName="Castro &amp; Naruse" authorityYear="2014" box="[572,664,328,352]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="okala" status="sp. nov.">
<emphasis id="7FCA033C4D3E5775026BFEE2FD67E46B" box="[572,664,328,352]" italics="true" pageId="2" pageNumber="171">L. okala</emphasis>
</taxonomicName>
<emphasis id="7FCA033C4D3E577502C8FEECFD22E46B" bold="true" box="[671,733,327,352]" pageId="2" pageNumber="171">
<taxonomicNameLabel id="64F9BE474D3E577502C8FEECFD22E46B" box="[671,733,327,352]" pageId="2" pageNumber="171" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
are evident from the general morphology of the dorsal surface of the carapace, posterior portion of the abdomen, and P1, P2, and P5, all of which were illustrated by Edmondson. The surface of the carapace and appendages were furthermore described as “densely covered with flat, wartlike granules” (
<bibRefCitation id="292FA2DF4D3E57750145FE1AFDE7E4C1" author="Edmondson, C. H." box="[274,536,433,458]" pageId="2" pageNumber="171" pagination="183 - 243" refId="ref6433" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of Bernice P. Bishop Museum, 20 (13), 183 - 243." type="journal article" year="1951">Edmondson 1951: 202</bibRefCitation>
), which is diagnostic of the new species. There are some differences, however. The slender teeth on the dorsal regions and outer margins of the carapace and on the pseudorostral spines (
<bibRefCitation id="292FA2DF4D3E577500C9FE5CFE95E71B" author="Edmondson, C. H." box="[158,362,503,528]" pageId="2" pageNumber="171" pagination="183 - 243" refId="ref6433" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of Bernice P. Bishop Museum, 20 (13), 183 - 243." type="journal article" year="1951">Edmondson 1951</bibRefCitation>
: fig. 10a, b) are more blunt and thick in the
<typeStatus id="9205618C4D3E5775033FFE5CFC35E71B" box="[872,970,503,528]" pageId="2" pageNumber="171" type="holotype">holotype</typeStatus>
of the new species (
<figureCitation id="D585C3AB4D3E577504E0FE5CFB05E71B" box="[1207,1274,503,528]" captionStart="FIGURE 1" captionStartId="3.[151,249,1983,2005]" captionTargetBox="[164,1423,298,1961]" captionTargetId="figure-95@3.[164,1423,298,1961]" captionTargetPageId="3" captionText="FIGURE 1. Latreillopsis okala n. sp. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, habitus, dorsal view; b, color in life (photograph by P. Fiene)." figureDoi="http://doi.org/10.5281/zenodo.5045614" httpUri="https://zenodo.org/record/5045614/files/figure.png" pageId="2" pageNumber="171">Fig. 1</figureCitation>
,
<figureCitation id="D585C3AB4D3E5775055DFE5CFAD7E71B" box="[1290,1320,503,528]" captionStart="FIGURE 3" captionStartId="5.[151,249,1877,1899]" captionTargetBox="[180,1414,213,1841]" captionTargetId="figure-71@5.[172,1414,193,1856]" captionTargetPageId="5" captionText="FIGURE 3. Latreillopsis okala n. sp. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, cephalothorax, dorsal view (wartlike granules are not drawn); b, left anterior part of ventral surface of cephalothorax (wart-like granules are not drawn); c, left G1, ventral view; d, left G2, ventral view. Scales: ad = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5045618" httpUri="https://zenodo.org/record/5045618/files/figure.png" pageId="2" pageNumber="171">3a</figureCitation>
). The two small teeth just anterior to the posterior margin shown in Edmondsons figure (
<bibRefCitation id="292FA2DF4D3E57750466FDB0FB00E73F" author="Edmondson, C. H." box="[1073,1279,539,564]" pageId="2" pageNumber="171" pagination="183 - 243" refId="ref6433" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of Bernice P. Bishop Museum, 20 (13), 183 - 243." type="journal article" year="1951">Edmondson 1951</bibRefCitation>
: fig. 10a) are absent in the
<typeStatus id="9205618C4D3E57750162FD95FE64E75C" box="[309,411,574,599]" pageId="2" pageNumber="171" type="holotype">holotype</typeStatus>
. The rostrum is slightly shorter than the pseudorostral spines in Edmondsons figure but much shorter than the pseudorostral spines in the
<typeStatus id="9205618C4D3E57750295FDCAFCDBE771" box="[706,804,609,634]" pageId="2" pageNumber="171" type="holotype">holotype</typeStatus>
of the new species. The rostrum and pseudorstral spines are cut off in the
<typeStatus id="9205618C4D3E57750103FD2FFE46E796" box="[340,441,644,669]" pageId="2" pageNumber="171" type="holotype">holotype</typeStatus>
, however, so the actual size relation cannot be ascertained. The pseudorostral spines, also incomplete, are slightly less curved in the
<typeStatus id="9205618C4D3E5775022FFD0CFD21E7CB" box="[632,734,679,704]" pageId="2" pageNumber="171" type="holotype">holotype</typeStatus>
. These differences can perhaps be explained by differences in size, Edmondsons specimens (cl
<quantity id="8A4672CB4D3E57750244FD60FDA8E7E9" box="[531,599,715,740]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="171" unit="mm" value="8.0">8 mm</quantity>
, not including the rostrum) being larger than the
<typeStatus id="9205618C4D3E577504D3FD60FB19E7EF" box="[1156,1254,715,740]" pageId="2" pageNumber="171" type="holotype">holotype</typeStatus>
(cl
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, not including the rostrum). The right cheliped of the female illustrated by
<bibRefCitation id="292FA2DF4D3E577503F0FD45FB85E60C" author="Edmondson, C. H." box="[935,1146,750,775]" pageId="2" pageNumber="171" pagination="183 - 243" refId="ref6433" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of Bernice P. Bishop Museum, 20 (13), 183 - 243." type="journal article" year="1951">Edmondson (1951</bibRefCitation>
: fig. 10c), is more slender than in the male
<typeStatus id="9205618C4D3E57750100FCBAFE46E621" box="[343,441,785,810]" pageId="2" pageNumber="171" type="holotype">holotype</typeStatus>
(
<figureCitation id="D585C3AB4D3E5775019DFCBAFDE7E621" box="[458,536,785,810]" captionStart="FIGURE 1" captionStartId="3.[151,249,1983,2005]" captionTargetBox="[164,1423,298,1961]" captionTargetId="figure-95@3.[164,1423,298,1961]" captionTargetPageId="3" captionText="FIGURE 1. Latreillopsis okala n. sp. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, habitus, dorsal view; b, color in life (photograph by P. Fiene)." figureDoi="http://doi.org/10.5281/zenodo.5045614" httpUri="https://zenodo.org/record/5045614/files/figure.png" pageId="2" pageNumber="171">Figs. 1</figureCitation>
,
<figureCitation id="D585C3AB4D3E5775027FFCBAFDC7E621" box="[552,568,785,810]" captionStart="FIGURE 2" captionStartId="4.[151,249,1280,1302]" captionTargetBox="[151,1436,193,1259]" captionTargetId="figure-387@4.[151,1436,193,1259]" captionTargetPageId="4" captionText="FIGURE 2. Latreillopsis okala n. sp. species. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, cephalothorax, ventral view; b, outer surface of propodus and dactylus, outer view." figureDoi="http://doi.org/10.5281/zenodo.5045616" httpUri="https://zenodo.org/record/5045616/files/figure.png" pageId="2" pageNumber="171">2</figureCitation>
), but the discrepancy is most probably the result of a sex difference. We are therefore confident that the
<typeStatus id="9205618C4D3E57750199FC9FFDCFE646" box="[462,560,820,845]" pageId="2" pageNumber="171" type="holotype">holotype</typeStatus>
of the new species and Edmondsons
<specimenCount id="5BB814A74D3E5775038CFC9EFB7EE646" box="[987,1153,820,845]" pageId="2" pageNumber="171" type="generic">two specimens</specimenCount>
are conspecific.
