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<document id="75926765B8AF160592553AE381340D6C" ID-CLB-Dataset="55975" ID-DOI="10.1093/zoolinnean/zlab077" ID-GBIF-Dataset="5e350c66-bf9d-4f32-b68f-16a6b329a76f" ID-ISSN="0024-4082" ID-Zenodo-Dep="6461343" ID-ZooBank="FFC8945F-6A97-4DB7-99CD-E08162DD5819" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1649833861871" checkinUser="plazi" docAuthor="Vilarino, Albane, Dias, Everton Santos &amp; Bispo, Pitágoras Da Conceição" docDate="2022" docId="8F15A94DFF99FF8FB9E6FCE30BECFB97" docLanguage="en" docName="zlab077.pdf" docOrigin="Zoological Journal of the Linnean Society 194 (4)" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Caenocentron rafamoralesi Vilarino &amp; Dias &amp; Bispo 2022, SP. NOV." docType="treatment" docVersion="7" lastPageNumber="1360" masterDocId="732CD135FF8BFF9CB937FD2E0956FFFD" masterDocTitle="Phylogeny indicates polyphyly in Cnodocentron (Trichoptera: Xiphocentronidae): biogeography and revision of New World species (Caenocentron)" masterLastPageNumber="1373" masterPageNumber="1341" pageNumber="1359" updateTime="1699328856895" updateUser="ExternalLinkService">
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<mods:title id="CD49D4EFCFB4E3793557F4F34949F5A3">Phylogeny indicates polyphyly in Cnodocentron (Trichoptera: Xiphocentronidae): biogeography and revision of New World species (Caenocentron)</mods:title>
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<mods:namePart id="D1DB2799EBDBE790A4BCF6AB5DD80FF0">Vilarino, Albane</mods:namePart>
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<mods:namePart id="55A836E7F46A39A77701DAD398231577">Dias, Everton Santos</mods:namePart>
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<mods:namePart id="A9442FD9001D15A3369437756E5D1BDD">Bispo, Pitágoras Da Conceição</mods:namePart>
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<mods:title id="7FC6D38805D5CC0B6E74E9F6C2DE8951">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="1AA466CE9BA820C92458C827520AD796">2022</mods:date>
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<mods:number id="B643357BC4F058F127D42111BF93F013">2021-12-23</mods:number>
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<mods:detail id="94BD3BC77E6EBCBF719FCB6F7125B21B" type="volume">
<mods:number id="8D6CD7EE7373ECB819DE7DD46AF2F139">194</mods:number>
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<mods:classification id="159A878D401174BB7BB3CD7CCAED4F26">journal article</mods:classification>
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<mods:identifier id="D713F6BA01233013AE66201FE47F3DEE" type="DOI">10.1093/zoolinnean/zlab077</mods:identifier>
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<treatment id="8F15A94DFF99FF8FB9E6FCE30BECFB97" ID-DOI="http://doi.org/10.5281/zenodo.6457193" ID-GBIF-Taxon="194598719" ID-Zenodo-Dep="6457193" LSID="urn:lsid:plazi:treatment:8F15A94DFF99FF8FB9E6FCE30BECFB97" httpUri="http://treatment.plazi.org/id/8F15A94DFF99FF8FB9E6FCE30BECFB97" lastPageId="19" lastPageNumber="1360" pageId="18" pageNumber="1359">
<subSubSection id="4FA64BD0FF99FF8EB9E6FCE30BEEFE18" box="[209,696,461,485]" pageId="18" pageNumber="1359" type="nomenclature">
<paragraph id="0703185BFF99FF8EB9E6FCE30BEEFE18" blockId="18.[209,696,461,485]" box="[209,696,461,485]" pageId="18" pageNumber="1359">
