treatments-xml/data/C3/60/A2/C360A203FFB2FFEAFF17F9B2FD95F85C.xml
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<document id="32D538EEB9324DC061DD77FA60F00A1A" ID-CLB-Dataset="24219" ID-DOI="10.11646/zootaxa.4729.2.7" ID-GBIF-Dataset="1221bec5-8abd-4b4f-b1dc-7d0fe8065ce0" ID-ISSN="1175-5326" ID-Zenodo-Dep="3632638" ID-ZooBank="8CC9339D-6709-4CC7-8E6E-832FC493CC1C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1580475164295" checkinUser="plazi" docAuthor="Dinesh, K. P., Vijayakumar, S. P., Ramesh, Vijay, Jayarajan, Aditi, Chandramouli, S. R. &amp; Shanker, Kartik" docDate="2020" docId="C360A203FFB2FFEAFF17F9B2FD95F85C" docLanguage="en" docName="zootaxa.4729.2.7.pdf" docOrigin="Zootaxa 4729 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Walkerana muduga Dinesh &amp; Vijayakumar &amp; Ramesh &amp; Jayarajan &amp; Chandramouli &amp; Shanker 2020, sp. nov." docType="treatment" docUuid="5AF718F0-CB6E-41F3-816E-5BF838DF83C9" docUuidSource="ZooBank" docVersion="8" lastPageNumber="271" masterDocId="3F59DA7BFFB1FFEFFF80FFB9FFC5FF98" masterDocTitle="A deeply divergent lineage of Walkerana (Anura: Ranixalidae) from the Western Ghats of Peninsular India" masterLastPageNumber="276" masterPageNumber="266" pageNumber="269" updateTime="1698779764623" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="39ED3E908D6D9ACBC0EAFD6DC8635141">A deeply divergent lineage of Walkerana (Anura: Ranixalidae) from the Western Ghats of Peninsular India</mods:title>
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<mods:namePart id="E301BC893CFE653713B4207B334B3D63">Dinesh, K. P.</mods:namePart>
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<mods:namePart id="691964BB0BDB81979E74001B94D736FA">Vijayakumar, S. P.</mods:namePart>
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<mods:namePart id="54E3FF635D94C23DC364D900768933CC">Ramesh, Vijay</mods:namePart>
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<mods:namePart id="63F9C940473198FCDF33BBDC7BE27DE5">Chandramouli, S. R.</mods:namePart>
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<paragraph id="4B761315FFB2FFECFF17F9B2FE2FF9BE" blockId="3.[151,490,1547,1610]" box="[151,490,1547,1574]" pageId="3" pageNumber="269">
<heading id="103EA479FFB2FFECFF17F9B2FE2FF9BE" bold="true" box="[151,490,1547,1574]" fontSize="11" level="1" pageId="3" pageNumber="269" reason="1">
<emphasis id="79BDCF07FFB2FFECFF17F9B2FE2FF9BE" bold="true" box="[151,490,1547,1574]" pageId="3" pageNumber="269">
<taxonomicName id="8CC96896FFB2FFECFF17F9B2FE40F9BE" authority="Dinesh &amp; Vijayakumar &amp; Ramesh &amp; Jayarajan &amp; Chandramouli &amp; Shanker, 2020" authorityName="Dinesh &amp; Vijayakumar &amp; Ramesh &amp; Jayarajan &amp; Chandramouli &amp; Shanker" authorityYear="2020" box="[151,389,1547,1574]" class="Insecta" family="Haglotettigoniidae" genus="Walkerana" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="269" phylum="Arthropoda" rank="species" species="muduga" status="sp. nov.">
<emphasis id="79BDCF07FFB2FFECFF17F9B2FE40F9BE" bold="true" box="[151,389,1547,1574]" italics="true" pageId="3" pageNumber="269">Walkerana muduga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="628E727CFFB2FFECFE0DF9B5FE2FF9BE" box="[397,490,1548,1574]" pageId="3" pageNumber="269" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
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<paragraph id="4B761315FFB2FFECFF17F989FE78F9D2" blockId="3.[151,490,1547,1610]" box="[151,445,1583,1610]" pageId="3" pageNumber="269">
(
<tableCitation id="064B26AEFFB2FFECFF1FF989FF36F9D2" box="[159,243,1584,1610]" captionStart="TABLE 1" captionStartId="6.[151,239,154,178]" captionTargetPageId="6" captionText="TABLE 1. Morphometric data (in mm) for the type series and reference collections of Walkerana muduga sp. nov." httpUri="http://table.plazi.org/id/1FB6439DFFB7FFE9FF17FF23FAA5FF2A" pageId="3" pageNumber="269" tableUuid="1FB6439DFFB7FFE9FF17FF23FAA5FF2A">Table 1</tableCitation>
,
<tableCitation id="064B26AEFFB2FFECFE81F989FED7F9D2" box="[257,274,1584,1610]" captionStart="TABLE 2" captionStartId="6.[151,239,1349,1373]" captionTargetPageId="6" captionText="TABLE 2. Extant valid species of Walkerana in the Western Ghats and GenBank accession numbers for the Ranixalidae taxa used in the construction of Bayesian tree" httpUri="http://table.plazi.org/id/1FB6439DFFB7FFE9FF17FAFCFDB9FA19" pageId="3" pageNumber="269" tableUuid="1FB6439DFFB7FFE9FF17FAFCFDB9FA19">2</tableCitation>
; Fig. 1,2,3,4,5)
</paragraph>
</subSubSection>
<subSubSection id="03D3409EFFB2FFECFF17F9C1FF0AF8BA" pageId="3" pageNumber="269" type="materials_examined">
<paragraph id="4B761315FFB2FFECFF17F9C1FD70F942" blockId="3.[151,1437,1656,2042]" pageId="3" pageNumber="269">
<materialsCitation id="FBA11948FFB2FFECFF17F9C1FD70F942" ID-GBIF-Occurrence="2561484859" collectingDate="2010-10" collectionCode="ZSI, WRC, SVL" collectorName="S. P. Vijayakumar" county="Palghat Gap." latitude="10.945" location="Elivalmala" longLatPrecision="77" longitude="76.644" municipality="Elivalmalai" pageId="3" pageNumber="269" specimenCode="CESF 1607" specimenCount="1" typeStatus="holotype">
<emphasis id="79BDCF07FFB2FFECFF17F9C1FEC8F90A" bold="true" box="[151,269,1656,1682]" pageId="3" pageNumber="269">
<typeStatus id="9472ADB7FFB2FFECFF17F9C1FECDF90A" box="[151,264,1656,1682]" pageId="3" pageNumber="269" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="2DD88BD0FFB2FFECFE94F9C1FEF8F90A" box="[276,317,1656,1682]" pageId="3" pageNumber="269">ZSI</collectionCode>
/