</paragraph>
<paragraph id="4D01DF2E4D3E57750090FCFCFDA2E1A6" blockId="2.[151,1437,151,2007]" pageId="2" pageNumber="171">
There are superficial similarities between
<taxonomicName id="8ABEA4AD4D3E577502FEFCF2FCF8E67B" authority="Castro &amp; Naruse, 2014" authorityName="Castro &amp; Naruse" authorityYear="2014" box="[681,775,856,880]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="okala" status="sp. nov.">
<emphasis id="7FCA033C4D3E577502FEFCF2FCF8E67B" box="[681,775,856,880]" italics="true" pageId="2" pageNumber="171">L. okala</emphasis>
</taxonomicName>
<emphasis id="7FCA033C4D3E57750346FCFCFCADE67B" bold="true" box="[785,850,855,880]" pageId="2" pageNumber="171">
<taxonomicNameLabel id="64F9BE474D3E57750346FCFCFCADE67B" box="[785,850,855,880]" pageId="2" pageNumber="171" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and the description and photographs of
<taxonomicName id="8ABEA4AD4D3E57750572FCF2FA63E67B" authorityName="Guinot &amp; Richer de Forges" authorityYear="1995" box="[1317,1436,857,880]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="7FCA033C4D3E57750572FCF2FA63E67B" box="[1317,1436,857,880]" italics="true" pageId="2" pageNumber="171">L. cornuta</emphasis>
</taxonomicName>
given by
<bibRefCitation id="292FA2DF4D3E57750156FCD0FD74E69F" author="Guinot, D. &amp; Richer de Forges, B." box="[257,651,891,916]" pageId="2" pageNumber="171" pagination="283 - 517" refId="ref6581" refString="Guinot, D. &amp; Richer de Forges, B. (1995) Crustacea Decapoda Brachyura: Revision de la famille des Homolidae de Haan, 1839. In: Crosnier, A. (Ed.), Resultats des campagnes MUSORSTOM, vol. 13. Memoires du Museum national d'Histoire naturelle (Paris), 163, pp. 283 - 517." type="journal article" year="1995">Guinot &amp; Richer de Forges (1995)</bibRefCitation>
. As noted by
<bibRefCitation id="292FA2DF4D3E57750370FCD0FB16E69F" author="Guinot, D. &amp; Richer de Forges, B." box="[807,1257,891,916]" pageId="2" pageNumber="171" pagination="283 - 517" refId="ref6581" refString="Guinot, D. &amp; Richer de Forges, B. (1995) Crustacea Decapoda Brachyura: Revision de la famille des Homolidae de Haan, 1839. In: Crosnier, A. (Ed.), Resultats des campagnes MUSORSTOM, vol. 13. Memoires du Museum national d'Histoire naturelle (Paris), 163, pp. 283 - 517." type="journal article" year="1995">Guinot &amp; Richer de Forges (1995: 415)</bibRefCitation>
, however, there are differences between
<taxonomicName id="8ABEA4AD4D3E577501FDFC34FDE0E6BD" authorityName="Guinot &amp; Richer de Forges" authorityYear="1995" box="[426,543,927,950]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="7FCA033C4D3E577501FDFC34FDE0E6BD" box="[426,543,927,950]" italics="true" pageId="2" pageNumber="171">L. cornuta</emphasis>
</taxonomicName>
and
<bibRefCitation id="292FA2DF4D3E57750200FC35FCBDE6BC" author="Edmondson, C. H." box="[599,834,926,951]" pageId="2" pageNumber="171" pagination="183 - 243" refId="ref6433" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of Bernice P. Bishop Museum, 20 (13), 183 - 243." type="journal article" year="1951">Edmondsons (1951)</bibRefCitation>
specimens, which are now identified as belonging to the new species. The rostrum is shorter in
<taxonomicName id="8ABEA4AD4D3E577502F5FC69FCE2E6D2" authorityName="Guinot &amp; Richer de Forges" authorityYear="1995" box="[674,797,962,985]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="7FCA033C4D3E577502F5FC69FCE2E6D2" box="[674,797,962,985]" italics="true" pageId="2" pageNumber="171">L. cornuta</emphasis>
</taxonomicName>
(
<bibRefCitation id="292FA2DF4D3E57750366FC6AFB39E6D1" author="Guinot, D. &amp; Richer de Forges, B." box="[817,1222,961,986]" pageId="2" pageNumber="171" pagination="283 - 517" refId="ref6581" refString="Guinot, D. &amp; Richer de Forges, B. (1995) Crustacea Decapoda Brachyura: Revision de la famille des Homolidae de Haan, 1839. In: Crosnier, A. (Ed.), Resultats des campagnes MUSORSTOM, vol. 13. Memoires du Museum national d'Histoire naturelle (Paris), 163, pp. 283 - 517." type="journal article" year="1995">Guinot &amp; Richer de Forges 1995</bibRefCitation>
: fig. 37e) than in Edmondsons specimens, there are four equidistant spinules on the P2P4 meri of
<taxonomicName id="8ABEA4AD4D3E577504C7FC4EFAF3E6F7" authorityName="Guinot &amp; Richer de Forges" authorityYear="1995" box="[1168,1292,997,1020]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="7FCA033C4D3E577504C7FC4EFAF3E6F7" box="[1168,1292,997,1020]" italics="true" pageId="2" pageNumber="171">L. cornuta</emphasis>
</taxonomicName>
(smooth in Edmondsons specimens), and four teeth on the subhepatic region, including two salient ones (three teeth, including a salient one in Edmondsons specimens). They also mentioned as an additional difference the absence or near absence of “tubercles” on the dorsal surface of Edmondsons specimens. This is an error because
<bibRefCitation id="292FA2DF4D3E57750541FBE5FEE1E181" author="Edmondson, C. H." pageId="2" pageNumber="171" pagination="183 - 243" refId="ref6433" refString="Edmondson, C. H. (1951) Some Central Pacific crustaceans. Occasional Papers of Bernice P. Bishop Museum, 20 (13), 183 - 243." type="journal article" year="1951">Edmondson (1951: 202)</bibRefCitation>
described the carapace of his specimens as covered with granules, a character they share with the
<typeStatus id="9205618C4D3E577500C0FB3FFF06E1A6" box="[151,249,1172,1197]" pageId="2" pageNumber="171" type="holotype">holotype</typeStatus>
of the new species (see above).
</paragraph>
<paragraph id="4D01DF2E4D3E57750090FB1CFDA4E35F" blockId="2.[151,1437,151,2007]" pageId="2" pageNumber="171">
The preceding differences between
<taxonomicName id="8ABEA4AD4D3E5775023DFB12FD1CE1DB" authorityName="Guinot &amp; Richer de Forges" authorityYear="1995" box="[618,739,1209,1232]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="7FCA033C4D3E5775023DFB12FD1CE1DB" box="[618,739,1209,1232]" italics="true" pageId="2" pageNumber="171">L. cornuta</emphasis>
</taxonomicName>
and Edmondsons specimens (i.e.
<taxonomicName id="8ABEA4AD4D3E577504D6FB12FB24E1DB" authority="Castro &amp; Naruse, 2014" authorityName="Castro &amp; Naruse" authorityYear="2014" box="[1153,1243,1208,1232]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="okala" status="sp. nov.">
<emphasis id="7FCA033C4D3E577504D6FB12FB24E1DB" box="[1153,1243,1208,1232]" italics="true" pageId="2" pageNumber="171">L okala</emphasis>
</taxonomicName>
<emphasis id="7FCA033C4D3E577504B0FB1CFAD4E1DB" bold="true" box="[1255,1323,1207,1232]" pageId="2" pageNumber="171">
<taxonomicNameLabel id="64F9BE474D3E577504B0FB1CFAD4E1DB" box="[1255,1323,1207,1232]" pageId="2" pageNumber="171" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
) are also observed when the description and illustrations of
<taxonomicName id="8ABEA4AD4D3E57750283FB77FCB5E1F8" authorityName="Guinot &amp; Richer de Forges" authorityYear="1995" box="[724,842,1244,1267]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="7FCA033C4D3E57750283FB77FCB5E1F8" box="[724,842,1244,1267]" italics="true" pageId="2" pageNumber="171">L. cornuta</emphasis>
</taxonomicName>
are compared with the
<typeStatus id="9205618C4D3E5775040FFB70FB45E1FF" box="[1112,1210,1243,1268]" pageId="2" pageNumber="171" type="holotype">holotype</typeStatus>
of the new species: rostrum that is shorter in
<taxonomicName id="8ABEA4AD4D3E577501EFFB54FDD1E01D" authorityName="Guinot &amp; Richer de Forges" authorityYear="1995" box="[440,558,1279,1302]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="7FCA033C4D3E577501EFFB54FDD1E01D" box="[440,558,1279,1302]" italics="true" pageId="2" pageNumber="171">L. cornuta</emphasis>
</taxonomicName>
(
<bibRefCitation id="292FA2DF4D3E57750268FB55FC44E01C" author="Guinot, D. &amp; Richer de Forges, B." box="[575,955,1278,1303]" pageId="2" pageNumber="171" pagination="283 - 517" refId="ref6581" refString="Guinot, D. &amp; Richer de Forges, B. (1995) Crustacea Decapoda Brachyura: Revision de la famille des Homolidae de Haan, 1839. In: Crosnier, A. (Ed.), Resultats des campagnes MUSORSTOM, vol. 13. Memoires du Museum national d'Histoire naturelle (Paris), 163, pp. 283 - 517." type="journal article" year="1995">Guinot &amp; Richer de Forges 1995</bibRefCitation>
: figs. 37e, f, 41C) than in the new species (
<figureCitation id="D585C3AB4D3E577500C8FA8AFF0FE031" box="[159,240,1313,1338]" captionStart="FIGURE 3" captionStartId="5.[151,249,1877,1899]" captionTargetBox="[180,1414,213,1841]" captionTargetId="figure-71@5.[172,1414,193,1856]" captionTargetPageId="5" captionText="FIGURE 3. Latreillopsis okala n. sp. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, cephalothorax, dorsal view (wartlike granules are not drawn); b, left anterior part of ventral surface of cephalothorax (wart-like granules are not drawn); c, left G1, ventral view; d, left G2, ventral view. Scales: ad = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5045618" httpUri="https://zenodo.org/record/5045618/files/figure.png" pageId="2" pageNumber="171">Fig. 3a</figureCitation>
), P2P4 meri bordered by spines (
<bibRefCitation id="292FA2DF4D3E57750221FA8AFC11E031" author="Guinot, D. &amp; Richer de Forges, B." box="[630,1006,1313,1338]" pageId="2" pageNumber="171" pagination="283 - 517" refId="ref6581" refString="Guinot, D. &amp; Richer de Forges, B. (1995) Crustacea Decapoda Brachyura: Revision de la famille des Homolidae de Haan, 1839. In: Crosnier, A. (Ed.), Resultats des campagnes MUSORSTOM, vol. 13. Memoires du Museum national d'Histoire naturelle (Paris), 163, pp. 283 - 517." type="journal article" year="1995">Guinot &amp; Richer de Forges 1995</bibRefCitation>
: figs. 37e) but unarmed (
<figureCitation id="D585C3AB4D3E5775055FFA8AFAB6E031" box="[1288,1353,1313,1338]" captionStart="FIGURE 1" captionStartId="3.[151,249,1983,2005]" captionTargetBox="[164,1423,298,1961]" captionTargetId="figure-95@3.[164,1423,298,1961]" captionTargetPageId="3" captionText="FIGURE 1. Latreillopsis okala n. sp. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, habitus, dorsal view; b, color in life (photograph by P. Fiene)." figureDoi="http://doi.org/10.5281/zenodo.5045614" httpUri="https://zenodo.org/record/5045614/files/figure.png" pageId="2" pageNumber="171">Fig. 1</figureCitation>
