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<mods:title id="1F42767CBF6D0BB3BCDA818ECEA54678">A new classification of the Chirostyloidea (Crustacea: Decapoda: Anomura)</mods:title>
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<mods:affiliation id="C1A9314E32BBD200B8071749EC7CA950">Australian Museum, 6 College Street, Sydney, NSW 2010 Australia. E-mail: shane. ahyong @ austmus. gov. au</mods:affiliation>
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<taxonomicName id="B7C13695C649FFC97C8DFE9EFDC6A211" ID-CoL="aa325b06-615d-45e3-8e9d-41b275c95cab" authority="Ortmann, 1892" authorityName="Ortmann" authorityYear="1892" box="[151,518,322,348]" class="Malacostraca" family="Chirostylidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="58" phylum="Arthropoda" rank="family">
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Chirostylidae
<bibRefCitation id="145030E7C649FFC97D5DFE9EFDC6A211" author="Ortmann, A." box="[327,518,322,348]" pageId="2" pageNumber="58" pagination="241 - 326" refId="ref6010" refString="Ortmann, A. (1892) Die Decapoden-Krebse des Strassbuger Museums IV. Die Abteilungen Galatheidea und Paguridea. Zoologische Jahrbucher. Abteilung fuer Systematic Oekologie und Geographie der Tiere, 6, 241 - 326, pls 11, 12." type="book chapter" year="1892">Ortmann, 1892</bibRefCitation>
</emphasis>
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<subSubSection id="38DB1E9DC649FFC97C8CFEB9FBA0A14A" pageId="2" pageNumber="58" type="description">
<paragraph id="707E4D16C649FFC97C8CFEB9FE97A232" blockId="2.[150,518,322,383]" box="[150,343,357,383]" pageId="2" pageNumber="58">
(
<figureCitation id="E8FA5193C649FFC97C85FEB9FE90A232" box="[159,336,357,383]" captionStart="FIGURE 2" captionStartId="4.[151,258,1730,1754]" captionTargetBox="[171,1426,225,1673]" captionTargetId="figure-150@4.[151,1436,197,1709]" captionTargetPageId="4" captionText="FIGURE 2. AF, mandibular dentition, ventral view. GL, right maxilliped 1 endopod and exopod, anterior view (epipod indicated by arrow). Eumunididae: A, G, Eumunida australis de Saint Laurent &amp; Macpherson, 1990b (♂, pcl 12.9 mm, NMNZ Cr.21765); B, H, Pseudomunida fragilis Haig, 1979 (♀ ovig., pcl 20.5 mm, MNHN-Ga7507). Chirostylidae: C, I, Chirostylus dolichopus Ortmann, 1892 (♂, pcl 2.7 mm, MNHN station LA REUNION, MD32, CP172); D, J, Gastroptychus rogeri Baba, 2000 (♂, pcl 23.2 mm, NIWA 14555); E, K, Uroptychodes epigaster Baba, 2004 (♀, pcl 3.4 mm, NMNZ Cr.21766); F, L, Uroptychus cardus Ahyong &amp; Poore, 2004 (♂, pcl 10.5 mm, NIWA 23086). Scale: AF, not to scale; G, H, J, L = 1.00 mm; I, K = 0.45 mm." figureDoi="http://doi.org/10.5281/zenodo.5301287" httpUri="https://zenodo.org/record/5301287/files/figure.png" pageId="2" pageNumber="58">Fig. 2CF, IL</figureCitation>
)
</paragraph>
<paragraph id="707E4D16C649FFC97C8DFE6CFB0DA28A" blockId="2.[151,1308,432,583]" box="[151,1229,432,455]" pageId="2" pageNumber="58">
Diptycinés A.
<bibRefCitation id="145030E7C649FFC97D28FE6CFD09A28A" author="Milne-Edwards, A. &amp; Bouvier, E. L." box="[306,713,432,455]" pageId="2" pageNumber="58" pagination="191 - 327" refId="ref5606" refString="Milne-Edwards, A. &amp; Bouvier, E. L. (1894) Considerations generales sur la famille des Galatheides. Annales des Sciences Naturelles, Zoologie, series 7, 16, 191 - 327, fig. 1 - 36." type="journal article" year="1894">Milne-Edwards &amp; Bouvier, 1894: 296</bibRefCitation>
, 312; 1897: 116 [vernacular name, unavailable].
