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<document id="4338CF846E108BA635238D74A498AC44" ID-CLB-Dataset="20868" ID-DOI="10.5281/zenodo.4689555" ID-GBIF-Dataset="4317b6e2-9539-4520-84ae-d776c1cb8a57" ID-ISSN="1638-9387" ID-Zenodo-Dep="4689555" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618437452689" checkinUser="felipe" docAuthor="Grave, Sammy De" docDate="2004" docId="AF198160FFBEFFCCFF2999E7FB8C6F11" docLanguage="en" docName="Zoosystema.26.3.503-509.pdf" docOrigin="Zoosystema 26 (3)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Batella praecipua Grave 2004, n. sp." docType="treatment" docVersion="6" lastPageNumber="508" masterDocId="5320F918FFBFFFC9FFD29C6AFF906B37" masterDocTitle="A new species of Batella (Crustacea, Decapoda, Alpheidae) from New Caledonia" masterLastPageNumber="509" masterPageNumber="503" pageNumber="504" updateTime="1698936579378" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="5E13000957B37AAA08F78F88991A097A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="87283A51E07D8EF21ACA8CDAA2B7B6C5">
<mods:title id="03036205509DBA0548D788BDE6A26FAF">A new species of Batella (Crustacea, Decapoda, Alpheidae) from New Caledonia</mods:title>
</mods:titleInfo>
<mods:name id="79CD9DF154309E844A81069CDEF21490" type="personal">
<mods:role id="DE6D833DECBEF9E4165BFEB64BB0CEE4">
<mods:roleTerm id="2B9C0CCDB446E36D169EBA3742B06F59">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D2B46FBA7BA1EE72BA7A75CBEBB0DE8A">Grave, Sammy De</mods:namePart>
<mods:affiliation id="A74F3C2EEF7165B895ECABFEEB81E591">Oxford University Museum of Natural History, Parks Road, Oxford OX 1 3 PW (United Kingdom) sammy. degrave @ oum. ox. ac. uk</mods:affiliation>
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<mods:typeOfResource id="A9F4202177DBC6740D09F3295E2347EC">text</mods:typeOfResource>
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<mods:title id="F45049ACBA12DA8E62A9ABDE5E6C2100">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="272359A6CE77859C68EB2256B9C335B5">
<mods:date id="E46BD07C15C1721C8184966C26BC5AE5">2004</mods:date>
<mods:detail id="1A129F20264294813C873CA110604026" type="volume">
<mods:number id="56893B71539EFB54D095C1A97FECCF10">26</mods:number>
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<mods:number id="B84B969E9722C3FE230BA485D2C00624">3</mods:number>
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<mods:classification id="3B107BF23155E73C97BCD5055B422F4B">journal article</mods:classification>
<mods:identifier id="588EEE4284559574D45EB6B2557E1C1D" type="CLB-Dataset">20868</mods:identifier>
<mods:identifier id="F204E9F3BF4B1DCFA4ED51529CC5D203" type="DOI">10.5281/zenodo.4689555</mods:identifier>
<mods:identifier id="8BB8593D051720063C151781BEBF89E6" type="GBIF-Dataset">4317b6e2-9539-4520-84ae-d776c1cb8a57</mods:identifier>
<mods:identifier id="3708AD813B6333BCD6244614B6BF7E89" type="ISSN">1638-9387</mods:identifier>
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<treatment id="AF198160FFBEFFCCFF2999E7FB8C6F11" ID-DOI="http://doi.org/10.5281/zenodo.6267707" ID-GBIF-Taxon="193036479" ID-Zenodo-Dep="6267707" LSID="urn:lsid:plazi:treatment:AF198160FFBEFFCCFF2999E7FB8C6F11" httpUri="http://treatment.plazi.org/id/AF198160FFBEFFCCFF2999E7FB8C6F11" lastPageId="5" lastPageNumber="508" pageId="1" pageNumber="504">
<subSubSection id="6FAA63FDFFBEFFC8FF2999E7FE6C6E91" box="[251,508,1419,1447]" pageId="1" pageNumber="504" type="nomenclature">
<paragraph id="270F3076FFBEFFC8FF2999E7FE6C6E91" blockId="1.[251,508,1419,1479]" box="[251,508,1419,1447]" pageId="1" pageNumber="504">
<heading id="7C47871AFFBEFFC8FF2999E7FE6C6E91" box="[251,508,1419,1447]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="504" reason="2">
<taxonomicName id="E0B04BF5FFBEFFC8FF2999E7FE2A6E92" ID-CoL="85MW3" authority="Grave, 2004" authorityName="Grave" authorityYear="2004" box="[251,442,1419,1447]" class="Malacostraca" family="Alpheidae" genus="Batella" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="praecipua" status="sp. nov.">
<emphasis id="15C4EC64FFBEFFC8FF2999E7FE2A6E92" bold="true" box="[251,442,1419,1447]" italics="true" pageId="1" pageNumber="504">Batella praecipua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0EF7511FFFBEFFC8FE1399E6FE6C6E91" box="[449,508,1420,1446]" pageId="1" pageNumber="504" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6FAA63FDFFBEFFC8FE9599C7FE206EF0" box="[327,432,1453,1479]" pageId="1" pageNumber="504" type="description">
<paragraph id="270F3076FFBEFFC8FE9599C7FE206EF0" blockId="1.[251,508,1419,1479]" box="[327,432,1453,1479]" pageId="1" pageNumber="504">
(
<figureCitation id="BF8B2CF3FFBEFFC8FE9C99C7FE366EF0" box="[334,422,1453,1479]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="2.[113,124,1644,1661]" captionStartId-1="3.[113,124,1644,1661]" captionStartId-2="4.[113,124,1274,1291]" captionTargetBox-0="[144,1186,274,1576]" captionTargetBox-1="[169,1153,211,1610]" captionTargetBox-2="[126,1189,212,1237]" captionTargetId-0="figure-24@2.[77,1246,224,1644]" captionTargetId-1="figure-15@3.[169,1153,211,1610]" captionTargetId-2="figure-24@4.[33,1225,173,1326]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIG. 1. — Batella praecipua n. sp.; A-D, paratype (OUMNH 2004-01-0001); A, lateral view; B, frontal region, dorsal view; C, frontal region, lateral view; D, caudal fan; E, F, holotype (MHNN Na 11396); E, frontal region, lateral view; F, dorsal view. Scale bar: A, E, F, 1 mm; B-D, 0.5 mm." captionText-1="FIG. 2. — Batella praecipua n. sp., paratype (OUMNH 2004-01-0001); A, mandible; B, third maxilliped; C, tip of third maxilliped; D, antennular peduncle; E, second pereiopod; F, tip of second pereiopod (denuded); G, third pereiopod; H, dactyl of third pereio- pod; I, dactyl of fourth pereiopod. Scale bar: A, C, 0.1 mm; B, D, E, G, 0.2 mm; F, 0.03 mm; H, I, 0.05 mm." captionText-2="FIG. 3. — Batella praecipua n. sp., holotype (MHNN Na 11396); A, outer branch of external flagellum of antenna; B, scaphocerite; C, detail of anterolateral corner; D, second pereiopod; E, chelae of second pereiopod (denuded); F, dactyl of third pereiopod. Scale bar: A, 0.1 mm; B, D, 0.8 mm; C, 0.02 mm; E, F, 0.05 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4689557" figureDoi-1="http://doi.org/10.5281/zenodo.4689559" figureDoi-2="http://doi.org/10.5281/zenodo.4689561" httpUri-0="https://zenodo.org/record/4689557/files/figure.png" httpUri-1="https://zenodo.org/record/4689559/files/figure.png" httpUri-2="https://zenodo.org/record/4689561/files/figure.png" pageId="1" pageNumber="504">Figs 1-3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="6FAA63FDFFBEFFC8FFA399B6FE756D90" pageId="1" pageNumber="504" type="materials_examined">
<paragraph id="270F3076FFBEFFC8FFA399B6FE756D90" blockId="1.[113,646,1500,1703]" pageId="1" pageNumber="504">
<collectionCode id="41A1A8B3FFBEFFC8FFA399B6FE946EC5" box="[113,260,1500,1523]" pageId="1" pageNumber="504">TYPE MATERIAL</collectionCode>
. —
<typeStatus id="F80B8ED4FFBEFFC8FEFD99B7FE1E6EC2" box="[303,398,1501,1525]" pageId="1" pageNumber="504" type="holotype">Holotype</typeStatus>
: MUSORSTOM 6,
<collectingCountry id="5FA770E6FFBEFFC8FD8599B6FF476D38" name="New Caledonia" pageId="1" pageNumber="504">New Caledonia</collectingCountry>
,
<geoCoordinate id="428456B1FFBEFFC8FF33999DFED86D38" box="[225,328,1527,1551]" degrees="22" direction="south" minutes="53" orientation="latitude" pageId="1" pageNumber="504" precision="15" seconds="8" value="-22.885555">22°53’8”S</geoCoordinate>
,
<geoCoordinate id="428456B1FFBEFFC8FE81999DFE596D38" box="[339,457,1527,1551]" degrees="167" direction="east" minutes="13" orientation="longitude" pageId="1" pageNumber="504" precision="15" seconds="9" value="167.21916">167°13’9”E</geoCoordinate>
,
<quantity id="E0489D93FFBEFFC8FE01999DFDD46D3A" box="[467,580,1526,1551]" metricMagnitude="2" metricUnit="m" metricValue="4.325" metricValueMax="4.4" metricValueMin="4.25" pageId="1" pageNumber="504" unit="m" value="432.5" valueMax="440.0" valueMin="425.0">425-440 m</quantity>
, “dans une éponge avec des sténopodides”, non ovigerous, post-orbital carapace length (pocl)
<quantity id="E0489D93FFBEFFC8FE1A9A43FDB46D08" box="[456,548,1576,1601]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="1" pageNumber="504" unit="mm" value="5.5">5.50 mm</quantity>
(
<collectionCode id="41A1A8B3FFBEFFC8FDE09A43FD166D76" box="[562,646,1577,1601]" country="France" httpUri="http://grbio.org/cool/1hrs-xwta" name="Musee d'Histoire Naturalle" pageId="1" pageNumber="504">MHNN</collectionCode>
Na 11396).