) in the new species, and four subhepatic teeth (
<bibRefCitation id="292FA2DF4D3E5775020EFAEFFC2DE056" author="Guinot, D. &amp; Richer de Forges, B." box="[601,978,1348,1373]" pageId="2" pageNumber="171" pagination="283 - 517" refId="ref6581" refString="Guinot, D. &amp; Richer de Forges, B. (1995) Crustacea Decapoda Brachyura: Revision de la famille des Homolidae de Haan, 1839. In: Crosnier, A. (Ed.), Resultats des campagnes MUSORSTOM, vol. 13. Memoires du Museum national d'Histoire naturelle (Paris), 163, pp. 283 - 517." type="journal article" year="1995">Guinot &amp; Richer de Forges 1995</bibRefCitation>
: figs. 37e, f, 41C) instead of three in the new species (
<figureCitation id="D585C3AB4D3E5775017BFACCFE81E08B" box="[300,382,1383,1408]" captionStart="FIGURE 3" captionStartId="5.[151,249,1877,1899]" captionTargetBox="[180,1414,213,1841]" captionTargetId="figure-71@5.[172,1414,193,1856]" captionTargetPageId="5" captionText="FIGURE 3. Latreillopsis okala n. sp. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, cephalothorax, dorsal view (wartlike granules are not drawn); b, left anterior part of ventral surface of cephalothorax (wart-like granules are not drawn); c, left G1, ventral view; d, left G2, ventral view. Scales: ad = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5045618" httpUri="https://zenodo.org/record/5045618/files/figure.png" pageId="2" pageNumber="171">Fig. 3b</figureCitation>
). Other differences are also evident. The third maxilliped merus of the new species has a large, rounded tubercle on its anterior half (
<figureCitation id="D585C3AB4D3E57750203FA20FD55E0AF" box="[596,682,1419,1444]" captionStart="FIGURE 3" captionStartId="5.[151,249,1877,1899]" captionTargetBox="[180,1414,213,1841]" captionTargetId="figure-71@5.[172,1414,193,1856]" captionTargetPageId="5" captionText="FIGURE 3. Latreillopsis okala n. sp. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, cephalothorax, dorsal view (wartlike granules are not drawn); b, left anterior part of ventral surface of cephalothorax (wart-like granules are not drawn); c, left G1, ventral view; d, left G2, ventral view. Scales: ad = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5045618" httpUri="https://zenodo.org/record/5045618/files/figure.png" pageId="2" pageNumber="171">Fig. 3b</figureCitation>
) in contrast with two well-developed but smaller tubercles, one anterior and a second on the posterior half, in
<taxonomicName id="8ABEA4AD4D3E577502E3FA04FCD4E0CD" authorityName="Guinot &amp; Richer de Forges" authorityYear="1995" box="[692,811,1455,1478]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="7FCA033C4D3E577502E3FA04FCD4E0CD" box="[692,811,1455,1478]" italics="true" pageId="2" pageNumber="171">L. cornuta</emphasis>
</taxonomicName>
(
<bibRefCitation id="292FA2DF4D3E5775036AFA05FB3EE0CC" author="Guinot, D. &amp; Richer de Forges, B." box="[829,1217,1454,1479]" pageId="2" pageNumber="171" pagination="283 - 517" refId="ref6581" refString="Guinot, D. &amp; Richer de Forges, B. (1995) Crustacea Decapoda Brachyura: Revision de la famille des Homolidae de Haan, 1839. In: Crosnier, A. (Ed.), Resultats des campagnes MUSORSTOM, vol. 13. Memoires du Museum national d'Histoire naturelle (Paris), 163, pp. 283 - 517." type="journal article" year="1995">Guinot &amp; Richer de Forges 1995</bibRefCitation>
: fig. 41C), and the carapace is granular (
<figureCitation id="D585C3AB4D3E577501D1FA7AFE2AE0E1" box="[390,469,1489,1514]" captionStart="FIGURE 3" captionStartId="5.[151,249,1877,1899]" captionTargetBox="[180,1414,213,1841]" captionTargetId="figure-71@5.[172,1414,193,1856]" captionTargetPageId="5" captionText="FIGURE 3. Latreillopsis okala n. sp. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, cephalothorax, dorsal view (wartlike granules are not drawn); b, left anterior part of ventral surface of cephalothorax (wart-like granules are not drawn); c, left G1, ventral view; d, left G2, ventral view. Scales: ad = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5045618" httpUri="https://zenodo.org/record/5045618/files/figure.png" pageId="2" pageNumber="171">Fig. 3a</figureCitation>
) in contrast to the absence of conspicuous granules in
<taxonomicName id="8ABEA4AD4D3E5775046EFA79FB51E0E2" authorityName="Guinot &amp; Richer de Forges" authorityYear="1995" box="[1081,1198,1490,1513]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="7FCA033C4D3E5775046EFA79FB51E0E2" box="[1081,1198,1490,1513]" italics="true" pageId="2" pageNumber="171">L. cornuta</emphasis>
</taxonomicName>
(
<bibRefCitation id="292FA2DF4D3E577504EBFA7AFEDAE306" author="Guinot, D. &amp; Richer de Forges, B." pageId="2" pageNumber="171" pagination="283 - 517" refId="ref6581" refString="Guinot, D. &amp; Richer de Forges, B. (1995) Crustacea Decapoda Brachyura: Revision de la famille des Homolidae de Haan, 1839. In: Crosnier, A. (Ed.), Resultats des campagnes MUSORSTOM, vol. 13. Memoires du Museum national d'Histoire naturelle (Paris), 163, pp. 283 - 517." type="journal article" year="1995">Guinot &amp; Richer de Forges 1995</bibRefCitation>
: fig. 41C). Although the adult male
<typeStatus id="9205618C4D3E577502EEFA5FFCE4E306" box="[697,795,1524,1549]" pageId="2" pageNumber="171" type="holotype">holotype</typeStatus>
of
<taxonomicName id="8ABEA4AD4D3E57750317FA5EFC63E306" authority="Castro &amp; Naruse, 2014" authorityName="Castro &amp; Naruse" authorityYear="2014" box="[832,924,1525,1549]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="okala" status="sp. nov.">
<emphasis id="7FCA033C4D3E57750317FA5EFC63E306" box="[832,924,1525,1549]" italics="true" pageId="2" pageNumber="171">L. okala</emphasis>
</taxonomicName>
<emphasis id="7FCA033C4D3E577503F4FA5FFC1EE306" bold="true" box="[931,993,1524,1549]" pageId="2" pageNumber="171">
<taxonomicNameLabel id="64F9BE474D3E577503F4FA5FFC1EE306" box="[931,993,1524,1549]" pageId="2" pageNumber="171" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, is less than half (6.0 × 4.0 mm) that of the
<typeStatus id="9205618C4D3E57750097F9BCFEDDE33B" box="[192,290,1559,1584]" pageId="2" pageNumber="171" type="holotype">holotype</typeStatus>
of
<taxonomicName id="8ABEA4AD4D3E57750110F9B2FE44E33B" authorityName="Guinot &amp; Richer de Forges" authorityYear="1995" box="[327,443,1561,1584]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="7FCA033C4D3E57750110F9B2FE44E33B" box="[327,443,1561,1584]" italics="true" pageId="2" pageNumber="171">L. cornuta</emphasis>
</taxonomicName>
(13 ×
<quantity id="8A4672CB4D3E57750251F9BCFD9FE33B" box="[518,608,1559,1584]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="2" pageNumber="171" unit="mm" value="9.2">9.2 mm</quantity>
), these morphological differences are regarded as not to be related to size, notwithstanding the scarcity of material.
</paragraph>
<paragraph id="4D01DF2E4D3E57750090F9F5FE35E241" blockId="2.[151,1437,151,2007]" pageId="2" pageNumber="171">
<taxonomicName id="8ABEA4AD4D3E57750090F9F5FE67E37D" authority="Castro &amp; Naruse, 2014" authorityName="Castro &amp; Naruse" authorityYear="2014" box="[199,408,1630,1654]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="okala" status="sp. nov.">
<emphasis id="7FCA033C4D3E57750090F9F5FE67E37D" box="[199,408,1630,1654]" italics="true" pageId="2" pageNumber="171">Latreillopsis okala</emphasis>
</taxonomicName>
<emphasis id="7FCA033C4D3E577501C8F9F5FE22E37C" bold="true" box="[415,477,1630,1655]" pageId="2" pageNumber="171">
<taxonomicNameLabel id="64F9BE474D3E577501C8F9F5FE22E37C" box="[415,477,1630,1655]" pageId="2" pageNumber="171" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
shares with the nine previously described species of
<taxonomicName id="8ABEA4AD4D3E57750479F9F5FB44E37D" authorityName="Henderson" authorityYear="1888" box="[1070,1211,1630,1654]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">
<emphasis id="7FCA033C4D3E57750479F9F5FB44E37D" box="[1070,1211,1630,1654]" italics="true" pageId="2" pageNumber="171">Latreillopsis</emphasis>
</taxonomicName>
(see
<bibRefCitation id="292FA2DF4D3E577504A2F9F4FF58E391" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." pageId="2" pageNumber="171" pagination="1 - 286" refId="ref6844" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286." type="journal article" year="2008">
Ng
<emphasis id="7FCA033C4D3E57750549F9F4FAA9E37D" box="[1310,1366,1630,1654]" italics="true" pageId="2" pageNumber="171">et al.</emphasis>
2008: 9
</bibRefCitation>
), a well-developed sub-hepatic region of the carapace, sculpted dorsal surface of the carapace, small lateral carapace teeth, absence of supraorbital teeth, subcheliform P5, moderately long pseudorostral spines without welldeveloped teeth, and reduced chelipeds that are shorter than the ambulatory legs. Having a small size can be added to the characters shared with
<taxonomicName id="8ABEA4AD4D3E57750188F940FD91E20F" authorityName="Henderson" authorityYear="1888" box="[479,622,1771,1796]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">Latreillopsis</taxonomicName>
, as re-described by
<bibRefCitation id="292FA2DF4D3E5775031BF940FAF5E20F" author="Guinot, D. &amp; Richer de Forges, B." box="[844,1290,1771,1796]" pageId="2" pageNumber="171" pagination="283 - 517" refId="ref6581" refString="Guinot, D. &amp; Richer de Forges, B. (1995) Crustacea Decapoda Brachyura: Revision de la famille des Homolidae de Haan, 1839. In: Crosnier, A. (Ed.), Resultats des campagnes MUSORSTOM, vol. 13. Memoires du Museum national d'Histoire naturelle (Paris), 163, pp. 283 - 517." type="journal article" year="1995">Guinot &amp; Richer de Forges (1995: 393)</bibRefCitation>
, even if only three small specimens of the new species are known. The small
<typeStatus id="9205618C4D3E57750329F8A5FC1FE22C" box="[894,992,1806,1831]" pageId="2" pageNumber="171" type="holotype">holotype</typeStatus>
of the new species is an adult, having fully chitinized G1 and G2.
</paragraph>
<paragraph id="4D01DF2E4D3E57740090F8FFFB54E5FC" blockId="2.[151,1437,151,2007]" lastBlockId="3.[151,1436,151,247]" lastPageId="3" lastPageNumber="172" pageId="2" pageNumber="171">
The nine known species of
<taxonomicName id="8ABEA4AD4D3E5775025AF8FEFD65E266" authorityName="Henderson" authorityYear="1888" box="[525,666,1877,1901]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">
<emphasis id="7FCA033C4D3E5775025AF8FEFD65E266" box="[525,666,1877,1901]" italics="true" pageId="2" pageNumber="171">Latreillopsis</emphasis>
</taxonomicName>
are:
<taxonomicName id="8ABEA4AD4D3E5775028EF8FEFB00E266" authority="Guinot &amp; Richer de Forges, 1995" authorityName="Guinot &amp; Richer de Forges" authorityYear="1995" box="[729,1279,1876,1901]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="antennata">
<emphasis id="7FCA033C4D3E5775028EF8FEFC94E267" box="[729,875,1877,1900]" italics="true" pageId="2" pageNumber="171">L. antennata</emphasis>
<bibRefCitation id="292FA2DF4D3E57750322F8FFFB00E266" author="Guinot, D. &amp; Richer de Forges, B." box="[885,1279,1876,1901]" pageId="2" pageNumber="171" pagination="283 - 517" refId="ref6581" refString="Guinot, D. &amp; Richer de Forges, B. (1995) Crustacea Decapoda Brachyura: Revision de la famille des Homolidae de Haan, 1839. In: Crosnier, A. (Ed.), Resultats des campagnes MUSORSTOM, vol. 13. Memoires du Museum national d'Histoire naturelle (Paris), 163, pp. 283 - 517." type="journal article" year="1995">Guinot &amp; Richer de Forges, 1995</bibRefCitation>
</taxonomicName>
(Chesterfield Islands,
<collectingCountry id="35A99FBE4D3E577500AEF8D3FE4FE29B" box="[249,432,1911,1936]" name="New Caledonia" pageId="2" pageNumber="171">New Caledonia</collectingCountry>
),
<taxonomicName id="8ABEA4AD4D3E5775019FF8D2FCD9E29B" authority="Henderson, 1888" authorityName="Henderson" authorityYear="1888" box="[456,806,1911,1936]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="bispinosa">
<emphasis id="7FCA033C4D3E5775019FF8D2FDA8E29B" box="[456,599,1912,1936]" italics="true" pageId="2" pageNumber="171">L. bispinosa</emphasis>
<bibRefCitation id="292FA2DF4D3E57750235F8DCFCD9E29B" author="Henderson, J. R." box="[610,806,1911,1936]" pageId="2" pageNumber="171" pagination="1 - 221" refId="ref6717" refString="Henderson, J. R. (1888) Report on the Anomura collected by H. M. S. Challenger during the years 1873 - 1876. In: Wyville Thomson, C. &amp; Murray, J. (Eds.), Report of the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 76 Under the Command of Captain George S. Nares, R. N., F. R. S. and the late Captain Frank Tourle Thompson, R. N., Zoology, 27 (69), i - xi + pp. 1 - 221 pp., pls. 1 - 21." type="book chapter" year="1888">Henderson, 1888</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="35A99FBE4D3E5775036DF8DCFC81E29B" box="[826,894,1911,1936]" name="Japan" pageId="2" pageNumber="171">Japan</collectingCountry>
,
<collectingCountry id="35A99FBE4D3E577503D8F8DCFBEDE29B" box="[911,1042,1911,1936]" name="Philippines" pageId="2" pageNumber="171">Philippines</collectingCountry>
),
<taxonomicName id="8ABEA4AD4D3E5775047EF8D2FED6E2BF" authority="Guinot &amp; Richer de Forges, 1995" authorityName="Guinot &amp; Richer de Forges" authorityYear="1995" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="7FCA033C4D3E5775047EF8D2FB5CE29B" box="[1065,1187,1913,1936]" italics="true" pageId="2" pageNumber="171">L. cornuta</emphasis>
<bibRefCitation id="292FA2DF4D3E577504F8F8DCFED6E2BF" author="Guinot, D. &amp; Richer de Forges, B." pageId="2" pageNumber="171" pagination="283 - 517" refId="ref6581" refString="Guinot, D. &amp; Richer de Forges, B. (1995) Crustacea Decapoda Brachyura: Revision de la famille des Homolidae de Haan, 1839. In: Crosnier, A. (Ed.), Resultats des campagnes MUSORSTOM, vol. 13. Memoires du Museum national d'Histoire naturelle (Paris), 163, pp. 283 - 517." type="journal article" year="1995">Guinot &amp; Richer de Forges, 1995</bibRefCitation>
</taxonomicName>
(South
<collectingCountry id="35A99FBE4D3E577501D5F830FE3AE2BF" box="[386,453,1947,1972]" name="China" pageId="2" pageNumber="171">China</collectingCountry>
Sea),
<taxonomicName id="8ABEA4AD4D3E5775025BF837FC12E2BF" authority="Guinot &amp; Richer de Forges, 1995" authorityName="Guinot &amp; Richer de Forges" authorityYear="1995" box="[524,1005,1947,1972]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="daviei">
<emphasis id="7FCA033C4D3E5775025BF837FD91E2B8" box="[524,622,1947,1971]" italics="true" pageId="2" pageNumber="171">L. daviei</emphasis>
<bibRefCitation id="292FA2DF4D3E57750222F830FC12E2BF" author="Guinot, D. &amp; Richer de Forges, B." box="[629,1005,1947,1972]" pageId="2" pageNumber="171" pagination="283 - 517" refId="ref6581" refString="Guinot, D. &amp; Richer de Forges, B. (1995) Crustacea Decapoda Brachyura: Revision de la famille des Homolidae de Haan, 1839. In: Crosnier, A. (Ed.), Resultats des campagnes MUSORSTOM, vol. 13. Memoires du Museum national d'Histoire naturelle (Paris), 163, pp. 283 - 517." type="journal article" year="1995">Guinot &amp; Richer de Forges, 1995</bibRefCitation>
</taxonomicName>
(
<collectingRegion id="8F7A11CC4D3E577503ABF830FB7AE2BF" box="[1020,1157,1947,1972]" country="Australia" name="Queensland" pageId="2" pageNumber="171">Queensland</collectingRegion>
,
<collectingCountry id="35A99FBE4D3E577504C7F830FB04E2BF" box="[1168,1275,1947,1972]" name="Australia" pageId="2" pageNumber="171">Australia</collectingCountry>
),
<taxonomicName id="8ABEA4AD4D3E57750559F837FDDEE2DC" authority="Guinot &amp; Richer de Forges, 1981" authorityName="Guinot &amp; Richer de Forges" authorityYear="1981" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="7FCA033C4D3E57750559F837FA63E2B8" box="[1294,1436,1947,1971]" italics="true" pageId="2" pageNumber="171">L. gracilipes</emphasis>
<bibRefCitation id="292FA2DF4D3E577500C0F815FDDEE2DC" author="Guinot, D. &amp; Richer de Forges, B." box="[151,545,1982,2007]" pageId="2" pageNumber="171" pagination="523 - 581" refId="ref6524" refString="Guinot, D. &amp; Richer de Forges, B. (1981) Homolidae, rares ou nouveaux, de l'Indo-Pacifique (Crustacea, Decapoda, Brachyura). Bulletin du Museum national d'Histoire naturelle (Paris), ser. 4, 3, sect. A (2), 523 - 581." type="journal article" year="1981">Guinot &amp; Richer de Forges, 1981</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="35A99FBE4D3E57750263F815FD6AE2DC" box="[564,661,1982,2007]" name="Vanuatu" pageId="2" pageNumber="171">Vanuatu</collectingCountry>
,
<collectingCountry id="35A99FBE4D3E577502F4F814FCA6E2DC" box="[675,857,1982,2007]" name="New Caledonia" pageId="2" pageNumber="171">New Caledonia</collectingCountry>
),
<taxonomicName id="8ABEA4AD4D3E57750327F814FB17E2DC" authority="Sakai, 1936" authorityName="Sakai" authorityYear="1936" box="[880,1256,1982,2007]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="laciniata">
<emphasis id="7FCA033C4D3E57750327F814FC09E2DD" box="[880,1014,1982,2006]" italics="true" pageId="2" pageNumber="171">L. laciniata</emphasis>
Sakai, 1936 (
<collectingCountry id="35A99FBE4D3E577504CCF815FB1FE2DC" box="[1179,1248,1982,2007]" name="Japan" pageId="2" pageNumber="171">Japan</collectingCountry>
)
</taxonomicName>
,
<taxonomicName id="8ABEA4AD4D3E577404AFF814FD7FE5BB" authority="Richer de Forges &amp; Ng, 2007" authorityName="Richer de Forges &amp; Ng" authorityYear="2007" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" lastPageId="3" lastPageNumber="172" order="Decapoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="mariveneae">
<emphasis id="7FCA033C4D3E577504AFF814FA63E2DD" box="[1272,1436,1983,2006]" italics="true" pageId="2" pageNumber="171">L. mariveneae</emphasis>
Richer de Forges &amp; Ng, 2007 (
<collectingCountry id="35A99FBE4D3F577401A3FF3CFD87E5BB" box="[500,632,151,176]" name="Philippines" pageId="3" pageNumber="172">Philippines</collectingCountry>
)
</taxonomicName>
,
<taxonomicName id="8ABEA4AD4D3F577402DBFF32FBF4E5BB" authority="Dai &amp; Chen, 1980" authorityName="Dai &amp; Chen" authorityYear="1980" box="[652,1035,151,176]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="172" phylum="Arthropoda" rank="species" species="tetraspinosa">
<emphasis id="7FCA033C4D3F577402DBFF32FCCAE5A4" box="[652,821,152,176]" italics="true" pageId="3" pageNumber="172">L. tetraspinosa</emphasis>
<bibRefCitation id="292FA2DF4D3F5774036BFF3CFBF4E5BB" author="Dai, A. Y. &amp; Chen, H. L." box="[828,1035,151,176]" pageId="3" pageNumber="172" pagination="39 - 41" refId="ref6342" refString="Dai, A. Y. &amp; Chen, H. L. (1980) One new species of Latreillopsis from South China Sea. Acta Zootaxonomica Sinica, 5 (1), 39 - 41." type="journal article" year="1980">Dai &amp; Chen, 1980</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="35A99FBE4D3F5774044DFF3CFBA3E5BB" box="[1050,1116,151,176]" name="Japan" pageId="3" pageNumber="172">Japan</collectingCountry>
to
<collectingCountry id="35A99FBE4D3F577404D6FF3CFABEE5BB" box="[1153,1345,151,176]" name="Solomon Islands" pageId="3" pageNumber="172">Solomon Islands</collectingCountry>
), and
<taxonomicName id="8ABEA4AD4D3F577405D2FF32FD09E5DF" authority="Richer de Forges &amp; Ng, 2008" authorityName="Richer de Forges &amp; Ng" authorityYear="2008" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="172" phylum="Arthropoda" rank="species" species="trispinosa">
<emphasis id="7FCA033C4D3F577405D2FF32FEF8E5D8" italics="true" pageId="3" pageNumber="172">L. trispinosa</emphasis>
Richer de Forges &amp;
<bibRefCitation id="292FA2DF4D3F577401A5FF10FDA5E5DF" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[498,602,187,212]" pageId="3" pageNumber="172" pagination="1 - 286" refId="ref6844" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286." type="journal article" year="2008">Ng, 2008</bibRefCitation>
(
<collectingCountry id="35A99FBE4D3F5774023EFF10FD11E5DF" box="[617,750,187,212]" name="Philippines" pageId="3" pageNumber="172">Philippines</collectingCountry>
)
</taxonomicName>
. “
<taxonomicName id="8ABEA4AD4D3F5774035AFF10FC3CE5DF" box="[781,963,187,212]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="170" phylum="Arthropoda" rank="species" species="aff">
<emphasis id="7FCA033C4D3F5774035AFF10FC65E5D8" box="[781,922,187,211]" italics="true" pageId="3" pageNumber="172">Latreillopsis</emphasis>
aff
</taxonomicName>
.
<emphasis id="7FCA033C4D3F57740399FF17FBB2E5D8" box="[974,1101,188,211]" italics="true" pageId="3" pageNumber="172">tetrapinosa</emphasis>
” (
<bibRefCitation id="292FA2DF4D3F5774043FFF10FEF5E5FC" author="Guinot, D. &amp; Richer de Forges, B." pageId="3" pageNumber="172" pagination="283 - 517" refId="ref6581" refString="Guinot, D. &amp; Richer de Forges, B. (1995) Crustacea Decapoda Brachyura: Revision de la famille des Homolidae de Haan, 1839. In: Crosnier, A. (Ed.), Resultats des campagnes MUSORSTOM, vol. 13. Memoires du Museum national d'Histoire naturelle (Paris), 163, pp. 283 - 517." type="journal article" year="1995">Guinot &amp; Richer de Forges 1995: 406</bibRefCitation>
, figs.
<quantity id="8A4672CB4D3F5774011AFF75FE86E5FC" box="[333,377,222,247]" metricMagnitude="-2" metricUnit="kg" metricValue="3.9" pageId="3" pageNumber="172" unit="g" value="39.0">39g</quantity>
, 42B, 43B) from Durban,
<collectingCountry id="35A99FBE4D3F577402F6FF75FCCFE5FC" box="[673,816,222,247]" name="South Africa" pageId="3" pageNumber="172">South Africa</collectingCountry>
may represent a separate species.
</paragraph>
</subSubSection>
<caption id="19C18FA64D3F577400C0F814FE3FE2F8" ID-DOI="http://doi.org/10.5281/zenodo.5045614" ID-Zenodo-Dep="5045614" httpUri="https://zenodo.org/record/5045614/files/figure.png" pageId="3" pageNumber="172" startId="3.[151,249,1983,2005]" targetBox="[164,1423,298,1961]" targetPageId="3">
<paragraph id="4D01DF2E4D3F577400C0F814FE3FE2F8" blockId="3.[151,1436,1983,2035]" pageId="3" pageNumber="172">
<emphasis id="7FCA033C4D3F577400C0F814FEECE2DF" bold="true" box="[151,275,1983,2005]" pageId="3" pageNumber="172">FIGURE 1.</emphasis>
<taxonomicName id="8ABEA4AD4D3F5774014DF814FE28E2DE" authority="Castro &amp; Naruse, 2014" authorityName="Castro &amp; Naruse" authorityYear="2014" box="[282,471,1983,2005]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="172" phylum="Arthropoda" rank="species" species="okala" status="sp. nov.">
<emphasis id="7FCA033C4D3F5774014DF814FE28E2DE" box="[282,471,1983,2005]" italics="true" pageId="3" pageNumber="172">Latreillopsis okala</emphasis>
</taxonomicName>
<emphasis id="7FCA033C4D3F57740188F814FDE7E2DF" bold="true" box="[479,536,1983,2004]" pageId="3" pageNumber="172">
<taxonomicNameLabel id="64F9BE474D3F57740188F814FDE7E2DF" box="[479,536,1983,2004]" pageId="3" pageNumber="172" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, habitus, dorsal view; b, color in life (photograph by P. Fiene).
</paragraph>
</caption>
<caption id="19C18FA64D38577300C0FAABFD0AE03E" ID-DOI="http://doi.org/10.5281/zenodo.5045616" ID-Zenodo-Dep="5045616" httpUri="https://zenodo.org/record/5045616/files/figure.png" pageId="4" pageNumber="173" startId="4.[151,249,1280,1302]" targetBox="[151,1436,193,1259]" targetPageId="4">
<paragraph id="4D01DF2E4D38577300C0FAABFD0AE03E" blockId="4.[151,1435,1280,1333]" pageId="4" pageNumber="173">
<emphasis id="7FCA033C4D38577300C0FAABFEECE01E" bold="true" box="[151,275,1280,1302]" pageId="4" pageNumber="173">FIGURE 2.</emphasis>
<taxonomicName id="8ABEA4AD4D385773014CFAABFE26E01D" authority="Castro &amp; Naruse, 2014" authorityName="Castro &amp; Naruse" authorityYear="2014" box="[283,473,1280,1302]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="173" phylum="Arthropoda" rank="species" species="okala" status="sp. nov.">
<emphasis id="7FCA033C4D385773014CFAABFE26E01D" box="[283,473,1280,1302]" italics="true" pageId="4" pageNumber="173">Latreillopsis okala</emphasis>
</taxonomicName>
<emphasis id="7FCA033C4D38577301B6FAABFDE4E01E" bold="true" box="[481,539,1280,1301]" pageId="4" pageNumber="173">
<taxonomicNameLabel id="64F9BE474D38577301B6FAABFDE4E01E" box="[481,539,1280,1301]" pageId="4" pageNumber="173" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
species. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, cephalothorax, ventral view; b, outer surface of propodus and dactylus, outer view.
</paragraph>
</caption>
<subSubSection id="05A48CA54D3857710090FAF7FEB6E736" lastPageId="6" lastPageNumber="175" pageId="4" pageNumber="173" type="diagnosis">
<paragraph id="4D01DF2E4D3857710090FAF7FBAAE46E" blockId="4.[151,1436,1372,2009]" lastBlockId="6.[151,1436,151,645]" lastPageId="6" lastPageNumber="175" pageId="4" pageNumber="173">
Diagnostic for
<taxonomicName id="8ABEA4AD4D3857730127FAF6FE33E07F" authority="Castro &amp; Naruse, 2014" authorityName="Castro &amp; Naruse" authorityYear="2014" box="[368,460,1372,1396]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="173" phylum="Arthropoda" rank="species" species="okala" status="sp. nov.">
<emphasis id="7FCA033C4D3857730127FAF6FE33E07F" box="[368,460,1372,1396]" italics="true" pageId="4" pageNumber="173">L. okala</emphasis>
</taxonomicName>
<emphasis id="7FCA033C4D3857730183FAF7FDECE07E" bold="true" box="[468,531,1372,1397]" pageId="4" pageNumber="173">
<taxonomicNameLabel id="64F9BE474D3857730183FAF7FDECE07E" box="[468,531,1372,1397]" pageId="4" pageNumber="173" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is the presence of conspicuous tubercles on the carapace and pereopods (
<figureCitation id="D585C3AB4D3857730507FAF7FA6FE07E" box="[1360,1424,1372,1397]" captionStart="FIGURE 1" captionStartId="3.[151,249,1983,2005]" captionTargetBox="[164,1423,298,1961]" captionTargetId="figure-95@3.[164,1423,298,1961]" captionTargetPageId="3" captionText="FIGURE 1. Latreillopsis okala n. sp. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, habitus, dorsal view; b, color in life (photograph by P. Fiene)." figureDoi="http://doi.org/10.5281/zenodo.5045614" httpUri="https://zenodo.org/record/5045614/files/figure.png" pageId="4" pageNumber="173">Fig. 1</figureCitation>
) in contrast to the relatively smooth surface of congeners. The new species belongs to the group of four species of
<taxonomicName id="8ABEA4AD4D38577300C0FA0FFEDBE0B7" authorityName="Henderson" authorityYear="1888" box="[151,292,1444,1468]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="173" phylum="Arthropoda" rank="genus">
<emphasis id="7FCA033C4D38577300C0FA0FFEDBE0B7" box="[151,292,1444,1468]" italics="true" pageId="4" pageNumber="173">Latreillopsis</emphasis>
</taxonomicName>
having weakly-developed accessory teeth or spines on the pseudorostal spines (see
<bibRefCitation id="292FA2DF4D385773048AFA0FFE81E0EA" author="Guinot, D. &amp; Richer de Forges, B." pageId="4" pageNumber="173" pagination="283 - 517" refId="ref6581" refString="Guinot, D. &amp; Richer de Forges, B. (1995) Crustacea Decapoda Brachyura: Revision de la famille des Homolidae de Haan, 1839. In: Crosnier, A. (Ed.), Resultats des campagnes MUSORSTOM, vol. 13. Memoires du Museum national d'Histoire naturelle (Paris), 163, pp. 283 - 517." type="journal article" year="1995">Guinot &amp; Richer de Forges 1995: 394</bibRefCitation>
). The new species differs from
<taxonomicName id="8ABEA4AD4D385773028DFA62FC7EE0EB" authorityName="Dai &amp; Chen" authorityYear="1980" box="[730,897,1481,1504]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="173" phylum="Arthropoda" rank="species" species="tetraspinosa">
<emphasis id="7FCA033C4D385773028DFA62FC7EE0EB" box="[730,897,1481,1504]" italics="true" pageId="4" pageNumber="173">L. tetraspinosa</emphasis>
</taxonomicName>
by having three short subhepatic teeth (
<figureCitation id="D585C3AB4D385773056BFA63FA72E0EA" box="[1340,1421,1480,1505]" captionStart="FIGURE 3" captionStartId="5.[151,249,1877,1899]" captionTargetBox="[180,1414,213,1841]" captionTargetId="figure-71@5.[172,1414,193,1856]" captionTargetPageId="5" captionText="FIGURE 3. Latreillopsis okala n. sp. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, cephalothorax, dorsal view (wartlike granules are not drawn); b, left anterior part of ventral surface of cephalothorax (wart-like granules are not drawn); c, left G1, ventral view; d, left G2, ventral view. Scales: ad = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5045618" httpUri="https://zenodo.org/record/5045618/files/figure.png" pageId="4" pageNumber="173">Fig. 3b</figureCitation>
), one rounded tubercle on the third maxilliped merus (
<figureCitation id="D585C3AB4D38577302A7FA47FCBCE30E" box="[752,835,1516,1541]" captionStart="FIGURE 3" captionStartId="5.[151,249,1877,1899]" captionTargetBox="[180,1414,213,1841]" captionTargetId="figure-71@5.[172,1414,193,1856]" captionTargetPageId="5" captionText="FIGURE 3. Latreillopsis okala n. sp. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, cephalothorax, dorsal view (wartlike granules are not drawn); b, left anterior part of ventral surface of cephalothorax (wart-like granules are not drawn); c, left G1, ventral view; d, left G2, ventral view. Scales: ad = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5045618" httpUri="https://zenodo.org/record/5045618/files/figure.png" pageId="4" pageNumber="173">Fig. 3b</figureCitation>
) and anterolateral margin of third maxillipeds merus that is sharp but not pointed (