</paragraph>
<paragraph id="707E4D16C649FFC97C8DFE0CFADCA2AA" blockId="2.[151,1308,432,583]" box="[151,1308,464,487]" pageId="2" pageNumber="58">
Diptyciens A.
<bibRefCitation id="145030E7C649FFC97D28FE0CFD0AA2AA" author="Milne-Edwards, A. &amp; Bouvier, E. L." box="[306,714,464,487]" pageId="2" pageNumber="58" pagination="191 - 327" refId="ref5606" refString="Milne-Edwards, A. &amp; Bouvier, E. L. (1894) Considerations generales sur la famille des Galatheides. Annales des Sciences Naturelles, Zoologie, series 7, 16, 191 - 327, fig. 1 - 36." type="journal article" year="1894">Milne-Edwards &amp; Bouvier, 1894: 299</bibRefCitation>
. —
<bibRefCitation id="145030E7C649FFC97EEFFE0CFC02A2AA" author="Bouvier, E. L." box="[757,962,464,487]" pageId="2" pageNumber="58" pagination="307 - 312" refId="ref4815" refString="Bouvier, E. L. (1896) Sur la famille des Chirostylidae, Ortmann, et sur la classification des Galatheidea [Crust.]. Bulletin de la Societe entomologique de France, 65, 307 - 312." type="journal article" year="1896">Bouvier, 1896: 312</bibRefCitation>
[vernacular name, unavailable].
</paragraph>
<paragraph id="707E4D16C649FFC97C8DFE2CFBA0A14A" blockId="2.[151,1308,432,583]" box="[151,1120,496,519]" pageId="2" pageNumber="58">
Diptycinae
<bibRefCitation id="145030E7C649FFC97D09FE2CFE23A14A" author="Bouvier, E. L." box="[275,483,496,519]" pageId="2" pageNumber="58" pagination="307 - 312" refId="ref4815" refString="Bouvier, E. L. (1896) Sur la famille des Chirostylidae, Ortmann, et sur la classification des Galatheidea [Crust.]. Bulletin de la Societe entomologique de France, 65, 307 - 312." type="journal article" year="1896">Bouvier, 1896: 312</bibRefCitation>
. — A.
<bibRefCitation id="145030E7C649FFC97E35FE2CFC7AA14A" author="Milne-Edwards, A. &amp; Bouvier, E. L." box="[559,954,496,519]" pageId="2" pageNumber="58" pagination="1 - 106" refId="ref5791" refString="Milne-Edwards, A. &amp; Bouvier, E. L. (1899) Crustaces Decapodes provenant des campagnes de l'Hirondelle (supplement) et de la Princesse-Alice (1891 - 1897). Resultats des Campagnes scientifiques Yacht Albert I Monaco, 13, 1 - 106." type="journal article" year="1899">Milne-Edwards &amp; Bouvier, 1899: 71</bibRefCitation>
, 87; 1900: 350.
</paragraph>
</subSubSection>
<subSubSection id="38DB1E9DC649FFC97C8DFDCCFDF1A10A" pageId="2" pageNumber="58" type="reference_group">
<paragraph id="707E4D16C649FFC97C8DFDCCFDCCA16A" blockId="2.[151,1308,432,583]" box="[151,524,528,551]" pageId="2" pageNumber="58">
<treatmentCitationGroup id="50D16A38C649FFC97C8DFDCCFDCCA16A" box="[151,524,528,551]" pageId="2" pageNumber="58">
<treatmentCitation id="F1606B07C649FFC97C8DFDCCFDC8A16A" author="Ortmann, A." box="[151,520,528,551]" page="244" pageId="2" pageNumber="58" year="1892">
<taxonomicName id="B7C13695C649FFC97C8DFDCCFDC8A16A" ID-CoL="aa325b06-615d-45e3-8e9d-41b275c95cab" authority="Ortmann, 1892: 244" authorityName="Ortmann" authorityPageNumber="244" authorityYear="1892" box="[151,520,528,551]" class="Malacostraca" family="Chirostylidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="58" phylum="Arthropoda" rank="family">
Chirostylidae
<bibRefCitation id="145030E7C649FFC97D37FDCCFDC8A16A" author="Ortmann, A." box="[301,520,528,551]" pageId="2" pageNumber="58" pagination="241 - 326" refId="ref6010" refString="Ortmann, A. (1892) Die Decapoden-Krebse des Strassbuger Museums IV. Die Abteilungen Galatheidea und Paguridea. Zoologische Jahrbucher. Abteilung fuer Systematic Oekologie und Geographie der Tiere, 6, 241 - 326, pls 11, 12." type="book chapter" year="1892">Ortmann, 1892: 244</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
<paragraph id="707E4D16C649FFC97C8DFDECFDF1A10A" blockId="2.[151,1308,432,583]" box="[151,561,560,583]" pageId="2" pageNumber="58">
Uroptychidae
<bibRefCitation id="145030E7C649FFC97D35FDECFE36A10A" author="Alcock, A." box="[303,502,560,583]" pageId="2" pageNumber="58" refId="ref4268" refString="Alcock, A. (1901) A descriptive catalogue of the Indian deep-sea Crustacea Decapoda Macrura and Anomala, in the Indian Museum. Being a revised account of the deep-sea species collected by the Royal Indian Marine Survey Ship Investigator. Trustees of the Indian Museum, Calcutta, 286 pp." type="book" year="1901">Alcock, 1901: 236</bibRefCitation>
, 278.