<typeStatus id="F80B8ED4FFBEFFC8FF209A28FEDE6D6D" box="[242,334,1602,1626]" pageId="1" pageNumber="504" type="paratype">Paratype</typeStatus>
:
<collectingCountry id="5FA770E6FFBEFFC8FE889A28FE6D6D6C" box="[346,509,1602,1627]" name="New Caledonia" pageId="1" pageNumber="504">New Caledonia</collectingCountry>
,
<geoCoordinate id="428456B1FFBEFFC8FDDB9A29FD116D6C" box="[521,641,1603,1627]" degrees="22" direction="south" minutes="52" orientation="latitude" pageId="1" pageNumber="504" precision="15" seconds="42" value="-22.878334">22°52’42”S</geoCoordinate>
, 167°11’90”E,
<quantity id="E0489D93FFBEFFC8FEDB9A36FEC16D45" box="[265,337,1627,1652]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="504" unit="m" value="400.0">400 m</quantity>
, rocky bottom with sponges,
<date id="530E16B6FFBEFFC8FFA39A1FFF4F6DBA" box="[113,223,1653,1677]" pageId="1" pageNumber="504" value="1995-05-12">
12.
<collectionCode id="41A1A8B3FFBEFFC8FF5D9A1FFF346DBA" box="[143,164,1653,1677]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="504" type="Museum">V</collectionCode>
.1995
</date>
, leg. A. A. Myers, non ovigerous, pocl
<quantity id="E0489D93FFBEFFC8FFA39AE5FF5C6D92" box="[113,204,1678,1703]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="504" unit="mm" value="2.6">2.60 mm</quantity>
(
<collectionCode id="41A1A8B3FFBEFFC8FF0B9AE5FED06D90" box="[217,320,1679,1703]" pageId="1" pageNumber="504">OUMNH</collectionCode>
2004-01-0001).
</paragraph>
</subSubSection>
<subSubSection id="6FAA63FDFFBEFFC8FD769CA1FC446AC6" pageId="1" pageNumber="504" type="etymology">
<paragraph id="270F3076FFBEFFC8FD769CA1FBD86A22" blockId="1.[676,1209,203,497]" pageId="1" pageNumber="504">
ETYMOLOGY. — From the Latin
<emphasis id="15C4EC64FFBEFFC8FC239CA6FBC56BD4" box="[1009,1109,204,227]" italics="true" pageId="1" pageNumber="504">praecipuus</emphasis>
, meaning peculiar. In reference to the frontal region of the carapace, exhibiting a non-tridentate structure.
</paragraph>
<paragraph id="270F3076FFBEFFC8FD769D4DFC446AC6" blockId="1.[676,1209,203,497]" pageId="1" pageNumber="504">
HABITAT. — Known only from
<collectingCountry id="5FA770E6FFBEFFC8FC369D4DFB106A08" box="[996,1152,295,319]" name="New Caledonia" pageId="1" pageNumber="504">New Caledonia</collectingCountry>
,
<quantity id="E0489D93FFBEFFC8FB599D4DFD756A60" metricMagnitude="2" metricUnit="m" metricValue="4.2" metricValueMax="4.4" metricValueMin="4.0" pageId="1" pageNumber="504" unit="m" value="420.0" valueMax="440.0" valueMin="400.0">400- 440 m</quantity>
deep. The
<typeStatus id="F80B8ED4FFBEFFC8FC849D2BFC3C6A6E" box="[854,940,321,345]" pageId="1" pageNumber="504" type="holotype">holotype</typeStatus>
was collected with unidentified stenopodidean shrimp from a sponge. No host association is recorded for the
<typeStatus id="F80B8ED4FFBEFFC8FC019D19FBB66ABC" box="[979,1062,371,395]" pageId="1" pageNumber="504" type="paratype">paratype</typeStatus>
specimen, but this latter sample also contained examples of
<taxonomicName id="E0B04BF5FFBEFFC8FD769DCCFB196A89" authority="Hayashi &amp; Ogawa, 1987" authorityName="Hayashi &amp; Ogawa" authorityYear="1987" box="[676,1161,422,446]" class="Malacostraca" family="Spongicolidae" genus="Spongicola" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="levigata">
<emphasis id="15C4EC64FFBEFFC8FD769DCCFCFE6A89" box="[676,878,422,446]" italics="true" pageId="1" pageNumber="504">Spongicola levigata</emphasis>
Hayashi &amp; Ogawa, 1987
</taxonomicName>
and
<taxonomicName id="E0B04BF5FFBEFFC8FD769DD5FBBA6AE0" authority="Bruce, 1991" authorityName="Bruce" authorityYear="1991" box="[676,1066,447,471]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="504" phylum="Arthropoda" rank="species" species="forcipulatus">
<emphasis id="15C4EC64FFBEFFC8FD769DD5FC346AE0" box="[676,932,447,471]" italics="true" pageId="1" pageNumber="504">Periclimenes forcipulatus</emphasis>
Bruce,1991
</taxonomicName>
, both known hexactinellid sponge associates.
</paragraph>
</subSubSection>
<subSubSection id="6FAA63FDFFBEFFCCFD769E46FD176871" lastPageId="5" lastPageNumber="508" pageId="1" pageNumber="504" type="description">
<paragraph id="270F3076FFBEFFC8FD769E46FCA56973" blockId="1.[676,1209,556,1703]" box="[676,821,556,582]" pageId="1" pageNumber="504">DESCRIPTION</paragraph>
<paragraph id="270F3076FFBEFFC8FD769E27FBCD6850" blockId="1.[676,1209,556,1703]" pageId="1" pageNumber="504">
Carapace smooth, lacking setae (
<figureCitation id="BF8B2CF3FFBEFFC8FBD09E27FBAC6950" box="[1026,1084,589,615]" captionStart="FIG" captionStartId="2.[113,124,1644,1661]" captionTargetBox="[144,1186,274,1576]" captionTargetId="figure-24@2.[77,1246,224,1644]" captionTargetPageId="2" captionText="FIG. 1. — Batella praecipua n. sp.; A-D, paratype (OUMNH 2004-01-0001); A, lateral view; B, frontal region, dorsal view; C, frontal region, lateral view; D, caudal fan; E, F, holotype (MHNN Na 11396); E, frontal region, lateral view; F, dorsal view. Scale bar: A, E, F, 1 mm; B-D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4689557" httpUri="https://zenodo.org/record/4689557/files/figure.png" pageId="1" pageNumber="504">Fig. 1</figureCitation>
A-C, E, F). Anterior margin trilobed; rostrum poorly developed, tip upturned, adrostral furrows shallow, falling short of eyes. Pterygostomial angle produced, rounded. Ventral margin of carapace straight, not fringed with setae; posterior margin with shallow cardiac notch. Inconspicuous tubercle present on gastric region. Eyes not visible in dorsal or lateral view, anteriorly exposed.
</paragraph>
<paragraph id="270F3076FFBEFFC8FD769F07FBCE6FF0" blockId="1.[676,1209,556,1703]" pageId="1" pageNumber="504">
Antennular peduncle (
<figureCitation id="BF8B2CF3FFBEFFC8FC449F07FC7C68B0" box="[918,1004,877,903]" captionStart="FIG" captionStartId="3.[113,124,1644,1661]" captionTargetBox="[169,1153,211,1610]" captionTargetId="figure-15@3.[169,1153,211,1610]" captionTargetPageId="3" captionText="FIG. 2. — Batella praecipua n. sp., paratype (OUMNH 2004-01-0001); A, mandible; B, third maxilliped; C, tip of third maxilliped; D, antennular peduncle; E, second pereiopod; F, tip of second pereiopod (denuded); G, third pereiopod; H, dactyl of third pereio- pod; I, dactyl of fourth pereiopod. Scale bar: A, C, 0.1 mm; B, D, E, G, 0.2 mm; F, 0.03 mm; H, I, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4689559" httpUri="https://zenodo.org/record/4689559/files/figure.png" pageId="1" pageNumber="504">Fig. 2D</figureCitation>
) short, second and third segments subequal in length, approximately half length of first segment; stylocerite well developed, slightly overreaching first segment, narrowing distally to a blunt point, ventral carina of first article poorly developed, statocyst well developed. External flagellum biramous, outer branch consisting of single segment (
<figureCitation id="BF8B2CF3FFBEFFC8FBDD9827FBE66F50" box="[1039,1142,1101,1127]" captionStart="FIG" captionStartId="3.[113,124,1644,1661]" captionTargetBox="[169,1153,211,1610]" captionTargetId="figure-15@3.[169,1153,211,1610]" captionTargetPageId="3" captionText="FIG. 2. — Batella praecipua n. sp., paratype (OUMNH 2004-01-0001); A, mandible; B, third maxilliped; C, tip of third maxilliped; D, antennular peduncle; E, second pereiopod; F, tip of second pereiopod (denuded); G, third pereiopod; H, dactyl of third pereio- pod; I, dactyl of fourth pereiopod. Scale bar: A, C, 0.1 mm; B, D, E, G, 0.2 mm; F, 0.03 mm; H, I, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4689559" httpUri="https://zenodo.org/record/4689559/files/figure.png" pageId="1" pageNumber="504">Figs 2D</figureCitation>
;
<figureCitation id="BF8B2CF3FFBEFFC8FB579827FB3B6F50" box="[1157,1195,1101,1127]" captionStart="FIG" captionStartId="4.[113,124,1274,1291]" captionTargetBox="[126,1189,212,1237]" captionTargetId="figure-24@4.[33,1225,173,1326]" captionTargetPageId="4" captionText="FIG. 3. — Batella praecipua n. sp., holotype (MHNN Na 11396); A, outer branch of external flagellum of antenna; B, scaphocerite; C, detail of anterolateral corner; D, second pereiopod; E, chelae of second pereiopod (denuded); F, dactyl of third pereiopod. Scale bar: A, 0.1 mm; B, D, 0.8 mm; C, 0.02 mm; E, F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4689561" httpUri="https://zenodo.org/record/4689561/files/figure.png" pageId="1" pageNumber="504">3A</figureCitation>
), longer free ramus approximately 0.4 times carapace length. Mesial flagellum slender, filiform, approximately 0.7 times carapace length.
</paragraph>
<paragraph id="270F3076FFBEFFCDFD7698A7FE616EF0" blockId="1.[676,1209,556,1703]" lastBlockId="4.[113,646,1388,1703]" lastPageId="4" lastPageNumber="507" pageId="1" pageNumber="504">
Carpocerite of antenna (
<figureCitation id="BF8B2CF3FFBEFFC8FC6398A7FBF56FD1" box="[945,1125,1228,1255]" captionStart="FIG" captionStartId="2.[113,124,1644,1661]" captionTargetBox="[144,1186,274,1576]" captionTargetId="figure-24@2.[77,1246,224,1644]" captionTargetPageId="2" captionText="FIG. 1. — Batella praecipua n. sp.; A-D, paratype (OUMNH 2004-01-0001); A, lateral view; B, frontal region, dorsal view; C, frontal region, lateral view; D, caudal fan; E, F, holotype (MHNN Na 11396); E, frontal region, lateral view; F, dorsal view. Scale bar: A, E, F, 1 mm; B-D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4689557" httpUri="https://zenodo.org/record/4689557/files/figure.png" pageId="1" pageNumber="504">Fig. 1A, B, E, F</figureCitation>
) falling short of antennular peduncle; basicerite with well developed ventral tooth; scaphocerite (
<figureCitation id="BF8B2CF3FFBEFFC8FB899967FB206E10" box="[1115,1200,1293,1319]" captionStart="FIG" captionStartId="4.[113,124,1274,1291]" captionTargetBox="[126,1189,212,1237]" captionTargetId="figure-24@4.[33,1225,173,1326]" captionTargetPageId="4" captionText="FIG. 3. — Batella praecipua n. sp., holotype (MHNN Na 11396); A, outer branch of external flagellum of antenna; B, scaphocerite; C, detail of anterolateral corner; D, second pereiopod; E, chelae of second pereiopod (denuded); F, dactyl of third pereiopod. Scale bar: A, 0.1 mm; B, D, 0.8 mm; C, 0.02 mm; E, F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4689561" httpUri="https://zenodo.org/record/4689561/files/figure.png" pageId="1" pageNumber="504">Fig. 3B</figureCitation>
) reaching just past antennular peduncle, oval shaped, anterior margin rounded, lateral tooth rounded (
<figureCitation id="BF8B2CF3FFBEFFC8FCC29907FCF86EB0" box="[784,872,1389,1415]" captionStart="FIG" captionStartId="4.[113,124,1274,1291]" captionTargetBox="[126,1189,212,1237]" captionTargetId="figure-24@4.[33,1225,173,1326]" captionTargetPageId="4" captionText="FIG. 3. — Batella praecipua n. sp., holotype (MHNN Na 11396); A, outer branch of external flagellum of antenna; B, scaphocerite; C, detail of anterolateral corner; D, second pereiopod; E, chelae of second pereiopod (denuded); F, dactyl of third pereiopod. Scale bar: A, 0.1 mm; B, D, 0.8 mm; C, 0.02 mm; E, F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4689561" httpUri="https://zenodo.org/record/4689561/files/figure.png" pageId="1" pageNumber="504">Fig. 3C</figureCitation>
), poorly developed, not reaching past lamina. Flagellum filiform, approximately equal to or slightly longer than carapace length Mouthparts typical for the genus. Mandible (
<figureCitation id="BF8B2CF3FFBEFFC8FD7E9987FC926D30" box="[684,770,1517,1543]" captionStart="FIG" captionStartId="3.[113,124,1644,1661]" captionTargetBox="[169,1153,211,1610]" captionTargetId="figure-15@3.[169,1153,211,1610]" captionTargetPageId="3" captionText="FIG. 2. — Batella praecipua n. sp., paratype (OUMNH 2004-01-0001); A, mandible; B, third maxilliped; C, tip of third maxilliped; D, antennular peduncle; E, second pereiopod; F, tip of second pereiopod (denuded); G, third pereiopod; H, dactyl of third pereio- pod; I, dactyl of fourth pereiopod. Scale bar: A, C, 0.1 mm; B, D, E, G, 0.2 mm; F, 0.03 mm; H, I, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4689559" httpUri="https://zenodo.org/record/4689559/files/figure.png" pageId="1" pageNumber="504">Fig. 2A</figureCitation>
) deeply bifurcated, palp absent; molar process cylindrical ending in tuft of spines; incisor process spatulate, distal part terminating in tuft of setae. Third maxilliped (
<figureCitation id="BF8B2CF3FFBEFFC8FC2F9A27FBE16D50" box="[1021,1137,1613,1639]" captionStart="FIG" captionStartId="3.[113,124,1644,1661]" captionTargetBox="[169,1153,211,1610]" captionTargetId="figure-15@3.[169,1153,211,1610]" captionTargetPageId="3" captionText="FIG. 2. — Batella praecipua n. sp., paratype (OUMNH 2004-01-0001); A, mandible; B, third maxilliped; C, tip of third maxilliped; D, antennular peduncle; E, second pereiopod; F, tip of second pereiopod (denuded); G, third pereiopod; H, dactyl of third pereio- pod; I, dactyl of fourth pereiopod. Scale bar: A, C, 0.1 mm; B, D, E, G, 0.2 mm; F, 0.03 mm; H, I, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4689559" httpUri="https://zenodo.org/record/4689559/files/figure.png" pageId="1" pageNumber="504">Fig. 2B, C</figureCitation>
) overreaching antennular peduncle; exopod falling short of antepenultimate segment; ultimate segment bearing seven semicircular rows of plumose setae on mesioventral side, single spine subdistally, tip provided with three serrated spines.
</paragraph>
<caption id="73CF60FEFFBDFFCBFFA39A06FEA16D90" ID-DOI="http://doi.org/10.5281/zenodo.4689557" ID-Zenodo-Dep="4689557" httpUri="https://zenodo.org/record/4689557/files/figure.png" pageId="2" pageNumber="505" startId="2.[113,124,1644,1661]" targetBox="[144,1186,274,1576]" targetPageId="2">
<paragraph id="270F3076FFBDFFCBFFA39A06FEA16D90" blockId="2.[113,1210,1644,1703]" pageId="2" pageNumber="505">
FIG. 1. —
<taxonomicName id="E0B04BF5FFBDFFCBFF169A06FEC56D4A" authority="Grave, 2004" authorityName="Grave" authorityYear="2004" box="[196,341,1644,1661]" class="Malacostraca" family="Alpheidae" genus="Batella" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="505" phylum="Arthropoda" rank="species" species="praecipua" status="sp. nov.">
<emphasis id="15C4EC64FFBDFFCBFF169A06FEC56D4A" box="[196,341,1644,1661]" italics="true" pageId="2" pageNumber="505">Batella praecipua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0EF7511FFFBDFFCBFE899A06FE1E6D4A" box="[347,398,1644,1661]" pageId="2" pageNumber="505" rank="species">n. sp.</taxonomicNameLabel>
;
<emphasis id="15C4EC64FFBDFFCBFE449A06FE286D4A" bold="true" box="[406,440,1644,1661]" pageId="2" pageNumber="505">A -D</emphasis>
, paratype (OUMNH 2004-01-0001);
<emphasis id="15C4EC64FFBDFFCBFD399A06FD686D4A" bold="true" box="[747,760,1644,1661]" pageId="2" pageNumber="505">A</emphasis>
, lateral view;
<emphasis id="15C4EC64FFBDFFCBFCBF9A06FCEA6D4A" bold="true" box="[877,890,1644,1661]" pageId="2" pageNumber="505">B</emphasis>
, frontal region, dorsal view;
<emphasis id="15C4EC64FFBDFFCBFBB99A06FBE96D4A" bold="true" box="[1131,1145,1644,1661]" pageId="2" pageNumber="505">C</emphasis>
, frontal region, lateral view;
<emphasis id="15C4EC64FFBDFFCBFEC89AEBFEB86DA5" bold="true" box="[282,296,1665,1682]" pageId="2" pageNumber="505">D</emphasis>
, caudal fan;
<emphasis id="15C4EC64FFBDFFCBFE459AEBFE336DA5" bold="true" box="[407,419,1665,1682]" pageId="2" pageNumber="505">E</emphasis>
,
<emphasis id="15C4EC64FFBDFFCBFE7C9AEBFE296DA5" bold="true" box="[430,441,1665,1682]" pageId="2" pageNumber="505">F</emphasis>
, holotype (MHNN Na 11396);
<emphasis id="15C4EC64FFBDFFCBFD6B9AEBFD556DA5" bold="true" box="[697,709,1665,1682]" pageId="2" pageNumber="505">E</emphasis>
, frontal region, lateral view;
<emphasis id="15C4EC64FFBDFFCBFC679AEBFC506DA5" bold="true" box="[949,960,1665,1682]" pageId="2" pageNumber="505">F</emphasis>
, dorsal view. Scale bar: A, E, F, 1 mm; B-D, 0.5 mm.
</paragraph>
</caption>
<caption id="73CF60FEFFBCFFCAFFA39A06FC766D90" ID-DOI="http://doi.org/10.5281/zenodo.4689559" ID-Zenodo-Dep="4689559" httpUri="https://zenodo.org/record/4689559/files/figure.png" pageId="3" pageNumber="506" startId="3.[113,124,1644,1661]" targetBox="[169,1153,211,1610]" targetPageId="3">
<paragraph id="270F3076FFBCFFCAFFA39A06FC766D90" blockId="3.[113,1211,1644,1703]" pageId="3" pageNumber="506">
FIG. 2. —
<taxonomicName id="E0B04BF5FFBCFFCAFF149A06FEC96D4A" authority="Grave, 2004" authorityName="Grave" authorityYear="2004" box="[198,345,1644,1661]" class="Malacostraca" family="Alpheidae" genus="Batella" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="506" phylum="Arthropoda" rank="species" species="praecipua" status="sp. nov.">
<emphasis id="15C4EC64FFBCFFCAFF149A06FEC96D4A" box="[198,345,1644,1661]" italics="true" pageId="3" pageNumber="506">Batella praecipua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0EF7511FFFBCFFCAFEB29A06FE046D4A" box="[352,404,1644,1661]" pageId="3" pageNumber="506" rank="species">n. sp.</taxonomicNameLabel>
, paratype (OUMNH 2004-01-0001);
<emphasis id="15C4EC64FFBCFFCAFD189A06FD476D4A" bold="true" box="[714,727,1644,1661]" pageId="3" pageNumber="506">A</emphasis>
, mandible;
<emphasis id="15C4EC64FFBCFFCAFCEE9A06FCD96D4A" bold="true" box="[828,841,1644,1661]" pageId="3" pageNumber="506">B</emphasis>
, third maxilliped;
<emphasis id="15C4EC64FFBCFFCAFC369A06FC626D4A" bold="true" box="[996,1010,1644,1661]" pageId="3" pageNumber="506">C</emphasis>
, tip of third maxilliped;
<emphasis id="15C4EC64FFBCFFCAFFA39AEBFFEF6DA5" bold="true" box="[113,127,1665,1682]" pageId="3" pageNumber="506">D</emphasis>
, antennular peduncle;
<emphasis id="15C4EC64FFBCFFCAFE929AEBFEDC6DA5" bold="true" box="[320,332,1665,1682]" pageId="3" pageNumber="506">E</emphasis>
, second pereiopod;
<emphasis id="15C4EC64FFBCFFCAFE2E9AEBFD976DA5" bold="true" box="[508,519,1665,1682]" pageId="3" pageNumber="506">F</emphasis>
, tip of second pereiopod (denuded);
<emphasis id="15C4EC64FFBCFFCAFC919AEBFCC16DA5" bold="true" box="[835,849,1665,1682]" pageId="3" pageNumber="506">G</emphasis>
, third pereiopod;
<emphasis id="15C4EC64FFBCFFCAFC3B9AEBFC676DA5" bold="true" box="[1001,1015,1665,1682]" pageId="3" pageNumber="506">H</emphasis>
, dactyl of third pereio- pod;
<emphasis id="15C4EC64FFBCFFCAFF4E9AFCFF326D90" bold="true" box="[156,162,1686,1703]" pageId="3" pageNumber="506">I</emphasis>
, dactyl of fourth pereiopod. Scale bar: A, C, 0.1 mm; B, D, E, G, 0.2 mm; F, 0.03 mm; H, I, 0.05 mm.
</paragraph>
</caption>
<caption id="73CF60FEFFBBFFCDFFA39890FDC56E01" ID-DOI="http://doi.org/10.5281/zenodo.4689561" ID-Zenodo-Dep="4689561" httpUri="https://zenodo.org/record/4689561/files/figure.png" pageId="4" pageNumber="507" startId="4.[113,124,1274,1291]" targetBox="[126,1189,212,1237]" targetPageId="4">
<paragraph id="270F3076FFBBFFCDFFA39890FDC56E01" blockId="4.[113,1211,1274,1334]" pageId="4" pageNumber="507">
FIG. 3. —
<taxonomicName id="E0B04BF5FFBBFFCDFF179890FEC66E3C" authority="Grave, 2004" authorityName="Grave" authorityYear="2004" box="[197,342,1274,1291]" class="Malacostraca" family="Alpheidae" genus="Batella" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="507" phylum="Arthropoda" rank="species" species="praecipua" status="sp. nov.">
<emphasis id="15C4EC64FFBBFFCDFF179890FEC66E3C" box="[197,342,1274,1291]" italics="true" pageId="4" pageNumber="507">Batella praecipua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0EF7511FFFBBFFCDFE8F9890FE006E3C" box="[349,400,1274,1291]" pageId="4" pageNumber="507" rank="species">n. sp.</taxonomicNameLabel>
, holotype (MHNN Na 11396);
<emphasis id="15C4EC64FFBBFFCDFD5C9890FD0B6E3C" bold="true" box="[654,667,1274,1291]" pageId="4" pageNumber="507">A</emphasis>
, outer branch of external flagellum of antenna;
<emphasis id="15C4EC64FFBBFFCDFBFD9890FBAC6E3C" bold="true" box="[1071,1084,1274,1291]" pageId="4" pageNumber="507">B</emphasis>
, scaphocerite;
<emphasis id="15C4EC64FFBBFFCDFFA3997AFFEF6E16" bold="true" box="[113,127,1296,1313]" pageId="4" pageNumber="507">C</emphasis>
, detail of anterolateral corner;
<emphasis id="15C4EC64FFBBFFCDFEAD997AFE1D6E16" bold="true" box="[383,397,1296,1313]" pageId="4" pageNumber="507">D</emphasis>
, second pereiopod;
<emphasis id="15C4EC64FFBBFFCDFDE8997AFDD66E16" bold="true" box="[570,582,1296,1313]" pageId="4" pageNumber="507">E</emphasis>
, chelae of second pereiopod (denuded);
<emphasis id="15C4EC64FFBBFFCDFC4F997AFC386E16" bold="true" box="[925,936,1296,1313]" pageId="4" pageNumber="507">F</emphasis>
, dactyl of third pereiopod. Scale bar: A, 0.1 mm; B, D, 0.8 mm; C, 0.02 mm; E, F, 0.05 mm.
</paragraph>
</caption>
<paragraph id="270F3076FFBBFFCDFFA399A7FF456D31" blockId="4.[113,646,1388,1703]" pageId="4" pageNumber="507">
First pereiopods lacking in both holo- and
<typeStatus id="F80B8ED4FFBBFFCDFFA39986FF416D31" box="[113,209,1516,1542]" pageId="4" pageNumber="507" type="paratype">paratype</typeStatus>
.
</paragraph>
<paragraph id="270F3076FFBBFFCDFFA39A67FD4F6ED1" blockId="4.[113,646,1388,1703]" lastBlockId="4.[676,1210,1389,1703]" pageId="4" pageNumber="507">
Second pereiopods (
<figureCitation id="BF8B2CF3FFBBFFCDFE999A67FE326D10" box="[331,418,1549,1575]" captionStart="FIG" captionStartId="3.[113,124,1644,1661]" captionTargetBox="[169,1153,211,1610]" captionTargetId="figure-15@3.[169,1153,211,1610]" captionTargetPageId="3" captionText="FIG. 2. — Batella praecipua n. sp., paratype (OUMNH 2004-01-0001); A, mandible; B, third maxilliped; C, tip of third maxilliped; D, antennular peduncle; E, second pereiopod; F, tip of second pereiopod (denuded); G, third pereiopod; H, dactyl of third pereio- pod; I, dactyl of fourth pereiopod. Scale bar: A, C, 0.1 mm; B, D, E, G, 0.2 mm; F, 0.03 mm; H, I, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4689559" httpUri="https://zenodo.org/record/4689559/files/figure.png" pageId="4" pageNumber="507">Figs 2E</figureCitation>
;
<figureCitation id="BF8B2CF3FFBBFFCDFE7F9A67FE426D10" box="[429,466,1549,1575]" captionStart="FIG" captionStartId="4.[113,124,1274,1291]" captionTargetBox="[126,1189,212,1237]" captionTargetId="figure-24@4.[33,1225,173,1326]" captionTargetPageId="4" captionText="FIG. 3. — Batella praecipua n. sp., holotype (MHNN Na 11396); A, outer branch of external flagellum of antenna; B, scaphocerite; C, detail of anterolateral corner; D, second pereiopod; E, chelae of second pereiopod (denuded); F, dactyl of third pereiopod. Scale bar: A, 0.1 mm; B, D, 0.8 mm; C, 0.02 mm; E, F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4689561" httpUri="https://zenodo.org/record/4689561/files/figure.png" pageId="4" pageNumber="507">3D</figureCitation>
) short, subequal in length to third maxilliped. Ischium and merus elongate; carpus five-segmented, distal segment longest, segment length ratio (proximal to distal) 1.0:0.5:0.5:0.6:1.2. Dactylus (
<figureCitation id="BF8B2CF3FFBBFFCDFE6B9AE7FD9D6D90" box="[441,525,1677,1703]" captionStart="FIG" captionStartId="3.[113,124,1644,1661]" captionTargetBox="[169,1153,211,1610]" captionTargetId="figure-15@3.[169,1153,211,1610]" captionTargetPageId="3" captionText="FIG. 2. — Batella praecipua n. sp., paratype (OUMNH 2004-01-0001); A, mandible; B, third maxilliped; C, tip of third maxilliped; D, antennular peduncle; E, second pereiopod; F, tip of second pereiopod (denuded); G, third pereiopod; H, dactyl of third pereio- pod; I, dactyl of fourth pereiopod. Scale bar: A, C, 0.1 mm; B, D, E, G, 0.2 mm; F, 0.03 mm; H, I, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4689559" httpUri="https://zenodo.org/record/4689559/files/figure.png" pageId="4" pageNumber="507">Figs 2F</figureCitation>
;
<figureCitation id="BF8B2CF3FFBBFFCDFDCB9AE7FDA96D90" box="[537,569,1677,1703]" captionStart="FIG" captionStartId="4.[113,124,1274,1291]" captionTargetBox="[126,1189,212,1237]" captionTargetId="figure-24@4.[33,1225,173,1326]" captionTargetPageId="4" captionText="FIG. 3. — Batella praecipua n. sp., holotype (MHNN Na 11396); A, outer branch of external flagellum of antenna; B, scaphocerite; C, detail of anterolateral corner; D, second pereiopod; E, chelae of second pereiopod (denuded); F, dactyl of third pereiopod. Scale bar: A, 0.1 mm; B, D, 0.8 mm; C, 0.02 mm; E, F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4689561" httpUri="https://zenodo.org/record/4689561/files/figure.png" pageId="4" pageNumber="507">3E</figureCitation>
) short, beak-like;
<quantity id="E0489D93FFBBFFCDFCC49907FCF76EB0" box="[790,871,1389,1415]" metricMagnitude="-3" metricUnit="m" metricValue="3.81" pageId="4" pageNumber="507" unit="in" value="0.15">0.15 in</quantity>
length of propodus; terminating in small spine; fixed finger terminating in two blunt spines; fingers with distal tufts of plumose setae.
</paragraph>
<paragraph id="270F3076FFBBFFCCFD769987FE7B6A50" blockId="4.[676,1210,1389,1703]" lastBlockId="5.[113,647,204,839]" lastPageId="5" lastPageNumber="508" pageId="4" pageNumber="507">
Third pereiopod (
<figureCitation id="BF8B2CF3FFBBFFCDFCB99987FC536D30" box="[875,963,1517,1543]" captionStart="FIG" captionStartId="3.[113,124,1644,1661]" captionTargetBox="[169,1153,211,1610]" captionTargetId="figure-15@3.[169,1153,211,1610]" captionTargetPageId="3" captionText="FIG. 2. — Batella praecipua n. sp., paratype (OUMNH 2004-01-0001); A, mandible; B, third maxilliped; C, tip of third maxilliped; D, antennular peduncle; E, second pereiopod; F, tip of second pereiopod (denuded); G, third pereiopod; H, dactyl of third pereio- pod; I, dactyl of fourth pereiopod. Scale bar: A, C, 0.1 mm; B, D, E, G, 0.2 mm; F, 0.03 mm; H, I, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4689559" httpUri="https://zenodo.org/record/4689559/files/figure.png" pageId="4" pageNumber="507">Fig. 2G</figureCitation>
) robust; ischium with two spines on inferior margin; merus unarmed, provided with row of short, simple setae on mesial margin, 2.1 times length of ischium; carpus unarmed, equal in length to ischium; propodus armed with eight spines along inferior margin and two spines mediodistally, equal in length to merus; dactylus (
<figureCitation id="BF8B2CF3FFBAFFCCFE499C87FE6A6A30" box="[411,506,237,263]" captionStart="FIG" captionStartId="3.[113,124,1644,1661]" captionTargetBox="[169,1153,211,1610]" captionTargetId="figure-15@3.[169,1153,211,1610]" captionTargetPageId="3" captionText="FIG. 2. — Batella praecipua n. sp., paratype (OUMNH 2004-01-0001); A, mandible; B, third maxilliped; C, tip of third maxilliped; D, antennular peduncle; E, second pereiopod; F, tip of second pereiopod (denuded); G, third pereiopod; H, dactyl of third pereio- pod; I, dactyl of fourth pereiopod. Scale bar: A, C, 0.1 mm; B, D, E, G, 0.2 mm; F, 0.03 mm; H, I, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4689559" httpUri="https://zenodo.org/record/4689559/files/figure.png" pageId="5" pageNumber="508">Figs 2H</figureCitation>
;
<figureCitation id="BF8B2CF3FFBAFFCCFDD59C87FDB56A30" box="[519,549,237,263]" captionStart="FIG" captionStartId="4.[113,124,1274,1291]" captionTargetBox="[126,1189,212,1237]" captionTargetId="figure-24@4.[33,1225,173,1326]" captionTargetPageId="4" captionText="FIG. 3. — Batella praecipua n. sp., holotype (MHNN Na 11396); A, outer branch of external flagellum of antenna; B, scaphocerite; C, detail of anterolateral corner; D, second pereiopod; E, chelae of second pereiopod (denuded); F, dactyl of third pereiopod. Scale bar: A, 0.1 mm; B, D, 0.8 mm; C, 0.02 mm; E, F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4689561" httpUri="https://zenodo.org/record/4689561/files/figure.png" pageId="5" pageNumber="508">3F</figureCitation>
) slightly curved, approximately 0.25 times length of propodus, unguis clearly demarcated, serrated on inferior margin, not biunguiculate.
</paragraph>
<paragraph id="270F3076FFBAFFCCFFA39D07FDDF6AF0" blockId="5.[113,647,204,839]" pageId="5" pageNumber="508">
Fourth and fifth pereiopods similar to third, unguis (
<figureCitation id="BF8B2CF3FFBAFFCCFF189DE7FE866A90" box="[202,278,397,423]" captionStart="FIG" captionStartId="3.[113,124,1644,1661]" captionTargetBox="[169,1153,211,1610]" captionTargetId="figure-15@3.[169,1153,211,1610]" captionTargetPageId="3" captionText="FIG. 2. — Batella praecipua n. sp., paratype (OUMNH 2004-01-0001); A, mandible; B, third maxilliped; C, tip of third maxilliped; D, antennular peduncle; E, second pereiopod; F, tip of second pereiopod (denuded); G, third pereiopod; H, dactyl of third pereio- pod; I, dactyl of fourth pereiopod. Scale bar: A, C, 0.1 mm; B, D, E, G, 0.2 mm; F, 0.03 mm; H, I, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4689559" httpUri="https://zenodo.org/record/4689559/files/figure.png" pageId="5" pageNumber="508">Fig. 2I</figureCitation>
) serrated on inferior margin, not biunguiculate; fifth pereiopod more slender.
</paragraph>
<paragraph id="270F3076FFBAFFCCFFA39DA7FE526AD1" blockId="5.[113,647,204,839]" box="[113,450,460,487]" pageId="5" pageNumber="508">
Pleopods of usual alpheid
<typeStatus id="F80B8ED4FFBAFFCCFE5E9DA6FE2E6AD1" box="[396,446,460,486]" pageId="5" pageNumber="508">type</typeStatus>
.
</paragraph>
<paragraph id="270F3076FFBAFFCCFFA39D87FE4A69B0" blockId="5.[113,647,204,839]" pageId="5" pageNumber="508">Uropodal endopod reaching to end of telson, exopod slightly shorter, protopod laterally round- ed, exopod with diaresis complete, ending in semi-acute lateral tooth, lateral spine well developed, overreaching distal margin.</paragraph>
<paragraph id="270F3076FFBAFFCCFFA39EE7FD176871" blockId="5.[113,647,204,839]" pageId="5" pageNumber="508">Telson approximately 1.3 length of sixth somite, lateral margins straight, narrowing distally; two pairs of dorsal spines, situated at 0.45 and 0.80 of telson length; posterior margin broadly rounded, laterally with two pairs of spines, mesial pair longest, medially with three to five plumose setae.</paragraph>
</subSubSection>
<subSubSection id="6FAA63FDFFBAFFCCFFA39F06FBAE6930" pageId="5" pageNumber="508" type="discussion">
<paragraph id="270F3076FFBAFFCCFFA39F06FF4568B3" blockId="5.[113,646,876,1703]" box="[113,213,876,902]" pageId="5" pageNumber="508">REMARKS</paragraph>
<paragraph id="270F3076FFBAFFCCFFA39FE7FF506EB0" blockId="5.[113,646,876,1703]" pageId="5" pageNumber="508">
Although both specimens lack the first pereiopods (a diagnostic feature in
<taxonomicName id="E0B04BF5FFBAFFCCFDD79FC7FDEA68F0" authorityName="Rafinesque" authorityYear="1815" box="[517,634,941,967]" class="Malacostraca" family="Alpheidae" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="508" phylum="Arthropoda" rank="family">Alpheidae</taxonomicName>
), they clearly belong to the genus
<taxonomicName id="E0B04BF5FFBAFFCCFDDB9FA7FDCC68D0" authorityName="Holthuis" authorityYear="1955" box="[521,604,973,999]" class="Malacostraca" family="Alpheidae" genus="Batella" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis id="15C4EC64FFBAFFCCFDDB9FA7FDCC68D0" box="[521,604,973,999]" italics="true" pageId="5" pageNumber="508">Batella</emphasis>
</taxonomicName>
, as demonstrated by the lack of a mandibular palp, the incisor of the mandible terminating in setae rather than serrations, and the tufts of long, plumose setae covering the tips of the fingers of the second pereiopods (
<bibRefCitation id="43214D87FFBAFFCCFE539807FD116FB0" author="MIYA Y. &amp; MIYAKE S." box="[385,641,1133,1159]" pageId="5" pageNumber="508" pagination="113 - 120" refId="ref2430" refString="MIYA Y. &amp; MIYAKE S. 1968. - Redefinition of the genus Batella (Crustacea, Decapoda, Alpheidae), with description of a new species from Kyushu, Japan. Ohmu 1 (5): 113 - 120." type="journal article" year="1968">Miya &amp; Miyake 1968</bibRefCitation>
;
<bibRefCitation id="43214D87FFBAFFCCFFA398E7FF6D6F90" author="CHACE F. A. JR." box="[113,253,1165,1191]" pageId="5" pageNumber="508" pagination="1 - 99" refId="ref2304" refString="CHACE F. A. JR. 1988. - The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, part 5: family Alpheidae. Smithsonian Contributions to Zoology 466: 1 - 99." type="journal article" year="1988">Chace 1988</bibRefCitation>
). Until now only two species were known in the genus:
<taxonomicName id="E0B04BF5FFBAFFCCFEA398C7FDC16FF2" baseAuthorityName="Bate" baseAuthorityYear="1888" box="[369,593,1196,1223]" class="Malacostraca" family="Alpheidae" genus="Batella" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="508" phylum="Arthropoda" rank="species" species="parvimanus">
<emphasis id="15C4EC64FFBAFFCCFEA398C7FDC16FF2" box="[369,593,1196,1223]" italics="true" pageId="5" pageNumber="508">Batella parvimanus</emphasis>
</taxonomicName>
and
<taxonomicName id="E0B04BF5FFBAFFCCFFA398A7FEAA6FD0" authorityName="Chace" authorityYear="1988" box="[113,314,1229,1255]" class="Malacostraca" family="Alpheidae" genus="Batella" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="508" phylum="Arthropoda" rank="species" species="leptocarpus">
<emphasis id="15C4EC64FFBAFFCCFFA398A7FEAA6FD0" box="[113,314,1229,1255]" italics="true" pageId="5" pageNumber="508">Batella leptocarpus</emphasis>
</taxonomicName>
. The third described species,
<taxonomicName id="E0B04BF5FFBAFFCCFFA39887FDDD6E30" authority="Miya &amp; Miyake, 1968" authorityName="Miya &amp; Miyake" authorityYear="1968" box="[113,589,1261,1287]" class="Malacostraca" family="Alpheidae" genus="Batella" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="508" phylum="Arthropoda" rank="species" species="bifurcata">
<emphasis id="15C4EC64FFBAFFCCFFA39887FEA36E30" box="[113,307,1261,1287]" italics="true" pageId="5" pageNumber="508">Batella bifurcata</emphasis>
<bibRefCitation id="43214D87FFBAFFCCFEEF9887FDDD6E30" author="MIYA Y. &amp; MIYAKE S." box="[317,589,1261,1287]" pageId="5" pageNumber="508" pagination="113 - 120" refId="ref2430" refString="MIYA Y. &amp; MIYAKE S. 1968. - Redefinition of the genus Batella (Crustacea, Decapoda, Alpheidae), with description of a new species from Kyushu, Japan. Ohmu 1 (5): 113 - 120." type="journal article" year="1968">Miya &amp; Miyake, 1968</bibRefCitation>
</taxonomicName>
, was considered a junior synonym of
<taxonomicName id="E0B04BF5FFBAFFCCFE3E9967FF256E72" baseAuthorityName="Bate" baseAuthorityYear="1888" class="Malacostraca" family="Alpheidae" genus="Batella" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="508" phylum="Arthropoda" rank="species" species="parvimanus">
<emphasis id="15C4EC64FFBAFFCCFE3E9967FF256E72" italics="true" pageId="5" pageNumber="508">Batella parvimanus</emphasis>
</taxonomicName>
by
<bibRefCitation id="43214D87FFBAFFCCFF309947FEE16E70" author="MIYA Y." box="[226,369,1325,1351]" pageId="5" pageNumber="508" pagination="217 - 219" refId="ref2388" refString="MIYA Y. 1984. - Batella bifurcata Miya &amp; Miyake, 1968, a junior synonym of B. parvimanus (Bate, 1888) (Decapoda, Alpheidae). Crustaceana 47 (2): 217 - 219." type="journal article" year="1984">Miya (1984)</bibRefCitation>
, as its distinctive characters were based on a misinterpretation of Bate’s figures.
</paragraph>
<paragraph id="270F3076FFBAFFCCFFA399E7FBAE6930" blockId="5.[113,646,876,1703]" lastBlockId="5.[676,1209,204,1063]" pageId="5" pageNumber="508">
The new species can at once be distinguished from the other species by the non-tridentate nature of the frontal part of the carapace, which is clearly tridentate in both
<taxonomicName id="E0B04BF5FFBAFFCCFE609986FDC36D32" baseAuthorityName="Bate" baseAuthorityYear="1888" box="[434,595,1516,1541]" class="Malacostraca" family="Alpheidae" genus="Batella" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="508" phylum="Arthropoda" rank="species" species="parvimanus">
<emphasis id="15C4EC64FFBAFFCCFE609986FDC36D32" box="[434,595,1516,1541]" italics="true" pageId="5" pageNumber="508">B. parvimanus</emphasis>
</taxonomicName>
and
<taxonomicName id="E0B04BF5FFBAFFCCFFA39A66FE956D10" authorityName="Chace" authorityYear="1988" box="[113,261,1548,1575]" class="Malacostraca" family="Alpheidae" genus="Batella" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="508" phylum="Arthropoda" rank="species" species="leptocarpus">
<emphasis id="15C4EC64FFBAFFCCFFA39A66FE956D10" box="[113,261,1548,1575]" italics="true" pageId="5" pageNumber="508">B. leptocarpus</emphasis>
</taxonomicName>
, although damaged in the unique specimen of the latter species (
<bibRefCitation id="43214D87FFBAFFCCFE319A47FDE96D70" author="CHACE F. A. JR." box="[483,633,1581,1607]" pageId="5" pageNumber="508" pagination="1 - 99" refId="ref2304" refString="CHACE F. A. JR. 1988. - The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, part 5: family Alpheidae. Smithsonian Contributions to Zoology 466: 1 - 99." type="journal article" year="1988">Chace 1988</bibRefCitation>
). Although it was at first assumed that this nontridentate nature was possibly due to damage to the specimens and potential regeneration, further morphological differences are apparent. The unguis of the ambulatory pereiopods is serrate and non-biunguiculate in
<taxonomicName id="E0B04BF5FFBAFFCCFC199D66FBC36A12" authority="Grave, 2004" authorityName="Grave" authorityYear="2004" box="[971,1107,268,293]" class="Malacostraca" family="Alpheidae" genus="Batella" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="508" phylum="Arthropoda" rank="species" species="praecipua" status="sp. nov.">
<emphasis id="15C4EC64FFBAFFCCFC199D66FBC36A12" box="[971,1107,268,293]" italics="true" pageId="5" pageNumber="508">B. praecipua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0EF7511FFFBAFFCCFB8E9D66FB0A6A11" box="[1116,1178,268,294]" pageId="5" pageNumber="508" rank="species">n. sp.</taxonomicNameLabel>
vs non-serrate and biunguiculate in
<taxonomicName id="E0B04BF5FFBAFFCCFBC99D46FB296A72" baseAuthorityName="Bate" baseAuthorityYear="1888" box="[1051,1209,300,325]" class="Malacostraca" family="Alpheidae" genus="Batella" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="508" phylum="Arthropoda" rank="species" species="parvimanus">
<emphasis id="15C4EC64FFBAFFCCFBC99D46FB296A72" box="[1051,1209,300,325]" italics="true" pageId="5" pageNumber="508">B. parvimanus</emphasis>
</taxonomicName>
and
<taxonomicName id="E0B04BF5FFBAFFCCFD059D26FCE16A50" authorityName="Chace" authorityYear="1988" box="[727,881,332,359]" class="Malacostraca" family="Alpheidae" genus="Batella" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="508" phylum="Arthropoda" rank="species" species="leptocarpus">
<emphasis id="15C4EC64FFBAFFCCFD059D26FCE16A50" box="[727,881,332,359]" italics="true" pageId="5" pageNumber="508">B. leptocarpus</emphasis>
</taxonomicName>
; the pterygostomial angle is rounded and only slightly produced in
<taxonomicName id="E0B04BF5FFBAFFCCFBB09D06FD4C6A92" authority="Grave, 2004" authorityName="Grave" authorityYear="2004" class="Malacostraca" family="Alpheidae" genus="Batella" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="508" phylum="Arthropoda" rank="species" species="praecipua" status="sp. nov.">
<emphasis id="15C4EC64FFBAFFCCFBB09D06FD4C6A92" italics="true" pageId="5" pageNumber="508">B. praecipua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0EF7511FFFBAFFCCFD369DE6FCB06A91" box="[740,800,396,422]" pageId="5" pageNumber="508" rank="species">n. sp.</taxonomicNameLabel>
vs triangularly produced in the other species; the uropods are near-equal in length to the telson, as opposed to being distinctly shorter than the telson as in the other species.
</paragraph>
</subSubSection>
<subSubSection id="6FAA63FDFFBAFFCCFD769E67FB8C6F11" pageId="5" pageNumber="508" type="reference_group">
<paragraph id="270F3076FFBAFFCCFD769E67FC476990" blockId="5.[676,1209,204,1063]" pageId="5" pageNumber="508">
<taxonomicName id="E0B04BF5FFBAFFCCFD769E67FCE16912" ID-CoL="85MW3" authority="Grave, 2004" authorityName="Grave" authorityYear="2004" box="[676,881,524,551]" class="Malacostraca" family="Alpheidae" genus="Batella" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="508" phylum="Arthropoda" rank="species" species="praecipua" status="sp. nov.">
<emphasis id="15C4EC64FFBAFFCCFD769E67FCE16912" box="[676,881,524,551]" italics="true" pageId="5" pageNumber="508">Batella praecipua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0EF7511FFFBAFFCCFCAE9E66FC2F6911" box="[892,959,524,550]" pageId="5" pageNumber="508" rank="species">n. sp.</taxonomicNameLabel>
further differs from
<taxonomicName id="E0B04BF5FFBAFFCCFD769E46FCAC6972" ID-CoL="85MW2" baseAuthorityName="Bate" baseAuthorityYear="1888" box="[676,828,556,581]" class="Malacostraca" family="Alpheidae" genus="Batella" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="508" phylum="Arthropoda" rank="species" species="parvimanus">
<emphasis id="15C4EC64FFBAFFCCFD769E46FCAC6972" box="[676,828,556,581]" italics="true" pageId="5" pageNumber="508">B. parvimanus</emphasis>
</taxonomicName>
in having only a single segmented outer branch of the external flagellum on the antennule, this character being unknown in
<taxonomicName id="E0B04BF5FFBAFFCCFD769EE6FC426990" ID-CoL="85MVZ" authority="(Chace 1988)" baseAuthorityName="Chace" baseAuthorityYear="1988" box="[676,978,652,679]" class="Malacostraca" family="Alpheidae" genus="Batella" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="508" phylum="Arthropoda" rank="species" species="leptocarpus">
<emphasis id="15C4EC64FFBAFFCCFD769EE6FCA46990" box="[676,820,652,679]" italics="true" pageId="5" pageNumber="508">B. leptocarpus</emphasis>
(
<bibRefCitation id="43214D87FFBAFFCCFC909EE7FC5A6990" author="CHACE F. A. JR." box="[834,970,653,679]" pageId="5" pageNumber="508" pagination="1 - 99" refId="ref2304" refString="CHACE F. A. JR. 1988. - The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, part 5: family Alpheidae. Smithsonian Contributions to Zoology 466: 1 - 99." type="journal article" year="1988">Chace 1988</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="270F3076FFBAFFCCFD769EC7FB8C6F11" blockId="5.[676,1209,204,1063]" pageId="5" pageNumber="508">
The
<typeStatus id="F80B8ED4FFBAFFCCFD0D9EC7FCDC69F0" box="[735,844,685,711]" pageId="5" pageNumber="508" type="holotype">holotype</typeStatus>
was collected from a sponge containing stenopodidean shrimp, whilst the sample containing the
<typeStatus id="F80B8ED4FFBAFFCCFC799E86FB9B6831" box="[939,1035,748,774]" pageId="5" pageNumber="508" type="paratype">paratype</typeStatus>
also contained two specimens of
<taxonomicName id="E0B04BF5FFBAFFCCFCB69F67FBA46810" box="[868,1076,781,807]" class="Malacostraca" family="Spongicolidae" genus="Spongicola" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="508" phylum="Arthropoda" rank="species" species="laevigata">
<emphasis id="15C4EC64FFBAFFCCFCB69F67FBA46810" box="[868,1076,781,807]" italics="true" pageId="5" pageNumber="508">Spongicola laevigata</emphasis>
</taxonomicName>
and a single specimen of
<taxonomicName id="E0B04BF5FFBAFFCCFCE39F47FBA46870" ID-CoL="9308c942-a357-4f96-905f-f40c3b6e2c8f" authorityName="Bruce" authorityYear="1991" box="[817,1076,813,839]" class="Malacostraca" family="Palaemonidae" genus="Periclimenes" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="508" phylum="Arthropoda" rank="species" species="forcipulatus">
<emphasis id="15C4EC64FFBAFFCCFCE39F47FBA46870" box="[817,1076,813,839]" italics="true" pageId="5" pageNumber="508">Periclimenes forcipulatus</emphasis>
</taxonomicName>
. As both of these species are known associates of hexactinellid sponges, this lends further credence to the hypothesis by
<bibRefCitation id="43214D87FFBAFFCCFC949FE6FC4B6890" author="BRUCE A. J." box="[838,987,908,935]" pageId="5" pageNumber="508" pagination="139 - 149" refId="ref2265" refString="BRUCE A. J. 1988. - Bannereus anomalus, new genus, new species, a deep-sea alpheid shrimp from the Coral Sea. Pacific Science 42 (3 - 4): 139 - 149." type="journal article" year="1988">Bruce (1988)</bibRefCitation>
that
<taxonomicName id="E0B04BF5FFBAFFCCFBC89FE7FBF56890" ID-CoL="858XS" authorityName="Holthuis" authorityYear="1955" box="[1050,1125,909,935]" class="Malacostraca" family="Alpheidae" genus="Batella" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis id="15C4EC64FFBAFFCCFBC89FE7FBF56890" box="[1050,1125,909,935]" italics="true" pageId="5" pageNumber="508">Batella</emphasis>
</taxonomicName>
species are all probably hexactinellid or other sponge associates. The specimens were collected from depths between 400 and
<quantity id="E0489D93FFBAFFCCFC679F87FC6F6F31" box="[949,1023,1004,1031]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="5" pageNumber="508" unit="m" value="450.0">450 m</quantity>
, representing the deepest record so far for the genus.
</paragraph>
</subSubSection>
</treatment>
</document>
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