<figureCitation id="D585C3AB4D38577301B4F9BBFDCAE322" box="[483,565,1552,1577]" captionStart="FIGURE 3" captionStartId="5.[151,249,1877,1899]" captionTargetBox="[180,1414,213,1841]" captionTargetId="figure-71@5.[172,1414,193,1856]" captionTargetPageId="5" captionText="FIGURE 3. Latreillopsis okala n. sp. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, cephalothorax, dorsal view (wartlike granules are not drawn); b, left anterior part of ventral surface of cephalothorax (wart-like granules are not drawn); c, left G1, ventral view; d, left G2, ventral view. Scales: ad = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5045618" httpUri="https://zenodo.org/record/5045618/files/figure.png" pageId="4" pageNumber="173">Fig. 3b</figureCitation>
), in contrast to four subhepatic teeth, 2 or 3 tubercles on the third maxilliped merus and ischium, and pointed, salient anterolateral margin of third maxillipeds merus in
<taxonomicName id="8ABEA4AD4D38577304CCF99EFABAE347" authorityName="Dai &amp; Chen" authorityYear="1980" box="[1179,1349,1589,1612]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="173" phylum="Arthropoda" rank="species" species="tetraspinosa">
<emphasis id="7FCA033C4D38577304CCF99EFABAE347" box="[1179,1349,1589,1612]" italics="true" pageId="4" pageNumber="173">L. tetraspinosa</emphasis>
</taxonomicName>
(
<bibRefCitation id="292FA2DF4D3857730503F99FFEE9E37A" author="Dai, A. Y. &amp; Chen, H. L." pageId="4" pageNumber="173" pagination="39 - 41" refId="ref6342" refString="Dai, A. Y. &amp; Chen, H. L. (1980) One new species of Latreillopsis from South China Sea. Acta Zootaxonomica Sinica, 5 (1), 39 - 41." type="journal article" year="1980">Dai &amp; Chen 1980</bibRefCitation>
: fig. 1;
<bibRefCitation id="292FA2DF4D385773013AF9F3FD19E37A" author="Guinot, D. &amp; Richer de Forges, B." box="[365,742,1624,1649]" pageId="4" pageNumber="173" pagination="283 - 517" refId="ref6581" refString="Guinot, D. &amp; Richer de Forges, B. (1995) Crustacea Decapoda Brachyura: Revision de la famille des Homolidae de Haan, 1839. In: Crosnier, A. (Ed.), Resultats des campagnes MUSORSTOM, vol. 13. Memoires du Museum national d'Histoire naturelle (Paris), 163, pp. 283 - 517." type="journal article" year="1995">Guinot &amp; Richer de Forges 1995</bibRefCitation>
: figs. 42C, D). The new species can be differentiated from
<taxonomicName id="8ABEA4AD4D38577305D2F9F2FEFCE39F" authorityName="Henderson" authorityYear="1888" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="173" phylum="Arthropoda" rank="species" species="bispinosa">
<emphasis id="7FCA033C4D38577305D2F9F2FEFCE39F" italics="true" pageId="4" pageNumber="173">L. bispinosa</emphasis>
</taxonomicName>
by having three short subhepatic teeth and a rostrum that is slightly shorter than the pseudorostral spines (
<figureCitation id="D585C3AB4D38577300C8F90BFEECE3B2" box="[159,275,1696,1721]" captionStart="FIGURE 3" captionStartId="5.[151,249,1877,1899]" captionTargetBox="[180,1414,213,1841]" captionTargetId="figure-71@5.[172,1414,193,1856]" captionTargetPageId="5" captionText="FIGURE 3. Latreillopsis okala n. sp. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, cephalothorax, dorsal view (wartlike granules are not drawn); b, left anterior part of ventral surface of cephalothorax (wart-like granules are not drawn); c, left G1, ventral view; d, left G2, ventral view. Scales: ad = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5045618" httpUri="https://zenodo.org/record/5045618/files/figure.png" pageId="4" pageNumber="173">Fig. 3a, b</figureCitation>
) in contrast with two long subhepatic teeth and pseudorostral spines that are much longer than the rostrum in
<taxonomicName id="8ABEA4AD4D3857730145F96EFE65E3D7" authorityName="Henderson" authorityYear="1888" box="[274,410,1732,1756]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="173" phylum="Arthropoda" rank="species" species="bispinosa">
<emphasis id="7FCA033C4D3857730145F96EFE65E3D7" box="[274,410,1732,1756]" italics="true" pageId="4" pageNumber="173">L. bispinosa</emphasis>
</taxonomicName>
(
<bibRefCitation id="292FA2DF4D38577301FEF96FFD6FE3D6" author="Henderson, J. R." box="[425,656,1732,1757]" pageId="4" pageNumber="173" pagination="1 - 221" refId="ref6717" refString="Henderson, J. R. (1888) Report on the Anomura collected by H. M. S. Challenger during the years 1873 - 1876. In: Wyville Thomson, C. &amp; Murray, J. (Eds.), Report of the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 76 Under the Command of Captain George S. Nares, R. N., F. R. S. and the late Captain Frank Tourle Thompson, R. N., Zoology, 27 (69), i - xi + pp. 1 - 221 pp., pls. 1 - 21." type="book chapter" year="1888">Henderson 1888: 22</bibRefCitation>
, pl. 2, fig. 3;
<bibRefCitation id="292FA2DF4D3857730371F96FFB65E3D6" author="Guinot, D. &amp; Richer de Forges, B." box="[806,1178,1732,1757]" pageId="4" pageNumber="173" pagination="283 - 517" refId="ref6581" refString="Guinot, D. &amp; Richer de Forges, B. (1995) Crustacea Decapoda Brachyura: Revision de la famille des Homolidae de Haan, 1839. In: Crosnier, A. (Ed.), Resultats des campagnes MUSORSTOM, vol. 13. Memoires du Museum national d'Histoire naturelle (Paris), 163, pp. 283 - 517." type="journal article" year="1995">Guinot &amp; Richer de Forges 1995</bibRefCitation>
: figs.35a, b, 41B, 42A, A1), and from
<taxonomicName id="8ABEA4AD4D385773011AF942FE1FE20B" authorityName="Guinot &amp; Richer de Forges" authorityYear="1981" box="[333,480,1768,1792]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="173" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="7FCA033C4D385773011AF942FE1FE20B" box="[333,480,1768,1792]" italics="true" pageId="4" pageNumber="173">L. gracilipes</emphasis>
</taxonomicName>
and
<taxonomicName id="8ABEA4AD4D3857730276F942FD77E20B" authorityName="Guinot &amp; Richer de Forges" authorityYear="1995" box="[545,648,1768,1792]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="173" phylum="Arthropoda" rank="species" species="daviei">
<emphasis id="7FCA033C4D3857730276F942FD77E20B" box="[545,648,1768,1792]" italics="true" pageId="4" pageNumber="173">L. daviei</emphasis>
</taxonomicName>
by not having the swollen subhepatic region and the conspicuous lateroposterior tooth of both species (
<bibRefCitation id="292FA2DF4D385773026FF8A7FCE0E22E" author="Davie, P. J. F. &amp; Short, J. W." box="[568,799,1804,1829]" pageId="4" pageNumber="173" pagination="157 - 187" refId="ref6381" refString="Davie, P. J. F. &amp; Short, J. W. (1989) Deepwater Brachyura (Crustacea: Decapoda) from southern Queensland, Australia with descriptions of four new species. Memoirs of the Queensland Museum, 27 (2), 157 - 187." type="journal article" year="1989">Davie &amp; Short 1989</bibRefCitation>
: fig. 3A, as
<taxonomicName id="8ABEA4AD4D38577303F1F8A6FBD1E22F" authorityName="Henderson" authorityYear="1888" box="[934,1070,1804,1828]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="173" phylum="Arthropoda" rank="species" species="bispinosa">
<emphasis id="7FCA033C4D38577303F1F8A6FBD1E22F" box="[934,1070,1804,1828]" italics="true" pageId="4" pageNumber="173">L. bispinosa</emphasis>
</taxonomicName>
, and
<bibRefCitation id="292FA2DF4D385773043BF8A7FF2CE242" author="Guinot, D. &amp; Richer de Forges, B." pageId="4" pageNumber="173" pagination="283 - 517" refId="ref6581" refString="Guinot, D. &amp; Richer de Forges, B. (1995) Crustacea Decapoda Brachyura: Revision de la famille des Homolidae de Haan, 1839. In: Crosnier, A. (Ed.), Resultats des campagnes MUSORSTOM, vol. 13. Memoires du Museum national d'Histoire naturelle (Paris), 163, pp. 283 - 517." type="journal article" year="1995">Guinot &amp; Richer de Forges 1995</bibRefCitation>
: figs. 34, 40a, b for
<taxonomicName id="8ABEA4AD4D38577301E7F89AFDEEE243" authorityName="Guinot &amp; Richer de Forges" authorityYear="1995" box="[432,529,1840,1864]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="173" phylum="Arthropoda" rank="species" species="daviei">
<emphasis id="7FCA033C4D38577301E7F89AFDEEE243" box="[432,529,1840,1864]" italics="true" pageId="4" pageNumber="173">L. daviei</emphasis>
</taxonomicName>
;
<bibRefCitation id="292FA2DF4D3857730277F89BFC46E242" author="Guinot, D. &amp; Richer de Forges, B." box="[544,953,1840,1865]" pageId="4" pageNumber="173" pagination="283 - 517" refId="ref6581" refString="Guinot, D. &amp; Richer de Forges, B. (1995) Crustacea Decapoda Brachyura: Revision de la famille des Homolidae de Haan, 1839. In: Crosnier, A. (Ed.), Resultats des campagnes MUSORSTOM, vol. 13. Memoires du Museum national d'Histoire naturelle (Paris), 163, pp. 283 - 517." type="journal article" year="1995">Guinot &amp; Richer de Forges 1995: 38</bibRefCitation>
b, and Richer de Forges &amp;
<bibRefCitation id="292FA2DF4D38577304B3F89AFAB6E242" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[1252,1353,1840,1865]" pageId="4" pageNumber="173" pagination="1 - 286" refId="ref6844" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286." type="journal article" year="2008">Ng 2008</bibRefCitation>
: fig. 23 for
<taxonomicName id="8ABEA4AD4D38577300E8F8FEFEB3E267" authorityName="Guinot &amp; Richer de Forges" authorityYear="1981" box="[191,332,1876,1900]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="173" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="7FCA033C4D38577300E8F8FEFEB3E267" box="[191,332,1876,1900]" italics="true" pageId="4" pageNumber="173">L. gracilipes</emphasis>
</taxonomicName>
).
<taxonomicName id="8ABEA4AD4D3857730132F8FFFD96E267" authorityName="Guinot &amp; Richer de Forges" authorityYear="1981" box="[357,617,1876,1900]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="173" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="7FCA033C4D3857730132F8FFFD96E267" box="[357,617,1876,1900]" italics="true" pageId="4" pageNumber="173">Latreillopsis gracilipes</emphasis>
</taxonomicName>
is also characterized by having three subhepatic teeth and one “granule” (
<bibRefCitation id="292FA2DF4D38577300C9F8D3FDB1E29A" author="Guinot, D. &amp; Richer de Forges, B." box="[158,590,1912,1937]" pageId="4" pageNumber="173" pagination="283 - 517" refId="ref6581" refString="Guinot, D. &amp; Richer de Forges, B. (1995) Crustacea Decapoda Brachyura: Revision de la famille des Homolidae de Haan, 1839. In: Crosnier, A. (Ed.), Resultats des campagnes MUSORSTOM, vol. 13. Memoires du Museum national d'Histoire naturelle (Paris), 163, pp. 283 - 517." type="journal article" year="1995">Guinot &amp; Richer de Forges 1995: 402</bibRefCitation>
). The new species can be differentiated from
<taxonomicName id="8ABEA4AD4D385773041CF8D2FB26E29B" authorityName="Richer de Forges &amp; Ng" authorityYear="2008" box="[1099,1241,1913,1936]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="173" phylum="Arthropoda" rank="species" species="trispinosa">
<emphasis id="7FCA033C4D385773041CF8D2FB26E29B" box="[1099,1241,1913,1936]" italics="true" pageId="4" pageNumber="173">L. trispinosa</emphasis>
</taxonomicName>
by its three short subhepatic teeth, third maxilliped merus with a large, rounded anterior tubercle and sharp anteroexternal angle (
<figureCitation id="D585C3AB4D38577300C8F86BFEF4E2D2" box="[159,267,1984,2009]" captionStart="FIGURE 3" captionStartId="5.[151,249,1877,1899]" captionTargetBox="[180,1414,213,1841]" captionTargetId="figure-71@5.[172,1414,193,1856]" captionTargetPageId="5" captionText="FIGURE 3. Latreillopsis okala n. sp. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, cephalothorax, dorsal view (wartlike granules are not drawn); b, left anterior part of ventral surface of cephalothorax (wart-like granules are not drawn); c, left G1, ventral view; d, left G2, ventral view. Scales: ad = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5045618" httpUri="https://zenodo.org/record/5045618/files/figure.png" pageId="4" pageNumber="173">Fig. 3a, b</figureCitation>
) in contrast to three conspicuously long subhepatic teeth and smooth third maxilliped merus with a rounded anteroexternal angle in
<taxonomicName id="8ABEA4AD4D3A5771025EFF32FD66E5A4" authorityName="Richer de Forges &amp; Ng" authorityYear="2008" box="[521,665,152,176]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="175" phylum="Arthropoda" rank="species" species="trispinosa">
<emphasis id="7FCA033C4D3A5771025EFF32FD66E5A4" box="[521,665,152,176]" italics="true" pageId="6" pageNumber="175">L. trispinosa</emphasis>
</taxonomicName>
(
<bibRefCitation id="292FA2DF4D3A577102FEFF3CFBD7E5BB" author="Guinot, D. &amp; Richer de Forges, B." box="[681,1064,151,176]" pageId="6" pageNumber="175" pagination="283 - 517" refId="ref6581" refString="Guinot, D. &amp; Richer de Forges, B. (1995) Crustacea Decapoda Brachyura: Revision de la famille des Homolidae de Haan, 1839. In: Crosnier, A. (Ed.), Resultats des campagnes MUSORSTOM, vol. 13. Memoires du Museum national d'Histoire naturelle (Paris), 163, pp. 283 - 517." type="journal article" year="1995">Guinot &amp; Richer de Forges 1995</bibRefCitation>
: fig. 41A, as
<taxonomicName id="8ABEA4AD4D3A57710492FF32FAAFE5BB" authorityName="Henderson" authorityYear="1888" box="[1221,1360,152,176]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="175" phylum="Arthropoda" rank="species" species="bispinosa">
<emphasis id="7FCA033C4D3A57710492FF32FAAFE5BB" box="[1221,1360,152,176]" italics="true" pageId="6" pageNumber="175">L. bispinosa</emphasis>
</taxonomicName>
forme
<taxonomicName id="8ABEA4AD4D3A577100C0FF16FEF8E5DF" authorityName="Richer de Forges &amp; Ng" authorityYear="2008" box="[151,263,189,212]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="175" phylum="Arthropoda" rank="species" species="trispinosa">
<emphasis id="7FCA033C4D3A577100C0FF16FEF8E5DF" box="[151,263,189,212]" italics="true" pageId="6" pageNumber="175">trispinosa</emphasis>
</taxonomicName>
). The anteroexternal angles of the third maxillipeds in the specimens photographed by Richer de Forges &amp;
<bibRefCitation id="292FA2DF4D3A577100E2FF4BFEDAE5F3" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[181,293,223,248]" pageId="6" pageNumber="175" pagination="1 - 286" refId="ref6844" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286." type="journal article" year="2008">Ng (2008</bibRefCitation>
: figs. 24C, 25C), however, are sharp but the angle was described as rounded by
<bibRefCitation id="292FA2DF4D3A577104EDFF74FE86E416" author="Guinot, D. &amp; Richer de Forges, B." pageId="6" pageNumber="175" pagination="283 - 517" refId="ref6581" refString="Guinot, D. &amp; Richer de Forges, B. (1995) Crustacea Decapoda Brachyura: Revision de la famille des Homolidae de Haan, 1839. In: Crosnier, A. (Ed.), Resultats des campagnes MUSORSTOM, vol. 13. Memoires du Museum national d'Histoire naturelle (Paris), 163, pp. 283 - 517." type="journal article" year="1995">Guinot &amp; Richer de Forges (1995: 399)</bibRefCitation>
. Furthermore, the ocular peduncle, described as short (“
<emphasis id="7FCA033C4D3A57710458FEAFFA87E417" box="[1039,1400,260,284]" italics="true" pageId="6" pageNumber="175">beaucoup plus courts que chez</emphasis>
<taxonomicName id="8ABEA4AD4D3A577105D3FEAFFEDEE44B" authorityName="Dai &amp; Chen" authorityYear="1980" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="175" phylum="Arthropoda" rank="species" species="tetraspinosa">L. tetraspinosa</taxonomicName>
”) by
<bibRefCitation id="292FA2DF4D3A57710134FE8CFCD5E44B" author="Guinot, D. &amp; Richer de Forges, B." box="[355,810,295,320]" pageId="6" pageNumber="175" pagination="283 - 517" refId="ref6581" refString="Guinot, D. &amp; Richer de Forges, B. (1995) Crustacea Decapoda Brachyura: Revision de la famille des Homolidae de Haan, 1839. In: Crosnier, A. (Ed.), Resultats des campagnes MUSORSTOM, vol. 13. Memoires du Museum national d'Histoire naturelle (Paris), 163, pp. 283 - 517." type="journal article" year="1995">Guinot &amp; Richer de Forges (1995: 399</bibRefCitation>
, fig. 41A, as
<taxonomicName id="8ABEA4AD4D3A5771039BFE82FBA7E44B" authorityName="Henderson" authorityYear="1888" box="[972,1112,296,320]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="175" phylum="Arthropoda" rank="species" species="bispinosa">
<emphasis id="7FCA033C4D3A5771039BFE82FBA7E44B" box="[972,1112,296,320]" italics="true" pageId="6" pageNumber="175">L. bispinosa</emphasis>
</taxonomicName>
forme
<taxonomicName id="8ABEA4AD4D3A577104F8FE83FAE0E434" authorityName="Richer de Forges &amp; Ng" authorityYear="2008" box="[1199,1311,296,319]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="175" phylum="Arthropoda" rank="species" species="trispinosa">
<emphasis id="7FCA033C4D3A577104F8FE83FAE0E434" box="[1199,1311,296,319]" italics="true" pageId="6" pageNumber="175">trispinosa</emphasis>
</taxonomicName>
) is clearly longer in the specimens photographed by Richer de Forges &amp;
<bibRefCitation id="292FA2DF4D3A5771031AFEE7FC43E46E" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[845,956,332,357]" pageId="6" pageNumber="175" pagination="1 - 286" refId="ref6844" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286." type="journal article" year="2008">Ng (2008</bibRefCitation>
: figs. 24, 25).
</paragraph>
<caption id="19C18FA64D39577200C0F8FEFCEDE2A3" ID-DOI="http://doi.org/10.5281/zenodo.5045618" ID-Zenodo-Dep="5045618" httpUri="https://zenodo.org/record/5045618/files/figure.png" pageId="5" pageNumber="174" startId="5.[151,249,1877,1899]" targetBox="[180,1414,213,1841]" targetPageId="5">
<paragraph id="4D01DF2E4D39577200C0F8FEFCEDE2A3" blockId="5.[151,1436,1877,1961]" pageId="5" pageNumber="174">
<emphasis id="7FCA033C4D39577200C0F8FEFEEDE260" bold="true" box="[151,274,1877,1899]" pageId="5" pageNumber="174">FIGURE 3.</emphasis>
<taxonomicName id="8ABEA4AD4D395772014FF8FEFE2BE260" authority="Castro &amp; Naruse, 2014" authorityName="Castro &amp; Naruse" authorityYear="2014" box="[280,468,1877,1899]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="174" phylum="Arthropoda" rank="species" species="okala" status="sp. nov.">
<emphasis id="7FCA033C4D395772014FF8FEFE2BE260" box="[280,468,1877,1899]" italics="true" pageId="5" pageNumber="174">Latreillopsis okala</emphasis>
</taxonomicName>
<emphasis id="7FCA033C4D395772018DF8FDFDECE260" bold="true" box="[474,531,1878,1899]" pageId="5" pageNumber="174">
<taxonomicNameLabel id="64F9BE474D395772018DF8FDFDECE260" box="[474,531,1878,1899]" pageId="5" pageNumber="174" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, cephalothorax, dorsal view (wartlike granules are not drawn); b, left anterior part of ventral surface of cephalothorax (wart-like granules are not drawn); c, left G1, ventral view; d, left G2, ventral view. Scales: ad = 1 mm.
</paragraph>
</caption>
<paragraph id="4D01DF2E4D3A57710090FEC4FEB6E736" blockId="6.[151,1436,151,645]" pageId="6" pageNumber="175">
There is limited information on the morphology of the G1 and G2 among the species of
<taxonomicName id="8ABEA4AD4D3A577104FBFEDBFAC7E483" authorityName="Henderson" authorityYear="1888" box="[1196,1336,368,392]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="175" phylum="Arthropoda" rank="genus">
<emphasis id="7FCA033C4D3A577104FBFEDBFAC7E483" box="[1196,1336,368,392]" italics="true" pageId="6" pageNumber="175">Latreillopsis</emphasis>
</taxonomicName>
. The G1 of
<taxonomicName id="8ABEA4AD4D3A577100E3FE3EFEF0E4A7" authority="Castro &amp; Naruse, 2014" authorityName="Castro &amp; Naruse" authorityYear="2014" box="[180,271,404,428]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="175" phylum="Arthropoda" rank="species" species="okala" status="sp. nov.">
<emphasis id="7FCA033C4D3A577100E3FE3EFEF0E4A7" box="[180,271,404,428]" italics="true" pageId="6" pageNumber="175">L. okala</emphasis>
</taxonomicName>
<emphasis id="7FCA033C4D3A57710141FE3FFEABE4A6" bold="true" box="[278,340,404,429]" pageId="6" pageNumber="175">
<taxonomicNameLabel id="64F9BE474D3A57710141FE3FFEABE4A6" box="[278,340,404,429]" pageId="6" pageNumber="175" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is triangular (
<figureCitation id="D585C3AB4D3A577101B8FE3FFDC0E4A6" box="[495,575,404,429]" captionStart="FIGURE 3" captionStartId="5.[151,249,1877,1899]" captionTargetBox="[180,1414,213,1841]" captionTargetId="figure-71@5.[172,1414,193,1856]" captionTargetPageId="5" captionText="FIGURE 3. Latreillopsis okala n. sp. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, cephalothorax, dorsal view (wartlike granules are not drawn); b, left anterior part of ventral surface of cephalothorax (wart-like granules are not drawn); c, left G1, ventral view; d, left G2, ventral view. Scales: ad = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5045618" httpUri="https://zenodo.org/record/5045618/files/figure.png" pageId="6" pageNumber="175">Fig. 3c</figureCitation>
), but straight, with rounded apex in
<taxonomicName id="8ABEA4AD4D3A57710386FE3EFBA1E4A7" authorityName="Guinot &amp; Richer de Forges" authorityYear="1981" box="[977,1118,404,428]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="175" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="7FCA033C4D3A57710386FE3EFBA1E4A7" box="[977,1118,404,428]" italics="true" pageId="6" pageNumber="175">L. gracilipes</emphasis>
</taxonomicName>
(
<bibRefCitation id="292FA2DF4D3A5771043BFE3FFF2CE4DB" author="Guinot, D. &amp; Richer de Forges, B." pageId="6" pageNumber="175" pagination="523 - 581" refId="ref6524" refString="Guinot, D. &amp; Richer de Forges, B. (1981) Homolidae, rares ou nouveaux, de l'Indo-Pacifique (Crustacea, Decapoda, Brachyura). Bulletin du Museum national d'Histoire naturelle (Paris), ser. 4, 3, sect. A (2), 523 - 581." type="journal article" year="1981">Guinot &amp; Richer de Forges 1981</bibRefCitation>
: fig. 6C) and
<taxonomicName id="8ABEA4AD4D3A5771013DFE12FE34E4DB" authorityName="Guinot &amp; Richer de Forges" authorityYear="1995" box="[362,459,440,464]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="175" phylum="Arthropoda" rank="species" species="daviei">
<emphasis id="7FCA033C4D3A5771013DFE12FE34E4DB" box="[362,459,440,464]" italics="true" pageId="6" pageNumber="175">L. daviei</emphasis>
</taxonomicName>
(
<bibRefCitation id="292FA2DF4D3A5771018DFE1CFD3DE4DB" author="Davie, P. J. F. &amp; Short, J. W." box="[474,706,439,464]" pageId="6" pageNumber="175" pagination="157 - 187" refId="ref6381" refString="Davie, P. J. F. &amp; Short, J. W. (1989) Deepwater Brachyura (Crustacea: Decapoda) from southern Queensland, Australia with descriptions of four new species. Memoirs of the Queensland Museum, 27 (2), 157 - 187." type="journal article" year="1989">Davie &amp; Short 1989</bibRefCitation>
: fig. 2a, as
<taxonomicName id="8ABEA4AD4D3A57710368FE12FC37E4DB" authorityName="Henderson" authorityYear="1888" box="[831,968,440,464]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="175" phylum="Arthropoda" rank="species" species="bispinosa">
<emphasis id="7FCA033C4D3A57710368FE12FC37E4DB" box="[831,968,440,464]" italics="true" pageId="6" pageNumber="175">L. bispinosa</emphasis>
</taxonomicName>
). The G2, with a truncated distal part that is folded in cross section (
<figureCitation id="D585C3AB4D3A577101E9FE77FDEFE4FE" box="[446,528,476,501]" captionStart="FIGURE 3" captionStartId="5.[151,249,1877,1899]" captionTargetBox="[180,1414,213,1841]" captionTargetId="figure-71@5.[172,1414,193,1856]" captionTargetPageId="5" captionText="FIGURE 3. Latreillopsis okala n. sp. Holotype male 6.0 mm × 4.0 mm (USNM 1220315): a, cephalothorax, dorsal view (wartlike granules are not drawn); b, left anterior part of ventral surface of cephalothorax (wart-like granules are not drawn); c, left G1, ventral view; d, left G2, ventral view. Scales: ad = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5045618" httpUri="https://zenodo.org/record/5045618/files/figure.png" pageId="6" pageNumber="175">Fig. 3d</figureCitation>
), is similar to the G2 of
<taxonomicName id="8ABEA4AD4D3A57710372FE76FC78E4FF" authorityName="Guinot &amp; Richer de Forges" authorityYear="1995" box="[805,903,476,500]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="175" phylum="Arthropoda" rank="species" species="daviei">
<emphasis id="7FCA033C4D3A57710372FE76FC78E4FF" box="[805,903,476,500]" italics="true" pageId="6" pageNumber="175">L. daviei</emphasis>
</taxonomicName>
(
<bibRefCitation id="292FA2DF4D3A577103C0FE77FB7FE4FE" author="Davie, P. J. F. &amp; Short, J. W." box="[919,1152,476,501]" pageId="6" pageNumber="175" pagination="157 - 187" refId="ref6381" refString="Davie, P. J. F. &amp; Short, J. W. (1989) Deepwater Brachyura (Crustacea: Decapoda) from southern Queensland, Australia with descriptions of four new species. Memoirs of the Queensland Museum, 27 (2), 157 - 187." type="journal article" year="1989">Davie &amp; Short 1989</bibRefCitation>
: fig. 2b, as
<taxonomicName id="8ABEA4AD4D3A57710555FE76FA74E4FF" authorityName="Henderson" authorityYear="1888" box="[1282,1419,476,500]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="175" phylum="Arthropoda" rank="species" species="bispinosa">
<emphasis id="7FCA033C4D3A57710555FE76FA74E4FF" box="[1282,1419,476,500]" italics="true" pageId="6" pageNumber="175">L. bispinosa</emphasis>
</taxonomicName>
), but different from the slightly curved and rounded distal part of the G2 of
<taxonomicName id="8ABEA4AD4D3A57710387FDAAFBA2E713" authorityName="Guinot &amp; Richer de Forges" authorityYear="1981" box="[976,1117,512,536]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="175" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="7FCA033C4D3A57710387FDAAFBA2E713" box="[976,1117,512,536]" italics="true" pageId="6" pageNumber="175">L. gracilipes</emphasis>
</taxonomicName>
(
<bibRefCitation id="292FA2DF4D3A5771043CFE54FF2CE736" author="Guinot, D. &amp; Richer de Forges, B." pageId="6" pageNumber="175" pagination="523 - 581" refId="ref6524" refString="Guinot, D. &amp; Richer de Forges, B. (1981) Homolidae, rares ou nouveaux, de l'Indo-Pacifique (Crustacea, Decapoda, Brachyura). Bulletin du Museum national d'Histoire naturelle (Paris), ser. 4, 3, sect. A (2), 523 - 581." type="journal article" year="1981">Guinot &amp; Richer de Forges 1981</bibRefCitation>
: fig. 6C1).
</paragraph>
</subSubSection>
<subSubSection id="05A48CA54D3A57710090FDE3FE8AE78E" pageId="6" pageNumber="175" type="discussion">
<paragraph id="4D01DF2E4D3A57710090FDE3FE8AE78E" blockId="6.[151,1436,151,645]" pageId="6" pageNumber="175">
<treatmentCitationGroup id="6DAEF8004D3A57710090FDE3FE8AE78E" pageId="6" pageNumber="175">
<taxonomicName id="8ABEA4AD4D3A57710090FDE3FE63E76B" ID-CoL="6P4HQ" authority="Castro &amp; Naruse, 2014" authorityName="Castro &amp; Naruse" authorityYear="2014" box="[199,412,584,608]" class="Malacostraca" family="Homolidae" genus="Latreillopsis" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="175" phylum="Arthropoda" rank="species" species="okala" status="sp. nov.">
<emphasis id="7FCA033C4D3A57710090FDE3FE63E76B" box="[199,412,584,608]" italics="true" pageId="6" pageNumber="175">Latreillopsis okala</emphasis>
</taxonomicName>
<emphasis id="7FCA033C4D3A577101F1FDECFE16E76B" bold="true" box="[422,489,583,608]" pageId="6" pageNumber="175">
<taxonomicNameLabel id="64F9BE474D3A577101F1FDECFE16E76B" box="[422,489,583,608]" pageId="6" pageNumber="175" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is the fifth species of
<taxonomicName id="8ABEA4AD4D3A577102A2FDECFC8EE76B" authorityName="H. Milne Edwards" authorityYear="1837" box="[757,881,583,608]" class="Malacostraca" family="Homolidae" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="175" phylum="Arthropoda" rank="family">Homolidae</taxonomicName>
reliably known from the Hawaiian Islands (see
<treatmentCitation id="CC1FF93F4D3A577100C0FDC7FEC2E78E" author="Castro, P." box="[151,317,620,645]" page="9" pageId="6" pageNumber="175" year="2011">
<bibRefCitation id="292FA2DF4D3A577100C0FDC7FEC2E78E" author="Castro, P." box="[151,317,620,645]" pageId="6" pageNumber="175" pagination="1 - 154" refId="ref6255" refString="Castro, P. (2011) Catalog of the anomuran and brachyuran crabs (Crustacea: Decapoda: Anomura, Brachyura) of the Hawaiian Islands. Zootaxa, 2947, 1 - 154." type="journal article" year="2011">Castro 2011: 9</bibRefCitation>
</treatmentCitation>
, 34).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
</treatment>
</document>