</paragraph>
</subSubSection>
<subSubSection id="38DB1E9DC649FFC97C8DFDA9FEB4A0F7" pageId="2" pageNumber="58" type="diagnosis">
<paragraph id="707E4D16C649FFC97C8DFDA9FEB4A0F7" blockId="2.[151,1436,629,1178]" pageId="2" pageNumber="58">
<emphasis id="42B59104C649FFC97C8DFDA9FEDCA1C2" bold="true" box="[151,284,629,655]" pageId="2" pageNumber="58">Diagnosis.</emphasis>
Carapace surface smooth, tuberculate or spinose but without transverse striae, posterolateral margin not distinctly defined or greatly inflated; rostrum variously shaped; supraocular spines absent. Anterolateral margin of abdominal somite 2 without prominent, anterolaterally directed spine. Sternite 3 not strongly produced anteriorly. Eyes well developed. Basal antennular article with distolateral spines. Antennal peduncle consisting of 5 articles; acicle present or absent. Mandibular cutting edge calcified, strongly serrated along its length. Maxilliped 1 without epipod; exopod flagellum present or absent, not annulated. Maxilliped 3 to pereopod 4 each with 2 arthrobranchs (well-developed or vestigial on maxilliped 3). Pereopod 5 with 1 arthrobranch only. Pereopods 24 with pleurobranch. Male pleopods 1 and 2 present. Male pleopods 35 vestigial or absent.
</paragraph>
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<paragraph id="707E4D16C649FFC97CDCFC19FC04A092" blockId="2.[151,1436,629,1178]" box="[198,964,965,991]" pageId="2" pageNumber="58">
<emphasis id="42B59104C649FFC97CDCFC19FE9AA092" bold="true" box="[198,346,965,991]" pageId="2" pageNumber="58">
<typeStatus id="AF7AF3B4C649FFC97CDCFC19FEC3A092" box="[198,259,965,991]" pageId="2" pageNumber="58">Type</typeStatus>
genus.
</emphasis>
<taxonomicName id="B7C13695C649FFC97D7BFC19FD62A092" ID-CoL="51b5ea20-f107-439a-8abb-083d473a756a" authority="Ortmann, 1892" authorityName="Ortmann" authorityYear="1892" box="[353,674,965,991]" class="Malacostraca" family="Chirostylidae" genus="Chirostylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="42B59104C649FFC97D7BFC19FE27A093" box="[353,487,965,990]" italics="true" pageId="2" pageNumber="58">Chirostylus</emphasis>
<bibRefCitation id="145030E7C649FFC97DF4FC19FD62A092" author="Ortmann, A." box="[494,674,965,991]" pageId="2" pageNumber="58" pagination="241 - 326" refId="ref6010" refString="Ortmann, A. (1892) Die Decapoden-Krebse des Strassbuger Museums IV. Die Abteilungen Galatheidea und Paguridea. Zoologische Jahrbucher. Abteilung fuer Systematic Oekologie und Geographie der Tiere, 6, 241 - 326, pls 11, 12." type="book chapter" year="1892">Ortmann, 1892</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
<paragraph id="707E4D16C649FFC97CDCFC36FCF4A767" blockId="2.[151,1436,629,1178]" pageId="2" pageNumber="58">
<emphasis id="42B59104C649FFC97CDCFC36FEADA749" bold="true" box="[198,365,1002,1028]" pageId="2" pageNumber="58">Composition.</emphasis>
<taxonomicName id="B7C13695C649FFC97D63FC36FD02A749" ID-CoL="51b5ea20-f107-439a-8abb-083d473a756a" authority="Ortmann, 1892" authorityName="Ortmann" authorityYear="1892" box="[377,706,1002,1028]" class="Malacostraca" family="Chirostylidae" genus="Chirostylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="42B59104C649FFC97D63FC36FE3FA74E" box="[377,511,1002,1027]" italics="true" pageId="2" pageNumber="58">Chirostylus</emphasis>