<heading id="5C4BAF37FF99FF8EB9E6FCE30BEEFE18" box="[209,696,461,485]" centered="true" fontSize="9" level="2" pageId="18" pageNumber="1359" reason="2">
<taxonomicName id="C0BC63D8FF99FF8EB9E6FCE30B18FE1E" authority="Vilarino &amp; Dias &amp; Bispo, 2022" authorityName="Vilarino &amp; Dias &amp; Bispo" authorityYear="2022" box="[209,590,461,485]" class="Insecta" family="Xiphocentronidae" genus="Caenocentron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="1368" phylum="Arthropoda" rank="species" species="rafamoralesi" status="sp. nov.">CAENOCENTRON RAFAMORALESI</taxonomicName>
<emphasis id="35C8C449FF99FF8EBB62FCFE0BEEFE18" bold="true" box="[597,696,461,485]" pageId="18" pageNumber="1359">
<taxonomicNameLabel id="2EFB7932FF99FF8EBB62FCFE0BEEFE18" box="[597,696,461,485]" pageId="18" pageNumber="1359" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="4FA64BD0FF99FF8EB87FFCDD0BB0FD9A" pageId="18" pageNumber="1359" type="description">
<paragraph id="0703185BFF99FF8EB87FFCDD0B17FDF1" blockId="18.[328,577,499,524]" box="[328,577,499,524]" pageId="18" pageNumber="1359">
(
<figureCitation id="9F8704DEFF99FF8EB867FCDD088AFDF1" box="[336,476,499,524]" captionStart="Figures 9–11" captionStartId="16.[145,234,1647,1669]" captionTargetBox="[146,1426,196,1605]" captionTargetId="figure-72@16.[145,1428,195,1607]" captionTargetPageId="16" captionText="Figures 9–11. Caenocentron female genitalia. 9. Caenocentron carlosdelarosai sp. nov. A, lateral; B, dorsal; C, detail of apex lateral; D, detail of apex ventral. 10. Caenocentron rafamoralesi sp. nov.A, lateral; B, dorsal; C, detail of apex lateral; D, detail of apex ventral. 11. Caenocentron yavapai. A, lateral; B, dorsal; C, detail of apex ventral (modified from Ruiter, 2006). Scale bar = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.6461366" httpUri="https://zenodo.org/record/6461366/files/figure.png" pageId="18" pageNumber="1359">FIGS 10A–D</figureCitation>
,
<figureCitation id="9F8704DEFF99FF8EB8DEFCDD0B6CFDF1" box="[489,570,499,524]" captionStart="Figure 13" captionStartId="18.[808,890,1347,1369]" captionTargetBox="[812,1421,196,1306]" captionTargetId="figure-566@18.[811,1422,195,1307]" captionTargetPageId="18" captionText="Figure 13. Caenocentron rafamoralesi sp. nov. A, wing venation. Male genitalia: B, lateral; C, dorsal; D, ventral." figureDoi="http://doi.org/10.5281/zenodo.6461370" httpUri="https://zenodo.org/record/6461370/files/figure.png" pageId="18" pageNumber="1359">13A–D</figureCitation>
)
</paragraph>
<paragraph id="0703185BFF99FF8EB9A6FF1A0BB0FD9A" blockId="18.[145,761,564,615]" pageId="18" pageNumber="1359">
<emphasis id="35C8C449FF99FF8EB9A6FF1A0BB0FD9A" italics="true" pageId="18" pageNumber="1359">
Zoobank registration: urn: lsid: zoobank. org:act:
<uuid id="731A228EFF99FF8EB9D4FF7C0BB0FD9A" box="[227,742,594,615]" pageId="18" pageNumber="1359">063702DF-0860-43F1-8DE8-757C1B08AD4A</uuid>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="4FA64BD0FF99FF8EB9A6FFBF0890FCDF" pageId="18" pageNumber="1359" type="materials_examined">
<paragraph id="0703185BFF99FF8EB9A6FFBF0890FCDF" blockId="18.[145,760,657,802]" pageId="18" pageNumber="1359">
<materialsCitation id="B7D41206FF99FF8EB9A6FFBF0890FCDF" ID-GBIF-Occurrence="3751569301" collectingDate="2015-04-30" collectionCode="BIOUG" collectorName="W. Hallwachs" country="Costa Rica" elevation="1220" latitude="10.9292" location="Sector Cacao" longLatPrecision="1" longitude="-85.464294" municipality="Area de Conservacion" pageId="18" pageNumber="1359" specimenCode="BIOUG36194" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="holotype">
<emphasis id="35C8C449FF99FF8EB9A6FFBF085DFD5B" box="[145,267,657,678]" italics="true" pageId="18" pageNumber="1359">
<typeStatus id="D807A6F9FF99FF8EB9A6FFBF0853FD5B" box="[145,261,657,678]" pageId="18" pageNumber="1359" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectingCountry id="7FAB58CBFF99FF8EB82BFFBF0887FD5B" box="[284,465,657,678]" name="Costa Rica" pageId="18" pageNumber="1359">COSTA RICA</collectingCountry>
,
<collectingRegion id="C578D6B9FF99FF8EB8D4FFBF0B29FD5B" box="[483,639,657,678]" country="Costa Rica" name="Guanacaste" pageId="18" pageNumber="1359">Guanacaste</collectingRegion>
,
<collectingMunicipality id="E7678221FF99FF8EBBA6FFBF0866FD3B" pageId="18" pageNumber="1359">Area de Conservacion</collectingMunicipality>
<collectingRegion id="C578D6B9FF99FF8EB80FFF9E0893FD38" box="[312,453,688,709]" country="Costa Rica" name="Guanacaste" pageId="18" pageNumber="1359">Guanacaste</collectingRegion>
,
<location id="02634E80FF99FF8EB8E5FF9E0B3BFD38" LSID="urn:lsid:plazi:treatment:8F15A94DFF99FF8FB9E6FCE30BECFB97:02634E80FF99FF8EB8E5FF9E0B3BFD38" box="[466,621,688,709]" country="Costa Rica" latitude="10.9292" longLatPrecision="1" longitude="-85.464294" municipality="Area de Conservacion" name="Sector Cacao" pageId="18" pageNumber="1359" stateProvince="Guanacaste">Sector Cacao</location>
,
<location id="02634E80FF99FF8EBB4CFF9E0BA2FD3B" LSID="urn:lsid:plazi:treatment:8F15A94DFF99FF8FB9E6FCE30BECFB97:02634E80FF99FF8EBB4CFF9E0BA2FD3B" box="[635,756,688,710]" country="Costa Rica" latitude="10.9292" longLatPrecision="1" longitude="-85.464294" municipality="Area de Conservacion" name="Derrumbe" pageId="18" pageNumber="1359" stateProvince="Guanacaste">Derrumbe</location>
,
<geoCoordinate id="62887E9CFF99FF8EB9A6FFE00865FD19" box="[145,307,718,740]" degrees="10" direction="north" minutes="55" orientation="latitude" pageId="18" pageNumber="1359" precision="1" seconds="45.12" value="10.9292">10°55’45.12’’N</geoCoordinate>
,
<geoCoordinate id="62887E9CFF99FF8EB80BFFE008B6FD19" box="[316,480,718,740]" degrees="85" direction="west" minutes="27" orientation="longitude" pageId="18" pageNumber="1359" precision="1" seconds="51.48" value="-85.464294">85°27’51.48’’W</geoCoordinate>
;
<quantity id="C044B5BEFF99FF8EB8DEFFE10B6AFD19" box="[489,572,719,740]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="18" pageNumber="1359" unit="m" value="1220.0">
<elevation id="8C91FF68FF99FF8EB8DEFFE10B6AFD19" box="[489,572,719,740]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="18" pageNumber="1359" unit="m" value="1220.0">1220 m</elevation>
</quantity>
a.s.l.,
<date id="73023E9BFF99FF8EBBB6FFE00BA2FD19" box="[641,756,718,740]" pageId="18" pageNumber="1359" value="2015-04-30">
<collectingDate id="6346C773FF99FF8EBBB6FFE00BA2FD19" box="[641,756,718,740]" pageId="18" pageNumber="1359" value="2015-04-30">30.iv.2015</collectingDate>
</date>
,
<location id="02634E80FF99FF8EB9A6FFC30885FCFE" LSID="urn:lsid:plazi:treatment:8F15A94DFF99FF8FB9E6FCE30BECFB97:02634E80FF99FF8EB9A6FFC30885FCFE" box="[145,467,749,771]" country="Costa Rica" latitude="10.9292" longLatPrecision="1" longitude="-85.464294" municipality="Area de Conservacion" name="Malaise trap, D. H. Janzen" pageId="18" pageNumber="1359" stateProvince="Guanacaste">Malaise trap, D. H. Janzen</location>
,
<collectorName id="AA497D8DFF99FF8EB8E8FFC00B29FCFE" box="[479,639,749,771]" pageId="18" pageNumber="1359">W. Hallwachs</collectorName>
leg., (
<specimenCount id="11BAD3D2FF99FF8EBBFFFFC40B8EFCFF" box="[712,728,746,770]" pageId="18" pageNumber="1359" type="male">♂</specimenCount>
in alcohol
<specimenCode id="571AB020FF99FF8EB9DFFE2208DFFCDF" box="[232,393,780,802]" pageId="18" pageNumber="1359">BIOUG36194</specimenCode>
-F08).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="4FA64BD0FF99FF8EB9A6FE640BA1FC26" pageId="18" pageNumber="1359" type="etymology">
<paragraph id="0703185BFF99FF8EB9A6FE640BA1FC26" blockId="18.[145,761,842,988]" pageId="18" pageNumber="1359">
<emphasis id="35C8C449FF99FF8EB9A6FE640845FCA2" box="[145,275,842,863]" italics="true" pageId="18" pageNumber="1359">Etymology:</emphasis>
This new species is named in honour of stream parataxonomist Rafael Angel Morales Cueto in recognition of three decades of his research and station management for Estación Biológica Maritza in Sector Orosí of Área de Conservación
<collectingRegion id="C578D6B9FF99FF8EB8DFFEEB0B38FC27" box="[488,622,965,986]" country="Costa Rica" name="Guanacaste" pageId="18" pageNumber="1359">Guanacaste</collectingRegion>
,
<collectingCountry id="7FAB58CBFF99FF8EBB4FFEEB0BA2FC26" box="[632,756,965,987]" name="Costa Rica" pageId="18" pageNumber="1359">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="4FA64BD0FF99FF8EB9A6F92A0BC5FB2F" pageId="18" pageNumber="1359" type="diagnosis">
<paragraph id="0703185BFF99FF8EB9A6F92A0BC5FB2F" blockId="18.[145,761,1028,1234]" pageId="18" pageNumber="1359">
<emphasis id="35C8C449FF99FF8EB9A6F92A0B0AFBE4" box="[145,604,1028,1049]" italics="true" pageId="18" pageNumber="1359">
Diagnosis:
<taxonomicName id="C0BC63D8FF99FF8EB82CF92A0B0AFBE4" authorityName="Vilarino &amp; Dias &amp; Bispo" authorityYear="2022" box="[283,604,1028,1049]" class="Insecta" family="Xiphocentronidae" genus="Caenocentron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="1359" phylum="Arthropoda" rank="species" species="rafamoralesi">Caenocentron rafamoralesi</taxonomicName>
</emphasis>
is similar to
<taxonomicName id="C0BC63D8FF99FF8EB9A6F90D0855FBC5" authorityName="MOULTON &amp; STEWART" authorityYear="1997" box="[145,259,1059,1080]" class="Insecta" family="Xiphocentronidae" genus="Caenocentron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="1359" phylum="Arthropoda" rank="species" species="yavapai">
<emphasis id="35C8C449FF99FF8EB9A6F90D0855FBC5" box="[145,259,1059,1080]" italics="true" pageId="18" pageNumber="1359">C. yavapai</emphasis>
</taxonomicName>
by having an elongate projection on the apical margin of sternum IX. It can be diagnosed by these projections on sternum IX forming a pair of elongate and narrow processes; by the acute spine-like apical projection of coxopodite; and by having the ventral projections of coxopodite short and not fused with each other.
</paragraph>
</subSubSection>
<subSubSection id="4FA64BD0FF99FF8FB9A6F9DE089BFD1B" lastPageId="19" lastPageNumber="1360" pageId="18" pageNumber="1359" type="description">
<paragraph id="0703185BFF99FF8EB9A6F9DE0B2DF9E7" blockId="18.[145,761,1264,1562]" pageId="18" pageNumber="1359">
<emphasis id="35C8C449FF99FF8EB9A6F9DE083DFAF8" box="[145,363,1264,1285]" italics="true" pageId="18" pageNumber="1359">
Adult (
<figureCitation id="9F8704DEFF99FF8EB9D9F9DE080FFAF8" box="[238,345,1264,1285]" captionStart="Figure 13" captionStartId="18.[808,890,1347,1369]" captionTargetBox="[812,1421,196,1306]" captionTargetId="figure-566@18.[811,1422,195,1307]" captionTargetPageId="18" captionText="Figure 13. Caenocentron rafamoralesi sp. nov. A, wing venation. Male genitalia: B, lateral; C, dorsal; D, ventral." figureDoi="http://doi.org/10.5281/zenodo.6461370" httpUri="https://zenodo.org/record/6461370/files/figure.png" pageId="18" pageNumber="1359">Fig. 13A</figureCitation>
):
</emphasis>
Forewing length 4.0
<emphasis id="35C8C449FF99FF8EBBBBF9DE0BCEFAF8" box="[652,664,1264,1285]" italics="true" pageId="18" pageNumber="1359">–</emphasis>
<quantity id="C044B5BEFF99FF8EBBAEF9DE0BA1FAF8" box="[665,759,1264,1285]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="18" pageNumber="1359" unit="mm" value="4.3">4.3 mm</quantity>
(
<emphasis id="35C8C449FF99FF8EB9ADF82109F8FAD9" box="[154,174,1295,1316]" italics="true" pageId="18" pageNumber="1359">N</emphasis>
=
<specimenCount id="11BAD3D2FF99FF8EB9FCF82109BCFAD9" box="[203,234,1292,1316]" pageId="18" pageNumber="1359" type="male">2♂</specimenCount>
). Colour (in alcohol) uniformly dark brown. Maxillary palp formula (I = II = III)-IV-V; segment IV shorter than sum of segments I-II-III; tibial spurs 2-4-
<quantity id="C044B5BEFF99FF8EB9F6F84509A6FA7D" box="[193,240,1386,1408]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="18" pageNumber="1359" unit="in" value="3.0">3 in</quantity>
males and 2-4-
<quantity id="C044B5BEFF99FF8EB887F84508B4FA7D" box="[432,482,1386,1408]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="18" pageNumber="1359" unit="in" value="4.0">4 in</quantity>
females, male hindleg apical spur unmodified. Venation: forewing fork II and IV present, three anal veins present; hindwing fork II and V present, transverse vein between R1 and SR present. Abdominal sternum V with anterolateral oval region with cuticle modified and reticulate.
</paragraph>
<caption id="53C348D3FF99FF8EBA1FF86D0C21FA8A" ID-DOI="http://doi.org/10.5281/zenodo.6461370" ID-Zenodo-Dep="6461370" httpUri="https://zenodo.org/record/6461370/files/figure.png" pageId="18" pageNumber="1359" startId="18.[808,890,1347,1369]" targetBox="[812,1421,196,1306]" targetPageId="18">
<paragraph id="0703185BFF99FF8EBA1FF86D0C21FA8A" blockId="18.[808,1424,1347,1399]" pageId="18" pageNumber="1359">
<emphasis id="35C8C449FF99FF8EBA1FF86D0AF2FAA5" bold="true" box="[808,932,1347,1369]" pageId="18" pageNumber="1359">Figure 13.</emphasis>
<taxonomicName id="C0BC63D8FF99FF8EBA85F86D0D8AFAA4" authority="Vilarino &amp; Dias &amp; Bispo, 2022" authorityName="Vilarino &amp; Dias &amp; Bispo" authorityYear="2022" box="[946,1244,1347,1369]" class="Insecta" family="Xiphocentronidae" genus="Caenocentron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="18" pageNumber="1359" phylum="Arthropoda" rank="species" species="rafamoralesi" status="sp. nov.">
<emphasis id="35C8C449FF99FF8EBA85F86D0D8AFAA4" box="[946,1244,1347,1369]" italics="true" pageId="18" pageNumber="1359">Caenocentron rafamoralesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2EFB7932FF99FF8EBDD2F86D0C60FAA4" box="[1253,1334,1347,1369]" pageId="18" pageNumber="1359" rank="species">sp. nov.</taxonomicNameLabel>
A, wing venation. Male genitalia: B, lateral; C, dorsal; D, ventral.
</paragraph>
</caption>
<paragraph id="0703185BFF99FF8FB9A6FB130BB5FF07" blockId="18.[145,761,1597,1895]" lastBlockId="19.[163,778,197,250]" lastPageId="19" lastPageNumber="1360" pageId="18" pageNumber="1359">
<emphasis id="35C8C449FF99FF8EB9A6FB1308B2F9AF" box="[145,484,1597,1619]" italics="true" pageId="18" pageNumber="1359">
Male genitalia (
<figureCitation id="9F8704DEFF99FF8EB867FB100882F9AF" box="[336,468,1597,1619]" captionStart="Figure 13" captionStartId="18.[808,890,1347,1369]" captionTargetBox="[812,1421,196,1306]" captionTargetId="figure-566@18.[811,1422,195,1307]" captionTargetPageId="18" captionText="Figure 13. Caenocentron rafamoralesi sp. nov. A, wing venation. Male genitalia: B, lateral; C, dorsal; D, ventral." figureDoi="http://doi.org/10.5281/zenodo.6461370" httpUri="https://zenodo.org/record/6461370/files/figure.png" pageId="18" pageNumber="1359">Fig. 13B–D</figureCitation>
):
</emphasis>
Tergum IX, in lateral view, narrow, height greater than length; in dorsal view, short. Sternum IX, in lateral view, rounded, about as long as high; anterior margin deltoid, tapering to narrow apodeme; apical margin with elongate process; dorsal margin slightly prominent; in ventral view, apical margin with two elongate medial processes, as long as sternum length. Segment X membranous, fused to paraproct. Paraproct, in lateral view, oblong, setose, dorsal margin, sclerotized, without any process, ventroapically subacute; in dorsal view, mid-dorsal margins distinctly sclerotized subapically, fused, with rows of setae mesally and laterally. Pre-anal appendage, in lateral view, slightly sinuous, enlarged subapically, tapered apically. Inferior appendage with coxopodite and harpago fused, dorsally with wide gap between the articles. Coxopodite, in lateral view, apical margin with narrow and acute projection; basomesal surface with short projection covered with brush of short setae; in ventral view, basomesal surface of each coxopodite not fused mesally, covered with short setae. Harpago, in lateral view, slender, basal third with short basal spine, many sparse spines present until midlength; in dorsal view, basally enlarged. Basal plate, in lateral view, directed anterad; in ventral view, elongate, acute. Phallus tubular, long and slender, base flared, reaching segment V; apex slightly enlarged.
</paragraph>
<paragraph id="0703185BFF98FF8FB994FC0C089BFD1B" blockId="19.[163,779,290,742]" pageId="19" pageNumber="1360">
<emphasis id="35C8C449FF98FF8FB994FC0C08A5FEC5" box="[163,499,290,312]" italics="true" pageId="19" pageNumber="1360">
Female genitalia (
<figureCitation id="9F8704DEFF98FF8FB85EFC0D08B2FEC5" box="[361,484,291,312]" captionStart="Figures 9–11" captionStartId="16.[145,234,1647,1669]" captionTargetBox="[146,1426,196,1605]" captionTargetId="figure-72@16.[145,1428,195,1607]" captionTargetPageId="16" captionText="Figures 9–11. Caenocentron female genitalia. 9. Caenocentron carlosdelarosai sp. nov. A, lateral; B, dorsal; C, detail of apex lateral; D, detail of apex ventral. 10. Caenocentron rafamoralesi sp. nov.A, lateral; B, dorsal; C, detail of apex lateral; D, detail of apex ventral. 11. Caenocentron yavapai. A, lateral; B, dorsal; C, detail of apex ventral (modified from Ruiter, 2006). Scale bar = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.6461366" httpUri="https://zenodo.org/record/6461366/files/figure.png" pageId="19" pageNumber="1360">Fig. 10A–D</figureCitation>
):
</emphasis>
Telescopically elongate, forming slender oviscapt. Segment VIII synscleritous, dorsally open, membranous, posterior margin with linear row of long setae; each anterolateral margin with narrow, elongate apodeme extending anteriorly until segment VI. Intersegmental membrane VIII–IX well developed. Segment IX tubular, slender, covered with annulated striations; longer than segment VIII; each anterolateral margin with narrow, elongate apodeme extending anteriorly to segment V. Gonopod VIII elongate until segment IX; in lateral view, appearing as narrow, sclerotized rim along IX segment; in ventral view, apex semi-membranous. Gonopod IX rod-like, sclerotized. Segment X small, covered with sensilla. Cercus narrow, digitiform.
</paragraph>
</subSubSection>
<subSubSection id="4FA64BD0FF98FF8FB994FE380A51FC5A" pageId="19" pageNumber="1360" type="discussion">
<paragraph id="0703185BFF98FF8FB994FE380A51FC5A" blockId="19.[163,779,790,935]" pageId="19" pageNumber="1360">
<emphasis id="35C8C449FF98FF8FB994FE380840FCD6" box="[163,278,790,811]" italics="true" pageId="19" pageNumber="1360">Remarks:</emphasis>
The
<emphasis id="35C8C449FF98FF8FB850FE3908CEFCD1" box="[359,408,791,812]" italics="true" pageId="19" pageNumber="1360">COI</emphasis>
sequences of
<taxonomicName id="C0BC63D8FF98FF8FBB7DFE390A5DFCD6" authorityName="Vilarino &amp; Dias &amp; Bispo" authorityYear="2022" box="[586,779,790,812]" class="Insecta" family="Xiphocentronidae" genus="Caenocentron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="19" pageNumber="1360" phylum="Arthropoda" rank="species" species="rafamoralesi">
<emphasis id="35C8C449FF98FF8FBB7DFE390A5DFCD6" box="[586,779,790,812]" italics="true" pageId="19" pageNumber="1360">C. rafamoralesi</emphasis>
</taxonomicName>
had a mean intraspecific genetic distance of 1%. The genetic distance between
<taxonomicName id="C0BC63D8FF98FF8FBB20FE7A0B82FC94" authorityName="Vilarino &amp; Dias &amp; Bispo" authorityYear="2022" box="[535,724,852,873]" class="Insecta" family="Xiphocentronidae" genus="Caenocentron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="19" pageNumber="1360" phylum="Arthropoda" rank="species" species="rafamoralesi">
<emphasis id="35C8C449FF98FF8FBB20FE7A0B7FFC94" box="[535,553,852,873]" italics="true" pageId="19" pageNumber="1360">C</emphasis>
.
<emphasis id="35C8C449FF98FF8FBB0DFE7A0B82FC94" box="[570,724,852,873]" italics="true" pageId="19" pageNumber="1360">rafamoralesi</emphasis>
</taxonomicName>
and
<taxonomicName id="C0BC63D8FF98FF8FB994FE5D0824FC7A" authorityName="Vilarino &amp; Dias &amp; Bispo" authorityYear="2022" box="[163,370,882,904]" class="Insecta" family="Xiphocentronidae" genus="Caenocentron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="19" pageNumber="1360" phylum="Arthropoda" rank="species" species="carlosdelarosai">
<emphasis id="35C8C449FF98FF8FB994FE5D0824FC7A" box="[163,370,882,904]" italics="true" pageId="19" pageNumber="1360">C. carlosdelarosai</emphasis>
</taxonomicName>
was 10%; and the genetic distance between
<taxonomicName id="C0BC63D8FF98FF8FB83FFEBF08E0FC5B" authorityName="Vilarino &amp; Dias &amp; Bispo" authorityYear="2022" box="[264,438,913,934]" class="Insecta" family="Xiphocentronidae" genus="Caenocentron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="19" pageNumber="1360" phylum="Arthropoda" rank="species" species="rafamoralesi">
<emphasis id="35C8C449FF98FF8FB83FFEBF08E0FC5B" box="[264,438,913,934]" italics="true" pageId="19" pageNumber="1360">C. rafamoralesi</emphasis>
</taxonomicName>
and
<taxonomicName id="C0BC63D8FF98FF8FB8D8FEBF0BD9FC5B" baseAuthorityName="MOSELY" baseAuthorityYear="1934" box="[495,655,913,934]" class="Insecta" family="Xiphocentronidae" genus="Caenocentron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="19" pageNumber="1360" phylum="Arthropoda" rank="species" species="trilineatum">
<emphasis id="35C8C449FF98FF8FB8D8FEBF0BD9FC5B" box="[495,655,913,934]" italics="true" pageId="19" pageNumber="1360">C. trilineatum</emphasis>
</taxonomicName>
was 9.6%.
</paragraph>
</subSubSection>
<subSubSection id="4FA64BD0FF98FF8FB994FEF60895FC13" box="[163,451,984,1006]" pageId="19" pageNumber="1360" type="distribution">
<paragraph id="0703185BFF98FF8FB994FEF60895FC13" blockId="19.[163,451,984,1006]" box="[163,451,984,1006]" pageId="19" pageNumber="1360">
<emphasis id="35C8C449FF98FF8FB994FEF60862FC10" box="[163,308,984,1005]" italics="true" pageId="19" pageNumber="1360">Distribution:</emphasis>
<collectingCountry id="7FAB58CBFF98FF8FB876FEF608E9FC13" box="[321,447,984,1006]" name="Costa Rica" pageId="19" pageNumber="1360">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="4FA64BD0FF98FF8FB994F9380BECFB97" pageId="19" pageNumber="1360" type="materials_examined">
<paragraph id="0703185BFF98FF8FB994F9380BECFB97" blockId="19.[163,779,1046,1130]" pageId="19" pageNumber="1360">
<emphasis id="35C8C449FF98FF8FB994F9380829FBD6" box="[163,383,1046,1067]" italics="true" pageId="19" pageNumber="1360">Material examined:</emphasis>
<typeStatus id="D807A6F9FF98FF8FB8A6F9390B50FBD1" box="[401,518,1047,1068]" pageId="19" pageNumber="1360" type="paratype">Paratypes</typeStatus>
: same data as
<typeStatus id="D807A6F9FF98FF8FBB9DF9380A5DFBD1" box="[682,779,1046,1068]" pageId="19" pageNumber="1360" type="holotype">holotype</typeStatus>
except:
<date id="73023E9BFF98FF8FB9C8F91B0821FBB6" box="[255,375,1077,1099]" pageId="19" pageNumber="1360" value="2015-04-23">23.iv.2015</date>
(
<specimenCount id="11BAD3D2FF98FF8FB8BEF91C08CFFBB7" box="[393,409,1074,1098]" pageId="19" pageNumber="1360" type="male">♂</specimenCount>
in alcohol BIOUG33413-E09); and
<date id="73023E9BFF98FF8FB9E3F97A0814FB94" box="[212,322,1108,1129]" pageId="19" pageNumber="1360" value="2014-10-30">30.x.2014</date>
(
<specimenCount id="11BAD3D2FF98FF8FB866F97F0836FB94" box="[337,352,1105,1129]" pageId="19" pageNumber="1360" type="female">♀</specimenCount>
in alcohol BIOUG31506-F08).
</paragraph>
</subSubSection>
</treatment>
</document>
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