118 lines
14 KiB
XML
118 lines
14 KiB
XML
<document id="5156F048AD724AADD63E4CAC5E64FDB2" ID-CLB-Dataset="33695" ID-DOI="http://dx.doi.org/10.3897/zookeys.537.6090" ID-GBIF-Dataset="ae220ec2-27f7-43de-b96d-3b5e0dd853b6" ID-PMC="PMC4714052" ID-Pensoft-Pub="1313-2970-537-131" ID-PubMed="26798242" ID-ZooBank="EA901A84CB134889A77C07E89EDA172E" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-537-131" ModsDocOrigin="ZooKeys 537" ModsDocTitle="Three new species of Hagnagora Druce, 1885 (Lepidoptera, Geometridae, Larentiinae) from Ecuador and Costa Rica and a concise revision of the genus" checkinTime="1451243721882" checkinUser="pensoft" docAuthor="Brehm, Gunnar" docDate="2015" docId="AF06576E5442E55008A0FA53E9736E34" docLanguage="en" docName="ZooKeys 537: 131-156" docOrigin="ZooKeys 537" docSource="http://dx.doi.org/10.3897/zookeys.537.6090" docTitle="Hagnagora anicata Felder & Rogenhofer 1875" docType="treatment" docVersion="6" lastPageNumber="136" masterDocId="FFCE8A69FFE9FFA2FFE1FFD8EE5D462F" masterDocTitle="Three new species of Hagnagora Druce, 1885 (Lepidoptera, Geometridae, Larentiinae) from Ecuador and Costa Rica and a concise revision of the genus" masterLastPageNumber="156" masterPageNumber="131" pageNumber="135" updateTime="1701384186686" updateUser="ExternalLinkService">
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<mods:mods id="B5F6222BA1CE2E4FF9C0A9C02E27B35F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="FCD11782364081D6852074E11FDADF6F">
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<mods:title id="BEC087735CE3C1B617A3696B21CB38A4">Three new species of Hagnagora Druce, 1885 (Lepidoptera, Geometridae, Larentiinae) from Ecuador and Costa Rica and a concise revision of the genus</mods:title>
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<mods:name id="B7FC355AAEE812BC15A6FA59776BC3D5" type="personal">
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<mods:roleTerm id="56B8A5B1FCADAAFCF64095C6D1657991">Author</mods:roleTerm>
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<mods:namePart id="813EF5BF23E138D518F88242CC588E3C">Brehm, Gunnar</mods:namePart>
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<mods:typeOfResource id="CBC1588AECDACE95512520139712739D">text</mods:typeOfResource>
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<mods:title id="1C890D816291CAE216D9DBEA76ED99B5">ZooKeys</mods:title>
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<mods:part id="C8032F9E9057654E1A116F6F7CA02D31">
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<mods:date id="2FD05628975E3C81F868E69860BA3691">2015</mods:date>
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<mods:detail id="36545D75A8596BE69F514013B7763E57" type="volume">
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<mods:number id="BC71847655AC79BA50189FAE02C23912">537</mods:number>
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</mods:detail>
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<mods:extent id="8F8B01B113AF202E3F0A555574159B73" unit="page">
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<mods:start id="4EA25F02EDA74D598B0BE064F728777F">131</mods:start>
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<mods:end id="2D38C1E50266BDDEED67679606B32A55">156</mods:end>
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<mods:location id="DEEB755FF9DF776AE9E801500E95D746">
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<mods:url id="A2FFE1C2CB313148778C332A690B0A44">http://dx.doi.org/10.3897/zookeys.537.6090</mods:url>
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</mods:location>
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<mods:classification id="5B469DEAD8DB6AC8126A42E656D260C6">journal article</mods:classification>
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<mods:identifier id="D6176CA09F358835E79A4E33AC5505BC" type="DOI">http://dx.doi.org/10.3897/zookeys.537.6090</mods:identifier>
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<mods:identifier id="F746C850EF52D084542BA1F750532234" type="Pensoft-Pub">1313-2970-537-131</mods:identifier>
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<mods:identifier id="3852163BF6EB8F15408374F8129921B5" type="ZooBank">EA901A84CB134889A77C07E89EDA172E</mods:identifier>
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</mods:mods>
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<treatment id="AF06576E5442E55008A0FA53E9736E34" ID-GBIF-Taxon="127897775" LSID="urn:lsid:plazi:treatment:AF06576E5442E55008A0FA53E9736E34" httpUri="http://treatment.plazi.org/id/AF06576E5442E55008A0FA53E9736E34" lastPageId="5" lastPageNumber="136" pageId="4" pageNumber="135">
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<subSubSection id="3DCB470BBA5695D1BBE793382C62FE66" pageId="4" pageNumber="135" type="multiple">
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<paragraph id="00DE5CB2F944165583AEF3F446DF245E" pageId="4" pageNumber="135">Taxon classification Animalia Lepidoptera Geometridae</paragraph>
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</subSubSection>
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<subSubSection id="E92F65319ABFE453CD15EFB74D040D07" pageId="4" pageNumber="135" type="nomenclature">
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<paragraph id="C139DF8CDD0BA08E879A698BD044382A" pageId="4" pageNumber="135">
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<taxonomicName id="73032C26E7A62F4C063991F19139B7E8" ID-CoL="934NS" LSID="BIN: BOLD:AAA7285" authority="Felder & Rogenhofer, 1875" authorityName="Felder & Rogenhofer" authorityYear="1875" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora anicata" order="Lepidoptera" pageId="4" pageNumber="135" phylum="Arthropoda" rank="species" species="anicata">Hagnagora anicata (Felder & Rogenhofer, 1875)</taxonomicName>
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Figs 7, 8
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</paragraph>
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</subSubSection>
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<subSubSection id="D21730F3A8DA3E82BC99A33F4C09634E" pageId="5" pageNumber="136" type="type locality">
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<paragraph id="0CDD91EA0B4B7FC268A69670097B30CA" pageId="5" pageNumber="136">
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<pageBreakToken id="1DFE6F6C31BC88B5AB20DF90CED744C1" pageId="5" pageNumber="136" start="start">Type</pageBreakToken>
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locality.
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</paragraph>
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<paragraph id="10CBEFF8BA82C01F57653A0C70ED19C0" pageId="5" pageNumber="136">
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[Colombia],
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<normalizedToken id="418619198835D067B07663B19F7DDAE1" originalValue="Bogotá">Bogota</normalizedToken>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="99987177A37F9A71D94342C8BBA4C926" pageId="5" pageNumber="136" type="remarks">
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<paragraph id="DDDABFC51D00C3805DADFE96DC8A60DB" pageId="5" pageNumber="136">Remarks.</paragraph>
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<paragraph id="BDA14541103F5E81C54A1724D18B53CB" pageId="5" pageNumber="136">
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<taxonomicName id="BF84BE618A8B7542A1EF77702CC63E21" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora anicata" order="Lepidoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="species" species="anicata">Hagnagora anicata</taxonomicName>
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was re-described with a description also of the male genitalia, by Sullivan (2013). The lectotype is illustrated in Fig. 7. A series of specimens collected in southern Ecuador (1999-2013) (Fig. 8) is indistinguishable from
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<taxonomicName id="963E9F06B5F4947D69A894A50CC1D4BB" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora anicata" order="Lepidoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="species" species="anicata">Hagnagora anicata</taxonomicName>
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and therefore regarded as conspecific. The female (Fig. 12) is larger than the male. A living specimen is shown in Fig. 41 in the typical resting position of these beautiful moths.
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</paragraph>
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<caption id="1B6500BE9859ADB0827B25E462FAF118" pageId="5" pageNumber="136">
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<paragraph id="166FEBD17011AC23D75353CED07820C8" pageId="5" pageNumber="136">
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Figures 7-11. 7
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<taxonomicName id="D33821AFD9404C790BD1A03332274680" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora anicata" order="Lepidoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="species" species="anicata">Hagnagora anicata</taxonomicName>
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(F&R), male lectotype a dorsal view b ventral view c valvae d aedeagus 8
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<taxonomicName id="80AF5905939815BF635073EBCEFCF1BE" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora anicata" order="Lepidoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="species" species="anicata">Hagnagora anicata</taxonomicName>
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(F&R), male from Ecuador as reference specimen with Barcode Index Number (BIN) a dorsal view b ventral view c valvae d aedeagus 9
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<taxonomicName id="A2FF549BE536F5809362F3E51A0870A0" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora richardi" order="Lepidoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="species" species="richardi">Hagnagora richardi</taxonomicName>
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sp. n., male holotype a dorsal view b ventral view c valvae d aedeagus 10
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<taxonomicName id="F620044DD0461EBDAC4E832B600F54C5" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora unnia" order="Lepidoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="species" species="unnia">Hagnagora unnia</taxonomicName>
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Sullivan, male a dorsal view b ventral view 11
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<taxonomicName id="DAE831EE7421309A0F8AF2A184699726" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora marionae" order="Lepidoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="species" species="marionae">Hagnagora marionae</taxonomicName>
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Brehm & Sullivan, male a dorsal view b ventral view.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="401952B1127E96B5415593EF22330C4E" pageId="5" pageNumber="136" type="distribution">
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<paragraph id="E3C6A2C81ACDD8452758F2B6D61AFB8D" pageId="5" pageNumber="136">Distribution.</paragraph>
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<paragraph id="93DC3D75012881D823AB21A13042AA16" pageId="5" pageNumber="136">
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Apart from its Colombian type locality,
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<taxonomicName id="6E2A02F7DFB8561AB1995F5C54FFB5BD" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora anicata" order="Lepidoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="species" species="anicata">Hagnagora anicata</taxonomicName>
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has recently been collected and barcoded from sites in southern Ecuador to central Bolivia at elevations ranging from 2000 to 2920 m a.s.l.
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</paragraph>
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</subSubSection>
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<subSubSection id="20D00F80B90859E2470D6F81A40F6368" pageId="5" pageNumber="136" type="diagnosis">
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<paragraph id="1E8A09534008A02B32756552C5F37BCE" pageId="5" pageNumber="136">Diagnosis.</paragraph>
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<paragraph id="D904321FB10BE5DF7B2948636B08DC7B" pageId="5" pageNumber="136">
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Most species of the
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<taxonomicName id="99582F001BD09277B7A155E4CB0D585D" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora anicata" order="Lepidoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="species" species="anicata">Hagnagora anicata</taxonomicName>
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clade are very similar, and the most reliable current method for diagnosis is the COI barcode.
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<taxonomicName id="94B7CF1BF8DE7A71F9240B7D5B4BE596" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora anicata" order="Lepidoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="species" species="anicata">Hagnagora anicata</taxonomicName>
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tends to be smaller than the other species occurring sympatrically, namely
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<taxonomicName id="506797C23EBFD427553C801270872DDE" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora richardi" order="Lepidoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="species" species="richardi">Hagnagora richardi</taxonomicName>
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and
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<taxonomicName id="090487F3360DB2D7EB940B88F43485EB" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora hedwigae" order="Lepidoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="species" species="hedwigae">Hagnagora hedwigae</taxonomicName>
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: The wing length of the male (holotype) is only 17.5 mm in comparison to 19 mm in the male holotype of
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<taxonomicName id="9DAFCEE00AC62275AD5644052A3E2391" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora richardi" order="Lepidoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="species" species="richardi">Hagnagora richardi</taxonomicName>
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. The structures of the female signum are also more complex than in
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<taxonomicName id="4335C4781DA101261CE8FC0782630F1C" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora richardi" order="Lepidoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="species" species="richardi">Hagnagora richardi</taxonomicName>
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, but similar to those in
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<taxonomicName id="71F7A457F871DD535CED6B54E8FF89D2" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora hedwigae" order="Lepidoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="species" species="hedwigae">Hagnagora hedwigae</taxonomicName>
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. The uncus of the male is smaller and shorter than in
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<taxonomicName id="799AEE140083EB675984C5D212665C95" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora richardi" order="Lepidoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="species" species="richardi">Hagnagora richardi</taxonomicName>
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. Aedeagi of the known males are (
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<taxonomicName id="65788DF94204663116D9C3447BBDC1A5" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora anicata" order="Lepidoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="species" species="anicata">Hagnagora anicata</taxonomicName>
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and
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<taxonomicName id="1CD7311DEF69ECEA6FEADAEB1C5EDC92" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora richardi" order="Lepidoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="species" species="richardi">Hagnagora richardi</taxonomicName>
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) similar. COI barcode: The minimum observed distance to the presumbably most closely related species (
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<taxonomicName id="E9E367E06FDCB1D9D097360402B6F345" class="Insecta" family="Geometridae" genus="Hagnagora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hagnagora hedwigae" order="Lepidoptera" pageId="5" pageNumber="136" phylum="Arthropoda" rank="species" species="hedwigae">Hagnagora hedwigae</taxonomicName>
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) is 3.1%.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |