381 lines
49 KiB
XML
381 lines
49 KiB
XML
<document id="D5D6D4E89F55F749F7062A400487D201" ID-DOI="10.5281/zenodo.5172580" ID-ISSN="1942-1354" ID-Zenodo-Dep="5172580" ID-ZooBank="F8317A32-DFD7-42EA-8BC6-DCB5A3659F6C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628270718788" checkinUser="felipe" docAuthor="Neita-Moreno, Jhon César, Alexander, Recursos Biológicos, Agustín, San, de, Villa, Leyva, Boyacá, Ratcliffe, Brett C., Collections, Systematics Research, of, University, Museum, Nebraska State, Hall, Nebraska, of, University, Nebraska, Lincoln, Ne & Usa" docDate="2017" docId="A24087A73966042AFF7FFF51FAD2FEEC" docLanguage="en" docName="InsectaMundi.0547.1-36.pdf" docOrigin="Insecta Mundi 2017 (547)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Tomarus bidentulus" docType="treatment" docVersion="4" lastPageNumber="5" masterDocId="5E79FFDF3963042CFFBFFF93FF92FF96" masterDocTitle="The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay" masterLastPageNumber="36" masterPageNumber="1" pageNumber="4" updateTime="1699038600670" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="660FEEE5D9E6693DCB92C26361194B4F">The genus Tomarus Erichson (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) in Argentina, Chile, and Uruguay</mods:title>
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<treatment id="A24087A73966042AFF7FFF51FAD2FEEC" ID-DOI="http://doi.org/10.5281/zenodo.5185984" ID-Zenodo-Dep="5185984" LSID="urn:lsid:plazi:treatment:A24087A73966042AFF7FFF51FAD2FEEC" httpUri="http://treatment.plazi.org/id/A24087A73966042AFF7FFF51FAD2FEEC" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4">
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<taxonomicName id="EDE94D3239660429FF7FFF51FD59FF4F" authority="(Fairmaire, 1892)" baseAuthorityName="Fairmaire" baseAuthorityYear="1892" box="[192,715,193,218]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="bidentulus">
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<emphasis id="189DEAA339660429FF7FFF51FE47FF4C" bold="true" box="[192,469,193,218]" italics="true" pageId="5" pageNumber="4">Tomarus bidentulus</emphasis>
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(
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<bibRefCitation id="4E784B4039660429FE59FF52FD50FF4F" author="Fairmaire, L." box="[486,706,193,217]" pageId="5" pageNumber="4" pagination="242 - 253" refId="ref14326" refString="Fairmaire, L. 1892. Descriptions de quelques Coleopteres argentins. Annales de la Societe Entomologique de Belgique 36: 242 - 253." type="journal article" year="1892">Fairmaire, 1892</bibRefCitation>
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(
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<figureCitation id="B2D22A3439660429FF76FF71FE8DFF6C" box="[201,287,226,250]" captionStart="Figure 1" captionStartId="19.[192,276,1846,1868]" captionTargetBox="[207,1426,292,1810]" captionTargetId="figure-64@19.[192,1441,192,1828]" captionTargetPageId="19" captionText="Figure 1. Habitus of Tomarus spp. a) Tomarus bidentulus (Fairmaire) dorsal view. b) Tomarus burmeisteri (Steinheil) dorsal view. c) Tomarus gyas Erichson dorsal view. d) Tomarus rosettae (Endrödi) dorsal view. e) Tomarus roigjunenti n. sp. dorsal view, holotype. f) Tomarus roigjunenti n. sp. dorsal view, allotype." figureDoi="http://doi.org/10.5281/zenodo.5172584" httpUri="https://zenodo.org/record/5172584/files/figure.png" pageId="5" pageNumber="4">Fig. 1a</figureCitation>
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,
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<figureCitation id="B2D22A3439660429FE92FF71FEFEFF6C" box="[301,364,226,250]" captionStart="Figure 2" captionStartId="20.[192,276,1817,1839]" captionTargetBox="[229,1331,230,1795]" captionTargetId="figure-18@20.[192,1440,192,1807]" captionTargetPageId="20" captionText="Figure 2. Habitus of Tomarus spp. a) Tomarus rosettae (Endrödi) dorsal view. b) Tomarus spinipenis n. sp. dorsal view, holotype. c) Tomarus spinipenis dorsal view, allotype. d) Tomarus burmeisteri (Steinheil) dorsal view. e) Tomarus villosus (Burmeister) ventral view. f) Tomarus gyas ventral view. g) Tomarus bidentulus (Fairmaire) abdomen segments, ventral view (female). h) Tomarus bidentulus abdomen segments, ventral view (male)." figureDoi="http://doi.org/10.5281/zenodo.5172588" httpUri="https://zenodo.org/record/5172588/files/figure.png" pageId="5" pageNumber="4">
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–h
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<figureCitation id="B2D22A3439660429FEC7FF71FE05FF6C" box="[376,407,226,250]" captionStart="Figure 3" captionStartId="21.[192,276,1727,1749]" captionTargetBox="[197,1361,257,1710]" captionTargetId="figure-124@21.[197,1375,192,1717]" captionTargetPageId="21" captionText="Figure 3. Types of Tomarus spp. a) T. bidentulus female holotype, dorsal view. b) T. burmeisteri male holotype, dorsal view. c) T. burmeisteri male holotype, lateral view. d) T. burmeisteri male holotype parameres, caudal view. e) T. burmeisteri male holotype parameres (lateral view). f) Label for T. burmeisteri holotype. g) T. gyas male holotype, dorsal view and labels and parameres, caudal view). h) T. maimon holotype, dorsal view. i) T. rosettae male holotype, dorsal view. j) T. villosus male holotype, dorsal view. k) T. villosus male holotype, dorsal view of head. l) T. villosus male holotype, parameres, caudal view. m) Label for T. villosus holotype." figureDoi="http://doi.org/10.5281/zenodo.5172591" httpUri="https://zenodo.org/record/5172591/files/figure.png" pageId="5" pageNumber="4">3a</figureCitation>
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<figureCitation id="B2D22A3439660429FE1BFF71FE56FF6C" box="[420,452,226,250]" captionStart="Figure 4" captionStartId="22.[192,276,1848,1870]" captionTargetBox="[216,1440,216,1825]" captionTargetId="figure-63@22.[192,1442,192,1842]" captionTargetPageId="22" captionText="Figure 4. Head, dorsal view of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172595" httpUri="https://zenodo.org/record/5172595/files/figure.png" pageId="5" pageNumber="4">4a</figureCitation>
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<figureCitation id="B2D22A3439660429FE6EFF72FE63FF6F" box="[465,497,225,249]" captionStart="Figure 5" captionStartId="23.[192,276,1853,1875]" captionTargetBox="[215,1440,192,1823]" captionTargetId="figure-63@23.[192,1443,192,1843]" captionTargetPageId="23" captionText="Figure 5. Maxilla, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172599" httpUri="https://zenodo.org/record/5172599/files/figure.png" pageId="5" pageNumber="4">5a</figureCitation>
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,
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<figureCitation id="B2D22A3439660429FE42FF72FDA9FF6F" box="[509,571,225,249]" captionStart="Figure 6" captionStartId="24.[192,276,1739,1761]" captionTargetBox="[211,1436,218,1720]" captionTargetId="figure-63@24.[192,1440,192,1730]" captionTargetPageId="24" captionText="Figure 6. Mandibles, ventral view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172601" httpUri="https://zenodo.org/record/5172601/files/figure.png" pageId="5" pageNumber="4">6a–b</figureCitation>
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,
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<figureCitation id="B2D22A3439660429FDF8FF71FDF5FF6F" box="[583,615,226,249]" captionStart="Figure 7" captionStartId="25.[192,276,1683,1705]" captionTargetBox="[231,1392,238,1664]" captionTargetId="figure-63@25.[192,1440,192,1671]" captionTargetPageId="25" captionText="Figure 7. Labium, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172605" httpUri="https://zenodo.org/record/5172605/files/figure.png" pageId="5" pageNumber="4">7a</figureCitation>
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,
|
||
<figureCitation id="B2D22A3439660429FDCBFF71FD01FF6C" box="[628,659,226,250]" captionStart="Figure 8" captionStartId="26.[192,276,1841,1863]" captionTargetBox="[275,1367,208,1829]" captionTargetId="figure-18@26.[269,1376,192,1840]" captionTargetPageId="26" captionText="Figure 8. Pronotum, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172607" httpUri="https://zenodo.org/record/5172607/files/figure.png" pageId="5" pageNumber="4">8a</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439660429FD1FFF71FD7FFF6C" box="[672,749,226,250]" captionStart="Figure 10" captionStartId="28.[192,275,1624,1646]" captionTargetBox="[220,1411,210,1601]" captionTargetId="figure-65@28.[192,1440,192,1613]" captionTargetPageId="28" captionText="Figure 10. Meso- and metatibia, dorsal view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172614" httpUri="https://zenodo.org/record/5172614/files/figure.png" pageId="5" pageNumber="4">10a–b</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439660429FD46FF71FCBAFF6C" box="[761,808,226,250]" captionStart="Figure 11" captionStartId="29.[192,276,1835,1857]" captionTargetBox="[244,1411,241,1815]" captionTargetId="figure-63@29.[192,1442,192,1821]" captionTargetPageId="29" captionText="Figure 11. Parameres, caudal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172618" httpUri="https://zenodo.org/record/5172618/files/figure.png" pageId="5" pageNumber="4">11a</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439660429FC8AFF71FCF1FF6C" box="[821,867,226,250]" captionStart="Figure 12" captionStartId="30.[192,274,1776,1798]" captionTargetBox="[210,1434,220,1735]" captionTargetId="figure-61@30.[192,1440,192,1769]" captionTargetPageId="30" captionText="Figure 12. Parameres, lateral view, of: a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172620" httpUri="https://zenodo.org/record/5172620/files/figure.png" pageId="5" pageNumber="4">12a</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439660429FCCFFF71FC0CFF6C" box="[880,926,226,250]" captionStart="Figure 13" captionStartId="31.[192,275,1823,1845]" captionTargetBox="[238,1403,236,1798]" captionTargetId="figure-63@31.[192,1440,192,1807]" captionTargetPageId="31" captionText="Figure 13. Parameres, anterior view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172622" httpUri="https://zenodo.org/record/5172622/files/figure.png" pageId="5" pageNumber="4">13a</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439660429FC14FF71FC48FF6C" box="[939,986,226,250]" captionStart="Figure 14" captionStartId="32.[192,276,1832,1854]" captionTargetBox="[358,1267,208,1798]" captionTargetId="figure-70@32.[349,1298,192,1821]" captionTargetPageId="32" captionText="Figure 14. Gonocoxite, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (Gx) gonocoxite), (Gs) gonocoxal sternite." figureDoi="http://doi.org/10.5281/zenodo.5172626" httpUri="https://zenodo.org/record/5172626/files/figure.png" pageId="5" pageNumber="4">14a</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439660429FC58FF72FB87FF6F" box="[999,1045,225,249]" captionStart="Figure 15" captionStartId="33.[192,276,1817,1839]" captionTargetBox="[238,1409,229,1784]" captionTargetId="figure-88@33.[192,1440,192,1807]" captionTargetPageId="33" captionText="Figure 15. Internal sac of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (AL) accessory lamella, (CL) copulatory lamella (CL), (Ra) raspula, (Esp) speculum, and (Fg) a complex, spike-like flagellum that varies in length and thickness." figureDoi="http://doi.org/10.5281/zenodo.5172630" httpUri="https://zenodo.org/record/5172630/files/figure.png" pageId="5" pageNumber="4">15a</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439660429FB9DFF72FBDCFF6F" box="[1058,1102,225,249]" captionStart="Figure 16" captionStartId="34.[192,276,1818,1840]" captionTargetBox="[227,1428,205,1804]" captionTargetId="figure-41@34.[192,1440,192,1807]" captionTargetPageId="34" captionText="Figure 16. Distribution of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. maimon. d) T. gyas." figureDoi="http://doi.org/10.5281/zenodo.5172632" httpUri="https://zenodo.org/record/5172632/files/figure.png" pageId="5" pageNumber="4">16a</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62F3653A39660429FF7FFEB1FC59FEAC" box="[192,971,289,314]" pageId="5" pageNumber="4" type="reference_group">
|
||
<paragraph id="2A5636B139660429FF7FFEB1FC59FEAC" blockId="5.[192,971,289,314]" box="[192,971,289,314]" pageId="5" pageNumber="4">
|
||
<treatmentCitationGroup id="0AF9119F39660429FF7FFEB1FC59FEAC" box="[192,971,289,314]" pageId="5" pageNumber="4">
|
||
<treatmentCitation id="AB4810A039660429FF7FFEB1FD38FEAF" author="Fairmaire, L." box="[192,682,289,314]" page="245" pageId="5" pageNumber="4" year="1892">
|
||
<taxonomicName id="EDE94D3239660429FF7FFEB1FD38FEAF" authority="Fairmaire 1892: 245" authorityName="Fairmaire" authorityPageNumber="245" authorityYear="1892" box="[192,682,289,314]" class="Insecta" family="Dynastidae" genus="Ligyrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="bidentulus">
|
||
<emphasis id="189DEAA339660429FF7FFEB1FE3BFEAC" box="[192,425,290,314]" italics="true" pageId="5" pageNumber="4">Ligyrus bidentulus</emphasis>
|
||
<bibRefCitation id="4E784B4039660429FE0FFEB1FD38FEAF" author="Fairmaire, L." box="[432,682,289,314]" pageId="5" pageNumber="4" pagination="242 - 253" refId="ref14326" refString="Fairmaire, L. 1892. Descriptions de quelques Coleopteres argentins. Annales de la Societe Entomologique de Belgique 36: 242 - 253." type="journal article" year="1892">Fairmaire 1892: 245</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
(original combination).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62F3653A39660429FF7FFEF2FE05FBCC" pageId="5" pageNumber="4" type="description">
|
||
<paragraph id="2A5636B139660429FF7FFEF2FB4EFC6C" blockId="5.[192,1441,353,1114]" pageId="5" pageNumber="4">
|
||
<emphasis id="189DEAA339660429FF7FFEF2FE16FEEF" bold="true" box="[192,388,353,377]" pageId="5" pageNumber="4">Redescription</emphasis>
|
||
.
|
||
<emphasis id="189DEAA339660429FE2BFEF1FE4CFEEC" bold="true" box="[404,478,354,378]" pageId="5" pageNumber="4">Male.</emphasis>
|
||
Length
|
||
<quantity id="ED119B5439660429FDF7FEF2FD6BFEEF" box="[584,761,353,377]" metricMagnitude="-2" metricUnit="m" metricValue="1.4449999999999998" metricValueMax="1.67" metricValueMin="1.22" pageId="5" pageNumber="4" unit="mm" value="14.45" valueMax="16.7" valueMin="12.2">12.2–16.7 mm</quantity>
|
||
; width
|
||
<quantity id="ED119B5439660429FCE9FEF2FC78FEEF" box="[854,1002,353,377]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.6" metricValueMin="4.4" pageId="5" pageNumber="4" unit="mm" value="6.5" valueMax="8.6" valueMin="4.4">4.4–8.6 mm</quantity>
|
||
. Color light to dark reddish brown.
|
||
<emphasis id="189DEAA339660429FF7FFE11FE9DFE0C" bold="true" box="[192,271,386,410]" pageId="5" pageNumber="4">Head:</emphasis>
|
||
Frons mostly impunctate between eyes and coarsely rugopunctate just behind frontoclypeal carina. Frontoclypeal region with transverse carina slightly separated at middle (
|
||
<figureCitation id="B2D22A3439660429FBE1FE31FB2AFE2C" box="[1118,1208,418,442]" captionStart="Figure 4" captionStartId="22.[192,276,1848,1870]" captionTargetBox="[216,1440,216,1825]" captionTargetId="figure-63@22.[192,1442,192,1842]" captionTargetPageId="22" captionText="Figure 4. Head, dorsal view of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172595" httpUri="https://zenodo.org/record/5172595/files/figure.png" pageId="5" pageNumber="4">Fig. 4a</figureCitation>
|
||
). Clypeus strongly rugopunctate to transversely rugose, tapering to slightly emarginate, broad apex with 2 reflexed teeth. Mandibles with 2 teeth, apical tooth acute, subapical notch deep (
|
||
<figureCitation id="B2D22A3439660429FC77FE71FBA8FE6F" box="[968,1082,481,506]" captionStart="Figure 6" captionStartId="24.[192,276,1739,1761]" captionTargetBox="[211,1436,218,1720]" captionTargetId="figure-63@24.[192,1440,192,1730]" captionTargetPageId="24" captionText="Figure 6. Mandibles, ventral view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172601" httpUri="https://zenodo.org/record/5172601/files/figure.png" pageId="5" pageNumber="4">Fig. 6a–b</figureCitation>
|
||
); labium densely setose, paraglossa undeveloped, apex convergent (
|
||
<figureCitation id="B2D22A3439660429FD2EFD91FD74FD8C" box="[657,742,514,538]" captionStart="Figure 7" captionStartId="25.[192,276,1683,1705]" captionTargetBox="[231,1392,238,1664]" captionTargetId="figure-63@25.[192,1440,192,1671]" captionTargetPageId="25" captionText="Figure 7. Labium, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172605" httpUri="https://zenodo.org/record/5172605/files/figure.png" pageId="5" pageNumber="4">Fig. 7a</figureCitation>
|
||
); maxilla with galea developed, with 5 teeth.
|
||
<emphasis id="189DEAA339660429FAB3FD91FA32FD8C" bold="true" box="[1292,1440,514,538]" pageId="5" pageNumber="4">Pronotum:</emphasis>
|
||
Surface crazed, sparsely punctate; punctures moderately large, umbilicate. Anterior margin with small, median tubercle followed by small, shallow fovea (
|
||
<figureCitation id="B2D22A3439660429FC91FDD1FC14FDCC" box="[814,902,578,602]" captionStart="Figure 8" captionStartId="26.[192,276,1841,1863]" captionTargetBox="[275,1367,208,1829]" captionTargetId="figure-18@26.[269,1376,192,1840]" captionTargetPageId="26" captionText="Figure 8. Pronotum, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172607" httpUri="https://zenodo.org/record/5172607/files/figure.png" pageId="5" pageNumber="4">Fig. 8a</figureCitation>
|
||
).
|
||
<emphasis id="189DEAA339660429FC23FDD1FC69FDCC" bold="true" box="[924,1019,578,602]" pageId="5" pageNumber="4">Elytra:</emphasis>
|
||
Surface with micropunctures and large punctures mixed (large punctures moderately dense, ocellate) and with 3 distinct pairs of punctures. First broad interval with or without large punctures.
|
||
<emphasis id="189DEAA339660429FCF9FD12FC46FD0F" bold="true" box="[838,980,641,665]" pageId="5" pageNumber="4">Pygidium:</emphasis>
|
||
Surface sparsely punctate, punctures becoming dense in basal angles; punctures small to moderate in size, weakly ocellate. In lateral view, surface of males regularly convex, surface in females weakly convex.
|
||
<emphasis id="189DEAA339660429FBA7FD51FBCDFD4C" bold="true" box="[1048,1119,706,730]" pageId="5" pageNumber="4">Legs:</emphasis>
|
||
Protibia tridentate, basal tooth slightly removed. Protarsus in male simple, not enlarged. Mesotibia with 1 basal and 1 medial transverse carina, each with spine-like setae (
|
||
<figureCitation id="B2D22A3439660429FD40FC91FC17FC8C" box="[767,901,770,794]" captionStart="Figure 10" captionStartId="28.[192,275,1624,1646]" captionTargetBox="[220,1411,210,1601]" captionTargetId="figure-65@28.[192,1440,192,1613]" captionTargetPageId="28" captionText="Figure 10. Meso- and metatibia, dorsal view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172614" httpUri="https://zenodo.org/record/5172614/files/figure.png" pageId="5" pageNumber="4">Fig. 10a–b</figureCitation>
|
||
). Metatibia with 1 basal, transverse carina interrupted at middle and with spine-like setae and 1 medial, complete, transverse carina; apex with about 29 spinules (
|
||
<figureCitation id="B2D22A3439660429FE14FCD1FD81FCCC" box="[427,531,834,858]" captionStart="Figure 10" captionStartId="28.[192,275,1624,1646]" captionTargetBox="[220,1411,210,1601]" captionTargetId="figure-65@28.[192,1440,192,1613]" captionTargetPageId="28" captionText="Figure 10. Meso- and metatibia, dorsal view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172614" httpUri="https://zenodo.org/record/5172614/files/figure.png" pageId="5" pageNumber="4">Fig. 10b</figureCitation>
|
||
). First metatarsomere triangularly expanded, apex subtruncate.
|
||
<emphasis id="189DEAA339660429FA85FCD1FA32FCCC" bold="true" box="[1338,1440,834,858]" pageId="5" pageNumber="4">Venter:</emphasis>
|
||
Prosternal process tall, thick; apex small, suboval, usually with long, slender setae. Mesosternal surface sparsely setose, slightly concave at middle. Metasternal surface densely punctate, punctures ocellate, setose; setae slender, long; lateral edge rugopunctate with long, slender setae. Abdominal ventrite VIII depressed at middle, (
|
||
<figureCitation id="B2D22A3439660429FDB6FC51FDEDFC4C" box="[521,639,962,986]" captionStart="Figure 1" captionStartId="19.[192,276,1846,1868]" captionTargetBox="[207,1426,292,1810]" captionTargetId="figure-64@19.[192,1441,192,1828]" captionTargetPageId="19" captionText="Figure 1. Habitus of Tomarus spp. a) Tomarus bidentulus (Fairmaire) dorsal view. b) Tomarus burmeisteri (Steinheil) dorsal view. c) Tomarus gyas Erichson dorsal view. d) Tomarus rosettae (Endrödi) dorsal view. e) Tomarus roigjunenti n. sp. dorsal view, holotype. f) Tomarus roigjunenti n. sp. dorsal view, allotype." figureDoi="http://doi.org/10.5281/zenodo.5172584" httpUri="https://zenodo.org/record/5172584/files/figure.png" pageId="5" pageNumber="4">
|
||
Fig.
|
||
<quantity id="ED119B5439660429FDFFFC51FDC9FC4C" box="[576,603,962,986]" metricMagnitude="-3" metricUnit="kg" metricValue="1.0" pageId="5" pageNumber="4" unit="g" value="1.0">1g</quantity>
|
||
–h
|
||
</figureCitation>
|
||
).
|
||
<emphasis id="189DEAA339660429FD2BFC51FCCDFC4F" bold="true" box="[660,863,961,986]" pageId="5" pageNumber="4">Male Genitalia</emphasis>
|
||
: Parameres (
|
||
<figureCitation id="B2D22A3439660429FBBBFC51FBF8FC4C" box="[1028,1130,962,986]" captionStart="Figure 11" captionStartId="29.[192,276,1835,1857]" captionTargetBox="[244,1411,241,1815]" captionTargetId="figure-63@29.[192,1442,192,1821]" captionTargetPageId="29" captionText="Figure 11. Parameres, caudal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172618" httpUri="https://zenodo.org/record/5172618/files/figure.png" pageId="5" pageNumber="4">Fig. 11a</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439660429FBC9FC51FB37FC4C" box="[1142,1189,962,986]" captionStart="Figure 12" captionStartId="30.[192,274,1776,1798]" captionTargetBox="[210,1434,220,1735]" captionTargetId="figure-61@30.[192,1440,192,1769]" captionTargetPageId="30" captionText="Figure 12. Parameres, lateral view, of: a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172620" httpUri="https://zenodo.org/record/5172620/files/figure.png" pageId="5" pageNumber="4">12a</figureCitation>
|
||
,
|
||
<figureCitation id="B2D22A3439660429FB0EFC51FB4CFC4C" box="[1201,1246,962,986]" captionStart="Figure 13" captionStartId="31.[192,275,1823,1845]" captionTargetBox="[238,1403,236,1798]" captionTargetId="figure-63@31.[192,1440,192,1807]" captionTargetPageId="31" captionText="Figure 13. Parameres, anterior view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172622" httpUri="https://zenodo.org/record/5172622/files/figure.png" pageId="5" pageNumber="4">13a</figureCitation>
|
||
) subtriangular, rounded apices curving outwards. Internal sac (
|
||
<figureCitation id="B2D22A3439660429FCB3FC71FCE2FC6F" box="[780,880,993,1018]" captionStart="Figure 15" captionStartId="33.[192,276,1817,1839]" captionTargetBox="[238,1409,229,1784]" captionTargetId="figure-88@33.[192,1440,192,1807]" captionTargetPageId="33" captionText="Figure 15. Internal sac of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (AL) accessory lamella, (CL) copulatory lamella (CL), (Ra) raspula, (Esp) speculum, and (Fg) a complex, spike-like flagellum that varies in length and thickness." figureDoi="http://doi.org/10.5281/zenodo.5172630" httpUri="https://zenodo.org/record/5172630/files/figure.png" pageId="5" pageNumber="4">Fig. 15a</figureCitation>
|
||
) lacking copulatory lamellae.
|
||
</paragraph>
|
||
<paragraph id="2A5636B139660429FF4FFB91FE05FBCC" blockId="5.[192,1441,353,1114]" pageId="5" pageNumber="4">
|
||
<emphasis id="189DEAA339660429FF4FFB91FEC7FB8C" bold="true" box="[240,341,1026,1050]" pageId="5" pageNumber="4">Female</emphasis>
|
||
. Length
|
||
<quantity id="ED119B5439660429FE7DFB92FDE2FB8F" box="[450,624,1025,1049]" metricMagnitude="-2" metricUnit="m" metricValue="1.825" metricValueMax="1.96" metricValueMin="1.69" pageId="5" pageNumber="4" unit="mm" value="18.25" valueMax="19.6" valueMin="16.9">16.9–19.6 mm</quantity>
|
||
; width
|
||
<quantity id="ED119B5439660429FD76FB92FCF8FB8F" box="[713,874,1025,1049]" metricMagnitude="-3" metricUnit="m" metricValue="9.45" metricValueMax="10.6" metricValueMin="8.3" pageId="5" pageNumber="4" unit="mm" value="9.45" valueMax="10.6" valueMin="8.3">8.3–10.6 mm</quantity>
|
||
. Similar to male, but disc of pronotum smooth, pygidium slightly concave, and gonocoxite smaller than gonocoxal sternite and lacking a semicircular furrow (
|
||
<figureCitation id="B2D22A3439660429FE9CFBD1FE18FBCC" box="[291,394,1090,1114]" captionStart="Figure 14" captionStartId="32.[192,276,1832,1854]" captionTargetBox="[358,1267,208,1798]" captionTargetId="figure-70@32.[349,1298,192,1821]" captionTargetPageId="32" captionText="Figure 14. Gonocoxite, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (Gx) gonocoxite), (Gs) gonocoxal sternite." figureDoi="http://doi.org/10.5281/zenodo.5172626" httpUri="https://zenodo.org/record/5172626/files/figure.png" pageId="5" pageNumber="4">Fig. 14a</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62F3653A39660429FF7FFB12FEB2FA8F" pageId="5" pageNumber="4" type="distribution">
|
||
<paragraph id="2A5636B139660429FF7FFB12FEC3FB2C" blockId="5.[192,1440,1153,1210]" pageId="5" pageNumber="4">
|
||
<emphasis id="189DEAA339660429FF7FFB12FEFEFB0F" bold="true" box="[192,364,1153,1177]" pageId="5" pageNumber="4">Distribution</emphasis>
|
||
.
|
||
<taxonomicName id="EDE94D3239660429FEC5FB11FDE0FB0C" box="[378,626,1154,1178]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="bidentulus">
|
||
<emphasis id="189DEAA339660429FEC5FB11FDE0FB0C" box="[378,626,1154,1178]" italics="true" pageId="5" pageNumber="4">Tomarus bidentulus</emphasis>
|
||
</taxonomicName>
|
||
occurs in
|
||
<collectingCountry id="52FE762139660429FD54FB11FCFAFB0C" box="[747,872,1154,1178]" name="Argentina" pageId="5" pageNumber="4">Argentina</collectingCountry>
|
||
and
|
||
<collectingCountry id="52FE762139660429FC1BFB11FB80FB0C" box="[932,1042,1154,1178]" name="Uruguay" pageId="5" pageNumber="4">Uruguay</collectingCountry>
|
||
(
|
||
<bibRefCitation id="4E784B4039660429FB9FFB11FB47FB0F" author="Endrodi, S." box="[1056,1237,1153,1178]" pageId="5" pageNumber="4" pagination="1 - 145" refId="ref14087" refString="Endrodi, S. 1969 a. Monographie der Dynastinae, 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). Entomologische Abhandlungen 37: 1 - 145." type="journal article" year="1969">Endrödi 1969a</bibRefCitation>
|
||
–b, 1985;
|
||
<bibRefCitation id="4E784B4039660429FAF5FB11FED6FB2C" author="Abadie, E. & P. Grossi & P. S. Wagner" pageId="5" pageNumber="4" refId="ref13766" refString="Abadie, E., P. Grossi, and P. S. Wagner. 2008. A field guide of the Dynastinae family of the south of South America. Published by the authors; Buenos Aires, Argentina. 119 p." type="book" year="2008">Abadie et al. 2008</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2A5636B139660429FF7FFB72FEB2FA8F" blockId="5.[192,1441,1249,1722]" pageId="5" pageNumber="4">
|
||
<emphasis id="189DEAA339660429FF7FFB72FE38FB6C" bold="true" box="[192,426,1249,1274]" pageId="5" pageNumber="4">Locality Records</emphasis>
|
||
(
|
||
<figureCitation id="B2D22A3439660429FE04FB71FDB0FB6F" box="[443,546,1249,1274]" captionStart="Figure 16" captionStartId="34.[192,276,1818,1840]" captionTargetBox="[227,1428,205,1804]" captionTargetId="figure-41@34.[192,1440,192,1807]" captionTargetPageId="34" captionText="Figure 16. Distribution of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. maimon. d) T. gyas." figureDoi="http://doi.org/10.5281/zenodo.5172632" httpUri="https://zenodo.org/record/5172632/files/figure.png" pageId="5" pageNumber="4">Fig. 16a</figureCitation>
|
||
).
|
||
<specimenCount id="3CEFFD3839660429FD88FB71FD78FB6C" box="[567,746,1250,1274]" pageId="5" pageNumber="4" type="generic">711 specimens</specimenCount>
|
||
examined from BCRC, CMNC, IADIZA, MACN, MLPA, URMU.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62F3653A39660429FF4FFAB2FD71F88C" pageId="5" pageNumber="4" type="materials_examined">
|
||
<paragraph id="2A5636B139660429FF4FFAB2FCA8F92C" blockId="5.[192,1441,1249,1722]" pageId="5" pageNumber="4">
|
||
<emphasis id="189DEAA339660429FF4FFAB2FE31FAAF" bold="true" box="[240,419,1313,1337]" pageId="5" pageNumber="4">
|
||
<collectingCountry id="52FE762139660429FF4FFAB2FE31FAAF" box="[240,419,1313,1337]" name="Argentina" pageId="5" pageNumber="4">ARGENTINA</collectingCountry>
|
||
</emphasis>
|
||
(676).
|
||
<collectingRegion id="E82DF85339660429FE4DFAB1FD51FAAC" box="[498,707,1314,1338]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="5" pageNumber="4">BUENOS AIRES</collectingRegion>
|
||
(46):
|
||
<taxonomicName id="EDE94D3239660429FCBBFAB1FC3BFAAC" authority="Blanca" authorityName="Blanca" box="[772,937,1314,1338]" class="Magnoliopsida" family="Asteraceae" genus="Bahia" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="4" phylum="Tracheophyta" rank="genus">Bahía Blanca</taxonomicName>
|
||
, Buenos Aires-Capital Federal, Necochea, San Blas, Villa Gessell, Villarica (
|
||
<quantity id="ED119B5439660429FDCCFAD1FD56FACC" box="[627,708,1346,1370]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="4" unit="km" value="30.0">30 km</quantity>
|
||
SW).
|
||
<collectingRegion id="E82DF85339660429FCABFAD1FC54FACC" box="[788,966,1346,1370]" country="Argentina" name="Catamarca" pageId="5" pageNumber="4">CATAMARCA</collectingRegion>
|
||
(5): Tinogasti-Fiambala.
|
||
<collectingRegion id="E82DF85339660429FAB7FAD1FAFFFACC" box="[1288,1389,1346,1370]" country="Argentina" name="Chaco" pageId="5" pageNumber="4">CHACO</collectingRegion>
|
||
(1): Resistencia.
|
||
<collectingRegion id="E82DF85339660429FEE3FAF1FE44FAEC" box="[348,470,1378,1402]" country="Argentina" name="Chubut" pageId="5" pageNumber="4">CHUBUT</collectingRegion>
|
||
(2): Dolavon.
|
||
<collectingRegion id="E82DF85339660429FD3DFAF1FD6EFAEC" box="[642,764,1378,1402]" country="Argentina" name="Chubut" pageId="5" pageNumber="4">CHUBUT</collectingRegion>
|
||
(2): Estación de Esquel, San Jorge.
|
||
<collectingRegion id="E82DF85339660429FB05FAF2FAD7FAEC" box="[1210,1349,1377,1402]" country="Argentina" name="CORDOBA" pageId="5" pageNumber="4">CÓRDOBA</collectingRegion>
|
||
(1): No Data.
|
||
<collectingRegion id="E82DF85339660429FEB4FA11FEF5FA0F" box="[267,359,1410,1433]" country="Argentina" name="Jujuy" pageId="5" pageNumber="4">JUJUY</collectingRegion>
|
||
(3): Posta de Lozano, No Data.
|
||
<collectingRegion id="E82DF85339660429FD58FA11FCE4FA0C" box="[743,886,1410,1434]" country="Argentina" name="La Pampa" pageId="5" pageNumber="4">LA PAMPA</collectingRegion>
|
||
(2): Chacharramendi.
|
||
<collectingRegion id="E82DF85339660429FB35FA11FA99FA0C" box="[1162,1291,1410,1434]" country="Argentina" name="La Rioja" pageId="5" pageNumber="4">LA RIOJA</collectingRegion>
|
||
(7): Jaguey, Mascasin, Patiquía, Tinogasta, Villa Union.
|
||
<collectingRegion id="E82DF85339660429FD71FA31FCCEFA2C" box="[718,860,1442,1466]" country="Argentina" name="Mendoza" pageId="5" pageNumber="4">MENDOZA</collectingRegion>
|
||
(30): Bardas Blancas, Cerro Quadral, Las Heras, La Valle, Mendoza Capital, Ov. Balleza, Ranquil Norte, El Nihuil-San Rafael.
|
||
<collectingRegion id="E82DF85339660429FBCDFA51FA92FA4C" box="[1138,1280,1474,1498]" country="Argentina" name="MISIONES" pageId="5" pageNumber="4">MISIONES</collectingRegion>
|
||
(2): Posadas.
|
||
<collectingRegion id="E82DF85339660429FF7FFA72FEC0FA6C" box="[192,338,1505,1530]" country="Argentina" name="Neuquen" pageId="5" pageNumber="4">NEUQUÉN</collectingRegion>
|
||
(72): Allere, Anelo, Bajada Marucho, Bajada Agrio, Collon-Curá, Covuco, Coyunca, Huncal, La Negra, La Pintada, Pampa Salado, Picun Leufu, Piedra del Agüila, Pilmatus, Planicie Banderita, Río Salado, Zalapa-Catuto.
|
||
<collectingRegion id="E82DF85339660429FDA9F9B2FD25F9AC" box="[534,695,1569,1594]" country="Argentina" name="RIO NEGRO" pageId="5" pageNumber="4">RÍO NEGRO</collectingRegion>
|
||
(469): Allén, Catriel, Cipolletti, Conesal, Fray Luis Beltrán, General Gómez, General Roca, Paso Flores, Piedra Aguila, Río Negro-Capital, San Antonio Oeste, Valcheta, Villa Regina.
|
||
<collectingRegion id="E82DF85339660429FE5BF9F1FDADF9EC" box="[484,575,1634,1658]" country="Argentina" name="Salta" pageId="5" pageNumber="4">SALTA</collectingRegion>
|
||
(32): Cafayate-Norte
|
||
<quantity id="ED119B5439660429FCF4F9F1FC1CF9EC" box="[843,910,1634,1658]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="4" unit="km" value="1.0">1 km</quantity>
|
||
., Los Lajitas, Salta-Capital.
|
||
<collectingRegion id="E82DF85339660429FB53F9F1FAE2F9EC" box="[1260,1392,1634,1658]" country="Argentina" name="San Luis" pageId="5" pageNumber="4">SAN LUIS</collectingRegion>
|
||
(1): Paso Patria.
|
||
<collectingRegion id="E82DF85339660429FEDCF911FD8BF90C" box="[355,537,1666,1690]" country="Argentina" name="SANTA CRUZ" pageId="5" pageNumber="4">SANTA CRUZ</collectingRegion>
|
||
(1): Santa Cruz-Capital.
|
||
<collectingRegion id="E82DF85339660429FCE7F912FC79F90C" box="[856,1003,1665,1690]" country="Argentina" name="Tucuman" pageId="5" pageNumber="4">TUCUMÁN</collectingRegion>
|
||
(1): Tafi de Valle.
|
||
<emphasis id="189DEAA339660429FB67F912FAFCF90F" bold="true" box="[1240,1390,1665,1689]" pageId="5" pageNumber="4">
|
||
<collectingCountry id="52FE762139660429FB67F912FAFCF90F" box="[1240,1390,1665,1689]" name="Uruguay" pageId="5" pageNumber="4">URUGUAY</collectingCountry>
|
||
</emphasis>
|
||
(7).
|
||
<collectingRegion id="E82DF85339660429FF7FF931FEE2F92C" box="[192,368,1698,1722]" country="Uruguay" name="Canelones" pageId="5" pageNumber="4">CANELONES</collectingRegion>
|
||
(7): Banda Oriental.
|
||
<emphasis id="189DEAA339660429FDC9F932FD6EF92F" bold="true" box="[630,764,1697,1721]" pageId="5" pageNumber="4">
|
||
<collectingRegion id="E82DF85339660429FDC9F932FD6EF92F" box="[630,764,1697,1721]" country="Uruguay" name="NO DATA" pageId="5" pageNumber="4">NO DATA</collectingRegion>
|
||
</emphasis>
|
||
(28).
|
||
</paragraph>
|
||
<paragraph id="2A5636B139660429FF7FF971FD71F88C" blockId="5.[192,1440,1761,1818]" pageId="5" pageNumber="4">
|
||
<emphasis id="189DEAA339660429FF7FF971FE6BF96F" bold="true" box="[192,505,1761,1786]" pageId="5" pageNumber="4">Temporal Distribution</emphasis>
|
||
. January (36), February (29), March (35), April (15), May (4) September (3), October (14), November (42), December (12).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="62F3653A3966042AFF7FF8D2FAD2FEEC" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4" type="diagnosis">
|
||
<paragraph id="2A5636B13966042AFF7FF8D2FCFFFE8F" blockId="5.[192,1440,1857,1914]" lastBlockId="6.[192,1440,194,282]" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4">
|
||
<emphasis id="189DEAA339660429FF7FF8D2FEDAF8CF" bold="true" box="[192,328,1857,1881]" pageId="5" pageNumber="4">Diagnosis</emphasis>
|
||
.
|
||
<taxonomicName id="EDE94D3239660429FEE5F8D1FDC5F8CC" box="[346,599,1858,1882]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="bidentulus">
|
||
<emphasis id="189DEAA339660429FEE5F8D1FDC5F8CC" box="[346,599,1858,1882]" italics="true" pageId="5" pageNumber="4">Tomarus bidentulus</emphasis>
|
||
</taxonomicName>
|
||
is distinctive because the apex of clypeus has the teeth distinctly separated (
|
||
<figureCitation id="B2D22A3439660429FEF4F8F1FE36F8EC" box="[331,420,1890,1914]" captionStart="Figure 4" captionStartId="22.[192,276,1848,1870]" captionTargetBox="[216,1440,216,1825]" captionTargetId="figure-63@22.[192,1442,192,1842]" captionTargetPageId="22" captionText="Figure 4. Head, dorsal view of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172595" httpUri="https://zenodo.org/record/5172595/files/figure.png" pageId="5" pageNumber="4">Fig. 4a</figureCitation>
|
||
); the pronotum has a small, subapical tubercle followed by a small fovea; and the surface on the disc is moderately punctate with the sides and anterior angles densely punctate (
|
||
<figureCitation id="B2D22A343965042AFACFFF51FF4DFF6C" captionStart="Figure 8" captionStartId="26.[192,276,1841,1863]" captionTargetBox="[275,1367,208,1829]" captionTargetId="figure-18@26.[269,1376,192,1840]" captionTargetPageId="26" captionText="Figure 8. Pronotum, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus." figureDoi="http://doi.org/10.5281/zenodo.5172607" httpUri="https://zenodo.org/record/5172607/files/figure.png" pageId="6" pageNumber="5">Fig. 8a</figureCitation>
|
||
).
|
||
<taxonomicName id="EDE94D323965042AFF48FF71FE63FF6C" box="[247,497,226,250]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="bidentulus">
|
||
<emphasis id="189DEAA33965042AFF48FF71FE63FF6C" box="[247,497,226,250]" italics="true" pageId="6" pageNumber="5">Tomarus bidentulus</emphasis>
|
||
</taxonomicName>
|
||
also resembles
|
||
<taxonomicName id="EDE94D323965042AFD08FF71FCF4FF6C" baseAuthorityName="Steinheil" baseAuthorityYear="1872" box="[695,870,226,250]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="burmeisteri">
|
||
<emphasis id="189DEAA33965042AFD08FF71FCF4FF6C" box="[695,870,226,250]" italics="true" pageId="6" pageNumber="5">T. burmeisteri</emphasis>
|
||
</taxonomicName>
|
||
but is easily distinguished by the presence of the pronotal tubercle, which is absent in
|
||
<taxonomicName id="EDE94D323965042AFD07FE91FCF4FE8C" baseAuthorityName="Steinheil" baseAuthorityYear="1872" box="[696,870,258,282]" class="Insecta" family="Dynastidae" genus="Tomarus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="burmeisteri">
|
||
<emphasis id="189DEAA33965042AFD07FE91FCF4FE8C" box="[696,870,258,282]" italics="true" pageId="6" pageNumber="5">T. burmeisteri</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="2A5636B13965042AFF7FFED2FAD2FEEC" blockId="6.[192,1440,321,378]" pageId="6" pageNumber="5">
|
||
<emphasis id="189DEAA33965042AFF7FFED2FEF6FECF" bold="true" box="[192,356,321,345]" pageId="6" pageNumber="5">Life History</emphasis>
|
||
. Adults are nocturnal and are attracted to lights. This is an infrequently collected species. Adult emergence is during the summer, which is the rainy season in
|
||
<collectingCountry id="52FE76213965042AFBAEFEF1FB1CFEEC" box="[1041,1166,354,378]" name="Argentina" pageId="6" pageNumber="5">Argentina</collectingCountry>
|
||
and
|
||
<collectingCountry id="52FE76213965042AFB74FEF1FAAEFEEC" box="[1227,1340,354,378]" name="Uruguay" pageId="6" pageNumber="5">Uruguay</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |