142 lines
14 KiB
XML
142 lines
14 KiB
XML
<document id="F9151784B7FFD59872332F8DC6B6A6DA" ID-DOI="10.11646/zootaxa.3905.1.10" ID-GBIF-Dataset="1b30f72e-6460-4037-baca-23fd7b4b159f" ID-ISSN="1175-5326" ID-Zenodo-Dep="235479" ID-ZooBank="1E6F3CE8-FAF9-443A-97A2-50388447AA06" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460927997643" checkinUser="plazi" docAuthor="Dow, Rory A. & Luke, Sarah H." docDate="2015" docId="6B168785C204FFA2FF2DFB07FC042FEE" docLanguage="en" docName="zt03905p150.pdf" docOrigin="Zootaxa 3905 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phaenandrogomphus treadawayi Muller & Hamalainen 1993, comb. nov." docType="treatment" docVersion="9" lastPageNumber="149" masterDocId="972FFFFDC200FFA6FFBAFFDFFFBB2957" masterDocTitle="Phaenandrogomphus safei, a new species from Sabah, northern Borneo (Odonata: Anisoptera: Gomphidae)" masterLastPageNumber="150" masterPageNumber="145" pageNumber="149" updateTime="1698362785536" updateUser="plazi">
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<mods:titleInfo id="7C0BA524966BBCB39B0630C71B514C1D">
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<mods:title id="A967239C81220605859802E968C9D217">Phaenandrogomphus safei, a new species from Sabah, northern Borneo (Odonata: Anisoptera: Gomphidae)</mods:title>
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<mods:name id="460C1663A1AA639A95DC7CFF0A2174FC" type="personal">
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<mods:namePart id="BAC98D2C93B5188DB57F8A3210081781">Dow, Rory A.</mods:namePart>
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<mods:name id="A74E912832D7E9F6808F3A46B418892D" type="personal">
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<mods:roleTerm id="0FCE31B8A5237E7CDB42E8E1B00BAA1B">Author</mods:roleTerm>
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<mods:namePart id="A47A0B93FB24285F76FCDCC1B28A574E">Luke, Sarah H.</mods:namePart>
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<mods:typeOfResource id="D3FEA561A03A24A9E5532E93B9E9F074">text</mods:typeOfResource>
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<mods:title id="B983C92E4084BFF7CE2843CB830AED80">Zootaxa</mods:title>
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<mods:part id="CA3DDA4E68241876A369F86692F3552D">
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<mods:date id="2AEFC15DCE0CA1C6CF8CC8FFE8A915C2">2015</mods:date>
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<mods:detail id="64388C807C66A0739D6C1B11C9B9323F" type="volume">
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<mods:number id="01AF61C42D1AA9E57CB76DF0120D8CC2">3905</mods:number>
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<mods:number id="8E850F5F7B0B8C04FE39AF69BDC49A03">1</mods:number>
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<mods:identifier id="E241E427F9FCB60B96596991A4851C25" type="DOI">10.11646/zootaxa.3905.1.10</mods:identifier>
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<treatment id="6B168785C204FFA2FF2DFB07FC042FEE" ID-DOI="http://doi.org/10.5281/zenodo.6118691" ID-GBIF-Taxon="119589955" ID-Zenodo-Dep="6118691" LSID="urn:lsid:plazi:treatment:6B168785C204FFA2FF2DFB07FC042FEE" httpUri="http://treatment.plazi.org/id/6B168785C204FFA2FF2DFB07FC042FEE" lastPageNumber="149" pageId="4" pageNumber="149">
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<subSubSection id="ABA56518C204FFA2FF2DFB07FF502C43" pageId="4" pageNumber="149" type="nomenclature">
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<paragraph id="E3003693C204FFA2FF2DFB07FB952DA4" blockId="4.[151,1070,1240,1300]" box="[151,1070,1240,1267]" pageId="4" pageNumber="149">
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<heading id="B84881FFC204FFA2FF2DFB07FB952DA4" bold="true" box="[151,1070,1240,1267]" fontSize="11" level="1" pageId="4" pageNumber="149" reason="1">
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<emphasis id="D1CBEA81C204FFA2FF2DFB07FB952DA4" bold="true" box="[151,1070,1240,1267]" pageId="4" pageNumber="149">