<collectionCode id="2DD88BD0FFB2FFECFEC6F9C1FE43F90A" box="[326,390,1656,1682]" country="Japan" httpUri="http://grbio.org/cool/i1ct-fvht" name="Wildlife Research Center of Kyoto University" pageId="3" pageNumber="269">WRC</collectionCode>
/A/2211 (
<specimenCode id="1B6FBB6EFFB2FFECFE6FF9C1FDB2F90B" box="[495,631,1656,1683]" collectionCode="CESF" pageId="3" pageNumber="269">CESF 1607</specimenCode>
), an adult male (
<collectionCode id="2DD88BD0FFB2FFECFCAFF9C1FCA0F90A" box="[815,869,1656,1682]" pageId="3" pageNumber="269">SVL</collectionCode>
<quantity id="8C31BEF0FFB2FFECFCECF9C1FC14F90A" box="[876,977,1656,1682]" metricMagnitude="-2" metricUnit="m" metricValue="2.79" pageId="3" pageNumber="269" unit="mm" value="27.9">27.9 mm</quantity>
) collected by
<collectorName id="E63C76C3FFB2FFECFBEFF9C1FAF5F90A" box="[1135,1328,1656,1682]" pageId="3" pageNumber="269">S.P. Vijayakumar</collectorName>
and team in
<date id="3F7735D5FFB2FFECFF34F925FE88F92E" box="[180,333,1692,1718]" pageId="3" pageNumber="269" value="2010-10">
<collectingDate id="2F33CC3DFFB2FFECFF34F925FE88F92E" box="[180,333,1692,1718]" pageId="3" pageNumber="269" value="2010-10">October 2010</collectingDate>
</date>
from Muthikulam, (
<geoCoordinate id="2EFD75D2FFB2FFECFDB3F925FD65F92E" box="[563,672,1691,1718]" degrees="10.945" direction="north" orientation="latitude" pageId="3" pageNumber="269" precision="55" value="10.945">N 10.945</geoCoordinate>
;
<geoCoordinate id="2EFD75D2FFB2FFECFD2CF925FCD6F92F" box="[684,787,1692,1719]" degrees="76.644" direction="east" orientation="longitude" pageId="3" pageNumber="269" precision="55" value="76.644">E 76.644</geoCoordinate>
) a high elevation site (1544 amsl) in the hill range
<location id="4E1645CEFFB2FFECFAD0F925FF08F942" LSID="urn:lsid:plazi:treatment:C360A203FFB2FFEAFF17F9B2FD95F85C:4E1645CEFFB2FFECFAD0F925FF08F942" county="Palghat Gap." latitude="10.945" longLatPrecision="77" longitude="76.644" municipality="Elivalmalai" name="Elivalmala" pageId="3" pageNumber="269">Elivalmala</location>
/
<collectingMunicipality id="AB12896FFFB2FFECFF63F979FEA1F942" box="[227,356,1728,1754]" pageId="3" pageNumber="269">Elivalmalai</collectingMunicipality>
located north of
<collectingCounty id="A2176B99FFB2FFECFDA4F979FD70F942" box="[548,693,1728,1754]" pageId="3" pageNumber="269">Palghat Gap.</collectingCounty>
</materialsCitation>
</paragraph>
<paragraph id="4B761315FFB2FFECFF47F95DFF0AF8BA" blockId="3.[151,1437,1656,2042]" pageId="3" pageNumber="269">
<materialsCitation id="FBA11948FFB2FFECFF47F95DFBE6F966" ID-GBIF-Occurrence="2561484858" box="[199,1059,1764,1791]" collectionCode="ZSI, WRC, SVL" pageId="3" pageNumber="269" specimenCode="CESF 1606" specimenCount="1" typeStatus="paratype">
<emphasis id="79BDCF07FFB2FFECFF47F95DFEF8F966" bold="true" box="[199,317,1764,1790]" pageId="3" pageNumber="269">
<typeStatus id="9472ADB7FFB2FFECFF47F95DFEFDF966" box="[199,312,1764,1790]" pageId="3" pageNumber="269" type="paratype">Paratype</typeStatus>
:
</emphasis>
<collectionCode id="2DD88BD0FFB2FFECFEC4F95DFEA8F966" box="[324,365,1764,1790]" pageId="3" pageNumber="269">ZSI</collectionCode>
/
<collectionCode id="2DD88BD0FFB2FFECFEF6F95DFE73F966" box="[374,438,1764,1790]" country="Japan" httpUri="http://grbio.org/cool/i1ct-fvht" name="Wildlife Research Center of Kyoto University" pageId="3" pageNumber="269">WRC</collectionCode>
/A/2212 (
<specimenCode id="1B6FBB6EFFB2FFECFDA0F95DFD6DF967" box="[544,680,1764,1791]" collectionCode="CESF" pageId="3" pageNumber="269">CESF 1606</specimenCode>
), an adult female (
<collectionCode id="2DD88BD0FFB2FFECFCF6F95DFC69F966" box="[886,940,1764,1790]" pageId="3" pageNumber="269">SVL</collectionCode>
<quantity id="8C31BEF0FFB2FFECFC33F95DFBDEF966" box="[947,1051,1764,1790]" metricMagnitude="-2" metricUnit="m" metricValue="4.49" pageId="3" pageNumber="269" unit="mm" value="44.9">44.9 mm</quantity>
)
</materialsCitation>
,
<materialsCitation id="FBA11948FFB2FFECFBAEF95DFF0EF8BA" ID-GBIF-Occurrence="2561484857" collectingDate="2010-10" collectionCode="SVL" collectorName="S. P. Vijayakumar" county="Palghat Gap." latitude="10.945" location="Elivalmala" longLatPrecision="77" longitude="76.644" municipality="Elivalmalai" pageId="3" pageNumber="269" specimenCount="1" typeStatus="paratype">collection data same as the holotype</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="03D3409EFFB2FFEBFF47F895FDEEFE87" lastPageId="4" lastPageNumber="270" pageId="3" pageNumber="269" type="etymology">
<paragraph id="4B761315FFB2FFECFF47F895FBE5F816" blockId="3.[151,1437,1656,2042]" pageId="3" pageNumber="269">
<emphasis id="79BDCF07FFB2FFECFF47F895FE97F8DE" bold="true" box="[199,338,1836,1862]" pageId="3" pageNumber="269">Etymology:</emphasis>
The new species is named after the “Mudugar” indigenous community of Palghat district,
<collectingRegion id="890DDDF7FFB2FFECFAD2F895FA58F8DE" box="[1362,1437,1836,1862]" country="India" name="Kerala" pageId="3" pageNumber="269">Kerala</collectingRegion>
who speak Muduga language (
<bibRefCitation id="2F586EE4FFB2FFECFE7BF8E9FD50F8F3" author="Menon, M. T." box="[507,661,1872,1899]" pageId="3" pageNumber="269" pagination="1 - 347" refId="ref6370" refString="Menon, M. T. (1996) The encyclopedia of Dravidian tribes. Vol. II. Thiruvananthapuram, Kerala. International School of Dravidian Linguistics, 1996, 1 - 347." type="journal article" year="1996">Menon, 1996</bibRefCitation>
). The species epithet is used as an invariable noun in apposition to the generic name. The suggested common name is Muduga mountain leaping frog.
</paragraph>
<paragraph id="4B761315FFB2FFEBFF47F82EFDEEFE87" blockId="3.[151,1437,1656,2042]" lastBlockId="4.[151,1437,153,575]" lastPageId="4" lastPageNumber="270" pageId="3" pageNumber="269">
<emphasis id="79BDCF07FFB2FFECFF47F82EFE63F82A" bold="true" box="[199,422,1943,1970]" pageId="3" pageNumber="269">Lineage diagnosis:</emphasis>
<taxonomicName id="8CC96896FFB2FFECFE2EF821FD4DF82A" authority="Dinesh &amp; Vijayakumar &amp; Ramesh &amp; Jayarajan &amp; Chandramouli &amp; Shanker, 2020" authorityName="Dinesh &amp; Vijayakumar &amp; Ramesh &amp; Jayarajan &amp; Chandramouli &amp; Shanker" authorityYear="2020" box="[430,648,1944,1970]" class="Insecta" family="Haglotettigoniidae" genus="Walkerana" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="269" phylum="Arthropoda" rank="species" species="muduga" status="sp. nov.">
<emphasis id="79BDCF07FFB2FFECFE2EF821FD4DF82A" box="[430,648,1944,1970]" italics="true" pageId="3" pageNumber="269">Walkerana muduga</emphasis>
</taxonomicName>
<emphasis id="79BDCF07FFB2FFECFD10F821FD2FF82A" bold="true" box="[656,746,1944,1970]" pageId="3" pageNumber="269">
<taxonomicNameLabel id="628E727CFFB2FFECFD10F821FD2FF82A" box="[656,746,1944,1970]" pageId="3" pageNumber="269" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be diagnosed phylogenetically as a member of the
<taxonomicName id="8CC96896FFB2FFECFAE8F821FF20F84E" class="Insecta" family="Haglotettigoniidae" genus="Walkerana" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="269" phylum="Arthropoda" rank="genus">
<emphasis id="79BDCF07FFB2FFECFAE8F821FF20F84E" italics="true" pageId="3" pageNumber="269">Walkerana</emphasis>
</taxonomicName>
clade (
<figureCitation id="D3F20F90FFB2FFECFEBEF805FE46F84E" box="[318,387,1980,2006]" captionStart="FIGURE 2" captionStartId="3.[151,250,1081,1106]" captionTargetBox="[159,1428,182,1056]" captionTargetId="figure@3.[151,1436,174,1062]" captionTargetPageId="3" captionText="FIGURE 2. Bayesian tree of the genus Walkerana with posterior probability values indicated on the respective nodes." figureDoi="http://doi.org/10.5281/zenodo.3632642" httpUri="https://zenodo.org/record/3632642/files/figure.png" pageId="3" pageNumber="269">Fig. 2</figureCitation>
), and exhibits 5.7% genetic divergence from
<taxonomicName id="8CC96896FFB2FFECFC00F804FBE8F84E" baseAuthorityName="Boulenger" baseAuthorityYear="1882" box="[896,1069,1980,2006]" class="Insecta" family="Haglotettigoniidae" genus="Walkerana" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="269" phylum="Arthropoda" rank="species" species="leptodactyla">
<emphasis id="79BDCF07FFB2FFECFC00F804FBE8F84E" box="[896,1069,1980,2006]" italics="true" pageId="3" pageNumber="269">W. leptodactyla</emphasis>
</taxonomicName>
, 7.5% divergence from
<taxonomicName id="8CC96896FFB2FFECFABDF804FF3EF862" baseAuthorityName="Boulenger" baseAuthorityYear="1882" class="Amphibia" family="Ranixalidae" genus="Indirana" kingdom="Animalia" order="Anura" pageId="3" pageNumber="269" phylum="Chordata" rank="species" species="phrynoderma">
<emphasis id="79BDCF07FFB2FFECFABDF804FF3EF862" italics="true" pageId="3" pageNumber="269">W. phrynoderma</emphasis>
</taxonomicName>
and 11.7% divergence from
<taxonomicName id="8CC96896FFB2FFECFDC3F858FD18F862" baseAuthorityName="Günther" baseAuthorityYear="1876" box="[579,733,2016,2042]" class="Amphibia" family="Ranixalidae" genus="Indirana" kingdom="Animalia" order="Anura" pageId="3" pageNumber="269" phylum="Chordata" rank="species" species="diplosticta">
<emphasis id="79BDCF07FFB2FFECFDC3F858FD18F862" box="[579,733,2016,2042]" italics="true" pageId="3" pageNumber="269">W. diplosticta</emphasis>
</taxonomicName>
at the 16S rRNA fragment. The species is highly divergent in morphology from all known species (see comparison section). Phylogenetic analysis also shows another undescribed lineage from the adjacent massif (north of Elival Mala), exhibiting high genetic divergence of 3.8 % for 16S rRNA (CESF 1554). However, due to the availability of only a single poorly preserved specimen (CESF 1554), we defer the description of this lineage.
</paragraph>
</subSubSection>
<subSubSection id="03D3409EFFB5FFEAFF47FE91FB89FA80" lastPageId="5" lastPageNumber="271" pageId="4" pageNumber="270" type="description">
<paragraph id="4B761315FFB5FFEBFF47FE91FEDCFDA7" blockId="4.[151,1437,153,575]" pageId="4" pageNumber="270">
<emphasis id="79BDCF07FFB5FFEBFF47FE91FE22FEDB" bold="true" box="[199,487,296,323]" pageId="4" pageNumber="270">
Description of
<typeStatus id="9472ADB7FFB5FFEBFEFAFE90FE22FEDB" box="[378,487,297,323]" pageId="4" pageNumber="270" type="holotype">Holotype</typeStatus>
</emphasis>
ZSI/WRC/A/2211
<emphasis id="79BDCF07FFB5FFEBFD44FE91FC19FEDB" bold="true" box="[708,988,296,323]" pageId="4" pageNumber="270">
(CESF 1607) (
<figureCitation id="D3F20F90FFB5FFEBFCEEFE91FC70FEDB" box="[878,949,296,323]" captionStart="FIGURE 2" captionStartId="3.[151,250,1081,1106]" captionTargetBox="[159,1428,182,1056]" captionTargetId="figure@3.[151,1436,174,1062]" captionTargetPageId="3" captionText="FIGURE 2. Bayesian tree of the genus Walkerana with posterior probability values indicated on the respective nodes." figureDoi="http://doi.org/10.5281/zenodo.3632642" httpUri="https://zenodo.org/record/3632642/files/figure.png" pageId="4" pageNumber="270">Fig. 2</figureCitation>
,
<figureCitation id="D3F20F90FFB5FFEBFC44FE91FC16FEDB" box="[964,979,296,323]" captionStart="FIGURE 3" captionStartId="4.[151,250,1310,1335]" captionTargetBox="[151,1435,589,1291]" captionTargetId="figure@4.[151,1437,589,1291]" captionTargetPageId="4" captionText="FIGURE 3. Preserved holotype of Walkerana muduga sp. nov. (A) dorsal view. (B) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3632644" httpUri="https://zenodo.org/record/3632644/files/figure.png" pageId="4" pageNumber="270">3</figureCitation>
)
</emphasis>
. A medium sized species of
<taxonomicName id="8CC96896FFB5FFEBFAA4FE90FA59FEDB" box="[1316,1436,297,323]" class="Insecta" family="Haglotettigoniidae" genus="Walkerana" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="270" phylum="Arthropoda" rank="genus">
<emphasis id="79BDCF07FFB5FFEBFAA4FE90FA59FEDB" box="[1316,1436,297,323]" italics="true" pageId="4" pageNumber="270">Walkerana</emphasis>
</taxonomicName>
(SVL =
<quantity id="8C31BEF0FFB5FFEBFF72FEF4FE92FEFF" box="[242,343,333,359]" metricMagnitude="-2" metricUnit="m" metricValue="2.79" pageId="4" pageNumber="270" unit="mm" value="27.9">27.9 mm</quantity>
) with squat, raised body; head width (HW =
<quantity id="8C31BEF0FFB5FFEBFCD1FEF4FC70FEFF" box="[849,949,333,359]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="4" pageNumber="270" unit="mm" value="11.3">11.3 mm</quantity>
) sub equal to head length (HL =
<quantity id="8C31BEF0FFB5FFEBFAA8FEF4FA55FEFF" box="[1320,1424,333,359]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="4" pageNumber="270" unit="mm" value="10.3">10.3 mm</quantity>
); snout bluntly pointed (ES =
<quantity id="8C31BEF0FFB5FFEBFE56FEC8FDE8FE13" box="[470,557,369,395]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="4" pageNumber="270" unit="mm" value="4.7">4.7 mm</quantity>
) in both dorsal and ventral view and more than 1.2 times of eye diameter (ED =
<quantity id="8C31BEF0FFB5FFEBFF2EFE2CFECCFE37" box="[174,265,405,431]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="4" pageNumber="270" unit="mm" value="3.9">3.9 mm</quantity>
); canthus rostralis angular, loreal region concave, inter orbital space flat (IO = 3.0 mm) and sub equal to upper eye lid (UEW =
<quantity id="8C31BEF0FFB5FFEBFE1CFE00FE32FE4B" box="[412,503,441,467]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="4" pageNumber="270" unit="mm" value="2.4">2.4 mm</quantity>
); nostrils oval, nearer to tip of snout than to eye; eyes relatively smaller (EL/HL = 0.37), protruding out of the head; tympanum (TYH =
<quantity id="8C31BEF0FFB5FFEBFD72FE64FC8CFE6F" box="[754,841,477,503]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="4" pageNumber="270" unit="mm" value="2.2">2.2 mm</quantity>
) distinct below the supratympanic fold and 1.7 times of eye diameter; minute vomerine ridges present with 2 to 3 spinular teeth; tongue bifid with a raised papilla; symphysial knob weak.
</paragraph>
<caption id="1FB6439DFFB5FFEBFF17FAA7FBB3FAAF" ID-DOI="http://doi.org/10.5281/zenodo.3632644" ID-Zenodo-Dep="3632644" box="[151,1142,1310,1335]" httpUri="https://zenodo.org/record/3632644/files/figure.png" pageId="4" pageNumber="270" startId="4.[151,250,1310,1335]" targetBox="[151,1435,589,1291]" targetPageId="4">
<paragraph id="4B761315FFB5FFEBFF17FAA7FBB3FAAF" blockId="4.[151,1142,1310,1335]" box="[151,1142,1310,1335]" pageId="4" pageNumber="270">
<emphasis id="79BDCF07FFB5FFEBFF17FAA7FED6FAAF" bold="true" box="[151,275,1310,1335]" pageId="4" pageNumber="270">FIGURE 3.</emphasis>
Preserved holotype of
<taxonomicName id="8CC96896FFB5FFEBFE7EFAA6FD06FAAF" authority="Dinesh &amp; Vijayakumar &amp; Ramesh &amp; Jayarajan &amp; Chandramouli &amp; Shanker, 2020" authorityName="Dinesh &amp; Vijayakumar &amp; Ramesh &amp; Jayarajan &amp; Chandramouli &amp; Shanker" authorityYear="2020" box="[510,707,1311,1335]" class="Insecta" family="Haglotettigoniidae" genus="Walkerana" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="270" phylum="Arthropoda" rank="species" species="muduga" status="sp. nov.">
<emphasis id="79BDCF07FFB5FFEBFE7EFAA6FD06FAAF" box="[510,707,1311,1335]" italics="true" pageId="4" pageNumber="270">Walkerana muduga</emphasis>
</taxonomicName>
<emphasis id="79BDCF07FFB5FFEBFD49FAA6FCDFFAAF" bold="true" box="[713,794,1311,1335]" pageId="4" pageNumber="270">
<taxonomicNameLabel id="628E727CFFB5FFEBFD49FAA6FCDFFAAF" box="[713,794,1311,1335]" pageId="4" pageNumber="270" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) dorsal view. (B) ventral view.
</paragraph>
</caption>
<paragraph id="4B761315FFB5FFEBFF47FADFFBC6F997" blockId="4.[151,1437,1381,2020]" pageId="4" pageNumber="270">
Fore arm slender (UAL =
<quantity id="8C31BEF0FFB5FFEBFE66FADFFDF9FA18" box="[486,572,1382,1408]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="4" pageNumber="270" unit="mm" value="4.7">4.7 mm</quantity>
) and sub equal to hand (PAL =
<quantity id="8C31BEF0FFB5FFEBFC15FADCFC2BFA18" box="[917,1006,1381,1408]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="4" pageNumber="270" unit="mm" value="7.6">7.6 mm</quantity>
); fingers short and thin (dermal fringes distinct in the ventral view on the finger 2, 3 and 4); first finger shorter than second (f1 =
<quantity id="8C31BEF0FFB5FFEBFBF8FA33FB08FA3C" box="[1144,1229,1418,1444]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="4" pageNumber="270" unit="mm" value="3.7">3.7 mm</quantity>
and f2 =
<quantity id="8C31BEF0FFB5FFEBFAB6FA33FA55FA3C" box="[1334,1424,1418,1444]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="4" pageNumber="270" unit="mm" value="4.2">4.2 mm</quantity>
); finger tips with truncated enlarged discs with minute dorsoterminal grooves; fingers free without webbing between fingers; rounded subarticular tubercles moderate (one each on finger 1 and 2, and two each on finger 3 and 4); prepollex indistinct and palmar tubercle distinct, supernumerary tubercles absent.
</paragraph>
<paragraph id="4B761315FFB5FFEBFF47F9A0FCC9F97F" blockId="4.[151,1437,1381,2020]" pageId="4" pageNumber="270">
Hind limbs moderately long, strongly overlap when folded at right angles to the body and tibio-tarsal articulation reaches beyond the snout tip; femur length less than tibia length (FEL =
<quantity id="8C31BEF0FFB5FFEBFC69F987FB8AF9C0" box="[1001,1103,1598,1624]" metricMagnitude="-2" metricUnit="m" metricValue="1.49" pageId="4" pageNumber="270" unit="mm" value="14.9">14.9 mm</quantity>
; TBL =
<quantity id="8C31BEF0FFB5FFEBFB29F987FAD5F9C0" box="[1193,1296,1598,1624]" metricMagnitude="-2" metricUnit="m" metricValue="1.8199999999999998" pageId="4" pageNumber="270" unit="mm" value="18.2">18.2 mm</quantity>
); foot length is 1.8 times more than the tarsus length (FOL =
<quantity id="8C31BEF0FFB5FFEBFD36F9DBFCD9F9E4" box="[694,796,1634,1660]" metricMagnitude="-2" metricUnit="m" metricValue="1.49" pageId="4" pageNumber="270" unit="mm" value="14.9">14.9 mm</quantity>
, TAR =
<quantity id="8C31BEF0FFB5FFEBFCFDF9DBFC12F9E4" box="[893,983,1634,1660]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="4" pageNumber="270" unit="mm" value="8.1">8.1 mm</quantity>
), relative toe length I&lt;II&lt;V&lt;III&lt;IV; tips of toes with truncated enlarged discs having minute dorsoterminal grooves; webbing between toes reduced, webbing just touches the last subarticular tubercle of toe IV and V; inner metatarsal tubercle moderate, bulbous; outer metatarsal tubercle and supernumerary tubercles absent.
</paragraph>
<paragraph id="4B761315FFB5FFEBFF47F948FE0AF8CB" blockId="4.[151,1437,1381,2020]" pageId="4" pageNumber="270">Overall skin on the dorsum smooth glandular with scattered small ridged longitudinal glandular folds, folds are discontinuous without any pattern; supratympanic fold distinct; on the ventral side of the body, region of throat smooth and belly glandular.</paragraph>
<paragraph id="4B761315FFB5FFEBFF47F8E7FB2DF87B" blockId="4.[151,1437,1381,2020]" pageId="4" pageNumber="270">In life, colour on the dorsum light fleshy brown, base of the raised ridges dark brown and the inter orbital space with light brown blotches; lower lip barred, dark brown streak from the tip of snout till the end of the supratympanic fold in front of shoulders; side of the body uniform light brown without any markings; fore and hind limbs barred; ventrally, region of throat yellowish brown, belly whitish grey and region of thighs flesh-coloured.</paragraph>
<paragraph id="4B761315FFB4FFEAFF47FF20FE31FEDB" blockId="5.[151,1437,153,323]" pageId="5" pageNumber="271">In preservative, overall colour on the dorsum grayish brown on the dorsum between the inter-orbital space, horizontal light black band connecting the raised eye balls; upper and lower lips barred; a thin blackish band starting from the tip of snout, through the upper edge of canthus rostralis on either side till the posterior end of the supratympanic fold; sides of the body light gray, raised longitudinal glandular folds dark grey; fore and hind limbs barred; ventrally uniform cream white.</paragraph>
<caption id="1FB6439DFFB4FFEAFF17FB9EFB20FBD8" ID-DOI="http://doi.org/10.5281/zenodo.3632646" ID-Zenodo-Dep="3632646" box="[151,1253,1063,1088]" httpUri="https://zenodo.org/record/3632646/files/figure.png" pageId="5" pageNumber="271" startId="5.[151,250,1063,1088]" targetBox="[170,1416,365,1001]" targetPageId="5">
<paragraph id="4B761315FFB4FFEAFF17FB9EFB20FBD8" blockId="5.[151,1253,1063,1088]" box="[151,1253,1063,1088]" pageId="5" pageNumber="271">
<emphasis id="79BDCF07FFB4FFEAFF17FB9EFED6FBD8" bold="true" box="[151,275,1063,1088]" pageId="5" pageNumber="271">FIGURE 4.</emphasis>
Holotype of
<taxonomicName id="8CC96896FFB4FFEAFE19FB9EFD9AFBA7" authority="Dinesh &amp; Vijayakumar &amp; Ramesh &amp; Jayarajan &amp; Chandramouli &amp; Shanker, 2020" authorityName="Dinesh &amp; Vijayakumar &amp; Ramesh &amp; Jayarajan &amp; Chandramouli &amp; Shanker" authorityYear="2020" box="[409,607,1063,1087]" class="Insecta" family="Haglotettigoniidae" genus="Walkerana" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="271" phylum="Arthropoda" rank="species" species="muduga" status="sp. nov.">
<emphasis id="79BDCF07FFB4FFEAFE19FB9EFD9AFBA7" box="[409,607,1063,1087]" italics="true" pageId="5" pageNumber="271">Walkerana muduga</emphasis>
</taxonomicName>
<emphasis id="79BDCF07FFB4FFEAFDE5FB91FD70FBD8" bold="true" box="[613,693,1064,1088]" pageId="5" pageNumber="271">
<taxonomicNameLabel id="628E727CFFB4FFEAFDE5FB91FD70FBD8" box="[613,693,1064,1088]" pageId="5" pageNumber="271" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in life (not to scale). (A) dorsal view. (B) ventral view.
</paragraph>
</caption>
<paragraph id="4B761315FFB4FFEAFF47FBD4FD6FFB34" blockId="5.[151,1436,1133,1988]" pageId="5" pageNumber="271">
<emphasis id="79BDCF07FFB4FFEAFF47FBD4FDE1FB10" bold="true" box="[199,548,1133,1160]" pageId="5" pageNumber="271">Secondary sexual characters:</emphasis>
No visible external sexual dimorphism was noticeable except for the amplected pairs where males are smaller than the females.
</paragraph>
<paragraph id="4B761315FFB4FFEAFF47FB0CFB89FA80" blockId="5.[151,1436,1133,1988]" pageId="5" pageNumber="271">
<emphasis id="79BDCF07FFB4FFEAFF47FB0CFC91FB48" bold="true" box="[199,852,1205,1232]" pageId="5" pageNumber="271">
Additional information from
<typeStatus id="9472ADB7FFB4FFEAFDA9FB0FFD5AFB48" box="[553,671,1206,1232]" pageId="5" pageNumber="271" type="paratype">paratypes</typeStatus>
and variations
</emphasis>
: Morphological data are given in
<tableCitation id="064B26AEFFB4FFEAFB57FB0FFAE9FB48" box="[1239,1324,1206,1232]" captionStart="TABLE 1" captionStartId="6.[151,239,154,178]" captionTargetPageId="6" captionText="TABLE 1. Morphometric data (in mm) for the type series and reference collections of Walkerana muduga sp. nov." httpUri="http://table.plazi.org/id/1FB6439DFFB7FFE9FF17FF23FAA5FF2A" pageId="5" pageNumber="271" tableUuid="1FB6439DFFB7FFE9FF17FF23FAA5FF2A">Table 1</tableCitation>
.
<materialsCitation id="FBA11948FFB4FFEAFABAFB0FFE2BFA80" ID-GBIF-Occurrence="2561484856" collectionCode="ZSI, WRC, SVL" pageId="5" pageNumber="271" specimenCode="CESF 1606, CESF 1608" specimenCount="1" typeStatus="paratype">
<typeStatus id="9472ADB7FFB4FFEAFABAFB0FFA59FB48" box="[1338,1436,1206,1232]" pageId="5" pageNumber="271" type="paratype">Paratype</typeStatus>
<collectionCode id="2DD88BD0FFB4FFEAFF17FB63FF05FB6C" box="[151,192,1242,1268]" pageId="5" pageNumber="271">ZSI</collectionCode>
/
<collectionCode id="2DD88BD0FFB4FFEAFF49FB63FECCFB6C" box="[201,265,1242,1268]" country="Japan" httpUri="http://grbio.org/cool/i1ct-fvht" name="Wildlife Research Center of Kyoto University" pageId="5" pageNumber="271">WRC</collectionCode>
/A/2212 (
<specimenCode id="1B6FBB6EFFB4FFEAFEF2FB63FE3DFB6D" box="[370,504,1242,1269]" collectionCode="CESF" pageId="5" pageNumber="271">CESF 1606</specimenCode>
), an adult female (
<collectionCode id="2DD88BD0FFB4FFEAFD41FB63FD32FB6C" box="[705,759,1242,1268]" pageId="5" pageNumber="271">SVL</collectionCode>
<quantity id="8C31BEF0FFB4FFEAFD7DFB63FCA5FB6C" box="[765,864,1242,1268]" metricMagnitude="-2" metricUnit="m" metricValue="4.49" pageId="5" pageNumber="271" unit="mm" value="44.9">44.9 mm</quantity>
) and other reference collections (
<specimenCode id="1B6FBB6EFFB4FFEAFB50FB63FA90FB6C" box="[1232,1365,1242,1268]" collectionCode="CESF" pageId="5" pageNumber="271">CESF 1608</specimenCode>
, male,
<collectionCode id="2DD88BD0FFB4FFEAFF17FB47FF0EFA80" box="[151,203,1278,1304]" pageId="5" pageNumber="271">SVL</collectionCode>
<quantity id="8C31BEF0FFB4FFEAFF52FB47FEF2FA80" box="[210,311,1278,1304]" metricMagnitude="-2" metricUnit="m" metricValue="2.27" pageId="5" pageNumber="271" unit="mm" value="22.7">22.7 mm</quantity>
) were similar to
</materialsCitation>
<typeStatus id="9472ADB7FFB4FFEAFE75FB47FD92FA80" box="[501,599,1278,1304]" pageId="5" pageNumber="271" type="holotype">holotype</typeStatus>
in all the external morphological characters.
</paragraph>
</subSubSection>
<subSubSection id="03D3409EFFB4FFEAFF47FA98FD95F85C" pageId="5" pageNumber="271" type="distribution">
<paragraph id="4B761315FFB4FFEAFF47FA98FE5AFAF8" blockId="5.[151,1436,1133,1988]" pageId="5" pageNumber="271">
<emphasis id="79BDCF07FFB4FFEAFF47FA98FE9CFAA4" bold="true" box="[199,345,1313,1340]" pageId="5" pageNumber="271">Distribution</emphasis>
:
<taxonomicName id="8CC96896FFB4FFEAFEEAFA9BFD81FAA4" authority="Dinesh &amp; Vijayakumar &amp; Ramesh &amp; Jayarajan &amp; Chandramouli &amp; Shanker, 2020" authorityName="Dinesh &amp; Vijayakumar &amp; Ramesh &amp; Jayarajan &amp; Chandramouli &amp; Shanker" authorityYear="2020" box="[362,580,1314,1340]" class="Insecta" family="Haglotettigoniidae" genus="Walkerana" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="271" phylum="Arthropoda" rank="species" species="muduga" status="sp. nov.">
<emphasis id="79BDCF07FFB4FFEAFEEAFA9BFD81FAA4" box="[362,580,1314,1340]" italics="true" pageId="5" pageNumber="271">Walkerana muduga</emphasis>
</taxonomicName>
<emphasis id="79BDCF07FFB4FFEAFDCCFA9BFD63FAA4" bold="true" box="[588,678,1314,1340]" pageId="5" pageNumber="271">
<taxonomicNameLabel id="628E727CFFB4FFEAFDCCFA9BFD63FAA4" box="[588,678,1314,1340]" pageId="5" pageNumber="271" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is known from the Elivalmalai range, north of the Palghat Gap and south of Nilgiri Massif.
</paragraph>
<paragraph id="4B761315FFB4FFEAFF47FAD0FD86F888" blockId="5.[151,1436,1133,1988]" pageId="5" pageNumber="271">
<emphasis id="79BDCF07FFB4FFEAFF47FAD0FEAAFA1C" bold="true" box="[199,367,1385,1412]" pageId="5" pageNumber="271">Comparisons:</emphasis>
All the described species of
<taxonomicName id="8CC96896FFB4FFEAFD2FFAD3FCE2FA1C" box="[687,807,1386,1412]" class="Insecta" family="Haglotettigoniidae" genus="Walkerana" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="271" phylum="Arthropoda" rank="genus">
<emphasis id="79BDCF07FFB4FFEAFD2FFAD3FCE2FA1C" box="[687,807,1386,1412]" italics="true" pageId="5" pageNumber="271">Walkerana</emphasis>
</taxonomicName>
are known to occur in the high elevation ranges south of the Palghat Gap. There are no congeners for
<taxonomicName id="8CC96896FFB4FFEAFD0DFA37FCA2FA30" authority="Dinesh &amp; Vijayakumar &amp; Ramesh &amp; Jayarajan &amp; Chandramouli &amp; Shanker, 2020" authorityName="Dinesh &amp; Vijayakumar &amp; Ramesh &amp; Jayarajan &amp; Chandramouli &amp; Shanker" authorityYear="2020" box="[653,871,1422,1448]" class="Insecta" family="Haglotettigoniidae" genus="Walkerana" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="271" phylum="Arthropoda" rank="species" species="muduga" status="sp. nov.">
<emphasis id="79BDCF07FFB4FFEAFD0DFA37FCA2FA30" box="[653,871,1422,1448]" italics="true" pageId="5" pageNumber="271">Walkerana muduga</emphasis>
</taxonomicName>
<emphasis id="79BDCF07FFB4FFEAFCEEFA37FC0DFA30" bold="true" box="[878,968,1422,1448]" pageId="5" pageNumber="271">
<taxonomicNameLabel id="628E727CFFB4FFEAFCEEFA37FC0DFA30" box="[878,968,1422,1448]" pageId="5" pageNumber="271" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
north of the Palghat Gap. However, there is a deeply divergent lineage (CESF 1554) (see diagnosis) recovered from an adjacent massif.
<taxonomicName id="8CC96896FFB4FFEAFB42FA0BFA59FA54" authority="Dinesh &amp; Vijayakumar &amp; Ramesh &amp; Jayarajan &amp; Chandramouli &amp; Shanker, 2020" authorityName="Dinesh &amp; Vijayakumar &amp; Ramesh &amp; Jayarajan &amp; Chandramouli &amp; Shanker" authorityYear="2020" box="[1218,1436,1458,1484]" class="Insecta" family="Haglotettigoniidae" genus="Walkerana" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="271" phylum="Arthropoda" rank="species" species="muduga" status="sp. nov.">
<emphasis id="79BDCF07FFB4FFEAFB42FA0BFA59FA54" box="[1218,1436,1458,1484]" italics="true" pageId="5" pageNumber="271">Walkerana muduga</emphasis>
</taxonomicName>
<emphasis id="79BDCF07FFB4FFEAFF17FA6FFF34FA68" bold="true" box="[151,241,1494,1520]" pageId="5" pageNumber="271">
<taxonomicNameLabel id="628E727CFFB4FFEAFF17FA6FFF34FA68" box="[151,241,1494,1520]" pageId="5" pageNumber="271" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from the undescribed
<taxonomicName id="8CC96896FFB4FFEAFDBBFA6FFD76FA68" box="[571,691,1494,1520]" class="Insecta" family="Haglotettigoniidae" genus="Walkerana" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="271" phylum="Arthropoda" rank="genus">
<emphasis id="79BDCF07FFB4FFEAFDBBFA6FFD76FA68" box="[571,691,1494,1520]" italics="true" pageId="5" pageNumber="271">Walkerana</emphasis>
</taxonomicName>
sp. (CESF 1554) in having higher HL/SVL ratio of 0.370 to 0.405 (n=2) (vs. lower HL/SVL ratio of 0.342 (n=1)); higher HW/SVL ratio of 0.397 to 0.406 (n=2) (vs. lower HW/SVL ratio of 0.390 (n=1)); lower TYH/SVL ratio of 0.079 to 0.080 (n=2) (vs. higher TYH/SVL ratio of 0.083 (n=1)); higher IO/SVL ratio of 0.107 to 0.109 (n=2) (vs. lower IO/SVL ratio of 0.097 (n=1)); higher PAL/SVL ratio of 0.257 to 0.271 (n=2) (vs. lower PAL/SVL ratio of 0.234 (n=1)); higher f1/SVL ratio of 0.119 to 0.133 (n=2) (vs. lower f1/SVL ratio of 0.112 (n=1)); higher f2/SVL ratio of 0.131 to 0.151 (n=2) (vs. lower f2/SVL ratio of 0.123 (n=1)); lower FEL/SVL ratio of 0.535 to 0.536 (n=2) (vs. higher FEL/SVL ratio of 0.545 (n=1)); lower TAR/SVL ratio of 0.287 to 0.291 (n=2) (vs. higher TAR/SVL ratio of 0.295 (n=1)); higher FOL/SVL ratio of 0.536 to 0.565 (n=2) (vs. lower FOL/SVL ratio of 0.530 (n=1)).
</paragraph>
<paragraph id="4B761315FFB4FFEAFF47F8A3FD95F85C" blockId="5.[151,1436,1133,1988]" pageId="5" pageNumber="271">
<taxonomicName id="8CC96896FFB4FFEAFF47F8A3FE65F8AC" authority="Dinesh &amp; Vijayakumar &amp; Ramesh &amp; Jayarajan &amp; Chandramouli &amp; Shanker, 2020" authorityName="Dinesh &amp; Vijayakumar &amp; Ramesh &amp; Jayarajan &amp; Chandramouli &amp; Shanker" authorityYear="2020" box="[199,416,1818,1844]" class="Insecta" family="Haglotettigoniidae" genus="Walkerana" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="271" phylum="Arthropoda" rank="species" species="muduga" status="sp. nov.">
<emphasis id="79BDCF07FFB4FFEAFF47F8A3FE65F8AC" box="[199,416,1818,1844]" italics="true" pageId="5" pageNumber="271">Walkerana muduga</emphasis>
</taxonomicName>
<emphasis id="79BDCF07FFB4FFEAFE27F8A3FDC5F8AC" bold="true" box="[423,512,1818,1844]" pageId="5" pageNumber="271">
<taxonomicNameLabel id="628E727CFFB4FFEAFE27F8A3FDC5F8AC" box="[423,512,1818,1844]" pageId="5" pageNumber="271" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from species found south of the Palghat Gap (see key to the species of
<taxonomicName id="8CC96896FFB4FFEAFAE8F8A3FF20F8C0" class="Insecta" family="Haglotettigoniidae" genus="Walkerana" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="271" phylum="Arthropoda" rank="genus">
<emphasis id="79BDCF07FFB4FFEAFAE8F8A3FF20F8C0" italics="true" pageId="5" pageNumber="271">Walkerana</emphasis>
</taxonomicName>
by
<bibRefCitation id="2F586EE4FFB4FFEAFE8CF887FDC6F8C1" author="Dahanukar, N. &amp; Modak, N. &amp; Krutha, K. &amp; Nameer, P. O. &amp; Padhye, A. D. &amp; Molur, S." box="[268,515,1854,1881]" pageId="5" pageNumber="271" pagination="9221 - 9288" refId="ref5597" refString="Dahanukar, N., Modak, N., Krutha, K., Nameer, P. O., Padhye, A. D. &amp; Molur, S. (2016) Leaping frogs (Anura: Ranixalidae) of the Western Ghats of India: An integrated taxonomic review. Journal of Threatened Taxa, 8, 9221 - 9288. https: // doi. org / 10.11609 / jott. 2532.8.10.9221 - 9288" type="journal article" year="2016">
Dahanukar
<emphasis id="79BDCF07FFB4FFEAFE0DF886FE79F8C0" box="[397,444,1854,1880]" italics="true" pageId="5" pageNumber="271">et al</emphasis>
. 2016
</bibRefCitation>
) in having distinct canthus rostralis and smooth skin (vs. indistinct canthus rostralis and warty skin in
<taxonomicName id="8CC96896FFB4FFEAFEE2F8DAFDD9F8E4" baseAuthorityName="Boulenger" baseAuthorityYear="1882" box="[354,540,1890,1916]" class="Amphibia" family="Ranixalidae" genus="Indirana" kingdom="Animalia" order="Anura" pageId="5" pageNumber="271" phylum="Chordata" rank="species" species="phrynoderma">
<emphasis id="79BDCF07FFB4FFEAFEE2F8DAFDD9F8E4" box="[354,540,1890,1916]" italics="true" pageId="5" pageNumber="271">W. phrynoderma</emphasis>
</taxonomicName>
); lacking a distinct pair of black patches on the loin (vs. distinct black patch on the loin in
<taxonomicName id="8CC96896FFB4FFEAFE90F83EFE6CF838" baseAuthorityName="Günther" baseAuthorityYear="1876" box="[272,425,1926,1952]" class="Amphibia" family="Ranixalidae" genus="Indirana" kingdom="Animalia" order="Anura" pageId="5" pageNumber="271" phylum="Chordata" rank="species" species="diplosticta">
<emphasis id="79BDCF07FFB4FFEAFE90F83EFE6CF838" box="[272,425,1926,1952]" italics="true" pageId="5" pageNumber="271">W. diplosticta</emphasis>
</taxonomicName>
) and tympanum relatively smaller than the diameter of eye (vs. tympanum three-fourth of the diameter of eye in
<taxonomicName id="8CC96896FFB4FFEAFE12F812FDFAF85C" baseAuthorityName="Boulenger" baseAuthorityYear="1882" box="[402,575,1962,1988]" class="Insecta" family="Haglotettigoniidae" genus="Walkerana" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="271" phylum="Arthropoda" rank="species" species="leptodactyla">
<emphasis id="79BDCF07FFB4FFEAFE12F812FDFAF85C" box="[402,575,1962,1988]" italics="true" pageId="5" pageNumber="271">W. leptodactyla</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>