<bibRefCitation id="145030E7C649FFC97E10FC36FD02A749" author="Ortmann, A." box="[522,706,1002,1028]" pageId="2" pageNumber="58" pagination="241 - 326" refId="ref6010" refString="Ortmann, A. (1892) Die Decapoden-Krebse des Strassbuger Museums IV. Die Abteilungen Galatheidea und Paguridea. Zoologische Jahrbucher. Abteilung fuer Systematic Oekologie und Geographie der Tiere, 6, 241 - 326, pls 11, 12." type="book chapter" year="1892">Ortmann, 1892</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B7C13695C649FFC97EC9FC36FBFBA749" ID-CoL="837fb598-5282-46e3-83af-4066a8a581d5" authority="Caullery, 1896" authorityName="Caullery" authorityYear="1896" box="[723,1083,1002,1028]" class="Malacostraca" family="Chirostylidae" genus="Gastroptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="42B59104C649FFC97EC9FC36FCBDA74E" box="[723,893,1002,1027]" italics="true" pageId="2" pageNumber="58">Gastroptychus</emphasis>
<bibRefCitation id="145030E7C649FFC97F92FC36FBFBA749" author="Caullery, M." box="[904,1083,1002,1028]" pageId="2" pageNumber="58" pagination="365 - 419" refId="ref4884" refString="Caullery, M. (1896) Crustaces Schizopodes et Decapodes. In: Koehler, R., Resultats scientifiques de la Campagne du Caudan dans le Golfe de Gascogne, aout - septembre, 1895. Annales de l'Universite de Lyons, 26, 365 - 419, pls 13 - 17." type="journal article" year="1896">Caullery, 1896</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B7C13695C649FFC97851FC36FA57A749" ID-CoL="cbbe43a0-83c6-48c6-a062-efc24b5aa38a" authority="Stebbing, 1920" authorityName="Stebbing" authorityYear="1920" box="[1099,1431,1002,1028]" class="Malacostraca" family="Chirostylidae" genus="Hapaloptyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="42B59104C649FFC97851FC36FB14A74E" box="[1099,1236,1002,1027]" italics="true" pageId="2" pageNumber="58">Hapaloptyx</emphasis>
<bibRefCitation id="145030E7C649FFC978C5FC36FA57A749" author="Stebbing, T. R. R." box="[1247,1431,1002,1028]" pageId="2" pageNumber="58" pagination="231 - 272" refId="ref6638" refString="Stebbing, T. R. R. (1920) South African Crustacea (Part X of S. A. Crustacea, for the Marine Investigations in South Africa). Annals of the South African Museum, 17, 231 - 272." type="journal article" year="1920">Stebbing, 1920</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B7C13695C649FFC97C8DFBCCFE09A767" ID-CoL="154bfeaa-9065-4715-99d1-fb94056c7c8d" authority="Baba, 2004" authorityName="Baba" authorityYear="2004" box="[151,457,1040,1066]" class="Malacostraca" family="Chirostylidae" genus="Uroptychodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="42B59104C649FFC97C8DFBCCFEFAA764" box="[151,314,1040,1065]" italics="true" pageId="2" pageNumber="58">Uroptychodes</emphasis>
<bibRefCitation id="145030E7C649FFC97D5BFBCCFE09A767" author="Baba, K." box="[321,457,1040,1066]" pageId="2" pageNumber="58" pagination="97 - 116" refId="ref4504" refString="Baba, K. (2004) Uroptychodes, new genus of Chirostylidae (Crustacea: Decapoda: Anomura), with description of three new species. Scientia Marina, 68, 97 - 116." type="journal article" year="2004">Baba, 2004</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B7C13695C649FFC97DCFFBCCFCF0A767" ID-CoL="3eddc1a6-5e09-4db0-bcba-91283f2ee5df" authority="Henderson, 1888" authorityName="Henderson" authorityYear="1888" box="[469,816,1040,1066]" class="Malacostraca" family="Chirostylidae" genus="Uroptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="42B59104C649FFC97DCFFBCCFD9CA764" box="[469,604,1040,1065]" italics="true" pageId="2" pageNumber="58">Uroptychus</emphasis>
<bibRefCitation id="145030E7C649FFC97E7EFBCCFCF0A767" author="Henderson, J. R." box="[612,816,1040,1066]" pageId="2" pageNumber="58" pagination="1 - 221" refId="ref5286" refString="Henderson, J. R. (1888) Report on the Anomura collected by H. M. S. Challenger during the years 1873 - 76. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 76. Zoology, 27, 1 - 221, 21 pls." type="journal article" year="1888">Henderson, 1888</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="38DB1E9DC649FFC97CDCFBE9FE38A7D7" pageId="2" pageNumber="58" type="discussion">
<paragraph id="707E4D16C649FFC97CDCFBE9FE38A7D7" blockId="2.[151,1436,629,1178]" pageId="2" pageNumber="58">
<emphasis id="42B59104C649FFC97CDCFBE9FEFFA702" bold="true" box="[198,319,1077,1103]" pageId="2" pageNumber="58">Remarks.</emphasis>
Pleopods 35 are absent in most male chirostylids, but are vestigial in some species of
<taxonomicName id="B7C13695C649FFC97C8DFB87FE82A739" authorityName="Caullery" authorityYear="1896" box="[151,322,1115,1140]" class="Malacostraca" family="Chirostylidae" genus="Gastroptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="42B59104C649FFC97C8DFB87FE82A739" box="[151,322,1115,1140]" italics="true" pageId="2" pageNumber="58">Gastroptychus</emphasis>
</taxonomicName>
(e.g.,
<taxonomicName id="B7C13695C649FFC97D94FB87FD4BA739" authority="Baba, 2000" authorityName="Baba" authorityYear="2000" box="[398,651,1114,1140]" class="Malacostraca" family="Chirostylidae" genus="Gastroptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="58" phylum="Arthropoda" rank="species" species="rogeri">
<emphasis id="42B59104C649FFC97D94FB87FE39A739" box="[398,505,1115,1140]" italics="true" pageId="2" pageNumber="58">G. rogeri</emphasis>
<bibRefCitation id="145030E7C649FFC97E1BFB86FD4BA739" author="Baba, K." box="[513,651,1114,1140]" pageId="2" pageNumber="58" pagination="246 - 252" refId="ref4469" refString="Baba, K. (2000) Two new species of chirostylids (Decapoda: Anomura: Chirostylidae) from Tasmania. Journal of Crustacean Biology, special number 2, 246 - 252." type="journal article" year="2000">Baba, 2000</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="B7C13695C649FFC97ED0FB87FB25A739" authority="(Alcock &amp; Anderson, 1899)" baseAuthorityName="Alcock &amp; Anderson" baseAuthorityYear="1899" box="[714,1253,1114,1140]" class="Malacostraca" family="Chirostylidae" genus="Gastroptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="58" phylum="Arthropoda" rank="species" species="investigatoris">
<emphasis id="42B59104C649FFC97ED0FB87FC4DA739" box="[714,909,1115,1140]" italics="true" pageId="2" pageNumber="58">G. investigatoris</emphasis>
(
<bibRefCitation id="145030E7C649FFC97F87FB86FB1DA739" author="Alcock, A. &amp; Anderson, A. R. S." box="[925,1245,1114,1140]" pageId="2" pageNumber="58" pagination="1 - 27" refId="ref4322" refString="Alcock, A. &amp; Anderson, A. R. S. (1899) Natural history notes from H. M. Royal Indian Marine Survey Ship &quot; Investigator &quot;, Commander T. H. Heming, R. N., commanding. Series III, no. 2. An account of the deep-sea Crustacea dredged during the surveying season of 1897 - 98. Annals and Magazine of Natural History, series 7, 3, 1 - 27." type="journal article" year="1899">Alcock &amp; Anderson, 1899</bibRefCitation>
)
</taxonomicName>
, K. Baba, pers. com.; AM P53251,
<taxonomicName id="B7C13695C649FFC97D65FB5CFE27A7D4" authorityName="Baba" authorityYear="2000" box="[383,487,1152,1177]" class="Malacostraca" family="Chirostylidae" genus="Gastroptychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="58" phylum="Arthropoda" rank="species" species="rogeri">
<emphasis id="42B59104C649FFC97D65FB5CFE27A7D4" box="[383,487,1152,1177]" italics="true" pageId="2" pageNumber="58">G. rogeri</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>