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<taxonomicName id="24BF4D10C204FFA2FF2DFB07FC1A2DA4" ID-CoL="9X5PS" authority="Muller & Hamalainen, 1993" authorityName="Muller & Hamalainen" authorityYear="1993" box="[151,929,1240,1267]" class="Insecta" family="Gomphidae" genus="Phaenandrogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="149" phylum="Arthropoda" rank="species" species="treadawayi" status="comb. nov.">
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<emphasis id="D1CBEA81C204FFA2FF2DFB07FD9C2DA5" bold="true" box="[151,551,1240,1266]" italics="true" pageId="4" pageNumber="149">Phaenandrogomphus treadawayi</emphasis>
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(
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<bibRefCitation id="872E4B62C204FFA2FD8CFB06FC222DA4" author="Muller" box="[566,921,1241,1267]" pageId="4" pageNumber="149" refString="Muller, R. A. & Hamalainen, M. (1993) Onychogomphus treadawayi n. sp., eine neue Libellenart von der Insel Busuanga, Philippinen (Odonata: Gomphidae). Entomologische Zeitschrift, Essen, 103 (4), 41 - 45." type="journal article" year="1993">Müller & Hämäläinen, 1993</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="CAF857FAC204FFA2FC12FB06FB952DA4" box="[936,1070,1241,1267]" pageId="4" pageNumber="149">comb. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="E3003693C204FFA2FF2DFB24FF502C43" blockId="4.[151,1070,1240,1300]" box="[151,235,1275,1300]" pageId="4" pageNumber="149">
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(
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<figureCitation id="7B842A16C204FFA2FF25FB24FF582C43" box="[159,227,1275,1300]" captionStart="FIGURES 1 – 2" captionStartId="1.[151,264,612,635]" captionTargetBox="[174,1434,195,586]" captionTargetId="figure@1.[151,1436,195,588]" captionTargetPageId="1" captionText="FIGURES 1 – 2. Habitus: (1) Phaenandrogomphus safei holotype; (2) Phaenandrogomphus treadawayi holotype." httpUri="https://zenodo.org/record/235480/files/figure.png" pageId="4" pageNumber="149">Fig. 2</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="ABA56518C204FFA2FF2DFA9CFC642C2F" pageId="4" pageNumber="149" type="reference_group">
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<paragraph id="E3003693C204FFA2FF2DFA9CFC642C2F" blockId="4.[151,1436,1347,1400]" pageId="4" pageNumber="149">
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<treatmentCitationGroup id="C3AF11BDC204FFA2FF2DFA9CFC642C2F" pageId="4" pageNumber="149">
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<taxonomicName id="24BF4D10C204FFA2FF2DFA9CFCBE2C0E" ID-CoL="6SM8Q" authority="Muller & Hamalainen 1993: 41" authorityName="Muller & Hamalainen" authorityPageNumber="41" authorityYear="1993" box="[151,773,1347,1369]" class="Insecta" family="Gomphidae" genus="Onychogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="149" phylum="Arthropoda" rank="species" species="treadawayi">
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<emphasis id="D1CBEA81C204FFA2FF2DFA9CFE012C0E" box="[151,442,1347,1369]" italics="true" pageId="4" pageNumber="149">Onychogomphus treadawayi</emphasis>
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<treatmentCitation id="621E1082C204FFA2FE7BFA9CFCBE2C0E" author="Muller" box="[449,773,1347,1369]" page="41" pageId="4" pageNumber="149" year="1993">
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<bibRefCitation id="872E4B62C204FFA2FE7BFA9CFD5A2C0E" author="Muller" box="[449,737,1347,1369]" pageId="4" pageNumber="149" refString="Muller, R. A. & Hamalainen, M. (1993) Onychogomphus treadawayi n. sp., eine neue Libellenart von der Insel Busuanga, Philippinen (Odonata: Gomphidae). Entomologische Zeitschrift, Essen, 103 (4), 41 - 45." type="journal article" year="1993">Müller & Hämäläinen 1993</bibRefCitation>
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: 41
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</treatmentCitation>
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</taxonomicName>
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–45 (original description ♂, Busuanga);—Hämäläinen & Müller 1997: 268, 305 (listed);—
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<treatmentCitation id="621E1082C204FFA2FE71FABDFD472C2F" author="Karube" box="[459,764,1378,1400]" page="68" pageId="4" pageNumber="149" year="2014">
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<bibRefCitation id="872E4B62C204FFA2FE71FABDFD632C2F" author="Karube" box="[459,728,1378,1400]" pageId="4" pageNumber="149" refString="Karube, H. & Sasamoto, A. (2014) Borneogomphus teramotoi, a new genus and species of Gomphid dragonfly from Borneo (Odonata: Gomphidae: Onychogomphinae). Tombo, 56, 65 - 72." type="journal article" year="2014">Karube & Sasamoto 2014</bibRefCitation>
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: 68
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</treatmentCitation>
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, 69 (brief discussion).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="ABA56518C204FFA2FF2DFA7BFCB82CB6" pageId="4" pageNumber="149" type="materials_examined">
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<paragraph id="E3003693C204FFA2FF2DFA7BFCB82CB6" blockId="4.[151,1436,1444,1721]" pageId="4" pageNumber="149">
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<emphasis id="D1CBEA81C204FFA2FF2DFA7BFE3C2CEA" bold="true" box="[151,391,1444,1469]" pageId="4" pageNumber="149">
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Material.
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<typeStatus id="3C048831C204FFA2FEABFA7AFE392CEA" box="[273,386,1445,1469]" pageId="4" pageNumber="149" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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♂, The
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<collectingCountry id="9BA87603C204FFA2FE54FA7BFDD42CEA" box="[494,623,1444,1469]" name="Philippines" pageId="4" pageNumber="149">Philippines</collectingCountry>
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, Palawan region, Busuanga Island, Coron, Mabentangen River,
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<date id="97011053C204FFA2FAE0FA7AFF6A2CB6" pageId="4" pageNumber="149" value="1991-05-04" valueMax="1991-05-07" valueMin="1991-05-04">4–7 v 1991</date>
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, leg. T. Borromeo, in
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<collectionCode id="85AEAE56C204FFA2FE7CFA16FDA62CB7" LSID="urn:lsid:biocol.org:col:34992" box="[454,541,1481,1504]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="4" pageNumber="149">RMNH</collectionCode>
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, JvT number 11935.
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</paragraph>
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</subSubSection>
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<subSubSection id="ABA56518C204FFA2FF7DFA32FC042FEE" pageId="4" pageNumber="149" type="discussion">
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<paragraph id="E3003693C204FFA2FF7DFA32FC042FEE" blockId="4.[151,1436,1444,1721]" pageId="4" pageNumber="149">
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<emphasis id="D1CBEA81C204FFA2FF7DFA32FE802F52" bold="true" box="[199,315,1517,1541]" pageId="4" pageNumber="149">Remarks.</emphasis>
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The habitus of the
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<typeStatus id="3C048831C204FFA2FDA3FA32FDC02F51" box="[537,635,1517,1542]" pageId="4" pageNumber="149" type="holotype">holotype</typeStatus>
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is shown in
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<figureCitation id="7B842A16C204FFA2FCB7FA32FCE92F52" box="[781,850,1517,1542]" captionStart="FIGURES 1 – 2" captionStartId="1.[151,264,612,635]" captionTargetBox="[174,1434,195,586]" captionTargetId="figure@1.[151,1436,195,588]" captionTargetPageId="1" captionText="FIGURES 1 – 2. Habitus: (1) Phaenandrogomphus safei holotype; (2) Phaenandrogomphus treadawayi holotype." httpUri="https://zenodo.org/record/235480/files/figure.png" pageId="4" pageNumber="149">Fig. 2</figureCitation>
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.
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<bibRefCitation id="872E4B62C204FFA2FCE4FA32FB462F52" author="Muller" box="[862,1277,1517,1542]" pageId="4" pageNumber="149" refString="Muller, R. A. & Hamalainen, M. (1993) Onychogomphus treadawayi n. sp., eine neue Libellenart von der Insel Busuanga, Philippinen (Odonata: Gomphidae). Entomologische Zeitschrift, Essen, 103 (4), 41 - 45." type="journal article" year="1993">
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Müller & Hämäläinen (1993:
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<figureCitation id="7B842A16C204FFA2FB0BFA32FB4F2F52" box="[1201,1268,1517,1542]" captionStart="FIGURES 1 – 2" captionStartId="1.[151,264,612,635]" captionTargetBox="[174,1434,195,586]" captionTargetId="figure@1.[151,1436,195,588]" captionTargetPageId="1" captionText="FIGURES 1 – 2. Habitus: (1) Phaenandrogomphus safei holotype; (2) Phaenandrogomphus treadawayi holotype." httpUri="https://zenodo.org/record/235480/files/figure.png" pageId="4" pageNumber="149">Fig. 2</figureCitation>
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)
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</bibRefCitation>
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gave a lateral view of the accessory genitalia of
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<taxonomicName id="24BF4D10C204FFA2FDA8F9CEFD112F7E" box="[530,682,1553,1577]" class="Insecta" family="Gomphidae" genus="Onychogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="149" phylum="Arthropoda" rank="species" species="treadawayi">
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<emphasis id="D1CBEA81C204FFA2FDA8F9CEFD112F7E" box="[530,682,1553,1577]" italics="true" pageId="4" pageNumber="149">P. treadawayi</emphasis>
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</taxonomicName>
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but no ventral view of the terminal segment of the penis. In fact the terminal segment of the penis has the same distinctive, bifurcate appearance in ventral view as that of
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<emphasis id="D1CBEA81C204FFA2FAA1F9E9FAD32F1A" box="[1307,1384,1589,1613]" italics="true" pageId="4" pageNumber="149">P. s a f e i</emphasis>
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and mainland species of
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<emphasis id="D1CBEA81C204FFA2FE33F986FD3B2F26" box="[393,640,1625,1649]" italics="true" pageId="4" pageNumber="149">
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<taxonomicName id="24BF4D10C204FFA2FE33F986FDC72F26" box="[393,636,1625,1649]" class="Insecta" family="Gomphidae" genus="Phaenandrogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="149" phylum="Arthropoda" rank="genus">Phaenandrogomphus</taxonomicName>
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.
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</emphasis>
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Because of the form of the penis and the shape of the epiproct,
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<taxonomicName id="24BF4D10C204FFA2FA3DF986FEAF2FC2" class="Insecta" family="Gomphidae" genus="Onychogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="149" phylum="Arthropoda" rank="species" species="treadawayi">
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<emphasis id="D1CBEA81C204FFA2FA3DF986FEAF2FC2" italics="true" pageId="4" pageNumber="149">P. treadawayi</emphasis>
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</taxonomicName>
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is here included in
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<taxonomicName id="24BF4D10C204FFA2FE56F9A2FD612FC2" box="[492,730,1661,1685]" class="Insecta" family="Gomphidae" genus="Phaenandrogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="149" phylum="Arthropoda" rank="genus">
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<emphasis id="D1CBEA81C204FFA2FE56F9A2FD612FC2" box="[492,730,1661,1685]" italics="true" pageId="4" pageNumber="149">Phaenandrogomphus</emphasis>
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</taxonomicName>
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. It is most closely related to
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<emphasis id="D1CBEA81C204FFA2FBA3F9A1FBDE2FC2" box="[1049,1125,1661,1685]" italics="true" pageId="4" pageNumber="149">P. s a f e i</emphasis>
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; further discussion of these two species and their placement in
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<taxonomicName id="24BF4D10C204FFA2FD9AF97EFCB52FEE" box="[544,782,1697,1721]" class="Insecta" family="Gomphidae" genus="Phaenandrogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="149" phylum="Arthropoda" rank="genus">
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<emphasis id="D1CBEA81C204FFA2FD9AF97EFCB52FEE" box="[544,782,1697,1721]" italics="true" pageId="4" pageNumber="149">Phaenandrogomphus</emphasis>
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</taxonomicName>
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is given below.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |