337 lines
48 KiB
XML
337 lines
48 KiB
XML
<document id="BE765292475D45EE0DDC46678F80E0C1" ID-CLB-Dataset="21863" ID-DOI="10.5852/ejt.2016.241" ID-GBIF-Dataset="f694842b-3af3-4198-b01e-13d0b5466b5d" ID-ISSN="2118-9773" ID-Zenodo-Dep="3850299" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590443252253" checkinUser="valdenar" docAuthor="Frolov, Andrey V. & Akhmetova, Lilia A." docDate="2016" docId="C4027A0914045559D333F924FA80B7F4" docLanguage="en" docName="ejt-2016-241.pdf.imf" docOrigin="European Journal of Taxonomy 241" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Orphnus (Phornus) Paulian 1948" docType="treatment" docVersion="4" lastPageNumber="4" masterDocId="383B02711405555AD167FFB9FFF7B60D" masterDocTitle="Revision of the subgenus Orphnus (Phornus) (Coleoptera, Scarabaeidae, Orphninae)" masterLastPageNumber="20" masterPageNumber="1" pageNumber="2" updateTime="1698841278433" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="8646CDEFAD5955DD3D2B8DF3816065B2">Revision of the subgenus Orphnus (Phornus) (Coleoptera, Scarabaeidae, Orphninae)</mods:title>
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<mods:namePart id="9E176D0DECBEE239D811AE5A207D6430">Frolov, Andrey V.</mods:namePart>
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<mods:affiliation id="37C3AA0A201043A49FEA1D567B88BDA9">Laboratory of Insect Systematics, Zoological Institute, Russian Academy of Sciences, Universitetskaya nab., 1, St. - Petersburg, 199034 Russia. Federal University of Mato Grosso, Av. Fernando Corrêa da Costa, 2367, Boa Esperança, 78060 - 900 Cuiabá, MT, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="8BF8697A12136ACB4C78D1E0BA3F07F7" type="email">frolov@scarabaeoidea.com</mods:nameIdentifier>
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<mods:namePart id="F9AAE80D997F36E2A4E1CB9562E674C5">Akhmetova, Lilia A.</mods:namePart>
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<mods:affiliation id="EC6AF2F968B1A471C9030DF547DBAD9E">Laboratory of Insect Systematics, Zoological Institute, Russian Academy of Sciences, Universitetskaya nab., 1, St. - Petersburg, 199034 Russia. Federal University of Mato Grosso, Av. Fernando Corrêa da Costa, 2367, Boa Esperança, 78060 - 900 Cuiabá, MT, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="FE3E7F221ABEEC09DFF959F974ED925F" type="email">akhmetova@scarabaeoidea.com</mods:nameIdentifier>
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<mods:title id="2033BEF60AE3C1FBECD5FFFB9992F3A2">European Journal of Taxonomy</mods:title>
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<mods:date id="A4745FF0E9724C77D5883A7927DAB2A6">2016</mods:date>
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<treatment id="C4027A0914045559D333F924FA80B7F4" ID-DOI="http://doi.org/10.5281/zenodo.3854778" ID-GBIF-Taxon="195118202" ID-Zenodo-Dep="3854778" LSID="urn:lsid:plazi:treatment:C4027A0914045559D333F924FA80B7F4" httpUri="http://treatment.plazi.org/id/C4027A0914045559D333F924FA80B7F4" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">
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<subSubSection id="04B198941404555BD333F924FC28B0B5" box="[596,991,1693,1720]" pageId="1" pageNumber="2" type="nomenclature">
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<paragraph id="4C14CB1F1404555BD333F924FC28B0B5" blockId="1.[596,991,1693,1720]" box="[596,991,1693,1720]" pageId="1" pageNumber="2">
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<heading id="175C7C731404555BD333F924FC28B0B5" box="[596,991,1693,1720]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="2" reason="2">
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Subgenus
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<taxonomicName id="8BABB09C1404555BD3A9F924FC28B0B5" authority="Paulian, 1948" authorityName="Paulian" authorityYear="1948" box="[718,991,1693,1720]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="subGenus" subGenus="Phornus">
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<emphasis id="7EDF170D1404555BD3A9F924FCC1B0B5" bold="true" box="[718,822,1693,1720]" italics="true" pageId="1" pageNumber="2">Phornus</emphasis>
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<bibRefCitation id="283AB6EE1404555BD259F924FC28B0B5" author="Paulian R." box="[830,991,1693,1720]" pageId="1" pageNumber="2" pagination="1 - 75" refId="ref8779" refString="Paulian R. 1948. Revision des Orphnus Africains (Coleoptera, Scarabaeidae). Annales de la Societe entomologique de France 117: 1 - 75." type="journal article" year="1948">Paulian, 1948</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="04B198941404555BD1DAF966FE0AB0F7" box="[189,509,1759,1786]" pageId="1" pageNumber="2" type="reference_group">
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<paragraph id="4C14CB1F1404555BD1DAF966FE0AB0F7" blockId="1.[189,509,1759,1786]" box="[189,509,1759,1786]" pageId="1" pageNumber="2">
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<treatmentCitationGroup id="6CBBEC311404555BD1DAF966FE0AB0F7" box="[189,509,1759,1786]" pageId="1" pageNumber="2">
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<treatmentCitation id="CD0AED0E1404555BD1DAF966FE0EB0F7" author="Paulian R." box="[189,505,1759,1786]" page="15" pageId="1" pageNumber="2" year="1948">
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<taxonomicName id="8BABB09C1404555BD1DAF966FE0EB0F7" authority="Paulian, 1948: 15" authorityName="Paulian" authorityYear="1948" box="[189,505,1759,1786]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="subGenus" subGenus="Phornus">
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<emphasis id="7EDF170D1404555BD1DAF966FED7B0F4" box="[189,288,1759,1785]" italics="true" pageId="1" pageNumber="2">Phornus</emphasis>
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<bibRefCitation id="283AB6EE1404555BD04FF966FE0EB0F7" author="Paulian R." box="[296,505,1759,1786]" pageId="1" pageNumber="2" pagination="1 - 75" refId="ref8779" refString="Paulian R. 1948. Revision des Orphnus Africains (Coleoptera, Scarabaeidae). Annales de la Societe entomologique de France 117: 1 - 75." type="journal article" year="1948">Paulian, 1948: 15</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="04B198941404555BD1DAF898FC84B167" pageId="1" pageNumber="2" type="type_taxon">
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<paragraph id="4C14CB1F1404555BD1DAF898FEAFB137" blockId="1.[189,344,1824,1851]" box="[189,344,1824,1851]" pageId="1" pageNumber="2">
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<heading id="175C7C731404555BD1DAF898FEAFB137" bold="true" box="[189,344,1824,1851]" fontSize="11" level="3" pageId="1" pageNumber="2" reason="3">
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<emphasis id="7EDF170D1404555BD1DAF898FEAFB137" bold="true" box="[189,344,1824,1851]" pageId="1" pageNumber="2">
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<typeStatus id="931075BD1404555BD1DAF898FF0DB136" box="[189,250,1825,1851]" pageId="1" pageNumber="2">Type</typeStatus>
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species
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="4C14CB1F1404555BD1DAF8F6FC84B167" blockId="1.[189,883,1871,1898]" box="[189,883,1871,1898]" pageId="1" pageNumber="2">
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<taxonomicName id="8BABB09C1404555BD1DAF8F6FD90B164" authority="Paulian, 1948" authorityName="Paulian" authorityYear="1948" box="[189,615,1871,1897]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="strangulatus">
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<emphasis id="7EDF170D1404555BD1DAF8F6FE49B164" box="[189,446,1871,1897]" italics="true" pageId="1" pageNumber="2">Orphnus strangulatus</emphasis>
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<bibRefCitation id="283AB6EE1404555BD0A2F8F6FD90B164" author="Paulian R." box="[453,615,1871,1897]" pageId="1" pageNumber="2" pagination="1 - 75" refId="ref8779" refString="Paulian R. 1948. Revision des Orphnus Africains (Coleoptera, Scarabaeidae). Annales de la Societe entomologique de France 117: 1 - 75." type="journal article" year="1948">Paulian, 1948</bibRefCitation>
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</taxonomicName>
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(original designation).
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</paragraph>
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</subSubSection>
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<subSubSection id="04B1989414075558D1DAFEB2FBCAB447" pageId="2" pageNumber="3" type="diagnosis">
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<paragraph id="4C14CB1F14075558D1DAFEB2FEC1B728" blockId="2.[189,310,267,293]" box="[189,310,267,293]" pageId="2" pageNumber="3">
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<heading id="175C7C7314075558D1DAFEB2FEC1B728" bold="true" box="[189,310,267,293]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
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<emphasis id="7EDF170D14075558D1DAFEB2FEC1B728" bold="true" box="[189,310,267,293]" pageId="2" pageNumber="3">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="4C14CB1F14075558D1DAFE80FBCAB447" blockId="2.[189,1399,313,586]" pageId="2" pageNumber="3">
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Medium sized beetles (body length 10.0–
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<quantity id="8B5366FA14075558D3C7FE80FCFCB759" box="[672,779,313,340]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="2" pageNumber="3" unit="mm" value="14.5">14.5 mm</quantity>
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). Colour uniformly brown to black. Dorsal surface of body more or less densely punctate, punctation of head and pronotum denser than of elytra. Elytra with flat to slightly convex intervals, first five to seven elytral striae more or less marked as slightly depressed lines. Head armature of males as short tubercle or transverse frontoclypeal carina. Pronotal disc of males slightly flattened anteromedially in some species. Stridulatory field in males with sparse coarse carinae, separated by 1/40 to
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<emphasis id="7EDF170D14075558D09AFE5EFDECB40F" bold="true" box="[509,539,487,514]" pageId="2" pageNumber="3">1⁄30</emphasis>
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length of stridulatory field in central ¼
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<superScript id="BBDE665714075558D287FE51FC19B7FB" attach="left" box="[992,1006,488,502]" fontSize="6" pageId="2" pageNumber="3">rd</superScript>
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(
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<figureCitation id="D490D79A14075558D29DFE50FBACB409" box="[1018,1115,489,516]" captionStart="Fig" captionStartId="4.[189,232,1860,1886]" captionTargetBox="[228,1350,266,1814]" captionTargetId="figure@4.[227,1357,265,1825]" captionTargetPageId="4" captionText="Fig. 1. — A–M. Orphnus giganteus Paulian, 1948. A–B. Habitus. C–D. Head in dorsal and apical view. E–G. Antennae. H. Aedeagus in lateral view. I. Parameres in dorsal view. J. Abdomen in dorsal view. K–L. Stridulatory field. M. Habitus. — N. O. renaudi sp. nov., holotype, habitus. — O. O. ferrierei sp. nov., holotype, habitus. — P. O. valeriae sp. nov., holotype, habitus. — Q. O. parastrangulatus sp. nov., holotype, habitus. — R. O. strangulatus Paulian, 1948, habitus. M–R = not to scale." figureDoi="http://doi.org/10.5281/zenodo.3850303" httpUri="https://zenodo.org/record/3850303/files/figure.png" pageId="2" pageNumber="3">Figs 1K</figureCitation>
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,
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<figureCitation id="D490D79A14075558D501FE53FB75B409" box="[1126,1154,490,516]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[203,1399,636,1876]" captionTargetId="figure@7.[189,1400,635,1888]" captionTargetPageId="7" captionText="Fig. 2. Orphnus renaudi sp. nov. A–B. Habitus. C–D. Head in dorsal and apical view. E. Aedeagus in lateral view. F. Parameres in dorsal view. G. Labels of holotype. H. Abdomen in dorsal view. I. Stridulatory field. A, C–D, G. Holotype. B, E–F, H–I. Paratypes." figureDoi="http://doi.org/10.5281/zenodo.3850307" httpUri="https://zenodo.org/record/3850307/files/figure.png" pageId="2" pageNumber="3">2I</figureCitation>
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). Sides of abdominal sternite 2 with more or less developed accessory plate (
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<figureCitation id="D490D79A14075558D225FDB5FC54B42A" box="[834,931,524,551]" captionStart="Fig" captionStartId="4.[189,232,1860,1886]" captionTargetBox="[228,1350,266,1814]" captionTargetId="figure@4.[227,1357,265,1825]" captionTargetPageId="4" captionText="Fig. 1. — A–M. Orphnus giganteus Paulian, 1948. A–B. Habitus. C–D. Head in dorsal and apical view. E–G. Antennae. H. Aedeagus in lateral view. I. Parameres in dorsal view. J. Abdomen in dorsal view. K–L. Stridulatory field. M. Habitus. — N. O. renaudi sp. nov., holotype, habitus. — O. O. ferrierei sp. nov., holotype, habitus. — P. O. valeriae sp. nov., holotype, habitus. — Q. O. parastrangulatus sp. nov., holotype, habitus. — R. O. strangulatus Paulian, 1948, habitus. M–R = not to scale." figureDoi="http://doi.org/10.5281/zenodo.3850303" httpUri="https://zenodo.org/record/3850303/files/figure.png" pageId="2" pageNumber="3">Figs 1G</figureCitation>
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,
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<figureCitation id="D490D79A14075558D2D7FDB4FC20B42A" box="[944,983,525,551]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[203,1399,636,1876]" captionTargetId="figure@7.[189,1400,635,1888]" captionTargetPageId="7" captionText="Fig. 2. Orphnus renaudi sp. nov. A–B. Habitus. C–D. Head in dorsal and apical view. E. Aedeagus in lateral view. F. Parameres in dorsal view. G. Labels of holotype. H. Abdomen in dorsal view. I. Stridulatory field. A, C–D, G. Holotype. B, E–F, H–I. Paratypes." figureDoi="http://doi.org/10.5281/zenodo.3850307" httpUri="https://zenodo.org/record/3850307/files/figure.png" pageId="2" pageNumber="3">2H</figureCitation>
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). Parameres with rounded apices in dorsal view, without lateral teeth. Endophallus without sclerotized armature.
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</paragraph>
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</subSubSection>
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<subSubSection id="04B1989414075558D1DAFDC8FB0AB5FF" pageId="2" pageNumber="3" type="discussion">
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<paragraph id="4C14CB1F14075558D1DAFDC8FEAAB486" blockId="2.[189,349,625,651]" box="[189,349,625,651]" pageId="2" pageNumber="3">
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||
<heading id="175C7C7314075558D1DAFDC8FEAAB486" bold="true" box="[189,349,625,651]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
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||
<emphasis id="7EDF170D14075558D1DAFDC8FEAAB486" bold="true" box="[189,349,625,651]" pageId="2" pageNumber="3">Composition</emphasis>
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</heading>
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</paragraph>
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<paragraph id="4C14CB1F14075558D1DAFD19FDDBB567" blockId="2.[189,1399,672,874]" pageId="2" pageNumber="3">
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In the present work we treat the subgenus as comprising six species: two originally described by
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<bibRefCitation id="283AB6EE14075558D1DAFD7AFE99B4D0" author="Paulian R." box="[189,366,707,733]" pageId="2" pageNumber="3" pagination="1 - 75" refId="ref8779" refString="Paulian R. 1948. Revision des Orphnus Africains (Coleoptera, Scarabaeidae). Annales de la Societe entomologique de France 117: 1 - 75." type="journal article" year="1948">
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||
Paulian (
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<date id="3815EDDF14075558D040FD7AFE93B4D0" box="[295,356,707,733]" pageId="2" pageNumber="3" value="1948">1948</date>
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)
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||
</bibRefCitation>
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and four new species.
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<bibRefCitation id="283AB6EE14075558D3E0FD7AFCBEB4D0" author="Petrovitz R." box="[647,841,707,733]" pageId="2" pageNumber="3" pagination="1 - 46" refId="ref8807" refString="Petrovitz R. 1971. Beitrag zur Kenntnis der Gattung Orphnus M'Leay (Orphninae, Scarabaeidae, Coleoptera). Revue de zoologie et botanique africaines 84: 1 - 46." type="journal article" year="1971">
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Petrovitz (
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<date id="3815EDDF14075558D264FD7AFCB7B4D0" box="[771,832,707,733]" pageId="2" pageNumber="3" value="1971">1971</date>
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||
)
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||
</bibRefCitation>
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described two further species in the subgenus
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<taxonomicName id="8BABB09C14075558D1DAFD5FFED7B50D" authorityName="Paulian" authorityYear="1948" box="[189,288,742,768]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subGenus" subGenus="Phornus">
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<emphasis id="7EDF170D14075558D1DAFD5FFED7B50D" box="[189,288,742,768]" italics="true" pageId="2" pageNumber="3">Phornus</emphasis>
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||
</taxonomicName>
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||
,
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<taxonomicName id="8BABB09C14075558D049FD5EFE39B50D" authorityName="Petrovitz" authorityYear="1971" box="[302,462,742,769]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="planicollis">
|
||
<emphasis id="7EDF170D14075558D049FD5EFE39B50D" box="[302,462,742,769]" italics="true" pageId="2" pageNumber="3">O. planicollis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="8BABB09C14075558D362FD5EFD52B50D" authorityName="Petrovitz" authorityYear="1971" box="[517,677,743,769]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="compactus">
|
||
<emphasis id="7EDF170D14075558D362FD5EFD52B50D" box="[517,677,743,769]" italics="true" pageId="2" pageNumber="3">O. compactus</emphasis>
|
||
</taxonomicName>
|
||
, although he noted that they had no similarity to
|
||
<taxonomicName id="8BABB09C14075558D586FD5EFA81B50D" authorityName="Paulian" authorityYear="1948" box="[1249,1398,743,769]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="giganteus">
|
||
<emphasis id="7EDF170D14075558D586FD5EFA81B50D" box="[1249,1398,743,769]" italics="true" pageId="2" pageNumber="3">O. giganteus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="8BABB09C14075558D195FCB3FE5AB52E" authorityName="Paulian" authorityYear="1948" box="[242,429,777,804]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="strangulatus">
|
||
<emphasis id="7EDF170D14075558D195FCB3FE5AB52E" box="[242,429,777,804]" italics="true" pageId="2" pageNumber="3">O. strangulatus</emphasis>
|
||
</taxonomicName>
|
||
. These species have genital and stridulatory structures more similar to those of
|
||
<taxonomicName id="8BABB09C14075558D1DAFC94FDF0B54A" authority="(Fabricius, 1801)" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[189,519,812,839]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="7EDF170D14075558D1DAFC94FEC2B54A" box="[189,309,813,839]" italics="true" pageId="2" pageNumber="3">O. bicolor</emphasis>
|
||
(Fabricius, 1801)
|
||
</taxonomicName>
|
||
, the
|
||
<typeStatus id="931075BD14075558D35AFC94FD87B54A" box="[573,624,813,839]" pageId="2" pageNumber="3">type</typeStatus>
|
||
species of
|
||
<taxonomicName id="8BABB09C14075558D396FC94FCAFB54A" authorityName="MacLeay" authorityYear="1819" box="[753,856,813,839]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="7EDF170D14075558D396FC94FCAFB54A" box="[753,856,813,839]" italics="true" pageId="2" pageNumber="3">Orphnus</emphasis>
|
||
</taxonomicName>
|
||
(sensu stricto), and therefore we transfer them to the nominotypical subgenus.
|
||
</paragraph>
|
||
<paragraph id="4C14CB1F14075558D1DAFC28FB0AB5FF" blockId="2.[189,1399,913,1010]" pageId="2" pageNumber="3">
|
||
<taxonomicName id="8BABB09C14075558D1DAFC28FE6CB5A6" authorityName="Paulian" authorityYear="1948" box="[189,411,913,939]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="giganteus">
|
||
<emphasis id="7EDF170D14075558D1DAFC28FE6CB5A6" box="[189,411,913,939]" italics="true" pageId="2" pageNumber="3">Orphnus giganteus</emphasis>
|
||
</taxonomicName>
|
||
differs from other members of the subgenus in a number of characters (see description below) and its taxonomic position needs further clarification. Until additional comparative data are available, we follow
|
||
<bibRefCitation id="283AB6EE14075558D0D4FC61FD96B5FF" author="Paulian R." box="[435,609,984,1010]" pageId="2" pageNumber="3" pagination="1 - 75" refId="ref8779" refString="Paulian R. 1948. Revision des Orphnus Africains (Coleoptera, Scarabaeidae). Annales de la Societe entomologique de France 117: 1 - 75." type="journal article" year="1948">
|
||
Paulian (
|
||
<date id="3815EDDF14075558D37CFC61FDAFB5FF" box="[539,600,984,1010]" pageId="2" pageNumber="3" value="1948">1948</date>
|
||
)
|
||
</bibRefCitation>
|
||
and treat this species as a member of subgenus
|
||
<taxonomicName id="8BABB09C14075558D5F5FC61FB02B5FF" authorityName="Paulian" authorityYear="1948" box="[1170,1269,984,1010]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subGenus" subGenus="Phornus">
|
||
<emphasis id="7EDF170D14075558D5F5FC61FB02B5FF" box="[1170,1269,984,1010]" italics="true" pageId="2" pageNumber="3">Phornus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="04B1989414075558D1DAFBA0FB65B2A4" pageId="2" pageNumber="3" type="distribution">
|
||
<paragraph id="4C14CB1F14075558D1DAFBA0FEA1B23E" blockId="2.[189,342,1049,1075]" box="[189,342,1049,1075]" pageId="2" pageNumber="3">
|
||
<heading id="175C7C7314075558D1DAFBA0FEA1B23E" bold="true" box="[189,342,1049,1075]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
|
||
<emphasis id="7EDF170D14075558D1DAFBA0FEA1B23E" bold="true" box="[189,342,1049,1075]" pageId="2" pageNumber="3">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="4C14CB1F14075558D1DAFBF1FB65B2A4" blockId="2.[189,1399,1096,1193]" pageId="2" pageNumber="3">
|
||
Members of the subgenus
|
||
<taxonomicName id="8BABB09C14075558D363FBF1FD90B26F" authorityName="Paulian" authorityYear="1948" box="[516,615,1096,1122]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subGenus" subGenus="Phornus">
|
||
<emphasis id="7EDF170D14075558D363FBF1FD90B26F" box="[516,615,1096,1122]" italics="true" pageId="2" pageNumber="3">Phornus</emphasis>
|
||
</taxonomicName>
|
||
are distributed in Equatorial Africa and the majority of species apparently have allopatric ranges. Most of the known localities are south of the
|
||
<collectingCountry id="34BC8B8F14075558D5F5FBD2FB17B288" box="[1170,1248,1131,1157]" name="Democratic Republic of the Congo" pageId="2" pageNumber="3">Congo</collectingCountry>
|
||
Depression, notably on the Lunda and
|
||
<collectingRegion id="8E6F05FD14075558D097FB36FDA5B2A4" box="[496,594,1167,1193]" country="Democratic Republic of the Congo" name="Katanga" pageId="2" pageNumber="3">Katanga</collectingRegion>
|
||
plateaus and the Eastern Arc Mountains (
|
||
<figureCitation id="D490D79A14075558D55BFB37FB73B2A4" box="[1084,1156,1166,1193]" captionStart="Fig" captionStartId="16.[189,232,1966,1992]" captionTargetBox="[194,1393,1011,1923]" captionTargetId="figure@16.[189,1398,1009,1925]" captionTargetPageId="16" captionText="Fig. 6. Orphnus (Phornus) spp. Locality map. Orphnus parastrangulatus sp. nov. is not mapped, because there are no more or less precise locality data available for this species." figureDoi="http://doi.org/10.5281/zenodo.3850323" httpUri="https://zenodo.org/record/3850323/files/figure.png" pageId="2" pageNumber="3">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="04B1989414075559D1DAFB69FA80B7F4" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="key">
|
||
<paragraph id="4C14CB1F14075558D1DAFB69FD62B2E4" blockId="2.[189,661,1231,1258]" box="[189,661,1231,1258]" pageId="2" pageNumber="3">
|
||
<heading id="175C7C7314075558D1DAFB69FD62B2E4" bold="true" box="[189,661,1231,1258]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
|
||
<emphasis id="7EDF170D14075558D1DAFB69FD62B2E4" bold="true" box="[189,661,1231,1258]" pageId="2" pageNumber="3">
|
||
Key to the subgenus
|
||
<taxonomicName id="8BABB09C14075558D0A7FB76FDDFB2E7" authorityName="Paulian" authorityYear="1948" box="[448,552,1231,1258]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subGenus" subGenus="Phornus">
|
||
<emphasis id="7EDF170D14075558D0A7FB76FDDFB2E7" bold="true" box="[448,552,1231,1258]" italics="true" pageId="2" pageNumber="3">Phornus</emphasis>
|
||
</taxonomicName>
|
||
(males):
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<key id="383A9C2E14075559D1DAFB47FA80B7F4" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
|
||
<paragraph id="4C14CB1F14075558D1DAFB47FA80B336" blockId="2.[189,1399,1278,1340]" pageId="2" pageNumber="3">
|
||
<keyLead id="F75A682A14075558D1DAFB47FA80B336" pageId="2" pageNumber="3">
|
||
<keyStep id="F75FD3BA14075558D1DAFB47FA80B336" pageId="2" pageNumber="3">
|
||
1. Macropterous, humeral humps well developed (
|
||
<figureCitation id="D490D79A14075558D244FB47FC77B315" box="[803,896,1278,1304]" captionStart="Fig" captionStartId="4.[189,232,1860,1886]" captionTargetBox="[228,1350,266,1814]" captionTargetId="figure@4.[227,1357,265,1825]" captionTargetPageId="4" captionText="Fig. 1. — A–M. Orphnus giganteus Paulian, 1948. A–B. Habitus. C–D. Head in dorsal and apical view. E–G. Antennae. H. Aedeagus in lateral view. I. Parameres in dorsal view. J. Abdomen in dorsal view. K–L. Stridulatory field. M. Habitus. — N. O. renaudi sp. nov., holotype, habitus. — O. O. ferrierei sp. nov., holotype, habitus. — P. O. valeriae sp. nov., holotype, habitus. — Q. O. parastrangulatus sp. nov., holotype, habitus. — R. O. strangulatus Paulian, 1948, habitus. M–R = not to scale." figureDoi="http://doi.org/10.5281/zenodo.3850303" httpUri="https://zenodo.org/record/3850303/files/figure.png" pageId="2" pageNumber="3">Fig. 1A</figureCitation>
|
||
) …………………………………………2 – Brachypterous, humeral humps feebly marked to indistinct (
|
||
<figureCitation id="D490D79A14075558D2C7FA98FC0DB336" box="[928,1018,1313,1339]" captionStart="Fig" captionStartId="9.[189,232,1929,1955]" captionTargetBox="[196,1394,1107,1876]" captionTargetId="figure@9.[195,1395,1106,1887]" captionTargetPageId="9" captionText="Fig. 3. Orphnus valeriae sp. nov. A. Habitus. B–C. Head in dorsal and apical view. D. Aedeagus in lateral view. E. Parameres in dorsal view. F. Labels of holotype. G. Abdomen in dorsal view. H. Stridulatory field. A–F. Holotype. G–H. Paratypes." figureDoi="http://doi.org/10.5281/zenodo.3850311" httpUri="https://zenodo.org/record/3850311/files/figure.png" pageId="2" pageNumber="3">Fig. 3A</figureCitation>
|
||
) ………………………………3
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<keyStep id="F75FD3BA14075558D1DAFADAFA80B3EA" pageId="2" pageNumber="3">
|
||
<paragraph id="4C14CB1F14075558D1DAFADAFA80B3AD" blockId="2.[189,1399,1379,1511]" pageId="2" pageNumber="3">
|
||
<keyLead id="F75A682A14075558D1DAFADAFA80B3AD" pageId="2" pageNumber="3">
|
||
2. Pronotum as wide as elytra (
|
||
<figureCitation id="D490D79A14075558D328FADAFD41B370" box="[591,694,1379,1405]" captionStart="Fig" captionStartId="4.[189,232,1860,1886]" captionTargetBox="[228,1350,266,1814]" captionTargetId="figure@4.[227,1357,265,1825]" captionTargetPageId="4" captionText="Fig. 1. — A–M. Orphnus giganteus Paulian, 1948. A–B. Habitus. C–D. Head in dorsal and apical view. E–G. Antennae. H. Aedeagus in lateral view. I. Parameres in dorsal view. J. Abdomen in dorsal view. K–L. Stridulatory field. M. Habitus. — N. O. renaudi sp. nov., holotype, habitus. — O. O. ferrierei sp. nov., holotype, habitus. — P. O. valeriae sp. nov., holotype, habitus. — Q. O. parastrangulatus sp. nov., holotype, habitus. — R. O. strangulatus Paulian, 1948, habitus. M–R = not to scale." figureDoi="http://doi.org/10.5281/zenodo.3850303" httpUri="https://zenodo.org/record/3850303/files/figure.png" pageId="2" pageNumber="3">Fig. 1M</figureCitation>
|
||
), densely punctate; head wide, frons convex and densely punctate (
|
||
<figureCitation id="D490D79A14075558D03BFA3FFE6BB3AD" box="[348,412,1414,1440]" captionStart="Fig" captionStartId="4.[189,232,1860,1886]" captionTargetBox="[228,1350,266,1814]" captionTargetId="figure@4.[227,1357,265,1825]" captionTargetPageId="4" captionText="Fig. 1. — A–M. Orphnus giganteus Paulian, 1948. A–B. Habitus. C–D. Head in dorsal and apical view. E–G. Antennae. H. Aedeagus in lateral view. I. Parameres in dorsal view. J. Abdomen in dorsal view. K–L. Stridulatory field. M. Habitus. — N. O. renaudi sp. nov., holotype, habitus. — O. O. ferrierei sp. nov., holotype, habitus. — P. O. valeriae sp. nov., holotype, habitus. — Q. O. parastrangulatus sp. nov., holotype, habitus. — R. O. strangulatus Paulian, 1948, habitus. M–R = not to scale." figureDoi="http://doi.org/10.5281/zenodo.3850303" httpUri="https://zenodo.org/record/3850303/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
|
||
C–D); eyes small, feebly visible in dorsal view …………
|
||
<taxonomicName id="8BABB09C14075558D55EFA3FFA80B3AD" authority="Paulian" authorityName="Paulian" authorityYear="1948" box="[1081,1399,1414,1440]" class="Insecta" family="Orphnidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="giganteus">
|
||
<emphasis id="7EDF170D14075558D55EFA3FFBA2B3AD" bold="true" box="[1081,1109,1414,1440]" pageId="2" pageNumber="3">
|
||
<emphasis id="7EDF170D14075558D55EFA3FFBB9B3AD" bold="true" box="[1081,1102,1414,1440]" italics="true" pageId="2" pageNumber="3">O</emphasis>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="7EDF170D14075558D53CFA3FFB27B3AD" bold="true" box="[1115,1232,1414,1440]" italics="true" pageId="2" pageNumber="3">giganteus</emphasis>
|
||
<bibRefCitation id="283AB6EE14075558D5B1FA3FFA80B3AD" author="Paulian R." box="[1238,1399,1414,1440]" pageId="2" pageNumber="3" pagination="1 - 75" refId="ref8779" refString="Paulian R. 1948. Revision des Orphnus Africains (Coleoptera, Scarabaeidae). Annales de la Societe entomologique de France 117: 1 - 75." type="journal article" year="1948">Paulian, 1948</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="4C14CB1F14075558D1DAFA13FA80B3EA" blockId="2.[189,1399,1379,1511]" pageId="2" pageNumber="3">
|
||
<keyLead id="F75A682A14075558D1DAFA13FA80B3EA" pageId="2" pageNumber="3">
|
||
– Pronotum narrower than elytra (
|
||
<figureCitation id="D490D79A14075558D30FFA10FD31B3CE" box="[616,710,1449,1475]" captionStart="Fig" captionStartId="4.[189,232,1860,1886]" captionTargetBox="[228,1350,266,1814]" captionTargetId="figure@4.[227,1357,265,1825]" captionTargetPageId="4" captionText="Fig. 1. — A–M. Orphnus giganteus Paulian, 1948. A–B. Habitus. C–D. Head in dorsal and apical view. E–G. Antennae. H. Aedeagus in lateral view. I. Parameres in dorsal view. J. Abdomen in dorsal view. K–L. Stridulatory field. M. Habitus. — N. O. renaudi sp. nov., holotype, habitus. — O. O. ferrierei sp. nov., holotype, habitus. — P. O. valeriae sp. nov., holotype, habitus. — Q. O. parastrangulatus sp. nov., holotype, habitus. — R. O. strangulatus Paulian, 1948, habitus. M–R = not to scale." figureDoi="http://doi.org/10.5281/zenodo.3850303" httpUri="https://zenodo.org/record/3850303/files/figure.png" pageId="2" pageNumber="3">Fig. 1N</figureCitation>
|
||
), sparsely punctate on disc; head narrower, frons concave and sparsely punctate (
|
||
<figureCitation id="D490D79A14075558D364FA75FDBAB3EB" box="[515,589,1484,1510]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[203,1399,636,1876]" captionTargetId="figure@7.[189,1400,635,1888]" captionTargetPageId="7" captionText="Fig. 2. Orphnus renaudi sp. nov. A–B. Habitus. C–D. Head in dorsal and apical view. E. Aedeagus in lateral view. F. Parameres in dorsal view. G. Labels of holotype. H. Abdomen in dorsal view. I. Stridulatory field. A, C–D, G. Holotype. B, E–F, H–I. Paratypes." figureDoi="http://doi.org/10.5281/zenodo.3850307" httpUri="https://zenodo.org/record/3850307/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
|
||
C–D); eyes large ………………………………
|
||
<taxonomicName id="8BABB09C14075558D5E1FA75FAFAB3EA" authority="Frolov & Akhmetova, 2016" authorityName="Frolov & Akhmetova" authorityYear="2016" box="[1158,1293,1484,1511]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="renaudi" status="sp. nov.">
|
||
<emphasis id="7EDF170D14075558D5E1FA75FAFAB3EA" bold="true" box="[1158,1293,1484,1511]" italics="true" pageId="2" pageNumber="3">O. renaudi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="65ECAA7614075558D47FFA74FA80B3EA" box="[1304,1399,1485,1511]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="F75FD3BA14075558D1DAF9B7FA81B0D5" pageId="2" pageNumber="3">
|
||
<paragraph id="4C14CB1F14075558D1DAF9B7FA80B063" blockId="2.[189,1399,1550,1752]" pageId="2" pageNumber="3">
|
||
<keyLead id="F75A682A14075558D1DAF9B7FA80B063" pageId="2" pageNumber="3">
|
||
3. Frontoclypeal process tubercle-shaped or horn-shaped, not sinuate medially (
|
||
<figureCitation id="D490D79A14075558D5F5F9B7FB28B025" box="[1170,1247,1550,1576]" captionStart="Fig" captionStartId="9.[189,232,1929,1955]" captionTargetBox="[196,1394,1107,1876]" captionTargetId="figure@9.[195,1395,1106,1887]" captionTargetPageId="9" captionText="Fig. 3. Orphnus valeriae sp. nov. A. Habitus. B–C. Head in dorsal and apical view. D. Aedeagus in lateral view. E. Parameres in dorsal view. F. Labels of holotype. G. Abdomen in dorsal view. H. Stridulatory field. A–F. Holotype. G–H. Paratypes." figureDoi="http://doi.org/10.5281/zenodo.3850311" httpUri="https://zenodo.org/record/3850311/files/figure.png" pageId="2" pageNumber="3">Figs 3</figureCitation>
|
||
B–C, 4H–I). Disc of pronotum somewhat flattened anteriorly; parameres more or less tapering apically (in lateral view) …………………………………………………………………………………………4
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="4C14CB1F14075558D1DAF9C1FA81B0D5" blockId="2.[189,1399,1550,1752]" pageId="2" pageNumber="3">
|
||
<keyLead id="F75A682A14075558D1DAF9C1FA81B0D5" pageId="2" pageNumber="3">
|
||
– Frontoclypeus with a low transverse keel, from ½ to almost whole head width (
|
||
<figureCitation id="D490D79A14075558D58BF9CEFACFB09F" box="[1260,1336,1655,1682]" captionStart="Fig" captionStartId="13.[189,232,1862,1888]" captionTargetBox="[190,1383,262,1815]" captionTargetId="figure@13.[189,1394,261,1821]" captionTargetPageId="13" captionText="Fig. 5. — A–H. O. parastrangulatus sp. nov. A. Habitus. B–C. Head in dorsal and apical view. D. Aedeagus in lateral view. E. Parameres in dorsal view. F. Label of holotype. G. Abdomen in dorsal view. H. Stridulatory field. — I–P. O. strangulatus Paulian, 1948. I. Habitus. J–K. Head in dorsal and apical view. L. Aedeagus in lateral view. M. Parameres in dorsal view. N. Label of holotype. O. Abdomen in dorsal view. P. Stridulatory field. A–H, I, N. Holotypes." figureDoi="http://doi.org/10.5281/zenodo.3850319" httpUri="https://zenodo.org/record/3850319/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation>
|
||
B–C, J–K); keel may be feebly sinuate medially; disc of pronotum more or less convex, not flattened anteriorly; parameres rounded apically (in lateral view)……………………………………………5
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="F75FD3BA14075558D1DAF946FA80B1C7" pageId="2" pageNumber="3">
|
||
<paragraph id="4C14CB1F14075558D1DAF946FA80B16D" blockId="2.[189,1399,1791,1994]" pageId="2" pageNumber="3">
|
||
<keyLead id="F75A682A14075558D1DAF946FA80B16D" pageId="2" pageNumber="3">
|
||
4. Pronotum larger (
|
||
<figureCitation id="D490D79A14075558D0DFF946FDF9B114" box="[440,526,1791,1817]" captionStart="Fig" captionStartId="4.[189,232,1860,1886]" captionTargetBox="[228,1350,266,1814]" captionTargetId="figure@4.[227,1357,265,1825]" captionTargetPageId="4" captionText="Fig. 1. — A–M. Orphnus giganteus Paulian, 1948. A–B. Habitus. C–D. Head in dorsal and apical view. E–G. Antennae. H. Aedeagus in lateral view. I. Parameres in dorsal view. J. Abdomen in dorsal view. K–L. Stridulatory field. M. Habitus. — N. O. renaudi sp. nov., holotype, habitus. — O. O. ferrierei sp. nov., holotype, habitus. — P. O. valeriae sp. nov., holotype, habitus. — Q. O. parastrangulatus sp. nov., holotype, habitus. — R. O. strangulatus Paulian, 1948, habitus. M–R = not to scale." figureDoi="http://doi.org/10.5281/zenodo.3850303" httpUri="https://zenodo.org/record/3850303/files/figure.png" pageId="2" pageNumber="3">Fig. 1P</figureCitation>
|
||
), more densely punctate, punctures on disc somewhat elongated (
|
||
<figureCitation id="D490D79A14075558D46DF946FA9FB114" box="[1290,1384,1791,1817]" captionStart="Fig" captionStartId="9.[189,232,1929,1955]" captionTargetBox="[196,1394,1107,1876]" captionTargetId="figure@9.[195,1395,1106,1887]" captionTargetPageId="9" captionText="Fig. 3. Orphnus valeriae sp. nov. A. Habitus. B–C. Head in dorsal and apical view. D. Aedeagus in lateral view. E. Parameres in dorsal view. F. Labels of holotype. G. Abdomen in dorsal view. H. Stridulatory field. A–F. Holotype. G–H. Paratypes." figureDoi="http://doi.org/10.5281/zenodo.3850311" httpUri="https://zenodo.org/record/3850311/files/figure.png" pageId="2" pageNumber="3">Fig. 3A</figureCitation>
|
||
); pronotum without longitudinal medial stria basally; frontoclypeal process horn-shaped; parameres more strongly sclerotized, with acute apices (in lateral view,
|
||
<figureCitation id="D490D79A14075558D2A3F8FCFBD6B16D" box="[964,1057,1861,1888]" captionStart="Fig" captionStartId="9.[189,232,1929,1955]" captionTargetBox="[196,1394,1107,1876]" captionTargetId="figure@9.[195,1395,1106,1887]" captionTargetPageId="9" captionText="Fig. 3. Orphnus valeriae sp. nov. A. Habitus. B–C. Head in dorsal and apical view. D. Aedeagus in lateral view. E. Parameres in dorsal view. F. Labels of holotype. G. Abdomen in dorsal view. H. Stridulatory field. A–F. Holotype. G–H. Paratypes." figureDoi="http://doi.org/10.5281/zenodo.3850311" httpUri="https://zenodo.org/record/3850311/files/figure.png" pageId="2" pageNumber="3">Fig. 3D</figureCitation>
|
||
) ………
|
||
<taxonomicName id="8BABB09C14075558D5ECF8FFFAE6B16D" authority="Frolov & Akhmetova, 2016" authorityName="Frolov & Akhmetova" authorityYear="2016" box="[1163,1297,1861,1888]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="valeriae" status="sp. nov.">
|
||
<emphasis id="7EDF170D14075558D5ECF8FFFAE6B16D" bold="true" box="[1163,1297,1861,1888]" italics="true" pageId="2" pageNumber="3">O. valeriae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="65ECAA7614075558D47DF8FFFA80B16D" box="[1306,1399,1862,1888]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="4C14CB1F14075558D1DAF8D0FA80B1C7" blockId="2.[189,1399,1791,1994]" pageId="2" pageNumber="3">
|
||
<keyLead id="F75A682A14075558D1DAF8D0FA80B1C7" pageId="2" pageNumber="3">
|
||
– Pronotum smaller (
|
||
<figureCitation id="D490D79A14075558D0A8F8D0FDDAB18E" box="[463,557,1897,1923]" captionStart="Fig" captionStartId="4.[189,232,1860,1886]" captionTargetBox="[228,1350,266,1814]" captionTargetId="figure@4.[227,1357,265,1825]" captionTargetPageId="4" captionText="Fig. 1. — A–M. Orphnus giganteus Paulian, 1948. A–B. Habitus. C–D. Head in dorsal and apical view. E–G. Antennae. H. Aedeagus in lateral view. I. Parameres in dorsal view. J. Abdomen in dorsal view. K–L. Stridulatory field. M. Habitus. — N. O. renaudi sp. nov., holotype, habitus. — O. O. ferrierei sp. nov., holotype, habitus. — P. O. valeriae sp. nov., holotype, habitus. — Q. O. parastrangulatus sp. nov., holotype, habitus. — R. O. strangulatus Paulian, 1948, habitus. M–R = not to scale." figureDoi="http://doi.org/10.5281/zenodo.3850303" httpUri="https://zenodo.org/record/3850303/files/figure.png" pageId="2" pageNumber="3">Fig. 1O</figureCitation>
|
||
), more sparsely punctate with rounded punctures (
|
||
<figureCitation id="D490D79A14075558D5EAF8D0FB1AB18E" box="[1165,1261,1897,1923]" captionStart="Fig" captionStartId="11.[189,232,1928,1954]" captionTargetBox="[194,1388,717,1886]" captionTargetId="figure@11.[193,1393,716,1887]" captionTargetPageId="11" captionText="Fig. 4. Orphnus ferrierei sp. nov. A, C–D. Habitus. B. Labels of holotype. E. Abdomen in dorsal view. F. Aedeagus in lateral view. G. Parameres in dorsal view. H–I. Head in dorsal and apical view. J. Stridulatory field. A–B, F–I. Holotype. C–E. Paratypes." figureDoi="http://doi.org/10.5281/zenodo.3850315" httpUri="https://zenodo.org/record/3850315/files/figure.png" pageId="2" pageNumber="3">Fig. 4A</figureCitation>
|
||
); pronotum with distinct medial longitudinal stria basally; frontoclypeal process tubercle-shaped; parameres less sclerotized, with angulate apices (in lateral view,
|
||
<figureCitation id="D490D79A14075558D208F816FC31B1C4" box="[879,966,1967,1993]" captionStart="Fig" captionStartId="11.[189,232,1928,1954]" captionTargetBox="[194,1388,717,1886]" captionTargetId="figure@11.[193,1393,716,1887]" captionTargetPageId="11" captionText="Fig. 4. Orphnus ferrierei sp. nov. A, C–D. Habitus. B. Labels of holotype. E. Abdomen in dorsal view. F. Aedeagus in lateral view. G. Parameres in dorsal view. H–I. Head in dorsal and apical view. J. Stridulatory field. A–B, F–I. Holotype. C–E. Paratypes." figureDoi="http://doi.org/10.5281/zenodo.3850315" httpUri="https://zenodo.org/record/3850315/files/figure.png" pageId="2" pageNumber="3">Fig. 4F</figureCitation>
|
||
) ………………
|
||
<taxonomicName id="8BABB09C14075558D5EFF816FAE6B1C4" authority="Frolov & Akhmetova, 2016" authorityName="Frolov & Akhmetova" authorityYear="2016" box="[1160,1297,1967,1993]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="ferrierei" status="sp. nov.">
|
||
<emphasis id="7EDF170D14075558D5EFF816FAE6B1C4" bold="true" box="[1160,1297,1967,1993]" italics="true" pageId="2" pageNumber="3">O. ferrierei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="65ECAA7614075558D47DF809FA80B1C7" box="[1306,1399,1968,1994]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="F75FD3BA14065559D1DAFEB2FA80B7F4" pageId="3" pageNumber="4">
|
||
<paragraph id="4C14CB1F14065559D1DAFEB2FA80B761" blockId="3.[189,1399,267,505]" pageId="3" pageNumber="4">
|
||
<keyLead id="F75A682A14065559D1DAFEB2FA80B761" pageId="3" pageNumber="4">
|
||
5. Frontoclypeus with a low transverse keel, almost as wide as frontoclypeus (
|
||
<figureCitation id="D490D79A14065559D5E6FEB2FB3EB72B" box="[1153,1225,267,294]" captionStart="Fig" captionStartId="13.[189,232,1862,1888]" captionTargetBox="[190,1383,262,1815]" captionTargetId="figure@13.[189,1394,261,1821]" captionTargetPageId="13" captionText="Fig. 5. — A–H. O. parastrangulatus sp. nov. A. Habitus. B–C. Head in dorsal and apical view. D. Aedeagus in lateral view. E. Parameres in dorsal view. F. Label of holotype. G. Abdomen in dorsal view. H. Stridulatory field. — I–P. O. strangulatus Paulian, 1948. I. Habitus. J–K. Head in dorsal and apical view. L. Aedeagus in lateral view. M. Parameres in dorsal view. N. Label of holotype. O. Abdomen in dorsal view. P. Stridulatory field. A–H, I, N. Holotypes." figureDoi="http://doi.org/10.5281/zenodo.3850319" httpUri="https://zenodo.org/record/3850319/files/figure.png" pageId="3" pageNumber="4">Fig. 5</figureCitation>
|
||
B–C); anterior margin of clypeus somewhat rounded, keel-shaped; abdomen shorter and somewhat convex ventrally (in lateral view contour of abdominal sternites is somewhat rounded) …
|
||
<taxonomicName id="8BABB09C14065559D540FEEBFAEFB761" authority="Frolov & Akhmetova, 2016" authorityName="Frolov & Akhmetova" authorityYear="2016" box="[1063,1304,337,364]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="parastrangulatus" status="sp. nov.">
|
||
<emphasis id="7EDF170D14065559D540FEEBFAEFB761" bold="true" box="[1063,1304,337,364]" italics="true" pageId="3" pageNumber="4">O. parastrangulatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="65ECAA7614065559D479FEEBFA80B761" box="[1310,1399,338,364]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="4C14CB1F14065559D1DAFECCFA80B7F4" blockId="3.[189,1399,267,505]" pageId="3" pageNumber="4">
|
||
<keyLead id="F75A682A14065559D1DAFECCFA80B7F4" pageId="3" pageNumber="4">
|
||
– Frontoclypeus with a higher transverse keel, slightly sinuate medially, about half the length of frontoclypeus (
|
||
<figureCitation id="D490D79A14065559D0C4FE21FE19B7BE" box="[419,494,408,435]" captionStart="Fig" captionStartId="13.[189,232,1862,1888]" captionTargetBox="[190,1383,262,1815]" captionTargetId="figure@13.[189,1394,261,1821]" captionTargetPageId="13" captionText="Fig. 5. — A–H. O. parastrangulatus sp. nov. A. Habitus. B–C. Head in dorsal and apical view. D. Aedeagus in lateral view. E. Parameres in dorsal view. F. Label of holotype. G. Abdomen in dorsal view. H. Stridulatory field. — I–P. O. strangulatus Paulian, 1948. I. Habitus. J–K. Head in dorsal and apical view. L. Aedeagus in lateral view. M. Parameres in dorsal view. N. Label of holotype. O. Abdomen in dorsal view. P. Stridulatory field. A–H, I, N. Holotypes." figureDoi="http://doi.org/10.5281/zenodo.3850319" httpUri="https://zenodo.org/record/3850319/files/figure.png" pageId="3" pageNumber="4">Fig. 5</figureCitation>
|
||
J–K); anterior margin of clypeus somewhat rectangular, sharp; abdomen longer and almost flat (in lateral view contour of abdominal sternites is almost straight) …………… ………………………………………………………………………
|
||
<taxonomicName id="8BABB09C14065559D565FE66FA80B7F4" authority="Paulian, 1948" authorityName="Paulian" authorityYear="1948" box="[1026,1399,478,505]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="strangulatus">
|
||
<emphasis id="7EDF170D14065559D565FE66FB33B7F4" bold="true" box="[1026,1220,478,505]" italics="true" pageId="3" pageNumber="4">O. strangulatus</emphasis>
|
||
<bibRefCitation id="283AB6EE14065559D5B7FE67FA80B7F4" author="Paulian R." box="[1232,1399,478,505]" pageId="3" pageNumber="4" pagination="1 - 75" refId="ref8779" refString="Paulian R. 1948. Revision des Orphnus Africains (Coleoptera, Scarabaeidae). Annales de la Societe entomologique de France 117: 1 - 75." type="journal article" year="1948">Paulian, 1948</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |