263 lines
36 KiB
XML
263 lines
36 KiB
XML
<document id="0932FD331F330E860067C960FCAC233A" ID-DOI="10.11646/zootaxa.3905.1.10" ID-GBIF-Dataset="1b30f72e-6460-4037-baca-23fd7b4b159f" ID-ISSN="1175-5326" ID-Zenodo-Dep="235479" ID-ZooBank="1E6F3CE8-FAF9-443A-97A2-50388447AA06" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460927997643" checkinUser="plazi" docAuthor="Dow, Rory A. & Luke, Sarah H." docDate="2015" docId="6B168785C200FFA2FF2DF8E0FE4A2DD1" docLanguage="en" docName="zt03905p150.pdf" docOrigin="Zootaxa 3905 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phaenandrogomphus safei Dow & Luke, 2015, sp. nov." docType="treatment" docUuid="BACEB6EA-2878-42E1-B049-AC8CD54AC1C1" docUuidSource="ZooBank" docVersion="10" lastPageNumber="149" masterDocId="972FFFFDC200FFA6FFBAFFDFFFBB2957" masterDocTitle="Phaenandrogomphus safei, a new species from Sabah, northern Borneo (Odonata: Anisoptera: Gomphidae)" masterLastPageNumber="150" masterPageNumber="145" pageNumber="145" updateTime="1698362785536" updateUser="plazi">
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<mods:mods id="739785BE80BBACC31D9FEDCF3FB0C75F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="1F208289E12C33C5769029BB288E0D2B">
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<mods:title id="C012F0D4AC1B53582B108A3707406D20">Phaenandrogomphus safei, a new species from Sabah, northern Borneo (Odonata: Anisoptera: Gomphidae)</mods:title>
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<mods:name id="D93DD658F41037CE91D9A19680A8EB26" type="personal">
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<mods:namePart id="BFA5A28D886FB671C7EAE56A2F4CE0C4">Dow, Rory A.</mods:namePart>
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<mods:name id="3BFDD58215BF642321CD95DFA2025395" type="personal">
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<mods:role id="F141FED45035D7AEABFBBA5D477D9078">
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<mods:roleTerm id="C74FDE2170DC9839260034C1FE1F516E">Author</mods:roleTerm>
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<mods:namePart id="09F8221FBB59474877A0ED764C3CDB31">Luke, Sarah H.</mods:namePart>
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<mods:typeOfResource id="517A0C680F758A7AE50D1C037800AE8C">text</mods:typeOfResource>
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<mods:title id="45785032930277678ABD58DA126086C4">Zootaxa</mods:title>
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<mods:part id="B9556C234F32AC916E5564AAABCE62F6">
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<mods:date id="643DFDBD037A591D7AF8668CC6BB7FF8">2015</mods:date>
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<mods:detail id="29D6F661A28814BE06506A92732CD94A" type="volume">
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<mods:number id="D4319F5100CDDF9BF87D05CBEFD21D4F">3905</mods:number>
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<mods:detail id="BCC340A32148ECAF3DF0DBF762A35C81" type="issue">
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<mods:number id="25AE5C02200CC4F2F88E3559E39638C3">1</mods:number>
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<mods:classification id="EA16075D597C80FD367C880D02044052">journal article</mods:classification>
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<mods:identifier id="D55458206A5EAFD5FFA718EBFEF834D6" type="DOI">10.11646/zootaxa.3905.1.10</mods:identifier>
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<treatment id="6B168785C200FFA2FF2DF8E0FE4A2DD1" ID-DOI="http://doi.org/10.5281/zenodo.6118689" ID-GBIF-Taxon="119589956" ID-Zenodo-Dep="6118689" LSID="urn:lsid:zoobank.org:act:BACEB6EA-2878-42E1-B049-AC8CD54AC1C1" httpUri="http://treatment.plazi.org/id/6B168785C200FFA2FF2DF8E0FE4A2DD1" lastPageId="4" lastPageNumber="149" pageId="0" pageNumber="145">
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<subSubSection id="ABA56518C200FFA6FF2DF8E0FD222E2C" pageId="0" pageNumber="145" type="nomenclature">
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<paragraph id="E3003693C200FFA6FF2DF8E0FDFA2E0E" blockId="0.[151,665,1855,1915]" box="[151,577,1855,1881]" pageId="0" pageNumber="145">
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<heading id="B84881FFC200FFA6FF2DF8E0FDFA2E0E" bold="true" box="[151,577,1855,1881]" fontSize="11" level="1" pageId="0" pageNumber="145" reason="1">
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<emphasis id="D1CBEA81C200FFA6FF2DF8E0FDFA2E0E" bold="true" box="[151,577,1855,1881]" pageId="0" pageNumber="145">
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<taxonomicName id="24BF4D10C200FFA6FF2DF8E0FE672E0E" ID-CoL="9WYYR" box="[151,476,1855,1881]" class="Insecta" family="Gomphidae" genus="Phaenandrogomphus" kingdom="Animalia" order="Odonata" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="safei" status="sp. nov.">
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<emphasis id="D1CBEA81C200FFA6FF2DF8E0FE672E0E" bold="true" box="[151,476,1855,1881]" italics="true" pageId="0" pageNumber="145">Phaenandrogomphus safei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="CAF857FAC200FFA6FE59F8E0FDFA2E0E" box="[483,577,1855,1881]" pageId="0" pageNumber="145" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="E3003693C200FFA6FF2DF8BDFD222E2C" blockId="0.[151,665,1855,1915]" box="[151,665,1890,1915]" pageId="0" pageNumber="145">
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(
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<figureCitation id="7B842A16C200FFA6FF25F8BDFF572E2C" box="[159,236,1890,1915]" captionStart="FIGURES 1 – 2" captionStartId="1.[151,264,612,635]" captionTargetBox="[174,1434,195,586]" captionTargetId="figure@1.[151,1436,195,588]" captionTargetPageId="1" captionText="FIGURES 1 – 2. Habitus: (1) Phaenandrogomphus safei holotype; (2) Phaenandrogomphus treadawayi holotype." httpUri="https://zenodo.org/record/235480/files/figure.png" pageId="0" pageNumber="145">Figs. 1</figureCitation>
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,
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<figureCitation id="7B842A16C200FFA6FF40F8BCFEE52E2C" box="[250,350,1890,1915]" captionStart="FIGURES 3 – 6" captionStartId="1.[151,264,1680,1703]" captionTargetBox="[150,1436,684,1659]" captionTargetId="figure@1.[113,1474,684,1659]" captionTargetPageId="1" captionText="FIGURES 3 – 6. Phaenandrogomphus safei holotype: (3) head, anterior view; (4) head, dorsal view; (5) synthorax lateral view; (6) mesepisternum dorsal view." httpUri="https://zenodo.org/record/235481/files/figure.png" pageId="0" pageNumber="145">3, 4, 5, 6</figureCitation>
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,
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<figureCitation id="7B842A16C200FFA6FED0F8BCFE2E2E2C" box="[362,405,1891,1915]" captionStart="FIGURES 7 – 10" captionStartId="2.[151,264,1260,1283]" captionTargetBox="[157,1431,808,1220]" captionTargetId="figure@2.[153,1433,805,1224]" captionTargetPageId="2" captionText="FIGURES 7 – 10. Wings: (7) left forewing Phaenandrogomphus safei holotype; (8) right forewing Phaenandrogomphus asthenes holotype, flipped to left for comparison; (9) left hindwing Phaenandrogomphus safei holotype; (10) right hindwing Phaenandrogomphus asthenes holotype, flipped to left for comparison." httpUri="https://zenodo.org/record/235482/files/figure.png" pageId="0" pageNumber="145">7, 8</figureCitation>
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,
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<figureCitation id="7B842A16C200FFA6FE18F8BCFDA82E2C" box="[418,531,1891,1915]" captionStart="FIGURES 11 – 13" captionStartId="3.[151,264,1242,1265]" captionTargetBox="[151,1436,193,1200]" captionTargetId="figure@3.[151,1436,193,1203]" captionTargetPageId="3" captionText="FIGURES 11 – 13. Phaenandrogomphus safei holotype: (11) accessory genitalia, lateral view; (12) terminal segment of penis, ventral view; (13) penis vesicle, ventral-posterior view, posterior apex indicated by arrow." httpUri="https://zenodo.org/record/235483/files/figure.png" pageId="0" pageNumber="145">11, 12, 13</figureCitation>
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,
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<figureCitation id="7B842A16C200FFA6FDA5F8BCFD862E2C" box="[543,573,1891,1915]" captionStart="FIGURE 14" captionStartId="3.[151,250,1905,1927]" captionText="FIGURE 14. Phaenandrogomphus safei holotype, lateral view anal appendages." pageId="0" pageNumber="145">14</figureCitation>
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,
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<figureCitation id="7B842A16C200FFA6FDF3F8BDFD2B2E2C" box="[585,656,1890,1915]" captionStart="FIGURES 15 – 16" captionStartId="4.[151,264,561,584]" captionTargetBox="[151,1435,193,540]" captionTargetId="figure@4.[151,1436,193,541]" captionTargetPageId="4" captionText="FIGURES 15 – 16. Phaenandrogomphus safei holotype, anal appendages: (15) dorsal view; (16) ventral view." httpUri="https://zenodo.org/record/235484/files/figure.png" pageId="0" pageNumber="145">15, 16</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="ABA56518C200FFA6FF2DF876FC252EB0" pageId="0" pageNumber="145" type="materials_examined">
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<paragraph id="E3003693C200FFA6FF2DF876FC252EB0" blockId="0.[151,1436,1961,2023]" pageId="0" pageNumber="145">
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<emphasis id="D1CBEA81C200FFA6FF2DF876FE3E2E95" bold="true" box="[151,389,1961,1986]" pageId="0" pageNumber="145">
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Material.
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<typeStatus id="3C048831C200FFA6FEB5F875FE3B2E95" box="[271,384,1962,1986]" pageId="0" pageNumber="145" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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♂,
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<collectingCountry id="9BA87603C200FFA6FE09F875FDA62E94" box="[435,541,1962,1987]" name="Malaysia" pageId="0" pageNumber="145">Malaysia</collectingCountry>
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, Sabah, Kalabakan Forest Reserve, 4 ̊43’N, 117 ̊37’E,
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<date id="97011053C200FFA6FB19F874FA992E94" box="[1187,1314,1962,1987]" pageId="0" pageNumber="145" value="2013-04-29">29 iv 2013</date>
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, leg. S. H. Luke. To be deposited at the Forest Research Centre, Sepilok, Sabah.
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</paragraph>
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</subSubSection>
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<caption id="B7C0661BC201FFA7FF2DFDBBFAA82B2D" box="[151,1299,612,635]" httpUri="https://zenodo.org/record/235480/files/figure.png" pageId="1" pageNumber="146" targetBox="[174,1434,195,586]" targetPageId="1">
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<paragraph id="E3003693C201FFA7FF2DFDBBFAA82B2D" blockId="1.[151,1299,612,635]" box="[151,1299,612,635]" pageId="1" pageNumber="146">
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<emphasis id="D1CBEA81C201FFA7FF2DFDBBFE812B2E" bold="true" box="[151,314,612,635]" pageId="1" pageNumber="146">FIGURES 1–2.</emphasis>
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Habitus: (1)
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<taxonomicName id="24BF4D10C201FFA7FE7AFDBBFD752B2D" box="[448,718,612,634]" class="Insecta" family="Gomphidae" genus="Phaenandrogomphus" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="146" phylum="Arthropoda" rank="species" species="safei">
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<emphasis id="D1CBEA81C201FFA7FE7AFDBBFD752B2D" box="[448,718,612,634]" italics="true" pageId="1" pageNumber="146">Phaenandrogomphus safei</emphasis>
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</taxonomicName>
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holotype; (2)
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<taxonomicName id="24BF4D10C201FFA7FCE4FDBBFB162B2D" box="[862,1197,612,634]" class="Insecta" family="Gomphidae" genus="Phaenandrogomphus" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="146" phylum="Arthropoda" rank="species" species="treadawayi">
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<emphasis id="D1CBEA81C201FFA7FCE4FDBBFB162B2D" box="[862,1197,612,634]" italics="true" pageId="1" pageNumber="146">Phaenandrogomphus treadawayi</emphasis>
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</taxonomicName>
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holotype.
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</paragraph>
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</caption>
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<caption id="B7C0661BC201FFA7FF2DF94FFE6E2F91" httpUri="https://zenodo.org/record/235481/files/figure.png" pageId="1" pageNumber="146" targetBox="[150,1436,684,1659]" targetPageId="1">
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<paragraph id="E3003693C201FFA7FF2DF94FFE6E2F91" blockId="1.[151,1436,1680,1734]" pageId="1" pageNumber="146">
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<emphasis id="D1CBEA81C201FFA7FF2DF94FFE812FF1" bold="true" box="[151,314,1680,1703]" pageId="1" pageNumber="146">FIGURES 3–6.</emphasis>
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<taxonomicName id="24BF4D10C201FFA7FEFAF94EFDF52FF0" box="[320,590,1681,1703]" class="Insecta" family="Gomphidae" genus="Phaenandrogomphus" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="146" phylum="Arthropoda" rank="species" species="safei">
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<emphasis id="D1CBEA81C201FFA7FEFAF94EFDF52FF0" box="[320,590,1681,1703]" italics="true" pageId="1" pageNumber="146">Phaenandrogomphus safei</emphasis>
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</taxonomicName>
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holotype: (3) head, anterior view; (4) head, dorsal view; (5) synthorax lateral view; (6) mesepisternum dorsal view.
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</paragraph>
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</caption>
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<subSubSection id="ABA56518C201FFA7FF7DF932FC212E7D" pageId="1" pageNumber="146" type="etymology">
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<paragraph id="E3003693C201FFA7FF7DF932FC212E7D" blockId="1.[151,1437,1773,2014]" pageId="1" pageNumber="146">
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<emphasis id="D1CBEA81C201FFA7FF7DF932FEF52E52" bold="true" box="[199,334,1773,1797]" pageId="1" pageNumber="146">Etymology.</emphasis>
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<taxonomicName id="24BF4D10C201FFA7FEE2F932FE362E52" box="[344,397,1773,1797]" class="Insecta" family="Gomphidae" genus="Phaenandrogomphus" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="146" phylum="Arthropoda" rank="species" species="safei">
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<emphasis id="D1CBEA81C201FFA7FEE2F932FE362E52" box="[344,397,1773,1797]" italics="true" pageId="1" pageNumber="146">safei</emphasis>
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</taxonomicName>
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, a noun in the genitive case, formed from the acronym of the Stability of Altered Forest Ecosystems (SAFE) Project, where the
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<typeStatus id="3C048831C201FFA7FDE8F8CDFD392E7D" box="[594,642,1810,1834]" pageId="1" pageNumber="146">type</typeStatus>
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specimen was collected.
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</paragraph>
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</subSubSection>
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<subSubSection id="ABA56518C201FFA4FF7DF8EBFAA92EBE" lastPageId="2" lastPageNumber="147" pageId="1" pageNumber="146" type="description">
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<paragraph id="E3003693C201FFA4FF7DF8EBFDC829AF" blockId="1.[151,1437,1773,2014]" lastBlockId="2.[151,1437,151,752]" lastPageId="2" lastPageNumber="147" pageId="1" pageNumber="146">
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<emphasis id="D1CBEA81C201FFA7FF7DF8EBFD952E1A" bold="true" box="[199,558,1844,1869]" pageId="1" pageNumber="146">
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Description of
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<typeStatus id="3C048831C201FFA7FEC6F8EAFE582E1A" box="[380,483,1845,1869]" pageId="1" pageNumber="146" type="holotype">holotype</typeStatus>
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male.
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</emphasis>
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Head (
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<figureCitation id="7B842A16C201FFA7FD3FF8EAFD4D2E19" box="[645,758,1845,1870]" captionStart="FIGURES 3 – 6" captionStartId="1.[151,264,1680,1703]" captionTargetBox="[150,1436,684,1659]" captionTargetId="figure@1.[113,1474,684,1659]" captionTargetPageId="1" captionText="FIGURES 3 – 6. Phaenandrogomphus safei holotype: (3) head, anterior view; (4) head, dorsal view; (5) synthorax lateral view; (6) mesepisternum dorsal view." httpUri="https://zenodo.org/record/235481/files/figure.png" pageId="1" pageNumber="146">Figs. 3–4</figureCitation>
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): Labium pale grey centrally, whitish basally and laterally, black along free margin of median lobe and narrowly on apical outer margin of palps. Labrum black with pair of broad pale markings in upper half, narrowly divided centrally. Mandible bases pale, genae mostly olivaceous. Anteclypeus with lower margin slightly concave (
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<figureCitation id="7B842A16C201FFA7FD6AF87EFCAE2EED" box="[720,789,1953,1978]" captionStart="FIGURES 3 – 6" captionStartId="1.[151,264,1680,1703]" captionTargetBox="[150,1436,684,1659]" captionTargetId="figure@1.[113,1474,684,1659]" captionTargetPageId="1" captionText="FIGURES 3 – 6. Phaenandrogomphus safei holotype: (3) head, anterior view; (4) head, dorsal view; (5) synthorax lateral view; (6) mesepisternum dorsal view." httpUri="https://zenodo.org/record/235481/files/figure.png" pageId="1" pageNumber="146">Fig. 3</figureCitation>
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), greyish-green coloured. Postclypeus mostly black with pair of pale lateral marks. Ante- and postfrons not very sharply divided, black with green transverse stripe on anterior half of postfrons (
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<figureCitation id="7B842A16C202FFA4FE78FF48FDB229E7" box="[450,521,151,176]" captionStart="FIGURES 3 – 6" captionStartId="1.[151,264,1680,1703]" captionTargetBox="[150,1436,684,1659]" captionTargetId="figure@1.[113,1474,684,1659]" captionTargetPageId="1" captionText="FIGURES 3 – 6. Phaenandrogomphus safei holotype: (3) head, anterior view; (4) head, dorsal view; (5) synthorax lateral view; (6) mesepisternum dorsal view." httpUri="https://zenodo.org/record/235481/files/figure.png" pageId="2" pageNumber="147">Fig. 4</figureCitation>
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). Vertex and occiput mostly black to very dark brown, with poorly defined paler areas behind median ocellus and between eyes at level of their closest approach to one another. Pair of tubercles behind lateral ocelli; ocelli pale yellowish.
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</paragraph>
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<paragraph id="E3003693C202FFA4FF7DFEDBFAC92BB8" blockId="2.[151,1437,151,752]" pageId="2" pageNumber="147">
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Thorax: Prothorax almost entirely dark brown and black, except anterior pronotal lobe, where central part of anterior carina and area behind pale. Synthorax dark brown to black with pale yellow-green markings as follows (
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<figureCitation id="7B842A16C202FFA4FF25FE93FEB62832" box="[159,269,332,357]" captionStart="FIGURES 3 – 6" captionStartId="1.[151,264,1680,1703]" captionTargetBox="[150,1436,684,1659]" captionTargetId="figure@1.[113,1474,684,1659]" captionTargetPageId="1" captionText="FIGURES 3 – 6. Phaenandrogomphus safei holotype: (3) head, anterior view; (4) head, dorsal view; (5) synthorax lateral view; (6) mesepisternum dorsal view." httpUri="https://zenodo.org/record/235481/files/figure.png" pageId="2" pageNumber="147">Figs. 5–6</figureCitation>
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): short mesothoracic collar, narrowly divided at mid-dorsal carina, joined to narrow antehumeral stripes that extend beyond level of apex of antealar triangle, terminating just short of antealar carina. Small spot below antehumeral marking on mespisternum, near antealar carina. Stripe running length of mesepimeron. Metepisternum with stripe running below spiracle from metinfraepisternum, separated from ca triangular spot at antealar carina. A broad stripe occupying all of metepimeron except narrowly below metapleural suture. Mesinfraepisternum mostly dark brown, metinfraepisternum pale. Venter pale with brown stripe on either side. Legs robust and relatively short with posterior femur not reaching base of S1. Coxae mostly pale, trochanters brown and black, flexor surface of anterior femora, and distal ca one-third of flexor surface of middle femora pale, rest mostly black. Wings (
|
||
<figureCitation id="7B842A16C202FFA4FE6FFDB3FDF12BD3" box="[469,586,620,645]" captionStart="FIGURES 7 – 10" captionStartId="2.[151,264,1260,1283]" captionTargetBox="[157,1431,808,1220]" captionTargetId="figure@2.[153,1433,805,1224]" captionTargetPageId="2" captionText="FIGURES 7 – 10. Wings: (7) left forewing Phaenandrogomphus safei holotype; (8) right forewing Phaenandrogomphus asthenes holotype, flipped to left for comparison; (9) left hindwing Phaenandrogomphus safei holotype; (10) right hindwing Phaenandrogomphus asthenes holotype, flipped to left for comparison." httpUri="https://zenodo.org/record/235482/files/figure.png" pageId="2" pageNumber="147">Figs. 7–8</figureCitation>
|
||
): Sectors of arculus separated at origin with 2 cross veins before first bifurcation of superior sector in Fw and
|
||
<quantity id="24479B76C202FFA4FDE7FD4FFD332BFF" box="[605,648,655,680]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="2" pageNumber="147" unit="in" value="1.0">1 in</quantity>
|
||
Hw. Discoidal field with 2 rows of cells from origin to level of nodus in both wings. 13 (left) or 15 (right) Ax in Fw, 9 (left) or 10 (right) in Hw, 8 (left) or 9 (right) Px in Fw, 8 (left) or 9 (right) in Hw. Pt very dark brown, covering just over 4 underlying cells in Fw, ca 4-1/2 underlying cells in Hw.
|
||
</paragraph>
|
||
<caption id="B7C0661BC202FFA4FF2DFB33FCD22C17" httpUri="https://zenodo.org/record/235482/files/figure.png" pageId="2" pageNumber="147" targetBox="[157,1431,808,1220]" targetPageId="2">
|
||
<paragraph id="E3003693C202FFA4FF2DFB33FCD22C17" blockId="2.[151,1436,1260,1344]" pageId="2" pageNumber="147">
|
||
<emphasis id="D1CBEA81C202FFA4FF2DFB33FEF72C55" bold="true" box="[151,332,1260,1283]" pageId="2" pageNumber="147">FIGURES 7–10.</emphasis>
|
||
Wings: (7) left forewing
|
||
<taxonomicName id="24BF4D10C202FFA4FDD6FB32FCC42C54" box="[620,895,1261,1283]" class="Insecta" family="Gomphidae" genus="Phaenandrogomphus" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="147" phylum="Arthropoda" rank="species" species="safei">
|
||
<emphasis id="D1CBEA81C202FFA4FDD6FB32FCC42C54" box="[620,895,1261,1283]" italics="true" pageId="2" pageNumber="147">Phaenandrogomphus safei</emphasis>
|
||
</taxonomicName>
|
||
holotype; (8) right forewing
|
||
<taxonomicName id="24BF4D10C202FFA4FB7EFB32FF552C75" class="Insecta" family="Gomphidae" genus="Phaenandrogomphus" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="147" phylum="Arthropoda" rank="species" species="asthenes">
|
||
<emphasis id="D1CBEA81C202FFA4FB7EFB32FF552C75" italics="true" pageId="2" pageNumber="147">Phaenandrogomphus asthenes</emphasis>
|
||
</taxonomicName>
|
||
holotype, flipped to left for comparison; (9) left hindwing
|
||
<taxonomicName id="24BF4D10C202FFA4FCE9FAD3FBD82C75" box="[851,1123,1292,1314]" class="Insecta" family="Gomphidae" genus="Phaenandrogomphus" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="147" phylum="Arthropoda" rank="species" species="safei">
|
||
<emphasis id="D1CBEA81C202FFA4FCE9FAD3FBD82C75" box="[851,1123,1292,1314]" italics="true" pageId="2" pageNumber="147">Phaenandrogomphus safei</emphasis>
|
||
</taxonomicName>
|
||
holotype; (10) right hindwing
|
||
<taxonomicName id="24BF4D10C202FFA4FF2DFAF5FE772C17" box="[151,460,1322,1344]" class="Insecta" family="Gomphidae" genus="Phaenandrogomphus" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="147" phylum="Arthropoda" rank="species" species="asthenes">
|
||
<emphasis id="D1CBEA81C202FFA4FF2DFAF5FE772C17" box="[151,460,1322,1344]" italics="true" pageId="2" pageNumber="147">Phaenandrogomphus asthenes</emphasis>
|
||
</taxonomicName>
|
||
holotype, flipped to left for comparison.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E3003693C202FFA4FF7DFAB2FD8C2E91" blockId="2.[151,1437,1389,2026]" pageId="2" pageNumber="147">
|
||
Abdomen: Slender after base of S3, expanding moderately from base of S7, maximum width and height reached apical part of S8, then almost constant. Black with pale markings as follows (
|
||
<figureCitation id="7B842A16C202FFA4FB36FA4EFB692CFE" box="[1164,1234,1425,1450]" captionStart="FIGURES 1 – 2" captionStartId="1.[151,264,612,635]" captionTargetBox="[174,1434,195,586]" captionTargetId="figure@1.[151,1436,195,588]" captionTargetPageId="1" captionText="FIGURES 1 – 2. Habitus: (1) Phaenandrogomphus safei holotype; (2) Phaenandrogomphus treadawayi holotype." httpUri="https://zenodo.org/record/235480/files/figure.png" pageId="2" pageNumber="147">Fig. 1</figureCitation>
|
||
): S1 mostly pale yellowish laterally. S2 pale yellowish around and including auricle laterally, except free margin of auricle, another yellowish lateral mark placed posteriorly, narrow yellowish mid-dorsal stripe. S3–6 with small dorso-lateral markings at base, divided dorsally. S3–4 with small elongate oval yellow mark mid-dorsally, this reduced to small spot on S5 and absent on S6. S7 with more extensive yellow dorso-lateral marking at base, occupying slightly less than two-thirds of tergite and narrowly divided dorsally. S8–10 entirely black. Cerci (
|
||
<figureCitation id="7B842A16C202FFA4FBD4F99AFB4C2F09" box="[1134,1271,1605,1630]" captionStart-0="FIGURE 14" captionStart-1="FIGURES 15 – 16" captionStartId-0="3.[151,250,1905,1927]" captionStartId-1="4.[151,264,561,584]" captionTargetBox-1="[151,1435,193,540]" captionTargetId-1="figure@4.[151,1436,193,541]" captionTargetPageId-1="4" captionText-0="FIGURE 14. Phaenandrogomphus safei holotype, lateral view anal appendages." captionText-1="FIGURES 15 – 16. Phaenandrogomphus safei holotype, anal appendages: (15) dorsal view; (16) ventral view." httpUri-1="https://zenodo.org/record/235484/files/figure.png" pageId="2" pageNumber="147">Figs. 14–15</figureCitation>
|
||
) curved down and moderately dorso-ventrally expanded in apical ca one-third, bearing robust teeth on inner edge apically. Pale brown at base, darkening towards apices, especially on upper surface where almost black. Epiproct (
|
||
<figureCitation id="7B842A16C202FFA4FAB1F952FAD12FF1" box="[1291,1386,1677,1702]" captionStart="FIGURE 14" captionStartId="3.[151,250,1905,1927]" captionText="FIGURE 14. Phaenandrogomphus safei holotype, lateral view anal appendages." pageId="2" pageNumber="147">Figs. 14</figureCitation>
|
||
,
|
||
<figureCitation id="7B842A16C202FFA4FACDF952FA2F2FF1" box="[1399,1428,1677,1702]" captionStart="FIGURES 15 – 16" captionStartId="4.[151,264,561,584]" captionTargetBox="[151,1435,193,540]" captionTargetId="figure@4.[151,1436,193,541]" captionTargetPageId="4" captionText="FIGURES 15 – 16. Phaenandrogomphus safei holotype, anal appendages: (15) dorsal view; (16) ventral view." httpUri="https://zenodo.org/record/235484/files/figure.png" pageId="2" pageNumber="147">16</figureCitation>
|
||
) black, shorter than cerci, deeply divided, branches running approximately parallel on their inner margins for almost their entire length, apices pointed and slightly out-turned. In lateral view (
|
||
<figureCitation id="7B842A16C202FFA4FC66F90AFB952FB9" box="[988,1070,1749,1774]" captionStart="FIGURE 14" captionStartId="3.[151,250,1905,1927]" captionText="FIGURE 14. Phaenandrogomphus safei holotype, lateral view anal appendages." pageId="2" pageNumber="147">Fig. 14</figureCitation>
|
||
) branches curved downward for short distance from base, then straightening, with upper margin abruptly raised at this point, before curving upwards again at ca two-thirds length, finally straightening again apically, where upper margin gently concave before apex. Accessory genitalia as shown in
|
||
<figureCitation id="7B842A16C202FFA4FD24F89EFC9E2E0E" box="[670,805,1857,1882]" captionStart="FIGURES 11 – 13" captionStartId="3.[151,264,1242,1265]" captionTargetBox="[151,1436,193,1200]" captionTargetId="figure@3.[151,1436,193,1203]" captionTargetPageId="3" captionText="FIGURES 11 – 13. Phaenandrogomphus safei holotype: (11) accessory genitalia, lateral view; (12) terminal segment of penis, ventral view; (13) penis vesicle, ventral-posterior view, posterior apex indicated by arrow." httpUri="https://zenodo.org/record/235483/files/figure.png" pageId="2" pageNumber="147">Figs. 11–13</figureCitation>
|
||
, with anterior hamule moderate sized, directed posteroventrally, apical half narrow and hook shaped, penis with terminal segment (
|
||
<figureCitation id="7B842A16C202FFA4FBAAF8BAFB2C2E29" box="[1040,1175,1893,1918]" captionStart="FIGURES 11 – 13" captionStartId="3.[151,264,1242,1265]" captionTargetBox="[151,1436,193,1200]" captionTargetId="figure@3.[151,1436,193,1203]" captionTargetPageId="3" captionText="FIGURES 11 – 13. Phaenandrogomphus safei holotype: (11) accessory genitalia, lateral view; (12) terminal segment of penis, ventral view; (13) penis vesicle, ventral-posterior view, posterior apex indicated by arrow." httpUri="https://zenodo.org/record/235483/files/figure.png" pageId="2" pageNumber="147">Figs. 11–12</figureCitation>
|
||
) divided centrally and bearing cornua, penis vesicle moderately large, but not extending ventrally beyond level of hamuli, its posterior apex only weakly bifurcate (
|
||
<figureCitation id="7B842A16C202FFA4FE6CF872FD922E91" box="[470,553,1965,1990]" captionStart="FIGURES 11 – 13" captionStartId="3.[151,264,1242,1265]" captionTargetBox="[151,1436,193,1200]" captionTargetId="figure@3.[151,1436,193,1203]" captionTargetPageId="3" captionText="FIGURES 11 – 13. Phaenandrogomphus safei holotype: (11) accessory genitalia, lateral view; (12) terminal segment of penis, ventral view; (13) penis vesicle, ventral-posterior view, posterior apex indicated by arrow." httpUri="https://zenodo.org/record/235483/files/figure.png" pageId="2" pageNumber="147">Fig. 13</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E3003693C202FFA4FF7DF80EFAA92EBE" blockId="2.[151,1437,1389,2026]" box="[199,1298,2001,2026]" pageId="2" pageNumber="147">
|
||
<emphasis id="D1CBEA81C202FFA4FF7DF80EFECD2EBE" bold="true" box="[199,374,2001,2025]" pageId="2" pageNumber="147">Measurements</emphasis>
|
||
[mm]: Hw 26, abdomen without anal appendages 31.5, cerci ca 3, epiproct ca 2.7.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="B7C0661BC203FFA5FF2DFB05FB9D2C58" httpUri="https://zenodo.org/record/235483/files/figure.png" pageId="3" pageNumber="148" targetBox="[151,1436,193,1200]" targetPageId="3">
|
||
<paragraph id="E3003693C203FFA5FF2DFB05FB9D2C58" blockId="3.[151,1436,1242,1295]" pageId="3" pageNumber="148">
|
||
<emphasis id="D1CBEA81C203FFA5FF2DFB05FEE82DA7" bold="true" box="[151,339,1242,1265]" pageId="3" pageNumber="148">FIGURES 11–13.</emphasis>
|
||
<taxonomicName id="24BF4D10C203FFA5FEE0FB04FDD22DA6" box="[346,617,1243,1265]" class="Insecta" family="Gomphidae" genus="Phaenandrogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="148" phylum="Arthropoda" rank="species" species="safei">
|
||
<emphasis id="D1CBEA81C203FFA5FEE0FB04FDD22DA6" box="[346,617,1243,1265]" italics="true" pageId="3" pageNumber="148">Phaenandrogomphus safei</emphasis>
|
||
</taxonomicName>
|
||
holotype: (11) accessory genitalia, lateral view; (12) terminal segment of penis, ventral view; (13) penis vesicle, ventral-posterior view, posterior apex indicated by arrow.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B7C0661BC203FFA5FF2DF8AEFC712ED0" box="[151,970,1905,1927]" pageId="3" pageNumber="148">
|
||
<paragraph id="E3003693C203FFA5FF2DF8AEFC712ED0" blockId="3.[151,970,1905,1927]" box="[151,970,1905,1927]" pageId="3" pageNumber="148">
|
||
<emphasis id="D1CBEA81C203FFA5FF2DF8AEFEA42ED1" bold="true" box="[151,287,1905,1927]" pageId="3" pageNumber="148">FIGURE 14.</emphasis>
|
||
<taxonomicName id="24BF4D10C203FFA5FE9FF8AEFD8F2ED0" box="[293,564,1905,1927]" class="Insecta" family="Gomphidae" genus="Phaenandrogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="148" phylum="Arthropoda" rank="species" species="safei">
|
||
<emphasis id="D1CBEA81C203FFA5FE9FF8AEFD8F2ED0" box="[293,564,1905,1927]" italics="true" pageId="3" pageNumber="148">Phaenandrogomphus safei</emphasis>
|
||
</taxonomicName>
|
||
holotype, lateral view anal appendages.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B7C0661BC204FFA2FF2DFDEEFB552B1F" box="[151,1262,561,584]" httpUri="https://zenodo.org/record/235484/files/figure.png" pageId="4" pageNumber="149" targetBox="[151,1435,193,540]" targetPageId="4">
|
||
<paragraph id="E3003693C204FFA2FF2DFDEEFB552B1F" blockId="4.[151,1262,561,584]" box="[151,1262,561,584]" pageId="4" pageNumber="149">
|
||
<emphasis id="D1CBEA81C204FFA2FF2DFDEEFEE82B10" bold="true" box="[151,339,561,584]" pageId="4" pageNumber="149">FIGURES 15–16.</emphasis>
|
||
<taxonomicName id="24BF4D10C204FFA2FEE0FDEDFDDC2B1F" box="[346,615,562,584]" class="Insecta" family="Gomphidae" genus="Phaenandrogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="149" phylum="Arthropoda" rank="species" species="safei">
|
||
<emphasis id="D1CBEA81C204FFA2FEE0FDEDFDDC2B1F" box="[346,615,562,584]" italics="true" pageId="4" pageNumber="149">Phaenandrogomphus safei</emphasis>
|
||
</taxonomicName>
|
||
holotype, anal appendages: (15) dorsal view; (16) ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="ABA56518C204FFA2FF7DFDABFBEA2A87" pageId="4" pageNumber="149" type="diagnosis">
|
||
<paragraph id="E3003693C204FFA2FF7DFDABFBEA2A87" blockId="4.[151,1437,628,1158]" pageId="4" pageNumber="149">
|
||
<emphasis id="D1CBEA81C204FFA2FF7DFDABFEFA2BDA" bold="true" box="[199,321,628,653]" pageId="4" pageNumber="149">Diagnosis.</emphasis>
|
||
A moderately small gomphid (
|
||
<figureCitation id="7B842A16C204FFA2FD78FDABFC932BDA" box="[706,808,628,653]" captionStart="FIGURES 1 – 2" captionStartId="1.[151,264,612,635]" captionTargetBox="[174,1434,195,586]" captionTargetId="figure@1.[151,1436,195,588]" captionTargetPageId="1" captionText="FIGURES 1 – 2. Habitus: (1) Phaenandrogomphus safei holotype; (2) Phaenandrogomphus treadawayi holotype." httpUri="https://zenodo.org/record/235480/files/figure.png" pageId="4" pageNumber="149">Figure 1</figureCitation>
|
||
), brightly coloured in life. It differs from all other
|
||
<taxonomicName id="24BF4D10C204FFA2FF2DFD46FE3E2BE6" box="[151,389,665,689]" class="Insecta" family="Gomphidae" genus="Phaenandrogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="149" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D1CBEA81C204FFA2FF2DFD46FE3E2BE6" box="[151,389,665,689]" italics="true" pageId="4" pageNumber="149">Phaenandrogomphus</emphasis>
|
||
</taxonomicName>
|
||
species except
|
||
<taxonomicName id="24BF4D10C204FFA2FD87FD46FD632BE6" box="[573,728,665,689]" class="Insecta" family="Gomphidae" genus="Phaenandrogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="149" phylum="Arthropoda" rank="species" species="treadawayi">
|
||
<emphasis id="D1CBEA81C204FFA2FD87FD46FD632BE6" box="[573,728,665,689]" italics="true" pageId="4" pageNumber="149">P. treadawayi</emphasis>
|
||
</taxonomicName>
|
||
in having a smaller penis vesicle with posterior part of apex much less strongly bifurcated (compare with
|
||
<figureCitation id="7B842A16C204FFA2FD23FD63FD672B82" box="[665,732,700,725]" captionStart="FIGURES 3 – 6" captionStartId="1.[151,264,1680,1703]" captionTargetBox="[150,1436,684,1659]" captionTargetId="figure@1.[113,1474,684,1659]" captionTargetPageId="1" captionText="FIGURES 3 – 6. Phaenandrogomphus safei holotype: (3) head, anterior view; (4) head, dorsal view; (5) synthorax lateral view; (6) mesepisternum dorsal view." httpUri="https://zenodo.org/record/235481/files/figure.png" pageId="4" pageNumber="149">
|
||
Fig.
|
||
<quantity id="24479B76C204FFA2FD74FD62FD672B82" box="[718,732,701,725]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="4" pageNumber="149" unit="in" value="3.0">3</quantity>
|
||
</figureCitation>
|
||
in
|
||
<bibRefCitation id="872E4B62C204FFA2FCB8FD63FC732B82" author="Lieftinck" box="[770,968,700,725]" pageId="4" pageNumber="149" refString="Lieftinck, M. A. (1964) Some Gomphidae and their larvae, cheifly from the Malay peninsula (Odonata). Zoologische Verhandelingen, 69, 1 - 38." type="journal article" year="1964">Lieftinck (1964))</bibRefCitation>
|
||
. It also differs in the form of the anterior hamule, which in the mainland species, while ventrally directed at base, makes an abrupt turn posteriad shortly after it emerges from the genital fossa. Additional differences from the mainland species include the less extensive pale markings on thorax and abdomen and sectors of arculus well separated at origin.
|
||
<taxonomicName id="24BF4D10C204FFA2FBEBFCF6FB1A2A16" box="[1105,1185,809,833]" class="Insecta" family="Gomphidae" genus="Phaenandrogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="149" phylum="Arthropoda" rank="species" species="safei">
|
||
<emphasis id="D1CBEA81C204FFA2FBEBFCF6FB1A2A16" box="[1105,1185,809,833]" italics="true" pageId="4" pageNumber="149">P. safei</emphasis>
|
||
</taxonomicName>
|
||
can be separated from
|
||
<taxonomicName id="24BF4D10C204FFA2FF2DFC91FE892A32" box="[151,306,845,869]" class="Insecta" family="Gomphidae" genus="Phaenandrogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="149" phylum="Arthropoda" rank="species" species="treadawayi">
|
||
<emphasis id="D1CBEA81C204FFA2FF2DFC91FE892A32" box="[151,306,845,869]" italics="true" pageId="4" pageNumber="149">P. treadawayi</emphasis>
|
||
</taxonomicName>
|
||
by the accessory genitalia with terminal hooked part of anterior hamule longer and penis vesicle differently shaped in lateral view, more flattened terminal segment of the penis, the markings of the synthorax, the more downcurved and expanded apical part of the cerci with large apical teeth, and the epiproct which has the upper margin expanded abruptly and is upturned more apically than in
|
||
<taxonomicName id="24BF4D10C204FFA2FC08FC66FBF12A86" box="[946,1098,953,977]" class="Insecta" family="Gomphidae" genus="Phaenandrogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="149" phylum="Arthropoda" rank="species" species="treadawayi">
|
||
<emphasis id="D1CBEA81C204FFA2FC08FC66FBF12A86" box="[946,1098,953,977]" italics="true" pageId="4" pageNumber="149">P. treadawayi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="ABA56518C204FFA2FF7DFC02FE4A2DD1" pageId="4" pageNumber="149" type="discussion">
|
||
<paragraph id="E3003693C204FFA2FF7DFC02FE4A2DD1" blockId="4.[151,1437,628,1158]" pageId="4" pageNumber="149">
|
||
<emphasis id="D1CBEA81C204FFA2FF7DFC02FE802AA2" bold="true" box="[199,315,989,1013]" pageId="4" pageNumber="149">Remarks.</emphasis>
|
||
The
|
||
<typeStatus id="3C048831C204FFA2FEC3FC02FE602AA1" box="[377,475,989,1014]" pageId="4" pageNumber="149" type="holotype">holotype</typeStatus>
|
||
was found flying over a shallow stream, approximately
|
||
<quantity id="24479B76C204FFA2FBD0FC02FB1D2AA1" box="[1130,1190,989,1014]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="149" unit="km" value="2.0">2 km</quantity>
|
||
downstream from its source, and flowing through lowland dipterocarp forest that had previously been selectively logged several times. The stream channel was fairly open following a large flooding event in 2011 that caused long distance transport of logs and extensive erosion of stream banks. In life the pale markings of the synthorax were more blue-green than yellow and the eyes were blue.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |